treatments-xml/data/03/D6/B3/03D6B32E8209D42B3ECAF8B37419FC0E.xml
2024-06-21 12:22:17 +02:00

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<document id="C478BC85698E40BFD2080CE9059E164C" ID-CLB-Dataset="199797" ID-DOI="10.11646/zootaxa.5214.3.5" ID-GBIF-Dataset="9eef089e-6399-479b-9200-b2e091ccbb12" ID-ISSN="1175-5326" ID-Zenodo-Dep="7389401" ID-ZooBank="B9AD99D4-7061-4280-BFA2-F5339A9B9A55" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669970497842" checkinUser="plazi" docAuthor="Reynoso-Velasco, Daniel &amp; Sites, Robert W." docDate="2022" docId="03D6B32E8209D42B3ECAF8B37419FC0E" docLanguage="en" docName="zootaxa.5214.3.5.pdf" docOrigin="Zootaxa 5214 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ambrysus signoreti Stal" docType="treatment" docVersion="10" lastPageNumber="424" masterDocId="FFEFCB568208D4283E5DFFB77524FFE3" masterDocTitle="Revision of the Ambrysus signoreti Stål species complex (Heteroptera: Naucoridae Ambrysinae). Section 1." masterLastPageNumber="439" masterPageNumber="421" pageNumber="422" updateTime="1699413971213" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D4A899E8EA498A41785824A151C089FB">Revision of the Ambrysus signoreti Stål species complex (Heteroptera: Naucoridae Ambrysinae). Section 1.</mods:title>
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<mods:namePart id="936184403ACB82C03A0F0C2AC6ED3715">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="E5A51F37205A486BA0AF02D762830FDD">Enns Entomology Museum, Division of Plant Sciences, University of Missouri, Columbia, Missouri 65211, U. S. A. sitesr @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<treatment id="03D6B32E8209D42B3ECAF8B37419FC0E" ID-DOI="http://doi.org/10.5281/zenodo.7391435" ID-GBIF-Taxon="205698866" ID-Zenodo-Dep="7391435" LSID="urn:lsid:plazi:treatment:03D6B32E8209D42B3ECAF8B37419FC0E" httpUri="http://treatment.plazi.org/id/03D6B32E8209D42B3ECAF8B37419FC0E" lastPageId="3" lastPageNumber="424" pageId="1" pageNumber="422">
<subSubSection id="C36551B38209D4293ECAF8B377A2F8FC" box="[151,646,1795,1823]" pageId="1" pageNumber="422" type="nomenclature">
<paragraph id="8BC002388209D4293ECAF8B377A2F8FC" blockId="1.[151,646,1795,1823]" box="[151,646,1795,1823]" pageId="1" pageNumber="422">
<heading id="D088B5548209D4293ECAF8B377A2F8FC" bold="true" box="[151,646,1795,1823]" fontSize="11" level="1" pageId="1" pageNumber="422" reason="1">
<emphasis id="B90BDE2A8209D4293ECAF8B377A2F8FC" bold="true" box="[151,646,1795,1823]" pageId="1" pageNumber="422">
<taxonomicName id="4C7F79BB8209D4293ECAF8B3749DF8FD" ID-CoL="7QM5Z" authority="Stal" authorityName="Stal" box="[151,441,1795,1823]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A8209D4293ECAF8B374A5F8FD" bold="true" box="[151,385,1796,1823]" italics="true" pageId="1" pageNumber="422">Ambrysus signoreti</emphasis>
Stål
</taxonomicName>
species complex
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36551B38209D42B3ECAF8FB71AAFD82" lastPageId="3" lastPageNumber="424" pageId="1" pageNumber="422" type="discussion">
<paragraph id="8BC002388209D42A3ECAF8FB7483FF1A" blockId="1.[151,1437,1868,2039]" lastBlockId="2.[151,1437,151,609]" lastPageId="2" lastPageNumber="423" pageId="1" pageNumber="422">
<emphasis id="B90BDE2A8209D4293ECAF8FB7438F885" bold="true" box="[151,284,1868,1894]" pageId="1" pageNumber="422">Discussion.</emphasis>
It was
<bibRefCitation id="EFEE7FC98209D4293F33F8FB7775F884" author="Usinger, R. L." box="[366,593,1868,1895]" pageId="1" pageNumber="422" pagination="185 - 210" refId="ref12453" refString="Usinger, R. L. (1946) Notes and descriptions of Ambrysus Stal with an account of the life history of Ambrysus mormon Montd. (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 31, 185 - 210." type="journal article" year="1946">Usinger (1946: 203)</bibRefCitation>
who first made reference to the
<taxonomicName id="4C7F79BB8209D4293DE6F8FA713AF885" box="[955,1054,1869,1894]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A8209D4293DE6F8FA713AF885" box="[955,1054,1869,1894]" italics="true" pageId="1" pageNumber="422">signoreti</emphasis>
</taxonomicName>
complex when describing 16 new species in
<taxonomicName id="4C7F79BB8209D4293F54F8C77453F869" authorityName="Stal" authorityYear="1861" box="[265,375,1904,1930]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="B90BDE2A8209D4293F54F8C77453F869" box="[265,375,1904,1930]" italics="true" pageId="1" pageNumber="422">Ambrysus</emphasis>
</taxonomicName>
. It was in the description of his
<taxonomicName id="4C7F79BB8209D4293C8FF8C67643F869" authorityName="Usinger" authorityYear="1946" box="[722,871,1905,1931]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="species" species="sonorensis">
<emphasis id="B90BDE2A8209D4293C8FF8C67643F869" box="[722,871,1905,1931]" italics="true" pageId="1" pageNumber="422">A. sonorensis</emphasis>
</taxonomicName>
that he stated the species belonged to this complex. Unfortunately, he did not provide a list of the group's unifying features nor the constituent species. The succeeding mention of the complex was by
<bibRefCitation id="EFEE7FC98209D4293C59F80E77ECF830" author="La Rivers, I." box="[516,712,1976,2003]" pageId="1" pageNumber="422" pagination="277 - 338" refId="ref11563" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8 (7), 277 - 338." type="journal article" year="1951">La Rivers (1951)</bibRefCitation>
, in his work on the genus
<taxonomicName id="4C7F79BB8209D4293DA9F80F7146F831" authorityName="Stal" authorityYear="1861" box="[1012,1122,1976,2002]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="B90BDE2A8209D4293DA9F80F7146F831" box="[1012,1122,1976,2002]" italics="true" pageId="1" pageNumber="422">Ambrysus</emphasis>
</taxonomicName>
in the
