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<mods:title id="A47CE9A21028BFD9AEB204B184AFA3A6">Redescriptions of Neanthes Kinberg, 1865 (Annelida: Errantia: Nereididae) species from worldwide regions</mods:title>
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<mods:affiliation id="78F0199767FBBE526B8523DAFFEEF64E">Depto. Sistemática y Ecología Acuática, El Colegio de la Frontera Sur, Chetumal, Quintana Roo, 77000, Mexico. Research and Exhibitions Department, Iziko Museums of South Africa, 25 Queen Victoria St, Gardens, Cape Town, 8001, South Africa. South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030, Kuala Nerus, Terengganu, Malaysia.</mods:affiliation>
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<taxonomicName id="4C7F4D53FFB0F525FDF5FA2AFC17FA7F" ID-CoL="45Y4J" authority="(Kinberg, 1865)" baseAuthorityName="Kinberg" baseAuthorityYear="1865" box="[588,999,1486,1514]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="114" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFB0F525FDF5FA2AFCD4FA7C" bold="true" box="[588,804,1486,1513]" italics="true" pageId="13" pageNumber="114">Neanthes helenae</emphasis>
(
<bibRefCitation id="EFEE4B21FFB0F525FC8AFA2BFC2FFA7F" author="Kinberg J. G. H." box="[819,991,1487,1514]" pageId="13" pageNumber="114" pagination="167 - 179" refId="ref21148" refString="Kinberg J. G. H. 1865. Annulata nova. Nereidum dispositio nova. Ofversigt af Kongelige Vetenskaps- Akademiens Forhandlingar 22: 167 - 179. Available from https: // www. biodiversitylibrary. org / page / 32339443 [accessed 1 Jul. 2021]." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
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<paragraph id="8BC036D0FFB0F525FD5EFA16FCBDF999" blockId="13.[588,999,1486,1548]" box="[743,845,1522,1548]" pageId="13" pageNumber="114">
<figureCitation id="13442A55FFB0F525FD5EFA16FCBDF999" box="[743,845,1522,1548]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[189,232,1533,1559]" captionStartId-1="21.[189,232,1463,1489]" captionTargetBox-0="[319,1268,265,1493]" captionTargetBox-1="[346,1241,265,1423]" captionTargetId-0="figure-14@16.[319,1268,265,1493]" captionTargetId-1="figure-11@21.[346,1241,265,1423]" captionTargetPageId-0="16" captionTargetPageId-1="21" captionText-0="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." captionText-1="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.5121740" figureDoi-1="http://doi.org/10.5281/zenodo.5121742" httpUri-0="https://zenodo.org/record/5121740/files/figure.png" httpUri-1="https://zenodo.org/record/5121742/files/figure.png" pageId="13" pageNumber="114">Figs 34</figureCitation>
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<paragraph id="8BC036D0FFB0F525FF04F9D7FB43F9DB" blockId="13.[189,1203,1587,1649]" box="[189,1203,1587,1614]" pageId="13" pageNumber="114">
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<emphasis id="B90BEAC2FFB0F525FF04F9D7FE7CF9D8" box="[189,396,1587,1613]" italics="true" pageId="13" pageNumber="114">Neanthes helenae</emphasis>
<bibRefCitation id="EFEE4B21FFB0F525FE2DF9D7FD88F9DB" author="Kinberg J. G. H." box="[404,632,1587,1614]" pageId="13" pageNumber="114" pagination="167 - 179" refId="ref21148" refString="Kinberg J. G. H. 1865. Annulata nova. Nereidum dispositio nova. Ofversigt af Kongelige Vetenskaps- Akademiens Forhandlingar 22: 167 - 179. Available from https: // www. biodiversitylibrary. org / page / 32339443 [accessed 1 Jul. 2021]." type="journal article" year="1865">Kinberg, 1865: 172</bibRefCitation>
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[
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locality: Jamestown,
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<emphasis id="B90BEAC2FFB0F525FF04F9B3FEF9F9E5" box="[189,265,1623,1648]" italics="true" pageId="13" pageNumber="114">Nereis</emphasis>
(
<emphasis id="B90BEAC2FFB0F525FEA3F9B3FE77F9E4" box="[282,391,1623,1649]" italics="true" pageId="13" pageNumber="114">Neanthes</emphasis>
)
<emphasis id="B90BEAC2FFB0F525FE21F9B3FE05F9E5" box="[408,501,1623,1648]" italics="true" pageId="13" pageNumber="114">nanciae</emphasis>
<bibRefCitation id="EFEE4B21FFB0F525FE45F9B3FD43F9E4" author="Day J. H." box="[508,691,1622,1649]" pageId="13" pageNumber="114" pagination="434 - 451" refId="ref19757" refString="Day J. H. 1949. On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnean Society of London, Zoology 41 (281): 434 - 451. https: // doi. org / 10.1111 / j. 1096 - 3642.1940. tb 02416. x" type="journal article" year="1949">Day, 1949: 445</bibRefCitation>
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, fig. 4ah.
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<emphasis id="B90BEAC2FFB0F525FF04F97CFE7CF927" box="[189,396,1688,1714]" italics="true" pageId="13" pageNumber="114">Neanthes helenae</emphasis>
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<treatmentCitation id="0ADE10C1FFB0F525FE11F97DFD7CF926" author="Hartman O." box="[424,652,1688,1715]" page="250" pageId="13" pageNumber="114" year="1959">
<bibRefCitation id="EFEE4B21FFB0F525FE11F97DFD7CF926" author="Hartman O." box="[424,652,1688,1715]" pageId="13" pageNumber="114" pagination="1 - 353" refId="ref20606" refString="Hartman O. 1959. Catalogue of the polychaetous annelids of the world, 1. Occasional Papers of the Allan Hancock Foundation 23: 1 - 353." type="journal article" year="1959">Hartman 1959: 250</bibRefCitation>
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(species list). —
<treatmentCitation id="0ADE10C1FFB0F525FCE1F97CFBD8F926" author="Wilson R. S." box="[856,1064,1688,1715]" page="225" pageId="13" pageNumber="114" year="1984">
<bibRefCitation id="EFEE4B21FFB0F525FCE1F97CFBD8F926" author="Wilson R. S." box="[856,1064,1688,1715]" pageId="13" pageNumber="114" pagination="209 - 226" refId="ref22922" refString="Wilson R. S. 1984. Neanthes (Polychaeta: Nereididae) from Victoria with descriptions of two new species. Proceedings of the Royal Society of Victoria 96: 209 - 226." type="journal article" year="1984">Wilson 1984: 225</bibRefCitation>
</treatmentCitation>
(species list). —
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Villalobos-
<collectingRegion id="49BBF832FFB0F525FF53F95FFEA4F940" box="[234,340,1723,1749]" country="Mexico" name="Guerrero" pageId="13" pageNumber="114">Guerrero</collectingRegion>
&amp; Idris 2021: 559
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(table 1), 561 (table 2).
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<paragraph id="8BC036D0FFB0F525FF04F93AFD8AF888" blockId="13.[189,1398,1688,1821]" pageId="13" pageNumber="114">
<treatmentCitationGroup id="AB6F11FEFFB0F525FF04F93AFD86F888" pageId="13" pageNumber="114">
<taxonomicName id="4C7F4D53FFB0F525FF04F93AFE60F96D" ID-CoL="8N3C5" baseAuthorityName="Day" baseAuthorityYear="1949" box="[189,400,1758,1784]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="114" phylum="Annelida" rank="species" species="nanciae">
<emphasis id="B90BEAC2FFB0F525FF04F93AFE60F96D" box="[189,400,1758,1784]" italics="true" pageId="13" pageNumber="114">Neanthes nanciae</emphasis>
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<treatmentCitation id="0ADE10C1FFB0F525FE09F93BFD6BF96C" author="Hartman" box="[432,667,1758,1785]" page="250" pageId="13" pageNumber="114" year="1959">
<bibRefCitation id="EFEE4B21FFB0F525FE09F93BFD6BF96C" author="Hartman O." box="[432,667,1758,1785]" pageId="13" pageNumber="114" pagination="1 - 353" refId="ref20606" refString="Hartman O. 1959. Catalogue of the polychaetous annelids of the world, 1. Occasional Papers of the Allan Hancock Foundation 23: 1 - 353." type="journal article" year="1959">Hartman 1959: 250</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ADE10C1FFB0F525FD74F93AFC48F96C" author="Fauchald" box="[717,952,1758,1785]" page="410" pageId="13" pageNumber="114" year="1972">
<bibRefCitation id="EFEE4B21FFB0F525FD74F93AFC48F96C" author="Fauchald K." box="[717,952,1758,1785]" pageId="13" pageNumber="114" pagination="1 - 575" refId="ref20097" refString="Fauchald K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Foundation Monographs in Marine Biology 7: 1 - 575. Available from https: // repository. si. edu / handle / 10088 / 6207 [accessed 1 Jul. 2021]." type="journal article" year="1972">Fauchald 1972: 410</bibRefCitation>
</treatmentCitation>
(group IIB 2a). —
<treatmentCitation id="0ADE10C1FFB0F525FB1AF93AFA86F96D" author="Wilson" box="[1187,1398,1758,1785]" page="226" pageId="13" pageNumber="114" year="1984">
<bibRefCitation id="EFEE4B21FFB0F525FB1AF93AFA86F96D" author="Wilson R. S." box="[1187,1398,1758,1785]" pageId="13" pageNumber="114" pagination="209 - 226" refId="ref22922" refString="Wilson R. S. 1984. Neanthes (Polychaeta: Nereididae) from Victoria with descriptions of two new species. Proceedings of the Royal Society of Victoria 96: 209 - 226." type="journal article" year="1984">Wilson 1984: 226</bibRefCitation>
</treatmentCitation>
(group IIB 2a) (all in species list)
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection id="C365655BFFB0F526FF04F8A7FC4BFC8D" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114" type="materials_examined">
<paragraph id="8BC036D0FFB0F525FF04F8A7FE9CF8C8" blockId="13.[189,364,1859,1885]" box="[189,364,1859,1885]" pageId="13" pageNumber="114">
<heading id="D08881BCFFB0F525FF04F8A7FE9CF8C8" bold="true" box="[189,364,1859,1885]" fontSize="11" level="3" pageId="13" pageNumber="114" reason="3">
<emphasis id="B90BEAC2FFB0F525FF04F8A7FE9CF8C8" bold="true" box="[189,364,1859,1885]" pageId="13" pageNumber="114">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFB0F525FF53F895FEAAF81E" blockId="13.[189,1399,1905,2003]" box="[234,346,1905,1931]" pageId="13" pageNumber="114">
<emphasis id="B90BEAC2FFB0F525FF53F895FEAAF81E" bold="true" box="[234,346,1905,1931]" pageId="13" pageNumber="114">
<typeStatus id="54C48872FFB0F525FF53F895FEAAF81E" box="[234,346,1905,1931]" pageId="13" pageNumber="114" type="syntype">Syntypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC036D0FFB0F526FF04F871FDDDFED2" blockId="13.[189,1399,1905,2003]" lastBlockId="14.[189,1399,264,327]" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114">
<materialsCitation id="3B173C8DFFB0F525FF04F871FCA7F846" ID-GBIF-Occurrence="3326566302" collectionCode="SMNH Type-457" elevation="28" latitude="-16.0" location="ST HELENA" longLatPrecision="1283" longitude="-5.6666665" municipality="Jamestown" pageId="13" pageNumber="114" specimenCount="2" specimenCount-female="1" typeStatus="syntype">
<location id="8EA0600BFFB0F525FF04F871FEACF83A" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB0F525FF04F871FEACF83A" box="[189,348,1941,1967]" latitude="-16.0" longLatPrecision="1283" longitude="-5.6666665" municipality="Jamestown" name="ST HELENA" pageId="13" pageNumber="114">ST HELENA</location>
<specimenCount id="9D79FD59FFB0F525FEC1F871FE1AF83A" box="[376,490,1941,1967]" pageId="13" pageNumber="114">2 atokous</specimenCount>
(one ♀);
<collectingMunicipality id="6BA4ACAAFFB0F525FDDBF871FD18F83A" box="[610,744,1941,1967]" pageId="13" pageNumber="114">Jamestown</collectingMunicipality>
;
<geoCoordinate id="EE4B5017FFB0F525FD4CF877FCADF83A" box="[757,861,1939,1967]" degrees="16" direction="south" minutes="00" orientation="latitude" pageId="13" pageNumber="114" precision="925" value="-16.0">16°00 S</geoCoordinate>
,
<geoCoordinate id="EE4B5017FFB0F525FCD5F877FC20F83A" box="[876,976,1939,1967]" degrees="5" direction="west" minutes="40" orientation="longitude" pageId="13" pageNumber="114" precision="925" value="-5.6666665">5°40 W</geoCoordinate>
;
<elevation id="0052D1E3FFB0F525FC67F871FB8FF83A" box="[990,1151,1941,1968]" metricMagnitude="1" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="114" unit="m" value="27.5">
<quantity id="4C879B35FFB0F525FC67F871FBC1F83A" box="[990,1073,1941,1968]" metricMagnitude="1" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="114" unit="m" value="27.5">27.5 m</quantity>
depth
</elevation>
; Eugenie Expedition 185153 leg.; sta. 173; poor condition;
<collectionCode id="ED6EAE15FFB0F525FD35F85CFCA7F846" box="[652,855,1976,2003]" pageId="13" pageNumber="114">SMNH Type-457</collectionCode>
</materialsCitation>
<materialsCitation id="3B173C8DFFB0F526FCC9F85CFDDDFED2" ID-GBIF-Occurrence="3326566315" collectionCode="NHMUK" elevation="28" lastPageId="14" lastPageNumber="115" latitude="-16.0" location="ST HELENA" longLatPrecision="1283" longitude="-5.6666665" municipality="Jamestown" pageId="13" pageNumber="114" specimenCode="SMNH Type-457" specimenCount="1">
<specimenCount id="9D79FD59FFB0F525FCC9F85CFC34F846" box="[880,964,1976,2003]" pageId="13" pageNumber="114" type="generic">1 spec.</specimenCount>
, permanent slide of atokous ♀; same data as for syntypes; parapodium 24
<superScript id="7C0A9B98FFB3F526FDD1FEECFD85FE83" attach="none" box="[616,629,264,278]" fontSize="6" pageId="14" pageNumber="115">th</superScript>
;
<specimenCode id="DBD99EABFFB3F526FD3FFEEDFCA5FEB1" box="[646,853,265,292]" pageId="14" pageNumber="115">SMNH Type-457</specimenCode>
, housed at the
<collectionCode id="ED6EAE15FFB3F526FBBCFEEDFB84FEB6" box="[1029,1140,265,291]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="115" type="Museum">NHMUK</collectionCode>
[apparently loaned to Arthur Willey in this museum].
