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<document id="B5805EC00E554D669996F50FAFBCD7CC" ID-DOI="10.1080/00222930500087408" ID-ISSN="1464-5262" ID-Zenodo-Dep="5217766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629301816659" checkinUser="felipe" docAuthor="Ohtsuka, Susumu, Nishida, Shuhei &amp; Machida, Ryuji J." docDate="2005" docId="03E0481BFFCDFF8ED7815590FBFC2F62" docLanguage="en" docName="JNatHist.39.27.2483-2514.pdf" docOrigin="Journal of Natural History 39 (27)" docSource="http://dx.doi.org/10.1080/00222930500087408" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Paraugaptiloides mirandipes Ohtsuka &amp; Nishida &amp; Machida 2005, n. sp." docType="treatment" docVersion="4" lastPageNumber="2504" masterDocId="FFD93063FFDCFF99D63D5678FFAE2E01" masterDocTitle="Systematics and zoogeography of the deep-sea hyperbenthic family Arietellidae (Copepoda: Calanoida) collected from the Sulu Sea" masterLastPageNumber="2514" masterPageNumber="2483" pageNumber="2498" updateTime="1699067816347" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="2AF261B8741CF21944FAEAC89E7757D0">Systematics and zoogeography of the deep-sea hyperbenthic family Arietellidae (Copepoda: Calanoida) collected from the Sulu Sea</mods:title>
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<mods:namePart id="37703E549A800636A2A91539F6A3D17D">Ohtsuka, Susumu</mods:namePart>
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<mods:namePart id="5BBFFAEB1E1B9F7F8F81FB2EE52555A5">Nishida, Shuhei</mods:namePart>
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<mods:namePart id="7421F8EA5792040B03B315218E0B54A3">Machida, Ryuji J.</mods:namePart>
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<mods:title id="86100F0F56444B7184E6399638E3CB33">Journal of Natural History</mods:title>
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<mods:part id="EE8B20E58A1A72A24E87A0F3EFC7C97F">
<mods:date id="8EEAD9AEB5F57CFB6DC9AD7C2550F239">2005</mods:date>
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<mods:number id="DCCCAC2CF8DC34584161EA0418FA9F25">2005-06-30</mods:number>
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<mods:number id="E0F4A9F46850872BD8A5A66804292999">39</mods:number>
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<mods:identifier id="4831C0A0E125D585F12060224290A445" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03E0481BFFCDFF8ED7815590FBFC2F62" LSID="urn:lsid:plazi:treatment:03E0481BFFCDFF8ED7815590FBFC2F62" httpUri="http://treatment.plazi.org/id/03E0481BFFCDFF8ED7815590FBFC2F62" lastPageId="23" lastPageNumber="2504" pageId="17" pageNumber="2498">
<subSubSection id="C353AA86FFCDFF88D7815590FC2C2A01" box="[444,898,1000,1024]" pageId="17" pageNumber="2498" type="nomenclature">
<paragraph id="8BF6F90DFFCDFF88D7815590FC2C2A01" blockId="17.[444,898,1000,1057]" box="[444,898,1000,1024]" pageId="17" pageNumber="2498">
<heading id="D0BE4E61FFCDFF88D7815590FC2C2A01" box="[444,898,1000,1024]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="2498" reason="4">
<taxonomicName id="4C49828EFFCDFF88D7815590FC992A01" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[444,823,1000,1024]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptiloides" kingdom="Animalia" order="Calanoida" pageId="17" pageNumber="2498" phylum="Arthropoda" rank="species" species="mirandipes" status="sp. nov.">
<emphasis id="B93D251FFFCDFF88D7815590FC992A01" bold="true" box="[444,823,1000,1024]" italics="true" pageId="17" pageNumber="2498">Paraugaptiloides mirandipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E9864FFCDFF88D57C5590FC2C2A01" box="[833,898,1000,1024]" pageId="17" pageNumber="2498" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C353AA86FFCDFF8ED47C5271FBFC2F62" lastPageId="23" lastPageNumber="2504" pageId="17" pageNumber="2498" type="description">
<paragraph id="8BF6F90DFFCDFF88D47C5271FD472A20" blockId="17.[444,898,1000,1057]" box="[577,745,1033,1057]" pageId="17" pageNumber="2498">
(