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, where he designated seven additional complexes to accommodate the North American species in the genus. This author also failed to list the diagnostic features and members of the complex, except for
<taxonomicName id="4C7F79BB820AD42A3A79FF20700FFF52" authority="Montandon" authorityName="Montandon" authorityYear="1909" box="[1060,1323,151,177]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mormon">
<emphasis id="B90BDE2A820AD42A3A79FF207184FF53" box="[1060,1184,151,177]" italics="true" pageId="2" pageNumber="423">A. mormon</emphasis>
Montandon
</taxonomicName>
, which he considered to be the most distinctive element of the complex. Regarding the
<taxonomicName id="4C7F79BB820AD42A3A5DFF0C7147FF37" box="[1024,1123,187,212]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820AD42A3A5DFF0C7147FF37" box="[1024,1123,187,212]" italics="true" pageId="2" pageNumber="423">signoreti</emphasis>
</taxonomicName>
complex,
<bibRefCitation id="EFEE7FC9820AD42A3A86FF0C70B8FF36" author="La Rivers, I." box="[1243,1436,186,213]" pageId="2" pageNumber="423" pagination="277 - 338" refId="ref11563" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8 (7), 277 - 338." type="journal article" year="1951">La Rivers (1951)</bibRefCitation>
stated without specifics:
</paragraph>
<paragraph id="8BC00238820AD42A3E9AFEB47166FEFE" blockId="2.[151,1437,151,609]" box="[199,1090,259,285]" pageId="2" pageNumber="423">
&quot;
<emphasis id="B90BDE2A820AD42A3E8EFEB47166FEFE" box="[211,1090,259,285]" italics="true" pageId="2" pageNumber="423">is the most complex unit of the entire genus and the largest in point of species&quot;</emphasis>
</paragraph>
<paragraph id="8BC00238820AD42A3E9AFE9076BEFEA2" blockId="2.[151,1437,151,609]" box="[199,922,295,321]" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3E9AFE9076BEFEA2" box="[199,922,295,321]" italics="true" pageId="2" pageNumber="423">&quot;is difficult on purely technical grounds to sharply characterize&quot;</emphasis>
</paragraph>
<paragraph id="8BC00238820AD42A3E9AFEFC76D6FE86" blockId="2.[151,1437,151,609]" box="[199,1010,331,357]" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3E9AFEFC76D6FE86" box="[199,1010,331,357]" italics="true" pageId="2" pageNumber="423">&quot;displays a homogeneity of form and coloration, which is unmistakable&quot;</emphasis>
</paragraph>
<paragraph id="8BC00238820AD42A3E9AFED8763CFD82" blockId="2.[151,1437,151,609]" pageId="2" pageNumber="423">
Later,
<bibRefCitation id="EFEE7FC9820AD42A3F49FED874FAFE6A" author="La Rivers, I." box="[276,478,366,393]" pageId="2" pageNumber="423" pagination="232 - 237" refId="ref11648" refString="La Rivers, I. (1957) A new Ambrysus from Mexico (Hemiptera, Naucoridae). Entomological News, 68, 232 - 237." type="journal article" year="1957">La Rivers (1957)</bibRefCitation>
described
<taxonomicName id="4C7F79BB820AD42A3C3FFED877EBFE6A" authorityName="La Rivers" authorityYear="1957" box="[610,719,367,393]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="drakei">
<emphasis id="B90BDE2A820AD42A3C3FFED877EBFE6A" box="[610,719,367,393]" italics="true" pageId="2" pageNumber="423">A. drakei</emphasis>
</taxonomicName>
, from northwestern
<collectingCountry id="F36842A8820AD42A3D9EFED8713FFE6A" box="[963,1051,367,393]" name="Mexico" pageId="2" pageNumber="423">Mexico</collectingCountry>
, and assigned it to the
<taxonomicName id="4C7F79BB820AD42A3B67FED870B9FE6B" box="[1338,1437,367,392]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820AD42A3B67FED870B9FE6B" box="[1338,1437,367,392]" italics="true" pageId="2" pageNumber="423">signoreti</emphasis>
</taxonomicName>
complex. In this work, La Rivers finally listed the group's unifying attributes, which included: 1) broad body shape, 2) wide embolia, 3) prominent maculation, and 4) species often with spinose posterolateral corners of abdominal segments. Even with this list, La Rivers failed to explicitly mention the constituent species; only
<taxonomicName id="4C7F79BB820AD42A3A98FE6C705EFE16" authority="Stal" authorityName="Stal" box="[1221,1402,474,501]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820AD42A3A98FE6C7062FE17" box="[1221,1350,475,501]" italics="true" pageId="2" pageNumber="423">A. signoreti</emphasis>
Stål
</taxonomicName>
,
<taxonomicName id="4C7F79BB820AD42A3BD9FE6C744AFDFA" authority="Usinger" authorityName="Usinger" authorityYear="1946" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sonorensis">
<emphasis id="B90BDE2A820AD42A3BD9FE6C742AFDFB" italics="true" pageId="2" pageNumber="423">A. sonorensis</emphasis>
Usinger
</taxonomicName>
,
<taxonomicName id="4C7F79BB820AD42A3F25FE4874D7FDFB" authorityName="Montandon" authorityYear="1909" box="[376,499,511,537]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mormon">
<emphasis id="B90BDE2A820AD42A3F25FE4874D7FDFB" box="[376,499,511,537]" italics="true" pageId="2" pageNumber="423">A. mormon</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7F79BB820AD42A3C73FE487622FDFA" authority="La Rivers" authorityName="La Rivers" authorityYear="1957" box="[558,774,511,537]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="drakei">
<emphasis id="B90BDE2A820AD42A3C73FE4877B0FDFA" box="[558,660,511,537]" italics="true" pageId="2" pageNumber="423">A. drakei</emphasis>
La Rivers
</taxonomicName>
were known to belong to the complex. Based on the current concept of the
<taxonomicName id="4C7F79BB820AD42A3F63FD947485FDDF" box="[318,417,547,572]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820AD42A3F63FD947485FDDF" box="[318,417,547,572]" italics="true" pageId="2" pageNumber="423">signoreti</emphasis>
</taxonomicName>
complex, we know that at that time, at least 13 species belonging to the complex (most of them authored by La Rivers) had been already described.