</materialsCitation>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FE8AFCA7FE12" blockId="14.[189,855,365,392]" box="[189,855,365,392]" pageId="14" pageNumber="115">
<heading id="D08881BCFFB3F526FF04FE8AFCA7FE12" bold="true" box="[189,855,365,392]" fontSize="11" level="3" pageId="14" pageNumber="115" reason="3">
<emphasis id="B90BEAC2FFB3F526FF04FE8AFCA7FE12" bold="true" box="[189,855,365,392]" pageId="14" pageNumber="115">
Type material of
<taxonomicName id="4C7F4D53FFB3F526FE2AFE89FCA7FE12" authority="Day, 1949" authorityName="Day" authorityYear="1949" box="[403,855,365,392]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="115" phylum="Annelida" rank="species" species="nanciae" subGenus="Neanthes">
<emphasis id="B90BEAC2FFB3F526FE2AFE89FE11FE12" bold="true" box="[403,481,365,391]" italics="true" pageId="14" pageNumber="115">Nereis</emphasis>
(
<emphasis id="B90BEAC2FFB3F526FE4BFE89FD94FE1D" bold="true" box="[498,612,365,392]" italics="true" pageId="14" pageNumber="115">Neanthes</emphasis>
)
<emphasis id="B90BEAC2FFB3F526FDCCFE89FD25FE12" bold="true" box="[629,725,365,391]" italics="true" pageId="14" pageNumber="115">nanciae</emphasis>
<bibRefCitation id="EFEE4B21FFB3F526FD64FE8AFCA7FE12" author="Day J. H." box="[733,855,365,392]" pageId="14" pageNumber="115" pagination="434 - 451" refId="ref19757" refString="Day J. H. 1949. On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnean Society of London, Zoology 41 (281): 434 - 451. https: // doi. org / 10.1111 / j. 1096 - 3642.1940. tb 02416. x" type="journal article" year="1949">Day, 1949</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF53FE78FEACFE23" blockId="14.[189,1399,412,509]" box="[234,348,412,438]" pageId="14" pageNumber="115">
<emphasis id="B90BEAC2FFB3F526FF53FE78FEACFE23" bold="true" box="[234,348,412,438]" pageId="14" pageNumber="115">
<typeStatus id="54C48872FFB3F526FF53FE78FEACFE23" box="[234,348,412,438]" pageId="14" pageNumber="115" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FE5BFB95FE68" blockId="14.[189,1399,412,509]" pageId="14" pageNumber="115">
<materialsCitation id="3B173C8DFFB3F526FF04FE5BFB95FE68" ID-GBIF-Occurrence="3326566306" collectedFrom="upper intertidal" collectingDate="1945-05-07" collectingDateMax="1945-08-03" collectingDateMin="1945-05-07" collectionCode="NHMUK" collectorName="J. Colman" elevation="73" location="ST HELENA" pageId="14" pageNumber="115" specimenCode="NHMUK 1950.1.5.72-95" specimenCount="1">
<location id="8EA0600BFFB3F526FF04FE5BFEACFE4F" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB3F526FF04FE5BFEACFE4F" box="[189,348,447,474]" name="ST HELENA" pageId="14" pageNumber="115">ST HELENA</location>
<specimenCount id="9D79FD59FFB3F526FECEFE5BFE1EFE4C" box="[375,494,447,473]" pageId="14" pageNumber="115">1 atokous</specimenCount>
;
<location id="8EA0600BFFB3F526FE42FE5BFD82FE4C" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB3F526FE42FE5BFD82FE4C" box="[507,626,447,473]" name="Dockyard" pageId="14" pageNumber="115">Dockyard</location>
,
<location id="8EA0600BFFB3F526FDC6FE5BFCE5FE4C" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB3F526FDC6FE5BFCE5FE4C" box="[639,789,447,473]" name="Crown Point" pageId="14" pageNumber="115">Crown Point</location>
and
<location id="8EA0600BFFB3F526FCE8FE24FC27FE4F" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB3F526FCE8FE24FC27FE4F" box="[849,983,448,474]" name="Jamestown" pageId="14" pageNumber="115">Jamestown</location>
;
<quantity id="4C879B35FFB3F526FC5CFE5BFBD2FE4F" box="[997,1058,447,474]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="115" unit="m" value="73.0">
<elevation id="0052D1E3FFB3F526FC5CFE5BFBD2FE4F" box="[997,1058,447,474]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="115" unit="m" value="73.0">73 m</elevation>
</quantity>
depth; some date between
<date id="FFC11010FFB3F526FAD1FE24FE5CFE68" pageId="14" pageNumber="115" value="1945-05-07" valueMax="1945-08-03" valueMin="1945-05-07">
<collectingDate id="EF85E9F8FFB3F526FAD1FE24FE5CFE68" pageId="14" pageNumber="115" value="1945-05-07" valueMax="1945-08-03" valueMin="1945-05-07">7 May to 3 Aug. 1945</collectingDate>
</date>
;
<collectorName id="268A5306FFB3F526FE01FE07FDDFFE69" box="[440,559,482,509]" pageId="14" pageNumber="115">J. Colman</collectorName>
leg.;
<collectedFrom id="3863752DFFB3F526FDC8FE07FCD9FE69" box="[625,809,482,509]" pageId="14" pageNumber="115">upper intertidal</collectedFrom>
;
<specimenCode id="DBD99EABFFB3F526FC8DFE07FB91FE68" box="[820,1121,482,509]" pageId="14" pageNumber="115">NHMUK 1950.1.5.72-95</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF53FDC0FE97FDAB" blockId="14.[189,1325,548,610]" box="[234,359,548,574]" pageId="14" pageNumber="115">
<emphasis id="B90BEAC2FFB3F526FF53FDC0FE97FDAB" bold="true" box="[234,359,548,574]" pageId="14" pageNumber="115">
<typeStatus id="54C48872FFB3F526FF53FDC0FE97FDAB" box="[234,359,548,574]" pageId="14" pageNumber="115" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FDA3FADDFDF7" blockId="14.[189,1325,548,610]" box="[189,1325,583,610]" pageId="14" pageNumber="115">
<materialsCitation id="3B173C8DFFB3F526FF04FDA3FADDFDF7" ID-GBIF-Occurrence="3326566309" box="[189,1325,583,610]" collectedFrom="intertidal, Laurencia or Acanthophora zones" collectingDate="1970-10-23" collectionCode="NHMUK" collectorName="Smithsonian Tropical Research Institute &amp; A. A. Reimer" country="Panama" location="ST HELENA" municipality="Canal Zone" pageId="14" pageNumber="115" specimenCode="NHMUK 1950.1.5.72-95b" specimenCount="1" stateProvince="Colon">
<location id="8EA0600BFFB3F526FF04FDA3FEAAFDF4" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB3F526FF04FDA3FEAAFDF4" box="[189,346,583,609]" name="ST HELENA" pageId="14" pageNumber="115">ST HELENA</location>
<specimenCount id="9D79FD59FFB3F526FECAFDA3FDDCFDF4" box="[371,556,583,609]" pageId="14" pageNumber="115">several atokous</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBD99EABFFB3F526FC57FDACFAD9FDF7" box="[1006,1321,583,610]" pageId="14" pageNumber="115">NHMUK 1950.1.5.72-95b</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FD6CFE23FD37" blockId="14.[189,467,648,674]" box="[189,467,648,674]" pageId="14" pageNumber="115">
<heading id="D08881BCFFB3F526FF04FD6CFE23FD37" bold="true" box="[189,467,648,674]" fontSize="11" level="3" pageId="14" pageNumber="115" reason="3">
<emphasis id="B90BEAC2FFB3F526FF04FD6CFE23FD37" bold="true" box="[189,467,648,674]" pageId="14" pageNumber="115">Comparative material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF53FD53FCB9FD44" blockId="14.[189,1398,695,792]" box="[234,841,695,722]" pageId="14" pageNumber="115">
<emphasis id="B90BEAC2FFB3F526FF53FD53FCB9FD44" bold="true" box="[234,841,695,722]" pageId="14" pageNumber="115">
<typeStatus id="54C48872FFB3F526FF53FD53FEAAFD44" box="[234,346,695,721]" pageId="14" pageNumber="115" type="syntype">Syntypes</typeStatus>
of
<taxonomicName id="4C7F4D53FFB3F526FE3BFD53FCB9FD44" authority="(Day, 1957)" baseAuthorityName="Day" baseAuthorityYear="1957" box="[386,841,695,722]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="115" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFB3F526FE3BFD53FD45FD47" bold="true" box="[386,693,695,722]" italics="true" pageId="14" pageNumber="115">Neanthes indica brunnea</emphasis>
(
<bibRefCitation id="EFEE4B21FFB3F526FD7DFD53FCB1FD44" author="Day J. H." box="[708,833,695,721]" pageId="14" pageNumber="115" pagination="59 - 129" refId="ref19816" refString="Day J. H. 1957. The Polychaete Fauna of South Africa, 4. New species and records from Natal and Mocambique. Annals of the Natal Museum 14 (1): 59 - 129." type="journal article" year="1957">Day, 1957</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FD3EFC4BFC8D" blockId="14.[189,1398,695,792]" pageId="14" pageNumber="115">
<materialsCitation id="3B173C8DFFB3F526FF04FD3EFA94FD61" ID-GBIF-Occurrence="3326566310" box="[189,1380,730,757]" collectionCode="NHMUK" collectorName="J. H. Day" country="Mozambique" location="Morrumbene Estuary" pageId="14" pageNumber="115" specimenCode="NHMUK 1961.16.23" specimenCount="1" stateProvince="Inhambane">
<collectingCountry id="F3687640FFB3F526FF04FD3EFE77FD61" box="[189,391,730,756]" name="Mozambique" pageId="14" pageNumber="115">MOZAMBIQUE</collectingCountry>
<specimenCount id="9D79FD59FFB3F526FE18FD3EFDE6FD61" box="[417,534,730,756]" pageId="14" pageNumber="115">1 atokous</specimenCount>
;
<collectingRegion id="49BBF832FFB3F526FD9AFD3EFD58FD61" box="[547,680,730,756]" country="Mozambique" name="Inhambane" pageId="14" pageNumber="115">Inhambane</collectingRegion>
,
<location id="8EA0600BFFB3F526FD0DFD3EFC42FD60" LSID="urn:lsid:plazi:treatment:03D687C6FFB0F53EFDF5FA2AFEC9F89A:8EA0600BFFB3F526FD0DFD3EFC42FD60" box="[692,946,730,757]" country="Mozambique" name="Morrumbene Estuary" pageId="14" pageNumber="115" stateProvince="Inhambane">Morrumbene Estuary</location>
;
<collectorName id="268A5306FFB3F526FC06FD3FFBD7FD60" box="[959,1063,731,757]" pageId="14" pageNumber="115">J.H. Day</collectorName>
leg.;
<specimenCode id="DBD99EABFFB3F526FBD3FD3FFA94FD61" box="[1130,1380,730,757]" pageId="14" pageNumber="115">NHMUK 1961.16.23</specimenCode>
</materialsCitation>
<materialsCitation id="3B173C8DFFB3F526FF04FD1AFC47FC8D" ID-GBIF-Occurrence="3326566308" box="[189,951,765,792]" collectionCode="NHMUK" collectorName="J. H. Day" country="Mozambique" location="Inhambane" pageId="14" pageNumber="115" specimenCode="SAM-A-20983" specimenCount="1" stateProvince="Inhambane">
<specimenCount id="9D79FD59FFB3F526FF04FD1AFEC2FC82" box="[189,306,765,792]" pageId="14" pageNumber="115">1 atokous</specimenCount>
; same collection data as for preceding;
<specimenCode id="DBD99EABFFB3F526FCB8FD1AFC47FC8D" box="[769,951,766,792]" pageId="14" pageNumber="115">SAM-A-20983</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365655BFFB3F53CFF04FCDBFC71FA6D" lastPageId="20" lastPageNumber="121" pageId="14" pageNumber="115" type="description">
<paragraph id="8BC036D0FFB3F526FF04FCDBFEBEFCCC" blockId="14.[189,334,831,857]" box="[189,334,831,857]" pageId="14" pageNumber="115">
<heading id="D08881BCFFB3F526FF04FCDBFEBEFCCC" bold="true" box="[189,334,831,857]" fontSize="11" level="3" pageId="14" pageNumber="115" reason="3">
<emphasis id="B90BEAC2FFB3F526FF04FCDBFEBEFCCC" bold="true" box="[189,334,831,857]" pageId="14" pageNumber="115">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FC8AFB31FC5B" blockId="14.[187,1398,877,974]" pageId="14" pageNumber="115">
COLOUR AND MEASUREMENTS.