<figureCitation id="1372E588FFCDFF88D4745271FD4F2A20" box="[585,737,1033,1057]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStartId-0="18.[138,198,1557,1576]" captionStartId-1="19.[125,185,1558,1577]" captionStartId-2="20.[138,198,1583,1602]" captionStartId-3="21.[125,185,867,886]" captionTargetBox-0="[186,1137,176,1519]" captionTargetBox-1="[175,1125,143,1519]" captionTargetBox-2="[189,1133,202,1543]" captionTargetBox-3="[175,1124,137,830]" captionTargetId-0="figure-0@18.[182,1140,173,1521]" captionTargetId-1="figure-1@19.[170,1128,141,1522]" captionTargetId-2="figure-1@20.[182,1140,199,1547]" captionTargetId-3="figure-368@21.[170,1128,133,831]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionTargetPageId-3="21" captionText-0="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." captionText-1="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." captionText-2="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." captionText-3="Figure 10. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Leg 5, anterior; (B) leg 5, posterior; (C) tip of proximal element on third exopodal segment of left leg 5. Scale bars in mm." figureDoi-0="http://doi.org/10.5281/zenodo.5217780" figureDoi-1="http://doi.org/10.5281/zenodo.5217801" figureDoi-2="http://doi.org/10.5281/zenodo.5217803" figureDoi-3="http://doi.org/10.5281/zenodo.5217805" httpUri-0="https://zenodo.org/record/5217780/files/figure.png" httpUri-1="https://zenodo.org/record/5217801/files/figure.png" httpUri-2="https://zenodo.org/record/5217803/files/figure.png" httpUri-3="https://zenodo.org/record/5217805/files/figure.png" pageId="17" pageNumber="2498">Figures 710</figureCitation>
)
</paragraph>
<paragraph id="8BF6F90DFFCDFF88D6405247FEE22A56" blockId="17.[125,332,1087,1111]" box="[125,332,1087,1111]" pageId="17" pageNumber="2498">
<emphasis id="B93D251FFFCDFF88D6405247FEE22A56" box="[125,332,1087,1111]" italics="true" pageId="17" pageNumber="2498">Material examined</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFCDFF88D6405208FEEB2AE8" blockId="17.[125,1172,1136,1257]" pageId="17" pageNumber="2498">
<materialsCitation id="3B21F350FFCDFF88D6405208FEEF2AE8" collectingDate="2002-12-07" elevation="2440" latitude="8.89965" location="Adult" longLatPrecision="1" longitude="120.42515" municipality="central Sulu Sea" pageId="17" pageNumber="2498" specimenCount="1" typeStatus="holotype">
<location id="8E96AFD6FFCDFF88D6405208FF112A89" LSID="urn:lsid:plazi:treatment:03E0481BFFCDFF8ED7815590FBFC2F62:8E96AFD6FFCDFF88D6405208FF112A89" box="[125,191,1136,1160]" latitude="8.89965" longLatPrecision="1" longitude="120.42515" municipality="central Sulu Sea" name="Adult" pageId="17" pageNumber="2498">Adult</location>
male (
<typeStatus id="54F247AFFFCDFF88D7065208FE092A89" box="[315,423,1136,1160]" pageId="17" pageNumber="2498" type="holotype">holotype</typeStatus>
), collected from the
<collectingMunicipality id="6B926377FFCDFF88D4CD5208FC742A89" box="[752,986,1136,1160]" pageId="17" pageNumber="2498">central Sulu Sea</collectingMunicipality>
(
<geoCoordinate id="EE7D9FCAFFCDFF88D23C5208FB212A89" box="[1025,1167,1136,1160]" degrees="08" direction="north" minutes="52.669" orientation="latitude" pageId="17" pageNumber="2498" precision="1" value="8.877816">
08
<emphasis id="B93D251FFFCDFF88D21D5208FB842A89" bold="true" box="[1056,1066,1136,1160]" italics="true" pageId="17" pageNumber="2498">°</emphasis>
52.669N
</geoCoordinate>
,
<geoCoordinate id="EE7D9FCAFFCDFF88D64052E8FEBF2AA9" box="[125,273,1168,1192]" degrees="120" direction="east" minutes="25.289" orientation="longitude" pageId="17" pageNumber="2498" precision="1" value="120.421486">
120
<emphasis id="B93D251FFFCDFF88D69452E8FF1D2AA9" bold="true" box="[169,179,1168,1192]" italics="true" pageId="17" pageNumber="2498">°</emphasis>
25.289E
</geoCoordinate>
,
<geoCoordinate id="EE7D9FCAFFCDFF88D71A52E8FE1D2AA9" box="[295,435,1168,1192]" degrees="08" direction="north" minutes="53.979" orientation="latitude" pageId="17" pageNumber="2498" precision="1" value="8.89965">
08
<emphasis id="B93D251FFFCDFF88D77852E8FEE12AA9" bold="true" box="[325,335,1168,1192]" italics="true" pageId="17" pageNumber="2498">°</emphasis>
53.979N
</geoCoordinate>
,
<geoCoordinate id="EE7D9FCAFFCDFF88D7FF52E8FDF62AA9" box="[450,600,1168,1192]" degrees="120" direction="east" minutes="25.509" orientation="longitude" pageId="17" pageNumber="2498" precision="1" value="120.42515">
120