</paragraph>
<caption id="DF0052B0820AD42A3ECAFD3976E0FD4A" ID-Table-UUID="DF0052B0820AD42A3ECAFD3976E0FD4A" box="[151,964,654,681]" httpUri="http://table.plazi.org/id/DF0052B0820AD42A3ECAFD3976E0FD4A" pageId="2" pageNumber="423" startId="2.[151,244,654,680]" targetBox="[163,1224,696,1889]" targetIsTable="true" targetPageId="2">
<paragraph id="8BC00238820AD42A3ECAFD39769DFD4A" blockId="2.[151,964,654,681]" box="[151,953,654,681]" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3ECAFD397435FD4B" bold="true" box="[151,273,654,680]" pageId="2" pageNumber="423">TABLE 1.</emphasis>
Checklist of species in the
<taxonomicName id="4C7F79BB820AD42A3C19FD38766BFD4A" authority="Stal" authorityName="Stal" box="[580,847,654,681]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820AD42A3C19FD387638FD4B" box="[580,796,655,681]" italics="true" pageId="2" pageNumber="423">Ambrysus signoreti</emphasis>
Stål
</taxonomicName>
complex
</paragraph>
<paragraph id="8BC00238820AD42A3DE4FD3876E0FD4A" blockId="2.[151,964,654,681]" box="[953,964,655,681]" pageId="2" pageNumber="423">*</paragraph>
</caption>
<paragraph id="8BC00238820AD42A3FBFFD0E76F7F882" pageId="2" pageNumber="423">
<table id="F97FF098820A2BD73EFEFD0F71ECF882" box="[163,1224,696,1889]" gridcols="2" gridrows="7" pageId="2" pageNumber="423">
<tr id="354F007A820A2BD73EFEFD0F71ECFD32" box="[163,1224,696,721]" gridrow="0" pageId="2" pageNumber="423">
<th id="769E6906820A2BD73EFEFD0F7752FD32" box="[163,630,696,721]" gridcol="0" gridrow="0" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3FBFFD0E770BFD32" bold="true" box="[482,559,697,721]" italics="true" pageId="2" pageNumber="423">Species</emphasis>
</th>
<th id="769E6906820A2BD73DDBFD0F71ECFD32" box="[902,1224,696,721]" gridcol="1" gridrow="0" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3A14FD0F71ECFD33" bold="true" box="[1097,1224,696,720]" italics="true" pageId="2" pageNumber="423">Distribution</emphasis>
</th>
</tr>
<tr id="354F007A820A2BD73EFEFD5771ECFC79" box="[163,1224,736,922]" gridrow="1" pageId="2" pageNumber="423">
<td id="769E6906820A2BD73EFEFD577752FC79" box="[163,630,736,922]" gridcol="0" gridrow="1" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3EFEFD577421FD1A" bold="true" box="[163,261,736,761]" pageId="2" pageNumber="423">Section 1</emphasis>
<taxonomicName id="4C7F79BB820AD42A3EFEFCBE772CFCC2" authority="La Rivers, 1953" authorityName="La Rivers" authorityYear="1953" box="[163,520,777,801]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="cosmius">
<emphasis id="B90BDE2A820AD42A3EFEFCBE747BFCC2" box="[163,351,777,801]" italics="true" pageId="2" pageNumber="423">Ambrysus cosmius</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F38FCBD772CFCC2" author="La Rivers, I." box="[357,520,777,801]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFC8574D6FCA9" authority="Reynoso &amp; Sites" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2022" box="[163,498,818,842]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="morelosensis" status="sp. nov.">
<emphasis id="B90BDE2A820AD42A3EFEFC857466FCA9" box="[163,322,818,842]" italics="true" pageId="2" pageNumber="423">A. morelosensis</emphasis>
Reynoso &amp; Sites
</taxonomicName>
<emphasis id="B90BDE2A820AD42A3FA4FC857752FCAA" bold="true" box="[505,630,817,842]" pageId="2" pageNumber="423">
<taxonomicNameLabel id="A2386351820AD42A3FA4FC857752FCAA" box="[505,630,817,842]" pageId="2" pageNumber="423" rank="species">new species</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C7F79BB820AD42A3EFEFCED74D5FC91" authority="Reynoso &amp; Sites, 2019" authorityName="Reynoso-Velasco &amp; Sites" authorityYear="2019" box="[163,497,858,882]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B90BDE2A820AD42A3EFEFCED7425FC91" box="[163,257,858,882]" italics="true" pageId="2" pageNumber="423">A. schuhi</emphasis>
Reynoso &amp; Sites, 2019
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFC34749BFC79" authority="Usinger, 1946" authorityName="Usinger" authorityYear="1946" box="[163,447,898,922]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B90BDE2A820AD42A3EFEFC34740EFC79" box="[163,298,899,922]" italics="true" pageId="2" pageNumber="423">A. variegatus</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F6DFC35749BFC79" author="Usinger, R. L." box="[304,447,898,922]" pageId="2" pageNumber="423" pagination="185 - 210" refId="ref12453" refString="Usinger, R. L. (1946) Notes and descriptions of Ambrysus Stal with an account of the life history of Ambrysus mormon Montd. (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 31, 185 - 210." type="journal article" year="1946">Usinger, 1946</bibRefCitation>
</taxonomicName>
</td>
<td id="769E6906820A2BD73DDBFD5771ECFC79" box="[902,1224,736,922]" gridcol="1" gridrow="1" pageId="2" pageNumber="423">Mexico Mexico Mexico Costa Rica</td>
</tr>
<tr id="354F007A820A2BD73EFEFC6571ECFB6F" box="[163,1224,978,1164]" gridrow="2" pageId="2" pageNumber="423">
<td id="769E6906820A2BD73EFEFC657752FB6F" box="[163,630,978,1164]" gridcol="0" gridrow="2" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3EFEFC657421FC09" bold="true" box="[163,261,978,1002]" pageId="2" pageNumber="423">Section 2</emphasis>
<taxonomicName id="4C7F79BB820AD42A3EFEFC4C7704FBF0" authority="Reynoso &amp; Sites, 2019" authorityName="Reynoso &amp; Sites" authorityYear="2019" box="[163,544,1019,1044]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="brailovskyi">
<emphasis id="B90BDE2A820AD42A3EFEFC4C7414FBF0" box="[163,304,1019,1043]" italics="true" pageId="2" pageNumber="423">A. brailovskyi</emphasis>
Reynoso &amp; Sites, 2019
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFB9374F0FBD8" authority="La Rivers, 1962" authorityName="La Rivers" authorityYear="1962" box="[163,468,1059,1083]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="harmodius">