<typeStatus id="54C48872FFB3F526FDBEFC8AFD97FC1D" box="[519,615,878,904]" pageId="14" pageNumber="115" type="syntype">Syntype</typeStatus>
atokous, complete but fragmented into two parts, in poor condition,
<quantity id="4C879B35FFB3F526FF02FC75FED7FC3E" box="[187,295,913,940]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="14" pageNumber="115" unit="mm" value="26.5">26.5 mm</quantity>
TL,
<quantity id="4C879B35FFB3F526FEDCFC75FE32FC3E" box="[357,450,913,939]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="14" pageNumber="115" unit="mm" value="3.3">3.3 mm</quantity>
L15, 1 mm W15, with 89 chaetigers. Body pale yellow, lacking pigmentation patterns, stained with Shirlastain A to observe edges, scars and paragnaths (
<figureCitation id="13442A55FFB3F526FB88FC50FB45FC5B" box="[1073,1205,948,974]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="14" pageNumber="115">Fig. 3AD</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FC12FAB8FBC3" blockId="14.[187,1399,1013,1110]" pageId="14" pageNumber="115">
PROSTOMIUM. Campanulate, as long as wide (
<figureCitation id="13442A55FFB3F526FD06FC11FCE9FB9A" box="[703,793,1013,1039]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="14" pageNumber="115">Fig. 3B</figureCitation>
); anterior end broad, distally complete; anterolateral gap beside palpophore narrow, as wide as antennal diameter; dorsal groove distinct, shallow, running mid-subdistally. Nuchal organs exposed, broad, twice as wide as diameter of posterior pair of eyes.
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FB9AFE99FB48" blockId="14.[189,1398,1149,1245]" pageId="14" pageNumber="115">
PALPOPHORES. Sub-conical, 1.2 times wider than long (
<figureCitation id="13442A55FFB3F526FC9DFB99FC8CFB02" box="[804,892,1149,1175]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="14" pageNumber="115">Fig. 3B</figureCitation>
), as long as four-fifths of entire prostomium; with distinct sub-distal transverse groove. Palpostyles ovoid, thick, with diameter as wide as two-fifths of palpophore.
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FAE1FAA8FAD7" blockId="14.[189,1398,1285,1347]" pageId="14" pageNumber="115">
ANTENNAE. Tapered, thick, long, extending forwards beyond tip of palpophore and posteriorly to halflength of prostomium; antennae separated, with gap as wide as basal diameter of antennae (
<figureCitation id="13442A55FFB3F526FB57FACCFABAFAD7" box="[1262,1354,1320,1346]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="14" pageNumber="115">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04FA8EFF04F9A1" blockId="14.[188,1399,1386,1588]" pageId="14" pageNumber="115">
EYES. Paired eyes purplish but gradually fading, arranged in a trapezoid form; gap between both pairs two-thirds as wide as diameter of posterior pair of eyes (
<figureCitation id="13442A55FFB3F526FCF4FA69FC58FA32" box="[845,936,1421,1447]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="14" pageNumber="115">Fig. 3B</figureCitation>
); anterior pair of eyes oval, 1.6 times as wide as basal diameter of antennae, gap between both eyes 3.5 times as wide as diameter of eyes, with lens barely distinct, translucent, covering 20% of eye; posterior pair of eyes rounded, 1.7 times as wide as basal diameter of antennae, with lens distinct, translucent, placed in middle of eye and covering 20% of it.
</paragraph>
<paragraph id="8BC036D0FFB3F526FF04F9BFFAB2F90D" blockId="14.[189,1398,1627,1689]" pageId="14" pageNumber="115">APODOUS ANTERIOR SEGMENT. Segment 4 times wider than long, 1.5 times as long as chaetiger 1, with rounded anterior margin and ventrolateral projections; dorsum without marked transverse wrinkle.</paragraph>
<paragraph id="8BC036D0FFB3F526FF04F924FC73F838" blockId="14.[188,1399,1728,1965]" pageId="14" pageNumber="115">
TENTACULAR CIRRI. Cirri of right flank dehiscent, remaining slightly thickened, smooth (
<figureCitation id="13442A55FFB3F526FB1EF924FAF1F94F" box="[1191,1281,1728,1754]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="14" pageNumber="115">Fig. 3B</figureCitation>
); posterodorsal cirri broken, extending posteriorly to chaetiger 6, but according to original description, extending to chaetiger 10; antero-dorsal cirri broken, extending posteriorly to chaetiger 3; postero-ventral cirri extended over opposite side of prostomium; antero-ventral cirri slightly longer than postero-ventral cirri and extending beyond twice length of palpophore; dorsal cirrophores cylindrical, ventral cirrophores ring-shaped, postero-dorsal cirrophores 1.5 times as long as antero-dorsal cirrophores, antero-ventral cirrophores 1.2 times as wide as postero-ventral cirrophores.
</paragraph>
<paragraph id="8BC036D0FFB2F527FF04FEEEFB2BFCA6" blockId="15.[189,1401,265,820]" pageId="15" pageNumber="116">
PHARYNX. Everted, in poor condition, damaged, jaws and several areas missing, only areas V and VI remain. Pharynx of second
<typeStatus id="54C48872FFB2F527FDB8FEC9FDACFED2" box="[513,604,301,327]" pageId="15" pageNumber="116" type="syntype">syntype</typeStatus>
described here, everted, in poor condition, damaged, jaws missing; paragnaths on maxillary and oral rings amber, barely distinct but stained here (
<figureCitation id="13442A55FFB2F527FBD1FEABFB1AFEFF" box="[1128,1258,335,362]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3CD</figureCitation>
), consisting of conical, p-bars, and merged paragnaths; plate-like basements absent. Area I: 1, small cone; areas IIa: 31 and IIb: 38, three slightly regular rows of uneven cones in eyebrow-shaped patch, subdistal cones larger, fang-shaped in outer-most row (
<figureCitation id="13442A55FFB2F527FD2FFE5DFD05FE46" box="[662,757,441,467]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3C</figureCitation>
); area III: 18, four irregular rows of uneven cones in sub-oval patch, with three distinct laterally-isolated cones, distal cones smaller; areas IVa: missing and IVb: 26, L-shaped patch with proximal half consisting of two regular rows and distal half with four slightly regular rows of uneven cones and four long and slender merged paragnaths (34 times longer than wide) located near jaw; area V: 1 (1), small conical paragnath placed on same level between most distal and proximal paragnaths on area VI (
<figureCitation id="13442A55FFB2F527FD03FD8DFCE7FD16" box="[698,791,617,643]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3C</figureCitation>
); areas VIa: 7 (5) and VIb: 6 (5), oblique transverse row of uneven p-bars becoming shorter outwards (six in VIa, four in VIb;
<figureCitation id="13442A55FFB2F527FBECFD68FB41FD33" box="[1109,1201,652,678]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3C</figureCitation>
) and one single outermost cone (
<figureCitation id="13442A55FFB2F527FE38FD4BFE2EFD5F" box="[385,478,687,714]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3C</figureCitation>
); areas VIIVIII: 31, two bands of cones, with anterior band consisting of two transversely aligned rows (furrow row with one small cone and ridge row with one fang-shaped cone on each region) (
<figureCitation id="13442A55FFB2F527FE3DFD12FE2EFC85" box="[388,478,758,784]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3D</figureCitation>
), and posterior band with two transverse rows (
<figureCitation id="13442A55FFB2F527FBB1FD12FB96FC85" box="[1032,1126,758,784]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3D</figureCitation>
). Areas VIVVI ridge pattern, π- shaped. Gap between area VI and areas VIIVIII narrow, as wide as palpostyle.
</paragraph>
<paragraph id="8BC036D0FFB2F527FF04FCBFFD74FCE1" blockId="15.[189,644,858,885]" box="[189,644,858,885]" pageId="15" pageNumber="116">PAIRED OESOPHAGEAL CAECA. Unknown.</paragraph>
<paragraph id="8BC036D0FFB2F527FF04FC79FDCAFBB5" blockId="15.[188,1399,924,1056]" pageId="15" pageNumber="116">PARAPODIA. Without glandular, dorsal patches. Notopodia consisting of dorsal cirrus, dorsal ligule (distal and proximal), notopodial prechaetal lobe, and median ligule in biramous parapodia. Neuropodia consisting of neuroacicular ligule with inferior and postchaetal lobes, ventral ligule, and ventral cirrus; superior lobe absent throughout.</paragraph>
<paragraph id="8BC036D0FFB2F527FF04FBACFC7BFB7B" blockId="15.[189,1399,1095,1262]" pageId="15" pageNumber="116">
DORSAL CIRRI. Cirriform, long, extending markedly beyond distal region of dorsal ligule throughout (
<figureCitation id="13442A55FFB2F527FF7FFB8EFECDFB11" box="[198,317,1130,1156]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3EI</figureCitation>
); dorsal cirri 33.5 times as long as proximal region of dorsal ligule in anteriormost and anterior parapodia (
<figureCitation id="13442A55FFB2F527FE1AFB69FDEDFB3D" box="[419,541,1165,1192]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3EF</figureCitation>
), 22.5 times as long as that in following parapodia (
<figureCitation id="13442A55FFB2F527FB30FB69FAF0FB3D" box="[1161,1280,1165,1192]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3GI</figureCitation>
); attached basally to dorsal ligule in anteriormost parapodia (
<figureCitation id="13442A55FFB2F527FCAFFB55FC81FB5E" box="[790,881,1201,1227]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3E</figureCitation>
), medially in anterior, middle and posterior parapodia (
<figureCitation id="13442A55FFB2F527FEFBFB30FE44FB7B" box="[322,436,1236,1262]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3FI</figureCitation>
), two-thirds in posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFB2F527FF04FAF2FE0FF91A" blockId="15.[189,1401,1301,1679]" pageId="15" pageNumber="116">
DORSAL LIGULE. Proximal region even throughout, except slightly humped from middle parapodia towards posterior end; shorter than distal region of dorsal ligule in anteriormost parapodia (
<figureCitation id="13442A55FFB2F527FAB5FADCFA98FAC6" box="[1292,1384,1336,1363]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3E</figureCitation>
), as long as that in anterior, middle and posterior parapodia (
<figureCitation id="13442A55FFB2F527FC23FAB8FBE0FAE3" box="[922,1040,1372,1398]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3FI</figureCitation>
), 1.5 times as long as that in posteriormost parapodia; two irregular glandular patches covering partially proximal region of dorsal ligule in anteriormost and anterior parapodia (
<figureCitation id="13442A55FFB2F527FCB8FA46FC8FFA29" box="[769,895,1442,1468]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3EF</figureCitation>
) and entirely covering that in following parapodia (
<figureCitation id="13442A55FFB2F527FEFAFA21FE4CFA4A" box="[323,444,1477,1503]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3GI</figureCitation>
). Distal region well developed, becoming slightly longer towards posterior end; digitiform in anteriormost parapodia (
<figureCitation id="13442A55FFB2F527FDC5FA0CFD27F996" box="[636,727,1512,1539]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3E</figureCitation>
), bluntly rounded in anterior parapodia (
<figureCitation id="13442A55FFB2F527FB0CFA0CFAFDF996" box="[1205,1293,1512,1539]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3F</figureCitation>
), conical in middle parapodia (
<figureCitation id="13442A55FFB2F527FE0FF9E8FDC8F9B3" box="[438,568,1548,1574]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3GH</figureCitation>
), bluntly conical in remaining parapodia (
<figureCitation id="13442A55FFB2F527FBA6F9E8FB81F9B3" box="[1055,1137,1548,1574]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3I</figureCitation>
); as long as or slightly shorter than median ligule in anterior parapodia (
<figureCitation id="13442A55FFB2F527FCA1F9CBFC84F9DC" box="[792,884,1583,1609]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3F</figureCitation>
), longer than that in following parapodia; projecting beyond notoacicula throughout; one irregular glandular patch covering entirely distal region of dorsal ligule (
<figureCitation id="13442A55FFB2F527FEC6F991FE01F91A" box="[383,497,1653,1679]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3FI</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFB2F527FF04F953FD89F882" blockId="15.[188,1400,1719,1815]" pageId="15" pageNumber="116">
NOTOPODIAL PRECHAETAL LOBE. Present from parapodia 5 to parapodia 14 (
<figureCitation id="13442A55FFB2F527FB9BF953FB8CF944" box="[1058,1148,1719,1745]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3F</figureCitation>
); bluntly rounded, as long as median ligule in anterior parapodia, then reducing and tapering gradually to notoacicular process that disappears in about parapodia 32.