<emphasis id="B93D251FFFCDFF88D7D352E8FE562AA9" bold="true" box="[494,504,1168,1192]" italics="true" pageId="17" pageNumber="2498">°</emphasis>
25.509E
</geoCoordinate>
; depth
<quantity id="4CB154E8FFCDFF88D48F52E8FCFC2AA9" box="[690,850,1168,1192]" metricMagnitude="3" metricUnit="m" metricValue="2.44" metricValueMax="2.45" metricValueMin="2.43" pageId="17" pageNumber="2498" unit="m" value="2440.0" valueMax="2450.0" valueMin="2430.0">
<elevation id="00641E3EFFCDFF88D48F52E8FCFC2AA9" box="[690,850,1168,1192]" metricMagnitude="3" metricUnit="m" metricValue="2.44" metricValueMax="2.45" metricValueMin="2.43" pageId="17" pageNumber="2498" unit="m" value="2440.0" valueMax="2450.0" valueMin="2430.0">24302450 m</elevation>
</quantity>
)
<collectingMethod id="5208811AFFCDFF88D55E52E8FBEA2AA9" box="[867,1092,1168,1192]" pageId="17" pageNumber="2498">with NORPAC net</collectingMethod>
set on OBT;
<date id="FFF7DFCDFFCDFF88D6F152C9FE082AC8" box="[204,422,1201,1225]" pageId="17" pageNumber="2498" value="2002-12-07">
<collectingDate id="EFB32625FFCDFF88D6F152C9FE082AC8" box="[204,422,1201,1225]" pageId="17" pageNumber="2498" value="2002-12-07">7 December 2002</collectingDate>
</date>
; appendages mounted on glass slides, and body proper in vial; NHM 2005.160
</materialsCitation>
.
</paragraph>
<paragraph id="8BF6F90DFFCDFF88D640535AFF5B2B3B" blockId="17.[125,245,1314,1338]" box="[125,245,1314,1338]" pageId="17" pageNumber="2498">
<emphasis id="B93D251FFFCDFF88D640535AFF5B2B3B" box="[125,245,1314,1338]" italics="true" pageId="17" pageNumber="2498">Description</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFCDFF8CD640532BFD192DF2" blockId="17.[125,1172,1363,1679]" lastBlockId="21.[125,1173,955,1685]" lastPageId="21" lastPageNumber="2502" pageId="17" pageNumber="2498">
<emphasis id="B93D251FFFCDFF88D640532BFE992B6A" box="[125,311,1363,1387]" italics="true" pageId="17" pageNumber="2498">
Male (
<typeStatus id="54F247AFFFCDFF88D6FA532BFE852B6A" box="[199,299,1363,1387]" pageId="17" pageNumber="2498" type="holotype">holotype</typeStatus>
).
</emphasis>
Body (
<figureCitation id="1372E588FFCDFF88D7AF532BFD852B6A" box="[402,555,1363,1387]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="17" pageNumber="2498">Figure 7A, B</figureCitation>
) 5.26 mm, relatively weakly chitinized except for leg 5; prosome approximately 2.3 times as long as urosome. Cephalosome produced posterolaterally to cover anterior part of first pediger (
<figureCitation id="1372E588FFCDFF88D52F53ECFC272BAD" box="[786,905,1428,1452]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="17" pageNumber="2498">Figure 7B</figureCitation>
); rostrum (
<figureCitation id="1372E588FFCDFF88D22953ECFB252BAD" box="[1044,1163,1428,1452]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="17" pageNumber="2498">Figure 7C</figureCitation>
) produced posteriorly, with pair of long filaments (one accidentally lost during dissection). Second and third pedigers produced mediodorsally; fifth pediger produced posteriorly at each side into irregular processes furnished with glandular openings and hair-like sensillae; posterior margin of processes fringed with fine setules (
<figureCitation id="1372E588FFCDFF88D57A506DFB90282C" box="[839,1086,1557,1581]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="17" pageNumber="2498">Figure 7A, B, DG</figureCitation>
); large spermatophore visible, extending from first pediger to genital somite (
<figureCitation id="1372E588FFCDFF88D5A1504EFBBB284F" box="[924,1045,1590,1614]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="17" pageNumber="2498">Figure 7A</figureCitation>
). Urosome five-segmented; genital somite with gonopore opening at right posterolateral corner (
<figureCitation id="1372E588FFCDFF88D6B8500FFEB1288E" box="[133,287,1655,1679]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="17" pageNumber="2498">Figure 7A, B</figureCitation>
); second and third urosomites almost equal in length; anal somite relatively conspicuous; caudal rami (
<figureCitation id="1372E588FFC9FF8CD78655C3FDF72DD2" box="[443,601,955,979]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="21" pageNumber="2502">Figure 7A, H</figureCitation>
) symmetrical, fringed with long setules on both sides; rudimentary seta I present, seta V longest.