<emphasis id="B90BDE2A820AD42A3EFEFB93740FFBD8" box="[163,299,1059,1083]" italics="true" pageId="2" pageNumber="423">A. harmodius</emphasis>
La Rivers, 1962
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFBFB74D4FB87" authority="Reynoso &amp; Sites, 2019" authorityName="Reynoso &amp; Sites" authorityYear="2019" box="[163,496,1100,1124]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="henryi">
<emphasis id="B90BDE2A820AD42A3EFEFBFB7424FB87" box="[163,256,1100,1124]" italics="true" pageId="2" pageNumber="423">A. henryi</emphasis>
Reynoso &amp; Sites, 2019
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFBC37490FB6F" authority="La Rivers, 1953" authorityName="La Rivers" authorityYear="1953" box="[163,436,1140,1164]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B90BDE2A820AD42A3EFEFBC3742FFB6F" box="[163,267,1140,1164]" italics="true" pageId="2" pageNumber="423">A. inflatus</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F4CFBC37490FB6F" author="La Rivers, I." box="[273,436,1140,1164]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
</td>
<td id="769E6906820A2BD73DDBFC6571ECFB6F" box="[902,1224,978,1164]" gridcol="1" gridrow="2" pageId="2" pageNumber="423">Mexico Costa Rica Mexico Mexico</td>
</tr>
<tr id="354F007A820A2BD73EFEFB2B71ECFB3F" box="[163,1224,1180,1244]" gridrow="3" pageId="2" pageNumber="423">
<td id="769E6906820A2BD73EFEFB2B7752FB3F" box="[163,630,1180,1244]" gridcol="0" gridrow="3" pageId="2" pageNumber="423">
<taxonomicName id="4C7F79BB820AD42A3EFEFB2B74BAFB57" authority="Usinger, 1946" authorityName="Usinger" authorityYear="1946" box="[163,414,1180,1204]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B90BDE2A820AD42A3EFEFB2B742EFB57" box="[163,266,1180,1204]" italics="true" pageId="2" pageNumber="423">A. lunatus</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F4DFB2B74BAFB57" author="Usinger, R. L." box="[272,414,1180,1204]" pageId="2" pageNumber="423" pagination="185 - 210" refId="ref12453" refString="Usinger, R. L. (1946) Notes and descriptions of Ambrysus Stal with an account of the life history of Ambrysus mormon Montd. (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 31, 185 - 210." type="journal article" year="1946">Usinger, 1946</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFB727491FB3F" authority="Usinger, 1946" authorityName="Usinger" authorityYear="1946" box="[163,437,1220,1244]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="B90BDE2A820AD42A3EFEFB727404FB3F" box="[163,288,1220,1244]" italics="true" pageId="2" pageNumber="423">A. lundbladi</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F7BFB737491FB3F" author="Usinger, R. L." box="[294,437,1220,1244]" pageId="2" pageNumber="423" pagination="185 - 210" refId="ref12453" refString="Usinger, R. L. (1946) Notes and descriptions of Ambrysus Stal with an account of the life history of Ambrysus mormon Montd. (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 31, 185 - 210." type="journal article" year="1946">Usinger, 1946</bibRefCitation>
</taxonomicName>
</td>
<td id="769E6906820A2BD73DDBFB2B71ECFB3F" box="[902,1224,1180,1244]" gridcol="1" gridrow="3" pageId="2" pageNumber="423">Mexico and United States Mexico</td>
</tr>
<tr id="354F007A820A2BD73EFEFAA371ECF9FD" box="[163,1224,1300,1566]" gridrow="4" pageId="2" pageNumber="423">
<td id="769E6906820A2BD73EFEFAA37752F9FD" box="[163,630,1300,1566]" gridcol="0" gridrow="4" pageId="2" pageNumber="423">
<emphasis id="B90BDE2A820AD42A3EFEFAA37421FACF" bold="true" box="[163,261,1300,1324]" pageId="2" pageNumber="423">Section 3</emphasis>
<taxonomicName id="4C7F79BB820AD42A3EFEFA8A74E7FAB6" authority="Reynoso, 2021" authorityName="Reynoso" authorityYear="2021" box="[163,451,1341,1366]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="altiplanus">
<emphasis id="B90BDE2A820AD42A3EFEFA8A7400FAB6" box="[163,292,1341,1365]" italics="true" pageId="2" pageNumber="423">A. altiplanus</emphasis>
Reynoso, 2021
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFAD174FDFA9E" authority="Sites &amp; Shepard, 2015" authorityName="Sites &amp; Shepard" authorityYear="2015" box="[163,473,1381,1405]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="cayo">
<emphasis id="B90BDE2A820AD42A3EFEFAD175CBFA9E" box="[163,239,1382,1405]" italics="true" pageId="2" pageNumber="423">A. cayo</emphasis>
Sites &amp; Shepard, 2015
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFA397771FA45" authority="Polhemus &amp; Polhemus, 1981" authorityName="Polhemus &amp; Polhemus" authorityYear="1981" box="[163,597,1422,1446]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="colimanus">
<emphasis id="B90BDE2A820AD42A3EFEFA397403FA45" box="[163,295,1422,1446]" italics="true" pageId="2" pageNumber="423">A. colimanus</emphasis>
Polhemus &amp; Polhemus, 1981
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFA017494FA2D" authority="La Rivers, 1953" authorityName="La Rivers" authorityYear="1953" box="[163,432,1462,1486]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="dyticus">
<emphasis id="B90BDE2A820AD42A3EFEFA017423FA2D" box="[163,263,1462,1486]" italics="true" pageId="2" pageNumber="423">A. dyticus</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F50FA017494FA2D" author="La Rivers, I." box="[269,432,1462,1486]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEFA687487FA15" authority="La Rivers, 1953" authorityName="La Rivers. At" authorityYear="1953" box="[163,419,1502,1526]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="hydor">
<emphasis id="B90BDE2A820AD42A3EFEFA6875DEFA15" box="[163,250,1502,1526]" italics="true" pageId="2" pageNumber="423">A. hydor</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F5DFA687487FA15" author="La Rivers, I." box="[256,419,1502,1526]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF9B074F0F9FD" authority="La Rivers, 1953" authorityName="La Rivers" authorityYear="1953" box="[163,468,1542,1566]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="magniceps">