</paragraph>
<paragraph id="8BC036D0FFB2F527FF04F8DBFCBBF80A" blockId="15.[189,1399,1854,1951]" pageId="15" pageNumber="116">
MEDIAN LIGULE. Digitiform in anteriormost parapodia, bluntly rounded in anterior parapodia (
<figureCitation id="13442A55FFB2F527FAB7F8DAFA98F8CC" box="[1294,1384,1854,1881]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3F</figureCitation>
), conical in middle parapodia (
<figureCitation id="13442A55FFB2F527FDA6F886FD52F8E9" box="[543,674,1890,1916]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3GH</figureCitation>
), bluntly conical in following parapodia (
<figureCitation id="13442A55FFB2F527FB2EF886FB1AF8E9" box="[1175,1258,1890,1916]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="15" pageNumber="116">Fig. 3I</figureCitation>
), becoming narrower from middle parapodia towards posterior end.
</paragraph>
<caption id="DF006658FFADF538FF04FA19FBEDF874" ID-DOI="http://doi.org/10.5281/zenodo.5121740" ID-Zenodo-Dep="5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="16" pageNumber="117" startId="16.[189,232,1533,1559]" targetBox="[319,1268,265,1493]" targetPageId="16">
<paragraph id="8BC036D0FFADF538FF04FA19FBEDF874" blockId="16.[187,1399,1533,2017]" pageId="16" pageNumber="117">
<emphasis id="B90BEAC2FFADF538FF04FA19FEE2F982" bold="true" box="[189,274,1533,1559]" pageId="16" pageNumber="117">Fig. 3.</emphasis>
<taxonomicName id="4C7F4D53FFADF538FEA6FA19FD46F98D" authority="Kinberg, 1865" authorityName="Kinberg" authorityYear="1865" box="[287,694,1533,1560]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="117" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFADF538FEA6FA19FE05F982" box="[287,501,1533,1559]" italics="true" pageId="16" pageNumber="117">Neanthes helenae</emphasis>
<bibRefCitation id="EFEE4B21FFADF538FDBAFA19FD46F98D" author="Kinberg J. G. H." box="[515,694,1533,1560]" pageId="16" pageNumber="117" pagination="167 - 179" refId="ref21148" refString="Kinberg J. G. H. 1865. Annulata nova. Nereidum dispositio nova. Ofversigt af Kongelige Vetenskaps- Akademiens Forhandlingar 22: 167 - 179. Available from https: // www. biodiversitylibrary. org / page / 32339443 [accessed 1 Jul. 2021]." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
</taxonomicName>
.
<emphasis id="B90BEAC2FFADF538FD7EFA19FCBDF982" bold="true" box="[711,845,1533,1559]" pageId="16" pageNumber="117">AB, EQ</emphasis>
. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous.
<emphasis id="B90BEAC2FFADF538FD34F9C4FD35F9AF" bold="true" box="[653,709,1568,1594]" pageId="16" pageNumber="117">CD</emphasis>
. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous.
<emphasis id="B90BEAC2FFADF538FDBAF9A7FDE6F9C8" bold="true" box="[515,534,1603,1629]" pageId="16" pageNumber="117">A</emphasis>
. Entire body in dorsal view.
<emphasis id="B90BEAC2FFADF538FCC0F9A0FC7AF9CB" bold="true" box="[889,906,1604,1630]" pageId="16" pageNumber="117">B</emphasis>
. Anterior end in dorsal view.
<emphasis id="B90BEAC2FFADF538FB4EF9A7FAFAF9C8" bold="true" box="[1271,1290,1603,1629]" pageId="16" pageNumber="117">C</emphasis>
. Everted pharynx in dorsal view.
<emphasis id="B90BEAC2FFADF538FE67F983FE02F914" bold="true" box="[478,498,1639,1665]" pageId="16" pageNumber="117">D</emphasis>
. Everted pharynx in ventral view.
<emphasis id="B90BEAC2FFADF538FC28F983FC53F914" bold="true" box="[913,931,1639,1665]" pageId="16" pageNumber="117">E</emphasis>
. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.
<emphasis id="B90BEAC2FFADF538FD37F96EFD50F931" bold="true" box="[654,672,1674,1700]" pageId="16" pageNumber="117">F</emphasis>
. Right parapodium from chaetiger 8 in anterior view.
<emphasis id="B90BEAC2FFADF538FAA9F96EFAD7F931" bold="true" box="[1296,1319,1674,1700]" pageId="16" pageNumber="117">G</emphasis>
. Right parapodium from chaetiger 27 in anterior view.
<emphasis id="B90BEAC2FFADF538FD50F949FCF0F952" bold="true" box="[745,768,1709,1735]" pageId="16" pageNumber="117">H</emphasis>
. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing.
<emphasis id="B90BEAC2FFADF538FDBAF934FDFEF97F" bold="true" box="[515,526,1744,1770]" pageId="16" pageNumber="117">I</emphasis>
. Right parapodium from chaetiger 60 in anterior view.
<emphasis id="B90BEAC2FFADF538FB03F934FB39F97F" bold="true" box="[1210,1225,1744,1770]" pageId="16" pageNumber="117">J</emphasis>
. Homogomph spiniger from notochaetae (chaetiger 27).
<emphasis id="B90BEAC2FFADF538FD7BF910FD29F89B" bold="true" box="[706,729,1780,1806]" pageId="16" pageNumber="117">K</emphasis>
. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1).
<emphasis id="B90BEAC2FFADF538FEDBF8F3FE85F8A4" bold="true" box="[354,373,1815,1841]" pageId="16" pageNumber="117">L</emphasis>
. Heterogomph falciger from subacicular neurochaetae (chaetiger 1).
<emphasis id="B90BEAC2FFADF538FB1AF8F3FB4EF8A4" bold="true" box="[1187,1214,1815,1841]" pageId="16" pageNumber="117">M</emphasis>
. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).
<emphasis id="B90BEAC2FFADF538FC39F8DEFC65F8C1" bold="true" box="[896,917,1850,1876]" pageId="16" pageNumber="117">N</emphasis>
. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13).
<emphasis id="B90BEAC2FFADF538FF04F864FF3FF80F" bold="true" box="[189,207,1920,1946]" pageId="16" pageNumber="117">P</emphasis>
. Heterogomph falciger from subacicular neurochaetae (chaetiger 60).
<emphasis id="B90BEAC2FFADF538FBBDF864FBEBF80F" bold="true" box="[1028,1051,1920,1946]" pageId="16" pageNumber="117">Q</emphasis>
. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm.
</paragraph>
</caption>
<paragraph id="8BC036D0FFACF539FF04FEEDFB8FFED0" blockId="17.[189,1399,264,326]" pageId="17" pageNumber="118">
NEUROACICULAR LIGULE. Smaller than ventral ligule in anteriormost parapodia (
<figureCitation id="13442A55FFACF539FBD0FEECFB35FEB7" box="[1129,1221,264,290]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3E</figureCitation>
), as long as in following parapodia (
<figureCitation id="13442A55FFACF539FE04FECFFDDEFED0" box="[445,558,299,325]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3FI</figureCitation>
), 1.52 times as wide as ventral ligule throughout.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FE89FD58FE3F" blockId="17.[189,1399,365,426]" pageId="17" pageNumber="118">NEUROPODIAL INFERIOR LOBE. Slightly developed and longer than neuroacicular ligule in first four parapodia, absent in following chaetigers.</paragraph>
<paragraph id="8BC036D0FFACF539FF04FE36FC30FD9A" blockId="17.[189,1399,465,527]" pageId="17" pageNumber="118">
NEUROPODIAL POSTCHAETAL LOBE. Digitiform, longer than neuroacicular ligule in first 5 parapodia (
<figureCitation id="13442A55FFACF539FF7FFE10FED0FD9A" box="[198,288,500,527]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3E</figureCitation>
), then reducing gradually to disappear in parapodium 12.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FDD2FC87FD2F" blockId="17.[189,1400,566,698]" pageId="17" pageNumber="118">
VENTRAL LIGULE. Well developed throughout; digitiform, thick in anteriormost parapodia (
<figureCitation id="13442A55FFACF539FAB0FDD2FA98FDC5" box="[1289,1384,566,592]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3E</figureCitation>
), bluntly rounded, thick in anterior parapodia (
<figureCitation id="13442A55FFACF539FD6DFDBDFCDDFDE6" box="[724,813,601,627]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3F</figureCitation>
), digitiform in following chaetigers; smaller than median ligule throughout, more distinct in middle and posterior parapodia (
<figureCitation id="13442A55FFACF539FBECFD98FB20FD03" box="[1109,1232,636,662]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3HI</figureCitation>
); shorter than distal region of dorsal ligule in parapodia 1 and 2 (
<figureCitation id="13442A55FFACF539FCB6FD7BFC9AFD2F" box="[783,874,671,698]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FD05FCA9FC8A" blockId="17.[189,1399,737,799]" pageId="17" pageNumber="118">
VENTRAL CIRRI. Cirriform, slender; as long as ventral ligule in anteriormost parapodia (
<figureCitation id="13442A55FFACF539FB75FD05FAD8FD6E" box="[1228,1320,737,763]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3E</figureCitation>
), twothirds as long as that in following parapodia (
<figureCitation id="13442A55FFACF539FD68FCE0FCBCFC8A" box="[721,844,772,799]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3F, H</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FCA2FD44FC33" blockId="17.[189,1399,838,935]" pageId="17" pageNumber="118">
ACICULAE. Colour mostly faded by long-term preservation. Notoaciculae absent in first two chaetigers (
<figureCitation id="13442A55FFACF539FF7FFC8DFED4FC16" box="[198,292,873,899]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3E</figureCitation>
). Neuroaciculae extending similarly to distal end of notoaciculae throughout, with proximal half 1.31.5 times as wide as notoaciculae.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FC2AFEC9FBBB" blockId="17.[189,1400,973,1070]" pageId="17" pageNumber="118">
NOTOCHAETAE. All homogomph spinigers (
<figureCitation id="13442A55FFACF539FD0BFC29FCF7FC7D" box="[690,775,973,1000]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3J</figureCitation>
); 1013 spinigers present in anterior parapodia, 69 spinigers in middle parapodia, 45 spinigers in posterior parapodia and 3 spinigers in posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FBB1FC69FB69" blockId="17.[188,1399,1109,1276]" pageId="17" pageNumber="118">
SUPRACICULAR NEUROCHAETAE. Consisting of homogomph spinigers and heterogomph falcigers (
<figureCitation id="13442A55FFACF539FAB7FBB1FF22FB06" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3K L</figureCitation>
), both present throughout; 45 spinigers present in anteriormost parapodia, 67 spinigers in anterior parapodia, 45 spinigers in middle parapodia, 12 spinigers in posterior and posteriormost parapodia; 2 falcigers present in anteriormost parapodia, 23 falcigers in anterior parapodia, 12 falcigers in middle parapodia, 1 falciger in posterior and posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FAC0FB2BFA32" blockId="17.[188,1399,1315,1447]" pageId="17" pageNumber="118">
SUBACICULAR NEUROCHAETAE. Consisting of heterogomph spinigers (
<figureCitation id="13442A55FFACF539FC76FAC7FBDFFAAB" box="[975,1071,1315,1342]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3M</figureCitation>
) and heterogomph falcigers (
<figureCitation id="13442A55FFACF539FF7CFAA3FEBCFAF4" box="[197,332,1351,1377]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3NP</figureCitation>
), both present throughout; 3 spinigers present throughout, sometimes 4 spinigers in middle parapodia; 8 falcigers present in anteriormost parapodia, 812 falcigers in anterior parapodia, 36 falcigers in middle parapodia, 13 falcigers in posterior and posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04FA2BFA99F8F9" blockId="17.[186,1401,1486,1901]" pageId="17" pageNumber="118">
BLADES. Both homogomph (