</paragraph>
<caption id="DF36A985FFCEFF8BD6B7506DFF182893" ID-DOI="http://doi.org/10.5281/zenodo.5217780" ID-Zenodo-Dep="5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="18" pageNumber="2499" startId="18.[138,198,1557,1576]" targetBox="[186,1137,176,1519]" targetPageId="18">
<paragraph id="8BF6F90DFFCEFF8BD6B7506DFF182893" blockId="18.[138,1186,1557,1682]" pageId="18" pageNumber="2499">
Figure 7.
<taxonomicName id="4C49828EFFCEFF8BD6D5506DFE742829" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[232,474,1557,1576]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptiloides" kingdom="Animalia" order="Calanoida" pageId="18" pageNumber="2499" phylum="Arthropoda" rank="species" species="mirandipes" status="sp. nov.">
<emphasis id="B93D251FFFCEFF8BD6D5506DFE742829" box="[232,474,1557,1576]" italics="true" pageId="18" pageNumber="2499">Paraugaptiloides mirandipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E9864FFCEFF8BD7E3506DFDBD2829" box="[478,531,1557,1576]" pageId="18" pageNumber="2499" rank="species">n. sp.</taxonomicNameLabel>
, male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm.
</paragraph>
</caption>
<caption id="DF36A985FFCFFF8AD640506EFE272892" ID-DOI="http://doi.org/10.5281/zenodo.5217801" ID-Zenodo-Dep="5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="19" pageNumber="2500" startId="19.[125,185,1558,1577]" targetBox="[175,1125,143,1519]" targetPageId="19">
<paragraph id="8BF6F90DFFCFFF8AD640506EFE272892" blockId="19.[125,1173,1558,1683]" pageId="19" pageNumber="2500">
Figure 8.
<taxonomicName id="4C49828EFFCFFF8AD6DF506EFE742828" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[226,474,1558,1577]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptiloides" kingdom="Animalia" order="Calanoida" pageId="19" pageNumber="2500" phylum="Arthropoda" rank="species" species="mirandipes" status="sp. nov.">
<emphasis id="B93D251FFFCFFF8AD6DF506EFE742828" box="[226,474,1558,1577]" italics="true" pageId="19" pageNumber="2500">Paraugaptiloides mirandipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E9864FFCFFF8AD7D8506EFD8E2828" box="[485,544,1558,1577]" pageId="19" pageNumber="2500" rank="species">n. sp.</taxonomicNameLabel>
, male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm.
</paragraph>
</caption>
<caption id="DF36A985FFC8FF8DD6B75057FF182893" ID-DOI="http://doi.org/10.5281/zenodo.5217803" ID-Zenodo-Dep="5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="20" pageNumber="2501" startId="20.[138,198,1583,1602]" targetBox="[189,1133,202,1543]" targetPageId="20">
<paragraph id="8BF6F90DFFC8FF8DD6B75057FF182893" blockId="20.[138,1185,1583,1682]" pageId="20" pageNumber="2501">
Figure 9.
<taxonomicName id="4C49828EFFC8FF8DD6D25057FE462843" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[239,488,1583,1602]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptiloides" kingdom="Animalia" order="Calanoida" pageId="20" pageNumber="2501" phylum="Arthropoda" rank="species" species="mirandipes" status="sp. nov.">
<emphasis id="B93D251FFFC8FF8DD6D25057FE462843" box="[239,488,1583,1602]" italics="true" pageId="20" pageNumber="2501">Paraugaptiloides mirandipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E9864FFC8FF8DD7CE5057FD802843" box="[499,558,1583,1602]" pageId="20" pageNumber="2501" rank="species">n. sp.</taxonomicNameLabel>
, male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm.