<emphasis id="B90BDE2A820AD42A3EFEF9B0740FF9FD" box="[163,299,1543,1566]" italics="true" pageId="2" pageNumber="423">A. magniceps</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F6CF9B074F0F9FD" author="La Rivers, I." box="[305,468,1542,1566]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
</td>
<td id="769E6906820A2BD73DDBFAA371ECF9FD" box="[902,1224,1300,1566]" gridcol="1" gridrow="4" pageId="2" pageNumber="423">Mexico Belize Mexico Guatemala Mexico Mexico</td>
</tr>
<tr id="354F007A820A2BD73EFEF99871ECF8F3" box="[163,1224,1583,1808]" gridrow="5" pageId="2" pageNumber="423">
<td id="769E6906820A2BD73EFEF9987752F8F3" box="[163,630,1583,1808]" gridcol="0" gridrow="5" pageId="2" pageNumber="423">
<taxonomicName id="4C7F79BB820AD42A3EFEF99874C6F9A4" authority="La Rivers, 1951" authorityName="La Rivers" authorityYear="1951" box="[163,482,1583,1607]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B90BDE2A820AD42A3EFEF998741DF9A4" box="[163,313,1583,1607]" italics="true" pageId="2" pageNumber="423">A. occidentalis</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F62F99874C6F9A4" author="La Rivers, I." box="[319,482,1583,1607]" pageId="2" pageNumber="423" pagination="277 - 338" refId="ref11563" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8 (7), 277 - 338." type="journal article" year="1951">La Rivers, 1951</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF9E07493F98C" authority="La Rivers, 1953" authorityName="La Rivers" authorityYear="1953" box="[163,439,1623,1647]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="portheo">
<emphasis id="B90BDE2A820AD42A3EFEF9E0742AF98C" box="[163,270,1623,1647]" italics="true" pageId="2" pageNumber="423">A. portheo</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F49F9E07493F98C" author="La Rivers, I." box="[276,439,1623,1647]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF93774FCF974" authority="La Rivers, 1976" authorityName="La Rivers" authorityYear="1976" box="[163,472,1663,1687]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B90BDE2A820AD42A3EFEF9377414F974" box="[163,304,1663,1687]" italics="true" pageId="2" pageNumber="423">A. quadracies</emphasis>
La Rivers, 1976
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF91F74E5F95C" authority="La Rivers, 1962" authorityName="La Rivers" authorityYear="1962" box="[163,449,1703,1727]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="rotundus">
<emphasis id="B90BDE2A820AD42A3EFEF91F743CF95C" box="[163,280,1703,1727]" italics="true" pageId="2" pageNumber="423">A. rotundus</emphasis>
La Rivers, 1962
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF967749BF90B" authority="La Rivers, 1953" authorityName="La Rivers" authorityYear="1953" box="[163,447,1744,1768]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="scalenus">
<emphasis id="B90BDE2A820AD42A3EFEF9677432F90B" box="[163,278,1744,1768]" italics="true" pageId="2" pageNumber="423">A. scalenus</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F41F967749BF90B" author="La Rivers, I." box="[284,447,1744,1768]" pageId="2" pageNumber="423" pagination="1279 - 1349" refId="ref11603" refString="La Rivers, I. (1953) The Ambrysus of Mexico (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 35, 1279 - 1349. https: // doi. org / 10.5962 / bhl. part. 26731" type="journal article" year="1953">La Rivers, 1953</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF94F74A1F8F3" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[163,389,1784,1808]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820AD42A3EFEF94F743CF8F3" box="[163,280,1784,1808]" italics="true" pageId="2" pageNumber="423">A. signoreti</emphasis>
Stål, 1862
</taxonomicName>
</td>
<td id="769E6906820A2BD73DDBF99871ECF8F3" box="[902,1224,1583,1808]" gridcol="1" gridrow="5" pageId="2" pageNumber="423">Mexico and United States Mexico Mexico Mexico Mexico Mexico</td>
</tr>
<tr id="354F007A820A2BD73EFEF89771ECF882" box="[163,1224,1824,1889]" gridrow="6" pageId="2" pageNumber="423">
<td id="769E6906820A2BD73EFEF8977752F882" box="[163,630,1824,1889]" gridcol="0" gridrow="6" pageId="2" pageNumber="423">
<taxonomicName id="4C7F79BB820AD42A3EFEF89674B5F8DB" authority="Reynoso, 2019" authorityName="Reynoso" authorityYear="2019" box="[163,401,1824,1849]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sitesi">
<emphasis id="B90BDE2A820AD42A3EFEF89675D7F8DB" box="[163,243,1825,1848]" italics="true" pageId="2" pageNumber="423">A. sitesi</emphasis>
Reynoso, 2019
</taxonomicName>
<taxonomicName id="4C7F79BB820AD42A3EFEF8FE749BF883" authority="Usinger, 1946" authorityName="Usinger" authorityYear="1946" box="[163,447,1864,1889]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sonorensis">
<emphasis id="B90BDE2A820AD42A3EFEF8FE740EF883" box="[163,298,1865,1888]" italics="true" pageId="2" pageNumber="423">A. sonorensis</emphasis>
<bibRefCitation id="EFEE7FC9820AD42A3F6DF8FE749BF883" author="Usinger, R. L." box="[304,447,1864,1889]" pageId="2" pageNumber="423" pagination="185 - 210" refId="ref12453" refString="Usinger, R. L. (1946) Notes and descriptions of Ambrysus Stal with an account of the life history of Ambrysus mormon Montd. (Hemiptera, Naucoridae). The University of Kansas Science Bulletin, 31, 185 - 210." type="journal article" year="1946">Usinger, 1946</bibRefCitation>
</taxonomicName>
</td>
<td id="769E6906820A2BD73DDBF89771ECF882" box="[902,1224,1824,1889]" gridcol="1" gridrow="6" pageId="2" pageNumber="423">Guatemala and Mexico Mexico</td>
</tr>
</table>
</paragraph>
<tableNote id="769903B6820AD42A3ECAF8C777C3F817" pageId="2" pageNumber="423" targetBox="[163,1224,696,1889]" targetPageId="2">