<figureCitation id="13442A55FFACF539FDB3FA2AFDADFA7C" box="[522,605,1486,1513]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3J</figureCitation>
) and heterogomph (
<figureCitation id="13442A55FFACF539FCF4FA2AFC40FA7C" box="[845,944,1486,1513]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3M</figureCitation>
) spinigers with blades of medium and long size (
<emphasis id="B90BEAC2FFACF539FE8EFA16FE92F999" box="[311,354,1522,1548]" italics="true" pageId="17" pageNumber="118">B/A</emphasis>
ratio 6.611.2), finely serrated towards toothed edge, evenly spaced, except homogomph spinigers from middle parapodia towards posterior end with coarse proximal teeth.Blades of heterogomph falcigers in anteriormost parapodia slender, convex, long (
<emphasis id="B90BEAC2FFACF539FCD4F9DDFC69F9C6" box="[877,921,1593,1619]" italics="true" pageId="17" pageNumber="118">B/A</emphasis>
ratio 2.22.7), terminal tooth blunt with inconspicuous tendon (
<figureCitation id="13442A55FFACF539FE75F9BFFDD9F9E0" box="[460,553,1627,1653]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3N</figureCitation>
), serrations present in about two-fifths (0.600.65) of total blade length; blades in following parapodia thicker, convex, short (
<emphasis id="B90BEAC2FFACF539FC85F99BFC98F90C" box="[828,872,1663,1689]" italics="true" pageId="17" pageNumber="118">B/A</emphasis>
ratio 0.91.6), terminal tooth incurved with distinct tendon in supracicular fascicle (
<figureCitation id="13442A55FFACF539FD1AF946FCD6F929" box="[675,806,1698,1724]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3KL</figureCitation>
), blunt with inconspicuous tendon in subacicular fascicle (
<figureCitation id="13442A55FFACF539FE92F921FE5DF94A" box="[299,429,1733,1759]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3OP</figureCitation>
), serrations present in about one-fifth to one-quarter (0.220.28) of total blade length. Shaft of supracicular falcigers thicker than subacicular ones in middle and posterior parapodia; camerated, with cavity divided sub-distally into two longitudinal partitions, supracicular rarely divided into three. Fused heterogomph falcigers present in supracicular fascicle, fusing from parapodia 23, simple in parapodia 35 to parapodia 65, separating gradually to non-fused in posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFACF539FF04F870FCA4F845" blockId="17.[188,1399,1939,2000]" pageId="17" pageNumber="118">
PYGIDIUM. Distinctly projected, bluntly rounded, wider than long (
<figureCitation id="13442A55FFACF539FC03F877FBE6F838" box="[954,1046,1939,1965]" captionStart="Fig" captionStartId="16.[189,232,1533,1559]" captionTargetBox="[319,1268,265,1493]" captionTargetId="figure-14@16.[319,1268,265,1493]" captionTargetPageId="16" captionText="Fig. 3. Neanthes helenae Kinberg, 1865. AB, EQ. Complete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. CD. Incomplete syntype (SMNH Type-457) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B. Anterior end in dorsal view. C. Everted pharynx in dorsal view. D. Everted pharynx in ventral view. E. Right parapodium from chaetiger 1 in anterior view with ventral cirri missing.F. Right parapodium from chaetiger 8 in anterior view. G. Right parapodium from chaetiger 27 in anterior view. H. Right parapodium from chaetiger 42 in anterior view with ventral cirri missing. I. Right parapodium from chaetiger 60 in anterior view. J. Homogomph spiniger from notochaetae (chaetiger 27). K. Heterogomph spiniger from subacicular neurochaetae (chaetiger 1). L. Heterogomph falciger from subacicular neurochaetae (chaetiger 1). M. Heterogomph, fused falciger from supracicular neurochaetae (chaetiger 42).N. Heterogomph falciger from supracicular neurochaetae (chaetiger 60). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 13). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 60). Q. Posterior end in dorsal view. AD, Q: stained with Shirlastain A. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AB = 1 mm; CD, Q = 0.5 mm; EI = 0.1 mm; JP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121740" httpUri="https://zenodo.org/record/5121740/files/figure.png" pageId="17" pageNumber="118">Fig. 3Q</figureCitation>
), with anal cirri as long as last 11 chaetigers; cirrophores of anal cirri barely developed.
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FEECFBC9FEB7" blockId="18.[189,1081,264,291]" box="[189,1081,264,291]" pageId="18" pageNumber="119">
<heading id="D08881BCFFAFF53AFF04FEECFBC9FEB7" bold="true" box="[189,1081,264,291]" fontSize="11" level="3" pageId="18" pageNumber="119" reason="3">
<emphasis id="B90BEAC2FFAFF53AFF04FEECFBC9FEB7" bold="true" box="[189,1081,264,291]" pageId="18" pageNumber="119">
Description of
<typeStatus id="54C48872FFAFF53AFECCFEEDFE11FEB6" box="[373,481,265,291]" pageId="18" pageNumber="119" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D79FD59FFAFF53AFDA6FEEDFD20FEB6" box="[543,720,265,291]" pageId="18" pageNumber="119" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
of
<taxonomicName id="4C7F4D53FFAFF53AFD4EFEECFBC9FEB7" authorityName="Day" authorityYear="1949" box="[759,1081,264,291]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="18" pageNumber="119" phylum="Annelida" rank="species" species="nanciae" subGenus="Neanthes">
<emphasis id="B90BEAC2FFAFF53AFD4EFEECFCB5FEB7" bold="true" box="[759,837,264,290]" italics="true" pageId="18" pageNumber="119">Nereis</emphasis>
(
<emphasis id="B90BEAC2FFAFF53AFCEFFEECFC38FEB6" bold="true" box="[854,968,264,291]" italics="true" pageId="18" pageNumber="119">Neanthes</emphasis>
)
<emphasis id="B90BEAC2FFAFF53AFC60FEECFBC9FEB7" bold="true" box="[985,1081,264,290]" italics="true" pageId="18" pageNumber="119">nanciae</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FED3FB7CFE2E" blockId="18.[187,1399,311,443]" pageId="18" pageNumber="119">
COLOUR AND MEASUREMENTS. All complete atokous (
<specimenCount id="9D79FD59FFAFF53AFCB3FED3FC0BFEC7" box="[778,1019,311,338]" pageId="18" pageNumber="119" type="female" typeStatus="paratype">one paratype female</specimenCount>
), in good condition,
<quantity id="4C879B35FFAFF53AFB5DFED3FA87FEC7" box="[1252,1399,311,338]" metricMagnitude="-2" metricUnit="m" metricValue="2.125" metricValueMax="2.25" metricValueMin="2.0" pageId="18" pageNumber="119" unit="mm" value="21.25" valueMax="22.5" valueMin="20.0">2022.5 mm</quantity>
TL,
<quantity id="4C879B35FFAFF53AFF55FEBEFE88FEE0" box="[236,376,346,373]" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.6" metricValueMin="6.3" pageId="18" pageNumber="119" unit="mm" value="6.949999999999999" valueMax="7.6" valueMin="6.3">6.37.6 mm</quantity>
L15, 1
<quantity id="4C879B35FFAFF53AFE6CFEBEFDC0FEE0" box="[469,560,346,373]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="119" unit="mm" value="1.6">1.6 mm</quantity>
W15, with 6064 chaetigers. Body brownish, distal half of prostomium, apodous anterior segment and few anterior chaetigers with reddish-brown pigmentation (
<figureCitation id="13442A55FFAFF53AFB5CFE99FA98FE0D" box="[1253,1384,381,408]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4AB</figureCitation>
), smallest specimen forming faint X-shaped pattern on dorsum of anterior segments.
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FE07FD3BFDF3" blockId="18.[189,1400,482,614]" pageId="18" pageNumber="119">
PROSTOMIUM. Campanulate, as long as wide or slightly longer than wide (
<figureCitation id="13442A55FFAFF53AFB98FE06FB8FFE69" box="[1057,1151,482,508]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4B</figureCitation>
); anterior end broad, distally complete; anterolateral gap beside palpophore narrow, as wide as antennal diameter; dorsal groove distinct, shallow, running mid-subdistally. Nuchal organs deeply embedded, broad, 1.72 times as wide as diameter of posterior pair of eyes.
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FD6AFD65FD78" blockId="18.[189,1398,653,750]" pageId="18" pageNumber="119">
PALPOPHORES. Sub-conical, as wide as long to slightly wider as long (
<figureCitation id="13442A55FFAFF53AFC5DFD69FBB0FD32" box="[996,1088,653,679]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4B</figureCitation>
), subequal to entire length of prostomium; with distinct sub-distal transverse groove. Palpostyles ovoid, thick, with diameter as wide as two-fifths to half of palpophore.
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FCF1FD81FCE0" blockId="18.[189,1400,789,886]" pageId="18" pageNumber="119">
ANTENNAE. Tapered, thick, long, extending forwards beyond tip of palpophore and posteriorly to half to two-fifths length of prostomium; antennae separated, with gap subequal to three-quarters as wide as basal diameter of antennae (
<figureCitation id="13442A55FFAFF53AFDBEFCBFFD93FCE0" box="[519,611,859,885]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FC79FD57FBF2" blockId="18.[189,1399,925,1128]" pageId="18" pageNumber="119">
EYES. Paired eyes black, arranged in a trapezoid form; gap between both pairs one-quarter to threequarters as wide as diameter of posterior pair of eyes (
<figureCitation id="13442A55FFAFF53AFCFEFC24FC55FC4F" box="[839,933,960,986]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4B</figureCitation>
); anterior pair of eyes oval to rounded, 1.51.7 times as wide as basal diameter of antennae, gap between both eyes 33.5 times as wide as diameter of eyes, with lens barely distinct, translucent, covering 1020% of eye; posterior pair of eyes rounded, 1.21.5 times as wide as basal diameter of antennae, with lens distinct, translucent, placed in middle of eye and covering 1525% of it.
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FB6AFE6DFB7A" blockId="18.[188,1398,1166,1263]" pageId="18" pageNumber="119">
APODOUS ANTERIOR SEGMENT. Segment 3.54 times wider than long, 1.5 times as long as chaetiger 1, with rounded anterior margin and distinct ventrolateral projections (
<figureCitation id="13442A55FFAFF53AFC01FB55FBCEFB59" box="[952,1086,1201,1228]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4CD</figureCitation>
); dorsum without marked transverse wrinkle.
</paragraph>
<paragraph id="8BC036D0FFAFF53AFF04FAF3FC86FA75" blockId="18.[187,1399,1302,1505]" pageId="18" pageNumber="119">
TENTACULAR CIRRI. Thick, smooth (
<figureCitation id="13442A55FFAFF53AFDEAFAF2FD40FAA5" box="[595,688,1302,1328]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4B</figureCitation>
); postero-dorsal cirri extending posteriorly to chaetiger 78; antero-dorsal cirri extending posteriorly to chaetiger 3; postero-ventral cirri extended over opposite side of prostomium; antero-ventral cirri subequal or slightly longer than postero-ventral cirri and extending beyond 1.52 times length of palpophore; dorsal cirrophores cylindrical, ventral cirrophores ring-shaped, postero-dorsal cirrophores 1.5 times as long as antero-dorsal cirrophores, antero-ventral cirrophores subequal or slightly wider than postero-ventral cirrophores.