</paragraph>
</caption>
<caption id="DF36A985FFC9FF8CD640551BFCE72D91" ID-DOI="http://doi.org/10.5281/zenodo.5217805" ID-Zenodo-Dep="5217805" httpUri="https://zenodo.org/record/5217805/files/figure.png" pageId="21" pageNumber="2502" startId="21.[125,185,867,886]" targetBox="[175,1124,137,830]" targetPageId="21">
<paragraph id="8BF6F90DFFC9FF8CD640551BFCE72D91" blockId="21.[125,1172,867,912]" pageId="21" pageNumber="2502">
Figure 10.
<taxonomicName id="4C49828EFFC9FF8CD6D5551BFE752D77" authority="Ohtsuka &amp; Nishida &amp; Machida, 2005" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[232,475,867,886]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptiloides" kingdom="Animalia" order="Calanoida" pageId="21" pageNumber="2502" phylum="Arthropoda" rank="species" species="mirandipes" status="sp. nov.">
<emphasis id="B93D251FFFC9FF8CD6D5551BFE752D77" box="[232,475,867,886]" italics="true" pageId="21" pageNumber="2502">Paraugaptiloides mirandipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E9864FFC9FF8CD7DF551BFDB92D77" box="[482,535,867,886]" pageId="21" pageNumber="2502" rank="species">n. sp.</taxonomicNameLabel>
, male (holotype). (A) Leg 5, anterior; (B) leg 5, posterior; (C) tip of proximal element on third exopodal segment of left leg 5. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="8BF6F90DFFC9FF8CD6A55584FC932835" blockId="21.[125,1173,955,1685]" pageId="21" pageNumber="2502">
Left geniculate antennule (
<figureCitation id="1372E588FFC9FF8CD7F35584FDCA2A15" box="[462,612,1020,1044]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8AF</figureCitation>
) consisting of 19 segments, with first (IIV) and second (V) posteriorly fringed with fine setules; first to fifth (VIII) segments each with square plate-like structure posterodistally; geniculation positioned between segments 17 (XX) and 18 (XXIXXIII); large anterodistal process originating from compound segment XXIVXXV, reaching beyond antennulary tip; terminal three segments corresponding to ancestral XXIVXXVIII incompletely fused with fusion lines visible clearly. Fusion pattern and armature as follows: IIV (1)56+4ae (two elements missing in
<figureCitation id="1372E588FFC9FF8CD5D152C4FBCC2AD5" box="[1004,1122,1212,1236]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8A</figureCitation>
), V (2)52+ae, VI (3)52+ae, VII (4)52+ae, VIII (5)52+ae, IX (6)52+ae, X (7)51+process +ae, XI (8)52+ae, XII (9)51+process+ae, XIII (10)51+process+ae, XIV (11)52+ae, XV (12)51+process+ae, XVI (13)52+ae, XVII (14)52+ae, XVIII (15)52+ae, XIX (16)51+process (
<figureCitation id="1372E588FFC9FF8CD7765344FE6C2B55" box="[331,450,1340,1364]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8C</figureCitation>
)+spine (
<figureCitation id="1372E588FFC9FF8CD4145344FD0F2B55" box="[553,673,1340,1364]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8D</figureCitation>
)+ae, XX (17)51+process (
<figureCitation id="1372E588FFC9FF8CD5DA5344FBF52B55" box="[999,1115,1340,1364]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8E</figureCitation>
)+ae, XXIXXIII (18)52 (one rudimentary)+2 processes+ae, XXIVXXVIII (19)512+process +2ae (see
<figureCitation id="1372E588FFC9FF8CD6C65304FEC12B95" box="[251,367,1404,1428]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8F</figureCitation>
: XXIV51+1+process; XXV51+1+ae; XXVIXXVIII58+ae), Right antennule 23-segmented, reaching to anal somite (see
<figureCitation id="1372E588FFC9FF8CD4C153E4FCDF2BB5" box="[764,881,1436,1460]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="21" pageNumber="2502">Figure 7B</figureCitation>
). Antenna (
<figureCitation id="1372E588FFC9FF8CD5C153E4FB222BB5" box="[1020,1164,1436,1460]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="21" pageNumber="2502">Figure 7IK</figureCitation>
) with unarmed coxa; basis with inner seta terminally; endopod two-segmented, first segment unarmed, second segment with three inner setae of unequal length and one reduced and five well-developed terminal setae (
<figureCitation id="1372E588FFC9FF8CD4195384FD312815" box="[548,671,1532,1556]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="21" pageNumber="2502">Figure 7K</figureCitation>
); exopod (
<figureCitation id="1372E588FFC9FF8CD51C5384FC052815" box="[801,939,1532,1556]" captionStart="Figure 7" captionStartId="18.[138,198,1557,1576]" captionTargetBox="[186,1137,176,1519]" captionTargetId="figure-0@18.[182,1140,173,1521]" captionTargetPageId="18" captionText="Figure 7. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Habitus, dorsal; (B) habitus, lateral; (C) rostrum, anterior, right rostral filament missing; (D) right distal corner of prosome, dorsal; (E) left distal corner of prosome, dorsal; (F) right distal corner of prosome, lateral; (G) left distal corner of prosome, lateral; (H) left caudal ramus, dorsal; (I) antenna; (J) antennary exopod, setae omitted; (K) terminal portion of antennary endopod. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217780" httpUri="https://zenodo.org/record/5217780/files/figure.png" pageId="21" pageNumber="2502">Figure 7I, J</figureCitation>
) indistinctly sevensegmented, setal formula 0, 1, 1, 1, 1, 1, 3 (one vestigial).