<paragraph id="8BC00238820AD42A3ECAF8C7765FF86B" blockId="2.[151,922,1904,2036]" box="[151,891,1904,1928]" pageId="2" pageNumber="423">* Sections are based on characteristics of the female subgenital plate:</paragraph>
<paragraph id="8BC00238820AD42A3ECAF82376BEF84F" blockId="2.[151,922,1904,2036]" box="[151,922,1940,1964]" pageId="2" pageNumber="423">1 - posterolateral corners extending posteriorly further than central lobe.</paragraph>
<paragraph id="8BC00238820AD42A3ECAF80F76BEF833" blockId="2.[151,922,1904,2036]" box="[151,922,1976,2000]" pageId="2" pageNumber="423">2 - central lobe extending posteriorly further than posterolateral corners.</paragraph>
<paragraph id="8BC00238820AD42A3ECAF86B77C3F817" blockId="2.[151,922,1904,2036]" box="[151,743,2012,2036]" pageId="2" pageNumber="423">3 - posterolateral corners and central lobe at same level</paragraph>
</tableNote>
<paragraph id="8BC00238820AD42A3CBAF86B77CFF817" blockId="2.[151,922,1904,2036]" box="[743,747,2012,2036]" pageId="2" pageNumber="423">
<tableNote id="769903B6820AD42A3CBAF86B77CFF817" box="[743,747,2012,2036]" pageId="2" pageNumber="423" targetBox="[163,1224,696,1889]" targetPageId="2">.</tableNote>
</paragraph>
<paragraph id="8BC00238820BD42B3E9AFF2070A0FE4E" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="424">
It is only based on the recent studies that have focused on the characterization and circumscription of the genus
<taxonomicName id="4C7F79BB820BD42B3ECAFF0C7421FF36" authorityName="Stal" authorityYear="1861" box="[151,261,187,213]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="B90BDE2A820BD42B3ECAFF0C7421FF36" box="[151,261,187,213]" italics="true" pageId="3" pageNumber="424">Ambrysus</emphasis>
</taxonomicName>
and its nominotypical subgenus (
<bibRefCitation id="EFEE7FC9820BD42B3CD5FF0C76D4FF36" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[648,1008,187,213]" pageId="3" pageNumber="424" pagination="900 - 914" refId="ref12228" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2021) Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology, 46, 900 - 914. https: // doi. org / 10.1111 / syen. 12501" type="journal article" year="2021">Reynoso-Velasco &amp; Sites 2021</bibRefCitation>
), as well as the species complexes in this subgenus (
<bibRefCitation id="EFEE7FC9820BD42B3F62FF687789FF1A" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[319,685,223,249]" pageId="3" pageNumber="424" pagination="201 - 250" refId="ref11874" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2016 a) Revision of the Ambrysus guttatipennis Stal species complex (Heteroptera: Naucoridae: Cryphocricinae) with the description of six new species from Mexico. Zootaxa, 4154 (3), 201 - 250. https: // doi. org / 10.11646 / zootaxa. 4154.3.1" type="journal article" year="2016">Reynoso-Velasco &amp; Sites 2016a</bibRefCitation>
,
<bibRefCitation id="EFEE7FC9820BD42B3CE4FF687626FF1A" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[697,770,223,249]" pageId="3" pageNumber="424" pagination="87 - 114" refId="ref11937" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2016 b) Revision of the Ambrysus hybridus Montandon species complex (Heteroptera: Naucoridae: Cryphocricinae) with the description of a new species from Mexico. Zootaxa, 4169 (1), 87 - 114. https: // doi. org / 10.11646 / zootaxa. 4169.1.4" type="journal article" year="2016">2016b</bibRefCitation>
,
<bibRefCitation id="EFEE7FC9820BD42B3D53FF687671FF1A" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[782,853,223,249]" pageId="3" pageNumber="424" pagination="351 - 377" refId="ref12000" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2018 a) Revision of the Ambrysus pulchellus Montandon species complex (Heteroptera: Nepomorpha: Naucoridae). Zootaxa, 4442 (3), 351 - 377. https: // doi. org / 10.11646 / zootaxa. 4442.3.1" type="journal article" year="2018">2018a</bibRefCitation>
,
<bibRefCitation id="EFEE7FC9820BD42B3D3CFF68768FFF1A" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[865,939,223,249]" pageId="3" pageNumber="424" pagination="1 - 64" refId="ref12054" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2018 b) Revision of the Ambrysus pudicus Stal species complex (Heteroptera: Nepomorpha: Naucoridae) with the description of a new species from southern Mexico. Zootaxa, 4491 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4491.1.1" type="journal article" year="2018">2018b</bibRefCitation>
), that we know that based on characteristics of the male phallosoma the
<taxonomicName id="4C7F79BB820BD42B3F8AFEB4771EFEFF" box="[471,570,259,284]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820BD42B3F8AFEB4771EFEFF" box="[471,570,259,284]" italics="true" pageId="3" pageNumber="424">signoreti</emphasis>
</taxonomicName>
complex includes 24 species distributed in North America (see
<tableCitation id="C6FD3783820BD42B3B4AFEB4704FFEFE" box="[1303,1387,259,285]" captionStart="TABLE 1" captionStartId="2.[151,244,654,680]" captionTargetBox="[163,1224,696,1889]" captionTargetPageId="2" captionText="TABLE 1. Checklist of species in the Ambrysus signoreti Stål complex*" httpUri="http://table.plazi.org/id/DF0052B0820AD42A3ECAFD3976E0FD4A" pageId="3" pageNumber="424" tableUuid="DF0052B0820AD42A3ECAFD3976E0FD4A">Table 1</tableCitation>
). In addition to the morphological data, molecular evidence also supports a close relationship of species in this complex (see
<bibRefCitation id="EFEE7FC9820BD42B3E90FEFC7717FE86" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[205,563,331,357]" pageId="3" pageNumber="424" pagination="900 - 914" refId="ref12228" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2021) Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology, 46, 900 - 914. https: // doi. org / 10.1111 / syen. 12501" type="journal article" year="2021">Reynoso-Velasco &amp; Sites 2021</bibRefCitation>
).