</paragraph>
<paragraph id="8BC036D0FFAFF53BFF04F9ECFAB5FEFC" blockId="18.[186,1401,1543,1992]" lastBlockId="19.[189,1399,265,362]" lastPageId="19" lastPageNumber="120" pageId="18" pageNumber="119">
PHARYNX. Non-everted. Jaws with distal quarter brownish, remaining yellow amber, with 910 welldeveloped and sharp denticles (
<figureCitation id="13442A55FFAFF53AFD9AF9CFFD8CF9D0" box="[547,636,1579,1605]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4E</figureCitation>
); pulp cavity with two canals (
<figureCitation id="13442A55FFAFF53AFC6CF9CFFBDCF9D0" box="[981,1068,1579,1605]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4F</figureCitation>
). Reddish-brown paragnaths on maxillary and oral rings (
<figureCitation id="13442A55FFAFF53AFDB1F9AAFD7AF9FD" box="[520,650,1614,1640]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4GH</figureCitation>
), consisting of cones, p-bars, and merged paragnaths; plate-like basements absent. Area I: 12, cones; areas IIa: 2027 and IIb: 1927, two slightly regular rows of uneven cones in eyebrow-shaped patch, subdistal cones larger, fang-shaped in outer-most row (
<figureCitation id="13442A55FFAFF53AFAB2F970FA99F93B" box="[1291,1385,1684,1710]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4G</figureCitation>
); area III: 1828, two to four irregular rows of uneven cones in sub-oval or sub-rectangular patch, with two to four distinct laterally-isolated cones, distal cones smaller; areas IVa: 2445 and IVb: 2648, L-shaped patch (
<figureCitation id="13442A55FFAFF53AFE3DF91AFE11F88D" box="[388,481,1790,1816]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4G</figureCitation>
) with proximal half consisting of two regular rows and distal half with three or four slightly regular rows of uneven cones and four to six long and slender (34 times longer than wide) merged paragnaths located near jaw; area V: 1 (1), coarse conical paragnath placed slightly behind level of paragnaths on area VI (
<figureCitation id="13442A55FFAFF53AFD90F883FD77F817" box="[553,647,1895,1922]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4G</figureCitation>
); areas VIa: 58 and VIb: 57, oblique transverse row of uneven p-bars becoming shorter outwards (four to seven in VIa, four to six in VIb;
<figureCitation id="13442A55FFAFF53AFBEEF86FFB45F830" box="[1111,1205,1931,1957]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4H</figureCitation>
) and one single outermost cone (
<figureCitation id="13442A55FFAFF53AFE3EF84AFE19F85D" box="[391,489,1966,1992]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="18" pageNumber="119">Fig. 4H</figureCitation>
); areas VIIVIII: 2833, two bands of cones ventrally well separated, with anterior band consisting of two transversely aligned rows (furrow and ridge rows with one coarse cone on each region), and posterior band with two transverse slightly displaced rows (
<figureCitation id="13442A55FFAEF53BFBD3FEC8FB39FED3" box="[1130,1225,300,326]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4G</figureCitation>
). Areas VIV VI ridge pattern, π-shaped. Gap between area VI and areas VIIVIII narrow, as wide as palpostyle.
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FE75FD96FE3E" blockId="19.[189,614,401,427]" box="[189,614,401,427]" pageId="19" pageNumber="120">PAIRED OESOPHAGEAL CAECA. Present.</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FE37FDCAFDC3" blockId="19.[188,1399,466,598]" pageId="19" pageNumber="120">PARAPODIA. Without glandular, dorsal patches. Notopodia consisting of dorsal cirrus, dorsal ligule (distal and proximal), notopodial prechaetal lobe, and median ligule in biramous parapodia. Neuropodia consisting of neuroacicular ligule with inferior and postchaetal lobes, ventral ligule, and ventral cirrus; superior lobe absent throughout.</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FD9AFBF8FCB1" blockId="19.[189,1399,637,804]" pageId="19" pageNumber="120">
DORSAL CIRRI. Cirriform, long, extending markedly beyond distal region of dorsal ligule throughout (
<figureCitation id="13442A55FFAEF53BFF7FFD44FEB5FD2E" box="[198,325,672,699]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4IM</figureCitation>
); dorsal cirri 33.5 times as long as proximal region of dorsal ligule in anteriormost and anterior parapodia (
<figureCitation id="13442A55FFAEF53BFE1BFD20FDFFFD4B" box="[418,527,708,734]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4IJ</figureCitation>
), 22.5 times as long as that in following parapodia (
<figureCitation id="13442A55FFAEF53BFBC0FD20FAF1FD4B" box="[1145,1281,708,734]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4KM</figureCitation>
); attached basally to dorsal ligule in anteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFCA0FD03FC9EFC94" box="[793,878,743,769]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I</figureCitation>
), medially in anterior, middle and posterior parapodia (
<figureCitation id="13442A55FFAEF53BFEFAFCEEFE48FCB1" box="[323,440,778,804]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4JL</figureCitation>
), two-thirds in posteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFC2EFCEEFC0BFCB1" box="[919,1019,778,804]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4M</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FCA8FAA9FB37" blockId="19.[188,1401,843,1187]" pageId="19" pageNumber="120">
DORSAL LIGULE. Proximal region even towards posterior end, except slightly humped from middle parapodia towards posterior end; shorter than distal region of dorsal ligule in anteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFF71FC76FEEDFC39" box="[200,285,914,940]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I</figureCitation>
), as long as that in anterior, middle and posterior parapodia (
<figureCitation id="13442A55FFAEF53BFC45FC76FB83FC39" box="[1020,1139,914,940]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4JL</figureCitation>
), 1.5 times as long as that in posteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFDF8FC51FD56FC5A" box="[577,678,949,975]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4M</figureCitation>
); two irregular glandular patches covering partially proximal region of dorsal ligule in anteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFC94FC3CFC73FC67" box="[813,899,984,1010]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I</figureCitation>
) and entirely covering that in following parapodia (
<figureCitation id="13442A55FFAEF53BFEFBFC1FFE32FB83" box="[322,450,1019,1046]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4JM</figureCitation>
). Distal region well developed, becoming slightly longer towards posterior end; digitiform in anteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFDCFFBFBFD38FBAC" box="[630,712,1055,1081]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I</figureCitation>
), bluntly rounded in anterior parapodia (
<figureCitation id="13442A55FFAEF53BFB27FBFBFB02FBAC" box="[1182,1266,1055,1081]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4J</figureCitation>
), conical to bluntly conical in following parapodia (
<figureCitation id="13442A55FFAEF53BFD37FBA6FCE6FBC9" box="[654,790,1090,1116]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4KM</figureCitation>
); as long as or slightly shorter than median ligule in anterior and middle parapodia (
<figureCitation id="13442A55FFAEF53BFD8AFB81FD59FBEA" box="[563,681,1125,1151]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4JL</figureCitation>
), longer than that in following parapodia; projecting beyond notoacicula throughout; one irregular glandular patch covering entirely distal region of dorsal ligule.
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FB2EFCF5FABF" blockId="19.[189,1399,1226,1323]" pageId="19" pageNumber="120">
NOTOPODIAL PRECHAETAL LOBE. Present from parapodia 5 to parapodia 1416 (
<figureCitation id="13442A55FFAEF53BFBF0FB2EFB6EFB71" box="[1097,1182,1226,1252]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4J</figureCitation>
); bluntly rounded, as long as median ligule in anterior parapodia, then reducing and tapering gradually to notoacicular process that disappears in about parapodia 3437.
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FAB6FDCEFA27" blockId="19.[189,1400,1361,1458]" pageId="19" pageNumber="120">
MEDIAN LIGULE. Digitiform in anteriormost parapodia, bluntly rounded in anterior parapodia (
<figureCitation id="13442A55FFAEF53BFAAAFAB5FA99FAF9" box="[1299,1385,1361,1388]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4J</figureCitation>
), digitiform to bluntly conical in following parapodia (
<figureCitation id="13442A55FFAEF53BFCF3FA91FC26FA1A" box="[842,982,1397,1423]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4KM</figureCitation>
), becoming narrower from middle parapodia towards posterior end.
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04FA3EFEB6F9AF" blockId="19.[188,1399,1497,1594]" pageId="19" pageNumber="120">
NEUROACICULAR LIGULE. Smaller than ventral ligule in anteriormost and posteriormost parapodia (
<figureCitation id="13442A55FFAEF53BFA98FA3DFF29F982" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I, M</figureCitation>
), as long as in remaining parapodia (
<figureCitation id="13442A55FFAEF53BFD1AFA19FCECF982" box="[675,796,1533,1559]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4JL</figureCitation>
), parapodia 1.52 times as wide as ventral ligule throughout.
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04F986FD58F90B" blockId="19.[189,1398,1633,1694]" pageId="19" pageNumber="120">NEUROPODIAL INFERIOR LOBE. Slightly developed and longer than neuroacicular ligule in first four or five parapodia, absent in following chaetigers.</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04F922FCABF896" blockId="19.[189,1398,1734,1796]" pageId="19" pageNumber="120">
NEUROPODIAL POSTCHAETAL LOBE. Digitiform, longer than neuroacicular ligule in first 5 parapodia (
<figureCitation id="13442A55FFAEF53BFAA1F922FA98F975" box="[1304,1384,1734,1760]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I</figureCitation>
), then reducing gradually to disappear in parapodia 1012.
</paragraph>
<paragraph id="8BC036D0FFAEF53BFF04F8CFFE9EF83B" blockId="19.[189,1399,1834,1966]" pageId="19" pageNumber="120">
VENTRAL LIGULE. Well developed throughout; digitiform, thick in all parapodia, except bluntly rounded, thick in anterior parapodia (
<figureCitation id="13442A55FFAEF53BFDBCF8AAFDAAF8FD" box="[517,602,1870,1896]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4J</figureCitation>
); smaller than median ligule throughout, more distinct from middle parapodia towards posterior end (
<figureCitation id="13442A55FFAEF53BFDF0F895FD3FF81E" box="[585,719,1905,1931]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4LM</figureCitation>
); shorter than distal region of dorsal ligule in parapodia 1 and 2 (
<figureCitation id="13442A55FFAEF53BFEB7F870FE91F83B" box="[270,353,1940,1966]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="19" pageNumber="120">Fig. 4I</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FEEDFBD4FED3" blockId="20.[189,1399,265,327]" pageId="20" pageNumber="121">
VENTRAL CIRRI. Cirriform, slender; as long as ventral ligule in anteriormost parapodia (
<figureCitation id="13442A55FFA9F53CFB6AFEEDFAD9FEB6" box="[1235,1321,265,291]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4I</figureCitation>
), twothirds to three-quarters as long as that in following parapodia (
<figureCitation id="13442A55FFA9F53CFC20FEC8FBE8FED3" box="[921,1048,300,326]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4JM</figureCitation>
).
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FE8AFEA9FE64" blockId="20.[189,1399,366,497]" pageId="20" pageNumber="121">
ACICULAE. Mostly black with basal end uncoloured in anterior chaetigers, one-half paler in posterior chaetigers (
<figureCitation id="13442A55FFA9F53CFEF1FE75FE59FE3E" box="[328,425,401,427]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4L</figureCitation>
). Notoaciculae absent in first two chaetigers (
<figureCitation id="13442A55FFA9F53CFC64FE75FBC4FE3E" box="[989,1076,401,427]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4I</figureCitation>
). Neuroaciculae extending similarly to distal end of notoaciculae throughout, with proximal half 1.31.5 times as wide as notoaciculae.
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FDFDFA88FDC3" blockId="20.[189,1400,537,599]" pageId="20" pageNumber="121">NOTOCHAETAE. All homogomph spinigers; 1114 spinigers present in anterior parapodia, 79 spinigers in middle parapodia, 45 spinigers in posterior parapodia and 23 spinigers in posteriormost parapodia.</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FD99FD51FCB1" blockId="20.[188,1400,637,804]" pageId="20" pageNumber="121">
SUPRACICULAR NEUROCHAETAE. Consisting of homogomph spinigers and heterogomph falcigers (
<figureCitation id="13442A55FFA9F53CFAB5FD99FA99FD02" box="[1292,1385,637,663]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4N</figureCitation>
), both present throughout; 35 spinigers present in anteriormost parapodia, 57 spinigers in anterior parapodia, 35 spinigers in middle parapodia, 12 spinigers in posterior and posteriormost parapodia; 2 falcigers present in anteriormost parapodia, 13 falcigers in anterior and middle parapodia, 1 falciger in posterior and posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FCA8FA87FC5A" blockId="20.[189,1399,843,975]" pageId="20" pageNumber="121">
SUBACICULAR NEUROCHAETAE. Consisting of heterogomph spinigers and heterogomph falcigers (
<figureCitation id="13442A55FFA9F53CFAB7FCAFFF20FC1C" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4O P</figureCitation>
), both present throughout; 23 spinigers present throughout, except 1 spiniger in posteriormost parapodia; 56 falcigers present in anteriormost parapodia, 710 falcigers in anterior parapodia, 56 falcigers in middle parapodia, 4 falcigers in posterior parapodia, 23 falcigers in posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FC13FC19FA01" blockId="20.[187,1400,1014,1429]" pageId="20" pageNumber="121">
BLADES. Both homogomph and heterogomph spinigers with blades of medium and long size (
<emphasis id="B90BEAC2FFA9F53CFF7EFBFEFF02FBA1" box="[199,242,1050,1076]" italics="true" pageId="20" pageNumber="121">B/A</emphasis>
ratio 5.18.3), finely serrated towards toothed edge, evenly spaced, except homogomph spinigers from middle parapodia towards posterior end with coarse proximal teeth. Blades of heterogomph falcigers in anteriormost parapodia slender, convex, long (
<emphasis id="B90BEAC2FFA9F53CFCD4FB85FC69FBEE" box="[877,921,1121,1147]" italics="true" pageId="20" pageNumber="121">B/A</emphasis>
ratio 2.33.4), terminal tooth blunt with inconspicuous tendon (
<figureCitation id="13442A55FFA9F53CFE75FB67FDDAFB08" box="[460,554,1155,1181]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4O</figureCitation>
), serrations present in about half to two-thirds (0.520.66) of total blade length; blades in following parapodia thicker, convex, short (
<emphasis id="B90BEAC2FFA9F53CFCC5FB43FC57FB54" box="[892,935,1191,1217]" italics="true" pageId="20" pageNumber="121">B/A</emphasis>
ratio 0.651.3), terminal tooth incurved with distinct tendon in supracicular fascicle (
<figureCitation id="13442A55FFA9F53CFD6AFB2EFCDFFB71" box="[723,815,1226,1252]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4P</figureCitation>
), blunt with inconspicuous tendon in subacicular fascicle, serrations present in about one-sixth to one-third (0.160.38) of total blade length. Shaft of supracicular falcigers thicker than subacicular ones in middle and posterior parapodia; camerated, with cavity divided sub-distally into two longitudinal partitions. Fused heterogomph falcigers present in supracicular fascicle (
<figureCitation id="13442A55FFA9F53CFE79FAB2FDD0FAE4" box="[448,544,1366,1393]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4N</figureCitation>
), fusing from parapodia 2326, simple in parapodia 3435 to parapodia 5965, separating gradually to non-fused in posteriormost parapodia.