</paragraph>
<paragraph id="8BF6F90DFFC9FF8FD6A55045FE8F2C5E" blockId="21.[125,1173,955,1685]" lastBlockId="22.[138,1185,136,1341]" lastPageId="22" lastPageNumber="2503" pageId="21" pageNumber="2502">
Mandible (
<figureCitation id="1372E588FFC9FF8CD72B5045FE212854" box="[278,399,1597,1621]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="21" pageNumber="2502">Figure 8G</figureCitation>
) with two mono- and one tricuspid teeth and two patches of setules along cutting edge; patch of long setules present near base of palp; endopod absent, but possibly represented by round prominence; exopod five-segmented, setal formula with 1, 1, 1, 1, 2. Maxillule (
<figureCitation id="1372E588FFCAFF8FD75756F0FE4C2EA1" box="[362,482,136,160]" captionStart="Figure 8" captionStartId="19.[125,185,1558,1577]" captionTargetBox="[175,1125,143,1519]" captionTargetId="figure-1@19.[170,1128,141,1522]" captionTargetPageId="19" captionText="Figure 8. Paraugaptiloides mirandipes n. sp., male (holotype). (A) First to 12th segments of left geniculate antennule; (B) 13th to 19th segments of left antennule; (C) proximal spiniform element on antennulary segment XIX; (D) distal spiniform element on left geniculate antennulary segment XIX; (E) anterior spiniform element on left geniculate antennulary segment XX; (F) terminal segments of left geniculate antennule; (G) mandible; (H) maxillule. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217801" httpUri="https://zenodo.org/record/5217801/files/figure.png" pageId="22" pageNumber="2503">Figure 8H</figureCitation>
) with praecoxal arthrite bearing five serrate spines and one process; coxal endite with one serrate seta; coxal epipodite with eight setae, two of which relatively short; basis with two rows of long setules and vestigial seta; endopod onesegmented, bulbous, with two serrate unequal setae terminally; exopod one-segmented, lobate, with three long setae terminally. Maxilla (
<figureCitation id="1372E588FFCAFF8FD4E35770FCFB2F21" box="[734,853,264,288]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9A</figureCitation>
) with first praecoxal endite bearing two setae and one rudimentary element; second praecoxal to second coxal endites each with two serrate setae; basis bearing long spine (
<figureCitation id="1372E588FFCAFF8FD4C25730FCDA2F61" box="[767,884,328,352]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9B</figureCitation>
) with longitudinal row of fine spinules; endopod indistinctly four-segmented, all setae on which well developed, with setal formula 1, 3, 2, 2. Maxilliped (
<figureCitation id="1372E588FFCAFF8FD40157FFFD7D2F9E" box="[572,723,391,415]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9CE</figureCitation>
) with syncoxa bearing one middle and two subterminal serrate setae and subterminal patch of minute spinules; basis as long as syncoxa, with two serrate setae and longitudinal patch of fine spinules; first endopodal segment nearly incorporated into basis, with serrate seta; second to fifth endopodal segments bearing four, four, three and three setae, respectively; fifth endopodal segment slightly longer than preceding segment; terminal endopodal segment with setae
<emphasis id="B93D251FFFCAFF8FD278545FFBFD2C3F" box="[1093,1107,551,574]" italics="true" pageId="22" pageNumber="2503">a</emphasis>
and
<emphasis id="B93D251FFFCAFF8FD2A9545FFB0E2C3E" box="[1172,1184,551,575]" italics="true" pageId="22" pageNumber="2503">b</emphasis>
rudimentary.