<bibRefCitation id="EFEE7FC9820BD42B3C15FEFC7620FE86" author="La Rivers, I." box="[584,772,330,357]" pageId="3" pageNumber="424" pagination="277 - 338" refId="ref11563" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8 (7), 277 - 338." type="journal article" year="1951">La Rivers (1951</bibRefCitation>
,
<bibRefCitation id="EFEE7FC9820BD42B3D4FFEFD766FFE86" author="La Rivers, I." box="[786,843,330,357]" pageId="3" pageNumber="424" pagination="232 - 237" refId="ref11648" refString="La Rivers, I. (1957) A new Ambrysus from Mexico (Hemiptera, Naucoridae). Entomological News, 68, 232 - 237." type="journal article" year="1957">1957</bibRefCitation>
) assigned
<taxonomicName id="4C7F79BB820BD42B3D9BFEFC7160FE87" authorityName="Montandon" authorityYear="1909" box="[966,1092,331,357]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="mormon">
<emphasis id="B90BDE2A820BD42B3D9BFEFC7160FE87" box="[966,1092,331,357]" italics="true" pageId="3" pageNumber="424">A. mormon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F79BB820BD42B3A23FEFC71C3FE86" authorityName="La Rivers" authorityYear="1957" box="[1150,1255,331,357]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="drakei">
<emphasis id="B90BDE2A820BD42B3A23FEFC71C3FE86" box="[1150,1255,331,357]" italics="true" pageId="3" pageNumber="424">A. drakei</emphasis>
</taxonomicName>
to the
<taxonomicName id="4C7F79BB820BD42B3B67FEFC70B9FE87" box="[1338,1437,331,356]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820BD42B3B67FEFC70B9FE87" box="[1338,1437,331,356]" italics="true" pageId="3" pageNumber="424">signoreti</emphasis>
</taxonomicName>
complex; also, the former species was preliminarily assigned to this complex by
<bibRefCitation id="EFEE7FC9820BD42B3A19FED8704EFE6A" author="Reynoso-Velasco, D." box="[1092,1386,367,393]" pageId="3" pageNumber="424" pagination="304 - 316" refId="ref11756" refString="Reynoso-Velasco, D. (2021 a) A new species of Ambrysus Stal from the Central Mexican Plateau and new state records for species of the genus in the country (Heteroptera: Nepomorpha: Naucoridae). Zootaxa, 5005 (3), 304 - 316. https: // doi. org / 10.11646 / zootaxa. 5005.3.5" type="journal article" year="2021">Reynoso-Velasco (2021a)</bibRefCitation>
, but characteristics of the phallosoma clearly indicate the two species belong to the
<taxonomicName id="4C7F79BB820BD42B3A51FE247199FE4F" box="[1036,1213,403,429]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="guttatipennis">
<emphasis id="B90BDE2A820BD42B3A51FE247199FE4F" box="[1036,1213,403,429]" italics="true" pageId="3" pageNumber="424">A. guttatipennis</emphasis>
</taxonomicName>
species complex.
</paragraph>
<paragraph id="8BC00238820BD42B3E9AFE0071AAFD82" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="424">
For convenience, the species-rich
<taxonomicName id="4C7F79BB820BD42B3C13FE0077F6FE33" box="[590,722,439,465]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820BD42B3C13FE0077F6FE33" box="[590,722,439,465]" italics="true" pageId="3" pageNumber="424">A. signoreti</emphasis>
</taxonomicName>
complex is herein organized into three sections (see
<tableCitation id="C6FD3783820BD42B3B6AFE0070AFFE32" box="[1335,1419,439,465]" captionStart="TABLE 1" captionStartId="2.[151,244,654,680]" captionTargetBox="[163,1224,696,1889]" captionTargetPageId="2" captionText="TABLE 1. Checklist of species in the Ambrysus signoreti Stål complex*" httpUri="http://table.plazi.org/id/DF0052B0820AD42A3ECAFD3976E0FD4A" pageId="3" pageNumber="424" tableUuid="DF0052B0820AD42A3ECAFD3976E0FD4A">Table 1</tableCitation>
). This organization is artificial and does not necessarily indicate a close relationship among the species. It is based on conspicuous characteristics of the female subgenital plate: 1) posterolateral corners extending posteriorly slightly further than the central lobe (
<figureCitation id="13441EBD820BD42B3FBFFD94771FFDDE" box="[482,571,547,573]" captionStart="FIGURE 1" captionStartId="4.[152,255,537,561]" captionTargetBox="[151,1435,176,513]" captionTargetId="figure-19@4.[151,1435,176,513]" captionTargetPageId="4" captionText="FIGURE 1. Representative female subgenital plate of each of three sections in the Ambrysus signoreti Stål complex (terminal abdominal segments removed; solid line indicates level of posterolateral corners and dashed line level of central lobe). (A) posterolateral corners extending posteriorly further than central lobe in Ambrysus cosmius La Rivers, (B) central lobe extending posteriorly further than posterolateral corners in A. inflatus La Rivers, (C) posterolateral corners and central lobe almost at the same level in A. signoreti Stål. sgp = subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7389407" httpUri="https://zenodo.org/record/7389407/files/figure.png" pageId="3" pageNumber="424">Fig. 1A</figureCitation>
), 2) central lobe extending posteriorly further than the posterolateral corners (
<figureCitation id="13441EBD820BD42B3EC2FDF075D2FD82" box="[159,246,583,609]" captionStart="FIGURE 1" captionStartId="4.[152,255,537,561]" captionTargetBox="[151,1435,176,513]" captionTargetId="figure-19@4.[151,1435,176,513]" captionTargetPageId="4" captionText="FIGURE 1. Representative female subgenital plate of each of three sections in the Ambrysus signoreti Stål complex (terminal abdominal segments removed; solid line indicates level of posterolateral corners and dashed line level of central lobe). (A) posterolateral corners extending posteriorly further than central lobe in Ambrysus cosmius La Rivers, (B) central lobe extending posteriorly further than posterolateral corners in A. inflatus La Rivers, (C) posterolateral corners and central lobe almost at the same level in A. signoreti Stål. sgp = subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7389407" httpUri="https://zenodo.org/record/7389407/files/figure.png" pageId="3" pageNumber="424">Fig. 1B</figureCitation>
), and 3) posterolateral corners and central lobe almost at the same level (
<figureCitation id="13441EBD820BD42B3A74FDF071A4FD82" box="[1065,1152,583,609]" captionStart="FIGURE 1" captionStartId="4.[152,255,537,561]" captionTargetBox="[151,1435,176,513]" captionTargetId="figure-19@4.[151,1435,176,513]" captionTargetPageId="4" captionText="FIGURE 1. Representative female subgenital plate of each of three sections in the Ambrysus signoreti Stål complex (terminal abdominal segments removed; solid line indicates level of posterolateral corners and dashed line level of central lobe). (A) posterolateral corners extending posteriorly further than central lobe in Ambrysus cosmius La Rivers, (B) central lobe extending posteriorly further than posterolateral corners in A. inflatus La Rivers, (C) posterolateral corners and central lobe almost at the same level in A. signoreti Stål. sgp = subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7389407" httpUri="https://zenodo.org/record/7389407/files/figure.png" pageId="3" pageNumber="424">Fig. 1C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36551B3820BD42B3E9AFDDD7419FC0E" pageId="3" pageNumber="424" type="diagnosis">