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04FA58FC71FA6D" blockId="20.[188,1399,1467,1528]" pageId="20" pageNumber="121">
PYGIDIUM. Distinctly projected, bluntly rounded, as wide as long (
<figureCitation id="13442A55FFA9F53CFC01FA5FFBE5FA40" box="[952,1045,1467,1493]" captionStart="Fig" captionStartId="21.[189,232,1463,1489]" captionTargetBox="[346,1241,265,1423]" captionTargetId="figure-11@21.[346,1241,265,1423]" captionTargetPageId="21" captionText="Fig. 4. Neanthes helenae Kinberg, 1865. AC, IP. Holotype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. A. Entire body in dorsal view. B.Anterior end in dorsal view.C. Anterior region in ventral view, arrow indicates ventrolateral projection. I. Left parapodium from chaetiger 1 in posterior view. J. Left parapodium from chaetiger 7 in posterior view. K. Right parapodium from chaetiger 21 in anterior view. L. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule. M. Right parapodium from chaetiger 60 in anterior view. N. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30). O. Heterogomph falciger from subacicular neurochaetae (chaetiger 7). P. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). — DH, Q. Paratype of Nereis (Neanthes) nanciae Day, 1949 (NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous. D. Anterior region in lateral view, arrow indicates ventrolateral projection. E. Left jaw in dorsal and ventral views, respectively. F. Inner canals of jaws in ventral view. G. Non-everted pharynx in ventral view. H. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars. Q. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="20" pageNumber="121">Fig. 4Q</figureCitation>
), with anal cirri as long as last 1112 chaetigers; cirrophores of anal cirri barely developed.
</paragraph>
</subSubSection>
<subSubSection id="C365655BFFA9F53EFF04F9FBFCA1FA53" lastPageId="22" lastPageNumber="123" pageId="20" pageNumber="121" type="discussion">
<paragraph id="8BC036D0FFA9F53CFF04F9FBFEDFF9AC" blockId="20.[189,303,1567,1593]" box="[189,303,1567,1593]" pageId="20" pageNumber="121">
<heading id="D08881BCFFA9F53CFF04F9FBFEDFF9AC" bold="true" box="[189,303,1567,1593]" fontSize="11" level="3" pageId="20" pageNumber="121" reason="3">
<emphasis id="B90BEAC2FFA9F53CFF04F9FBFEDFF9AC" bold="true" box="[189,303,1567,1593]" pageId="20" pageNumber="121">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFA9F53CFF04F9A8FD46F829" blockId="20.[188,1400,1612,1980]" pageId="20" pageNumber="121">
<taxonomicName id="4C7F4D53FFA9F53CFF04F9A8FE7CF9F3" authorityName="Kinberg" authorityYear="1865" box="[189,396,1612,1638]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFA9F53CFF04F9A8FE7CF9F3" box="[189,396,1612,1638]" italics="true" pageId="20" pageNumber="121">Neanthes helenae</emphasis>
</taxonomicName>
was not compared in the original description by
<bibRefCitation id="EFEE4B21FFA9F53CFC7BF9A8FB89F9F2" author="Kinberg J. G. H." box="[962,1145,1612,1639]" pageId="20" pageNumber="121" pagination="167 - 179" refId="ref21148" refString="Kinberg J. G. H. 1865. Annulata nova. Nereidum dispositio nova. Ofversigt af Kongelige Vetenskaps- Akademiens Forhandlingar 22: 167 - 179. Available from https: // www. biodiversitylibrary. org / page / 32339443 [accessed 1 Jul. 2021]." type="journal article" year="1865">Kinberg (1865)</bibRefCitation>
, although he placed it within a group formed by two other species,
<taxonomicName id="4C7F4D53FFA9F53CFD74F98BFCC8F91D" box="[717,824,1646,1672]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="rigida">
<emphasis id="B90BEAC2FFA9F53CFD74F98BFCC8F91D" box="[717,824,1646,1672]" italics="true" pageId="20" pageNumber="121">N. rigida</emphasis>
</taxonomicName>
(Grube &amp; Ørsted in Grube, 1858) and
<taxonomicName id="4C7F4D53FFA9F53CFABAF98BFA86F91D" box="[1283,1398,1646,1672]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="vallata">
<emphasis id="B90BEAC2FFA9F53CFABAF98BFA86F91D" box="[1283,1398,1646,1672]" italics="true" pageId="20" pageNumber="121">N. vallata</emphasis>
</taxonomicName>
(Grube &amp; Kröyer in Grube, 1858), currently belonging to
<taxonomicName id="4C7F4D53FFA9F53CFC31F975FC24F93F" authorityName="Linnaeus" authorityYear="1758" box="[904,980,1681,1706]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="genus">
<emphasis id="B90BEAC2FFA9F53CFC31F975FC24F93F" box="[904,980,1681,1706]" italics="true" pageId="20" pageNumber="121">Nereis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFA9F53CFBADF975FE34F958" authority="(Salazar-Vallejo &amp; Eibye-Jacobsen 2012)" baseAuthorityName="Salazar-Vallejo &amp; Eibye-Jacobsen" baseAuthorityYear="2012" class="Polychaeta" family="Nereididae" genus="Perinereis" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="genus">
<emphasis id="B90BEAC2FFA9F53CFBADF975FB7DF93F" box="[1044,1165,1681,1706]" italics="true" pageId="20" pageNumber="121">Perinereis</emphasis>
(
<bibRefCitation id="EFEE4B21FFA9F53CFB19F974FE4CF958" author="Salazar-Vallejo S. I. &amp; Eibye-Jacobsen D." pageId="20" pageNumber="121" pagination="1391 - 1402" refId="ref21951" refString="Salazar-Vallejo S. I. &amp; Eibye-Jacobsen D. 2012. Annulata orstediana: publication dates, composition and annotated taxonomic list, with some comments on Hemipodus (Polychaeta: Glyceridae). Revista de Biologia Tropical 60 (3): 1391 - 1402. https: // doi. org / 10.15517 / rbt. v 60 i 3.1815" type="journal article" year="2012">Salazar-Vallejo &amp; Eibye-Jacobsen 2012</bibRefCitation>
)
</taxonomicName>
, respectively. Among all species of
<taxonomicName id="4C7F4D53FFA9F53CFCD6F957FC2BF958" authorityName="Kinberg" authorityYear="1865" box="[879,987,1715,1741]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="genus">
<emphasis id="B90BEAC2FFA9F53CFCD6F957FC2BF958" box="[879,987,1715,1741]" italics="true" pageId="20" pageNumber="121">Neanthes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFA9F53CFC52F957FB99F958" authorityName="Kinberg" authorityYear="1865" box="[1003,1129,1715,1741]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFA9F53CFC52F957FB99F958" box="[1003,1129,1715,1741]" italics="true" pageId="20" pageNumber="121">N. helenae</emphasis>
</taxonomicName>
from St Helena Island resembles
<taxonomicName id="4C7F4D53FFA9F53CFE85F932FD4BF97A" authority="Wu, Sun &amp; Yang, 1981" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[316,699,1749,1775]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFA9F53CFE85F932FE69F97A" box="[316,409,1749,1775]" italics="true" pageId="20" pageNumber="121">N. flava</emphasis>
Wu, Sun &amp; Yang, 1981
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFA9F53CFD72F932FC52F97A" baseAuthorityName="Day" baseAuthorityYear="1957" box="[715,930,1749,1775]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFA9F53CFD72F932FC52F97A" box="[715,930,1749,1775]" italics="true" pageId="20" pageNumber="121">N. indica brunnea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFA9F53CFC0AF932FB1AF97A" authority="(Savigny, 1822)" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[947,1258,1749,1775]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="nubila">
<emphasis id="B90BEAC2FFA9F53CFC0AF932FBD2F97A" box="[947,1058,1749,1775]" italics="true" pageId="20" pageNumber="121">N. nubila</emphasis>
(Savigny, 1822)
</taxonomicName>
likely from the North Atlantic (precise locality unknown), and
<taxonomicName id="4C7F4D53FFA9F53CFC9FF91CFB53F884" authority="(Fauvel, 1932)" baseAuthorityName="Fauvel" baseAuthorityYear="1932" box="[806,1187,1783,1809]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="121" phylum="Annelida" rank="species" species="talehsapensis">
<emphasis id="B90BEAC2FFA9F53CFC9FF91CFC19F884" box="[806,1001,1783,1809]" italics="true" pageId="20" pageNumber="121">N. talehsapensis</emphasis>
(
<bibRefCitation id="EFEE4B21FFA9F53CFC45F913FB6BF884" author="Fauvel P." box="[1020,1179,1783,1809]" pageId="20" pageNumber="121" pagination="1 - 262" refId="ref20199" refString="Fauvel P. 1932. Annelida Polychaeta of the Indian Museum, Calcutta. Memoirs of the Indian Museum 12: 1 - 262." type="journal article" year="1932">Fauvel, 1932</bibRefCitation>
)
</taxonomicName>
from the Gulf of
<collectingCountry id="F3687640FFA9F53CFF04F8FDFED9F8A6" box="[189,297,1817,1843]" name="Thailand" pageId="20" pageNumber="121">Thailand</collectingCountry>
. These species share proximal region of dorsal ligules of similar size throughout the body (or slightly enlarged in posterior parapodia), presence of neuropodial postchaetal and notopodial prechaetal lobes at least in some anterior chaetigers, absence of neuropodial superior lobes, areas VIIVIII with two well-defined bands of more than 20 paragnaths, and area I with no more than two paragnaths (see
<bibRefCitation id="EFEE4B21FFA9F53CFF04F845FDBBF829" author="Villalobos-Guerrero T. F. &amp; Idris I." box="[189,587,1953,1980]" pageId="20" pageNumber="121" pagination="556 - 594" refId="ref22785" refString="Villalobos-Guerrero T. F. &amp; Idris I. 2021. Reproductive morphology and redescriptions of some Neanthes Kinberg, 1865 (Annelida: Nereididae) species from the southeastern Asian seas, with comparative synoptic tables of accepted species. The European Zoological Journal 88 (1): 556 - 594. https: // doi. org / 10.1080 / 24750263.2021.1899318" type="journal article" year="2021">Villalobos-Guerrero &amp; Idris 2021</bibRefCitation>
: table 2).