</paragraph>
<paragraph id="8BF6F90DFFCAFF8FD698541FFE672D3F" blockId="22.[138,1185,136,1341]" pageId="22" pageNumber="2503">
Seta and spine formula of legs 14 (
<figureCitation id="1372E588FFCAFF8FD460541FFD472C7E" box="[605,745,615,639]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9FI</figureCitation>
) as in
<taxonomicName id="4C49828EFFCAFF8FD57C541FFB372C7E" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[833,1177,615,639]" class="Hexanauplia" family="Arietellidae" genus="Protoparamisophria" kingdom="Animalia" order="Calanoida" pageId="22" pageNumber="2503" phylum="Arthropoda" rank="species" species="biforaminis">
<emphasis id="B93D251FFFCAFF8FD57C541FFB372C7E" box="[833,1177,615,639]" italics="true" pageId="22" pageNumber="2503">Protoparamisophria biforaminis</emphasis>
</taxonomicName>
. Legs 14 with both rami three-segmented. Leg 1 (
<figureCitation id="1372E588FFCAFF8FD51754FFFC312C9E" box="[810,927,647,671]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9F</figureCitation>
) with basis bearing anterodistal row of minute spinules near base of endopod; first endopodal segment with patch of fine setules anterodistally; outer spines on second and third exopodal segments weakly chitinized. Legs 2 (
<figureCitation id="1372E588FFCAFF8FD782549FFD962CFE" box="[447,568,743,767]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9G</figureCitation>
) and 3 (
<figureCitation id="1372E588FFCAFF8FD4A1549FFCB82CFE" box="[668,790,743,767]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9H</figureCitation>
) similar except for shape of distal outer process on second endopodal segment. Leg 4 (
<figureCitation id="1372E588FFCAFF8FD4CF557FFCF12D1E" box="[754,863,775,799]" captionStart="Figure 9" captionStartId="20.[138,198,1583,1602]" captionTargetBox="[189,1133,202,1543]" captionTargetId="figure-1@20.[182,1140,199,1547]" captionTargetPageId="20" captionText="Figure 9. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Maxilla; (B) basal spine of maxilla; (C) maxilliped, some setae on endopod omitted; (D) fourth endopodal segment of maxilliped; (E) fifth endopodal segment of maxilliped; (F) leg 1, anterior; (G) leg 2, anterior; (H) leg 3, anterior; (I) leg 4, anterior. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217803" httpUri="https://zenodo.org/record/5217803/files/figure.png" pageId="22" pageNumber="2503">Figure 9I</figureCitation>
) with round prominence at inner distal corner of basis.
</paragraph>
<paragraph id="8BF6F90DFFCAFF8FD698553FFD5A2B3C" blockId="22.[138,1185,136,1341]" pageId="22" pageNumber="2503">
Leg 5 (
<figureCitation id="1372E588FFCAFF8FD6CB553FFEC62D5E" box="[246,360,839,863]" captionStart="Figure 10" captionStartId="21.[125,185,867,886]" captionTargetBox="[175,1124,137,830]" captionTargetId="figure-368@21.[170,1128,133,831]" captionTargetPageId="21" captionText="Figure 10. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Leg 5, anterior; (B) leg 5, posterior; (C) tip of proximal element on third exopodal segment of left leg 5. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217805" httpUri="https://zenodo.org/record/5217805/files/figure.png" pageId="22" pageNumber="2503">Figure 10</figureCitation>
) highly complex, enlarged, and heavily chitinized; coxae and intercoxal sclerite completely fused. Left leg with basis bearing plumose subterminal seta on posterior surface; endopod distinctly two-segmented, located near inner distal corner of basis, first segment diamond-shaped, second segment originating from inner midpoint of preceding segment; exopod three-segmented, first segment elongate, with minute element at outer distal corner; second segment massive, expanded outwards; third segment strongly curved, bearing proximal short seta, middle curved spine with spatulate tip (
<figureCitation id="1372E588FFCAFF8FD5EF527EFBF42A1F" box="[978,1114,1030,1054]" captionStart="Figure 10" captionStartId="21.[125,185,867,886]" captionTargetBox="[175,1124,137,830]" captionTargetId="figure-368@21.[170,1128,133,831]" captionTargetPageId="21" captionText="Figure 10. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Leg 5, anterior; (B) leg 5, posterior; (C) tip of proximal element on third exopodal segment of left leg 5. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217805" httpUri="https://zenodo.org/record/5217805/files/figure.png" pageId="22" pageNumber="2503">Figure 10C</figureCitation>
), and terminal bifurcate process. Right leg with basis bearing plumose seta on posterior surface; endopod one-segmented with broad base along inner margin of basis, furnished with fine setules subterminally on posterior surface; exopod indistinctly three-segmented, first segment curved inwards at mid-length, carrying two round processes on anterior surface and triangular outer distal process with fine seta; second segment extremely elongated, as long as basis and first exopodal segment combined, inserted on inner subterminal corner of preceding segment, and bearing short seta at mid-length of inner margin, distal patch of fine setules and subterminal round process with minute spinule at tip; third segment lamellar with three minute setae along distal margin.