<paragraph id="8BC00238820BD42B3E9AFDDD7419FC0E" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="424">
<emphasis id="B90BDE2A820BD42B3E9AFDDD7466FD67" bold="true" box="[199,322,618,644]" pageId="3" pageNumber="424">Diagnosis.</emphasis>
Similar to other species complexes, the most reliable unifying feature among species in the
<taxonomicName id="4C7F79BB820BD42B3BD9FDDC75DEFD4B" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820BD42B3BD9FDDC75DEFD4B" italics="true" pageId="3" pageNumber="424">A. signoreti</emphasis>
</taxonomicName>
complex is the male phallosoma and includes the presence of small protuberances (pointed or rounded), referred to as &quot;denticles,&quot; on the sclerotized and variously shaped endosomal sclerites (e.g.,
<figureCitation id="13441EBD820BD42B3A94FD047007FD2E" box="[1225,1315,691,717]" captionStart="FIGURE 2" captionStartId="5.[152,255,1766,1790]" captionTargetBox="[151,1435,309,1743]" captionTargetId="figure-79@5.[151,1435,309,1743]" captionTargetPageId="5" captionText="FIGURE 2. Structures of Ambrysus cosmius La Rivers. (A) dorsal habitus of male (yellow arrow indicates light mark at lateral margin and white arrows light mark along posterolateral angle of corium), (B) sternum VI and subgenital plate of female (terminal abdominal segments removed), (C) 6th and 7th abdominal terga of male (inset indicates angles measured on the accessory genitalic process), (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). agp = accessory genitalic process, cl = central lobe, les = left endosomal sclerite, ll = lateral lobe, pct = proctiger, plc = posterolateral corner, pmc = posteromesal corner, pp = pseudoparamere, pr = paramere, pyg = pygophore, res = right endosomal sclerite, sgp = subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7389409" httpUri="https://zenodo.org/record/7389409/files/figure.png" pageId="3" pageNumber="424">Fig. 2E</figureCitation>
). In a few species, the right sclerite is reduced, greatly reduced (
<figureCitation id="13441EBD820BD42B3CB3FD607666FD12" box="[750,834,726,753]" captionStart="FIGURE 5" captionStartId="10.[152,255,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-19@10.[151,1435,181,1615]" captionTargetPageId="10" captionText="FIGURE 5. Structures of Ambrysus morelosensis Reynoso &amp; Sites n. sp. (A) dorsal habitus of female holotype, (B) sternum VI and subgenital plate of female holotype (terminal abdominal segments digitally removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe, res = right endosomal sclerite." figureDoi="http://doi.org/10.5281/zenodo.7389421" httpUri="https://zenodo.org/record/7389421/files/figure.png" pageId="3" pageNumber="424">Fig. 5E</figureCitation>
) or absent; nonetheless, the protuberances are present on the left sclerite, which can also be reduced. Only one species,
<taxonomicName id="4C7F79BB820BD42B3D3DFD4C76C4FCF7" box="[864,992,763,789]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820BD42B3D3DFD4C76C4FCF7" box="[864,992,763,789]" italics="true" pageId="3" pageNumber="424">A. signoreti</emphasis>
</taxonomicName>
, lacks both sclerites and in consequence the protuberances; notwithstanding, molecular evidence clearly supports the inclusion of the species in this complex (
<bibRefCitation id="EFEE7FC9820BD42B3EC2FCF474DBFCBE" author="Reynoso-Velasco, D. &amp; Sites, R. W." box="[159,511,835,861]" pageId="3" pageNumber="424" pagination="900 - 914" refId="ref12228" refString="Reynoso-Velasco, D. &amp; Sites, R. W. (2021) Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology, 46, 900 - 914. https: // doi. org / 10.1111 / syen. 12501" type="journal article" year="2021">Reynoso-Velasco &amp; Sites 2021</bibRefCitation>
). Additionally, the great majority of species in this complex, including
<taxonomicName id="4C7F79BB820BD42B3B49FCF470B2FCBF" box="[1300,1430,835,861]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="B90BDE2A820BD42B3B49FCF470B2FCBF" box="[1300,1430,835,861]" italics="true" pageId="3" pageNumber="424">A. signoreti</emphasis>
</taxonomicName>
, exhibit a characteristic maculation, as suggested but not described by
<bibRefCitation id="EFEE7FC9820BD42B3D92FCD071BEFC62" author="La Rivers, I." box="[975,1178,870,897]" pageId="3" pageNumber="424" pagination="232 - 237" refId="ref11648" refString="La Rivers, I. (1957) A new Ambrysus from Mexico (Hemiptera, Naucoridae). Entomological News, 68, 232 - 237." type="journal article" year="1957">La Rivers (1957)</bibRefCitation>
. This feature includes extensive light-colored markings (sometimes spotted) at the lateral and posterior margins of the corium (e.g.,
<figureCitation id="13441EBD820BD42B3B2DFC3C75EFFC2A" captionStart="FIGURE 10" captionStartId="15.[152,255,1874,1898]" captionTargetBox="[151,1435,417,1851]" captionTargetId="figure-104@15.[151,1435,417,1851]" captionTargetPageId="15" captionText="FIGURE 10. Structures of Ambrysus variegatus Usinger. (A) dorsal habitus of female paratype (yellow and white arrows indicate light marks at lateral and posterior margins of corium, respectively), (B) sternum VI and subgenital plate of female (terminal abdominal segments removed), (C) 6th and 7th abdominal terga of male, (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). al = apical lobe." figureDoi="http://doi.org/10.5281/zenodo.7389435" httpUri="https://zenodo.org/record/7389435/files/figure.png" pageId="3" pageNumber="424">Fig. 10A</figureCitation>
). In some species, the marking on the posterior margin extends along the posterolateral angle of the corium (e.g.,
<figureCitation id="13441EBD820BD42B3E8AFC647414FC0E" box="[215,304,979,1005]" captionStart="FIGURE 2" captionStartId="5.[152,255,1766,1790]" captionTargetBox="[151,1435,309,1743]" captionTargetId="figure-79@5.[151,1435,309,1743]" captionTargetPageId="5" captionText="FIGURE 2. Structures of Ambrysus cosmius La Rivers. (A) dorsal habitus of male (yellow arrow indicates light mark at lateral margin and white arrows light mark along posterolateral angle of corium), (B) sternum VI and subgenital plate of female (terminal abdominal segments removed), (C) 6th and 7th abdominal terga of male (inset indicates angles measured on the accessory genitalic process), (D) 8th abdominal tergum of male, (E) ventral surface of male phallosoma, (F) genital capsule of male (phallosoma removed). agp = accessory genitalic process, cl = central lobe, les = left endosomal sclerite, ll = lateral lobe, pct = proctiger, plc = posterolateral corner, pmc = posteromesal corner, pp = pseudoparamere, pr = paramere, pyg = pygophore, res = right endosomal sclerite, sgp = subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7389409" httpUri="https://zenodo.org/record/7389409/files/figure.png" pageId="3" pageNumber="424">Fig. 2A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>