</paragraph>
<paragraph id="8BF6F90DFFCAFF8FD6B75315FF472B84" blockId="22.[138,233,1389,1413]" box="[138,233,1389,1413]" pageId="22" pageNumber="2503">
<emphasis id="B93D251FFFCAFF8FD6B75315FF472B84" box="[138,233,1389,1413]" italics="true" pageId="22" pageNumber="2503">Remarks</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFCAFF8FD6B753E6FC922894" blockId="22.[138,1185,1438,1685]" pageId="22" pageNumber="2503">
The new species is easily distinguishable from
<taxonomicName id="4C49828EFFCAFF8FD4F053E6FCE82BB4" authorityName="Bradford" authorityYear="1974" box="[717,838,1438,1461]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptilus" kingdom="Animalia" order="Calanoida" pageId="22" pageNumber="2503" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B93D251FFFCAFF8FD4F053E6FCE82BB4" box="[717,838,1438,1461]" italics="true" pageId="22" pageNumber="2503">P. magnus</emphasis>
</taxonomicName>
(features in parentheses) as follows: (1) prosomal posterior corners irregularly pointed (not irregular); (2) three inner setae on the second endopodal segment of the antenna (only two); (3) absence of the mandibular endopod (one-segmented, with two terminal setae); (4) absence of a vestigial element on the inner distal corner of coxa of leg 4 (present); (5) third exopodal segment of the left fifth leg bearing three heavily chitinized processes (only two); (6) endopod of the right fifth leg well developed (relatively reduced); (7) proximal two exopodal segments of the right fifth leg extremely elongated (relatively compact).
</paragraph>
<paragraph id="8BF6F90DFFCBFF8ED6A556F1FC002EC0" blockId="23.[125,1172,137,193]" pageId="23" pageNumber="2504">
Unidentified ciliates were attached near the distal patch of fine setules of the second exopodal segment of the right fifth leg (not illustrated in
<figureCitation id="1372E588FFCBFF8ED52156D1FC0C2EC0" box="[796,930,169,193]" captionStart="Figure 10" captionStartId="21.[125,185,867,886]" captionTargetBox="[175,1124,137,830]" captionTargetId="figure-368@21.[170,1128,133,831]" captionTargetPageId="21" captionText="Figure 10. Paraugaptiloides mirandipes n. sp., male (holotype). (A) Leg 5, anterior; (B) leg 5, posterior; (C) tip of proximal element on third exopodal segment of left leg 5. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.5217805" httpUri="https://zenodo.org/record/5217805/files/figure.png" pageId="23" pageNumber="2504">Figure 10A</figureCitation>
).
</paragraph>
<paragraph id="8BF6F90DFFCBFF8ED6405682FF402F13" blockId="23.[125,238,250,274]" box="[125,238,250,274]" pageId="23" pageNumber="2504">
<emphasis id="B93D251FFFCBFF8ED6405682FF402F13" box="[125,238,250,274]" italics="true" pageId="23" pageNumber="2504">Etymology</emphasis>
</paragraph>
<paragraph id="8BF6F90DFFCBFF8ED6405753FBFC2F62" blockId="23.[125,1172,299,355]" pageId="23" pageNumber="2504">
The specific name of the new species,
<taxonomicName id="4C49828EFFCBFF8ED4055753FD032F42" authorityName="Ohtsuka &amp; Nishida &amp; Machida" authorityYear="2005" box="[568,685,299,323]" class="Hexanauplia" family="Arietellidae" genus="Paraugaptiloides" kingdom="Animalia" order="Calanoida" pageId="23" pageNumber="2504" phylum="Arthropoda" rank="species" species="mirandipes">
<emphasis id="B93D251FFFCBFF8ED4055753FD032F42" box="[568,685,299,323]" italics="true" pageId="23" pageNumber="2504">mirandipes</emphasis>
</taxonomicName>
(Latin
<emphasis id="B93D251FFFCBFF8ED5395753FCC72F42" box="[772,873,299,323]" italics="true" pageId="23" pageNumber="2504">mirandus</emphasis>
, meaning surprising, plus Latin
<emphasis id="B93D251FFFCBFF8ED6F95734FF4A2F62" box="[196,228,332,355]" italics="true" pageId="23" pageNumber="2504">pes</emphasis>
meaning leg), refers to the highly complex, enlarged fifth legs of the male.
</paragraph>
</subSubSection>
</treatment>
</document>