</paragraph>
<caption id="DF006658FFA8F53DFF04FA53FB33F82B" ID-DOI="http://doi.org/10.5281/zenodo.5121742" ID-Zenodo-Dep="5121742" httpUri="https://zenodo.org/record/5121742/files/figure.png" pageId="21" pageNumber="122" startId="21.[189,232,1463,1489]" targetBox="[346,1241,265,1423]" targetPageId="21">
<paragraph id="8BC036D0FFA8F53DFF04FA53FB33F82B" blockId="21.[186,1399,1463,1983]" pageId="21" pageNumber="122">
<emphasis id="B90BEAC2FFA8F53DFF04FA53FEFCFA44" bold="true" box="[189,268,1463,1489]" pageId="21" pageNumber="122">Fig. 4.</emphasis>
<taxonomicName id="4C7F4D53FFA8F53DFEADFA53FD69FA47" authority="Kinberg, 1865" authorityName="Kinberg" authorityYear="1865" box="[276,665,1463,1490]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="122" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFA8F53DFEADFA53FE14FA44" box="[276,484,1463,1489]" italics="true" pageId="21" pageNumber="122">Neanthes helenae</emphasis>
<bibRefCitation id="EFEE4B21FFA8F53DFE55FA53FD69FA47" author="Kinberg J. G. H." box="[492,665,1463,1490]" pageId="21" pageNumber="122" pagination="167 - 179" refId="ref21148" refString="Kinberg J. G. H. 1865. Annulata nova. Nereidum dispositio nova. Ofversigt af Kongelige Vetenskaps- Akademiens Forhandlingar 22: 167 - 179. Available from https: // www. biodiversitylibrary. org / page / 32339443 [accessed 1 Jul. 2021]." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
</taxonomicName>
.
<emphasis id="B90BEAC2FFA8F53DFD1CFA53FCE9FA44" bold="true" box="[677,793,1463,1489]" pageId="21" pageNumber="122">AC, IP</emphasis>
. Holotype of
<taxonomicName id="4C7F4D53FFA8F53DFC07FA5CFA87FA44" authority="Day, 1949" authorityName="Day" authorityYear="1949" box="[958,1399,1463,1490]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="122" phylum="Annelida" rank="species" species="nanciae" subGenus="Neanthes">
<emphasis id="B90BEAC2FFA8F53DFC07FA5CFB06FA44" box="[958,1270,1463,1489]" italics="true" pageId="21" pageNumber="122">Nereis (Neanthes) nanciae</emphasis>
<bibRefCitation id="EFEE4B21FFA8F53DFB47FA5CFA87FA44" author="Day J. H." box="[1278,1399,1463,1490]" pageId="21" pageNumber="122" pagination="434 - 451" refId="ref19757" refString="Day J. H. 1949. On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnean Society of London, Zoology 41 (281): 434 - 451. https: // doi. org / 10.1111 / j. 1096 - 3642.1940. tb 02416. x" type="journal article" year="1949">Day, 1949</bibRefCitation>
</taxonomicName>
(NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous.
<emphasis id="B90BEAC2FFA8F53DFBAAFA3EFBD7FA61" bold="true" box="[1043,1063,1498,1524]" pageId="21" pageNumber="122">A</emphasis>
. Entire body in dorsal view.
<emphasis id="B90BEAC2FFA8F53DFF07FA1AFF21F98D" bold="true" box="[190,209,1534,1560]" pageId="21" pageNumber="122">B</emphasis>
.Anterior end in dorsal view.
<emphasis id="B90BEAC2FFA8F53DFDA3FA19FDDFF982" bold="true" box="[538,559,1533,1559]" pageId="21" pageNumber="122">C</emphasis>
. Anterior region in ventral view, arrow indicates ventrolateral projection.
<emphasis id="B90BEAC2FFA8F53DFF03F9C5FF35F9AE" bold="true" box="[186,197,1569,1595]" pageId="21" pageNumber="122">I</emphasis>
. Left parapodium from chaetiger 1 in posterior view.
<emphasis id="B90BEAC2FFA8F53DFC87F9C5FCBDF9AE" bold="true" box="[830,845,1569,1595]" pageId="21" pageNumber="122">J</emphasis>
. Left parapodium from chaetiger 7 in posterior view.
<emphasis id="B90BEAC2FFA8F53DFEBDF9A0FEEBF9CB" bold="true" box="[260,283,1604,1630]" pageId="21" pageNumber="122">K</emphasis>
. Right parapodium from chaetiger 21 in anterior view.
<emphasis id="B90BEAC2FFA8F53DFC0BF9A0FC34F9CB" bold="true" box="[946,964,1604,1630]" pageId="21" pageNumber="122">L</emphasis>
. Left parapodium from chaetiger 30 in posterior view with ventral cirri contracted and hidden behind the neuroacicular ligule.
<emphasis id="B90BEAC2FFA8F53DFAB1F983FAD4F914" bold="true" box="[1288,1316,1639,1665]" pageId="21" pageNumber="122">M</emphasis>
. Right parapodium from chaetiger 60 in anterior view.
<emphasis id="B90BEAC2FFA8F53DFCAFF96EFCDBF931" bold="true" box="[790,811,1674,1700]" pageId="21" pageNumber="122">N</emphasis>
. Heterogomph fused falciger from supracicular neurochaetae (chaetiger 30).
<emphasis id="B90BEAC2FFA8F53DFDA7F949FDC5F952" bold="true" box="[542,565,1709,1735]" pageId="21" pageNumber="122">O</emphasis>
. Heterogomph falciger from subacicular neurochaetae (chaetiger 7).
<emphasis id="B90BEAC2FFA8F53DFF04F935FF3FF97E" bold="true" box="[189,207,1745,1771]" pageId="21" pageNumber="122">P</emphasis>
. Heterogomph falciger from subacicular neurochaetae (chaetiger 30). —
<emphasis id="B90BEAC2FFA8F53DFB97F935FB7FF97F" bold="true" box="[1070,1167,1744,1771]" pageId="21" pageNumber="122">DH, Q</emphasis>
. Paratype of
<taxonomicName id="4C7F4D53FFA8F53DFA92F935FDD3F89B" authority="Day, 1949" authorityName="Day" authorityYear="1949" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="122" phylum="Annelida" rank="species" species="nanciae" subGenus="Neanthes">
<emphasis id="B90BEAC2FFA8F53DFA92F935FE53F89B" italics="true" pageId="21" pageNumber="122">Nereis (Neanthes) nanciae</emphasis>
<bibRefCitation id="EFEE4B21FFA8F53DFE13F910FDD3F89B" author="Day J. H." box="[426,547,1780,1806]" pageId="21" pageNumber="122" pagination="434 - 451" refId="ref19757" refString="Day J. H. 1949. On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnean Society of London, Zoology 41 (281): 434 - 451. https: // doi. org / 10.1111 / j. 1096 - 3642.1940. tb 02416. x" type="journal article" year="1949">Day, 1949</bibRefCitation>
</taxonomicName>
(NHMUK 1950.1.5.72-95) from Jamestown, St Helena Island, atokous.
<emphasis id="B90BEAC2FFA8F53DFF07F8F3FF23F8A4" bold="true" box="[190,211,1815,1841]" pageId="21" pageNumber="122">D</emphasis>
. Anterior region in lateral view, arrow indicates ventrolateral projection.
<emphasis id="B90BEAC2FFA8F53DFBFFF8F3FBAAF8A4" bold="true" box="[1094,1114,1815,1841]" pageId="21" pageNumber="122">E</emphasis>
. Left jaw in dorsal and ventral views, respectively.
<emphasis id="B90BEAC2FFA8F53DFDBEF8DEFDE8F8C1" bold="true" box="[519,536,1850,1876]" pageId="21" pageNumber="122">F</emphasis>
. Inner canals of jaws in ventral view.
<emphasis id="B90BEAC2FFA8F53DFC64F8DEFC03F8C1" bold="true" box="[989,1011,1850,1876]" pageId="21" pageNumber="122">G</emphasis>
. Non-everted pharynx in ventral view.
<emphasis id="B90BEAC2FFA8F53DFEB8F8BAFEE8F8ED" bold="true" box="[257,280,1886,1912]" pageId="21" pageNumber="122">H</emphasis>
. Area VI of oral ring of non-everted pharynx in ventral view, arrows indicate p-bars.
<emphasis id="B90BEAC2FFA8F53DFB5EF8B9FB0EF8E2" bold="true" box="[1255,1278,1885,1911]" pageId="21" pageNumber="122">Q</emphasis>
. Posterior end in dorsal view. Roman numerals refer to the areas and Arabic numbers to chaetigers. Scale bars: AC, Q = 1 mm; DE, G = 0.5 mm; F, JL = 0.2 mm; HI, M = 0.1 mm; NP = 20 µm.
</paragraph>
</caption>
<paragraph id="8BC036D0FFABF53EFF04FEEEFAC9FC5B" blockId="22.[187,1399,265,974]" pageId="22" pageNumber="123">
Moreover,
<taxonomicName id="4C7F4D53FFABF53EFEFBFEEEFE34FEB6" authorityName="Kinberg" authorityYear="1865" box="[322,452,265,291]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFABF53EFEFBFEEEFE34FEB6" box="[322,452,265,291]" italics="true" pageId="22" pageNumber="123">N. helenae</emphasis>
</taxonomicName>
is distinguishable from them all by the following diagnostic features: (I) the presence of p-bars on area VI, in contrast to its absence in those species; (II) the presence of ventrolateral projections on the apodous segment, in comparison to its absence in those species; (III) the presence of simple chaetae (fused falcigers) in supracicular fascicle, in contrast to its absence in
<taxonomicName id="4C7F4D53FFABF53EFB19FE94FB0CFE1C" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[1184,1276,367,393]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFB19FE94FB0CFE1C" box="[1184,1276,367,393]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFABF53EFAB2FE94FEEFFE3E" baseAuthorityName="Day" baseAuthorityYear="1957" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFAB2FE94FEEFFE3E" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFABF53EFE95FE76FE67FE3E" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[300,407,401,427]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nubila">
<emphasis id="B90BEAC2FFABF53EFE95FE76FE67FE3E" box="[300,407,401,427]" italics="true" pageId="22" pageNumber="123">N. nubila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFE74FE76FD7CFE3E" baseAuthorityName="Fauvel" baseAuthorityYear="1932" box="[461,652,401,427]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="talehsapensis">
<emphasis id="B90BEAC2FFABF53EFE74FE76FD7CFE3E" box="[461,652,401,427]" italics="true" pageId="22" pageNumber="123">N. talehsapensis</emphasis>
</taxonomicName>
; (IV) one paragnath on area V, in comparison to none in
<taxonomicName id="4C7F4D53FFABF53EFAADFE76FA9DFE3E" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[1300,1389,401,427]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFAADFE76FA9DFE3E" box="[1300,1389,401,427]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFABF53EFF04FE50FE64FE5B" baseAuthorityName="Day" baseAuthorityYear="1957" box="[189,404,436,462]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFF04FE50FE64FE5B" box="[189,404,436,462]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFABF53EFE1AFE50FDE1FE5B" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[419,529,436,462]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nubila">
<emphasis id="B90BEAC2FFABF53EFE1AFE50FDE1FE5B" box="[419,529,436,462]" italics="true" pageId="22" pageNumber="123">N. nubila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFDF5FE50FCFDFE5B" baseAuthorityName="Fauvel" baseAuthorityYear="1932" box="[588,781,436,462]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="talehsapensis">
<emphasis id="B90BEAC2FFABF53EFDF5FE50FCFDFE5B" box="[588,781,436,462]" italics="true" pageId="22" pageNumber="123">N. talehsapensis</emphasis>
</taxonomicName>
; (V) the paragnaths disposed in a transverse row on area VI, in contrast to those in a rounded or oval patch in
<taxonomicName id="4C7F4D53FFABF53EFCF6FE33FC58FE65" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[847,936,470,496]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFCF6FE33FC58FE65" box="[847,936,470,496]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFABF53EFC0EFE33FB79FE65" baseAuthorityName="Day" baseAuthorityYear="1957" box="[951,1161,470,496]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFC0EFE33FB79FE65" box="[951,1161,470,496]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
and
<emphasis id="B90BEAC2FFABF53EFB79FE33FAC2FE65" box="[1216,1330,470,496]" italics="true" pageId="22" pageNumber="123">
<taxonomicName id="4C7F4D53FFABF53EFB79FE33FADFFE65" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[1216,1327,470,496]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nubila">N. nubila</taxonomicName>
,
</emphasis>
and a cross or irregular patch in
<taxonomicName id="4C7F4D53FFABF53EFE51FE1DFD57FD87" baseAuthorityName="Fauvel" baseAuthorityYear="1932" box="[488,679,504,530]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="talehsapensis">
<emphasis id="B90BEAC2FFABF53EFE51FE1DFD57FD87" box="[488,679,504,530]" italics="true" pageId="22" pageNumber="123">N. talehsapensis</emphasis>
</taxonomicName>
; (VI) the dorsal cirri extending markedly beyond distal region of dorsal ligule at least in anterior parapodia, in comparison to those that are subequal to or not extending beyond end in
<taxonomicName id="4C7F4D53FFABF53EFED0FDD9FE34FDC3" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[361,452,572,598]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFED0FDD9FE34FDC3" box="[361,452,572,598]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F4D53FFABF53EFE6AFDD9FD57FDC3" baseAuthorityName="Day" baseAuthorityYear="1957" box="[467,679,572,598]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFE6AFDD9FD57FDC3" box="[467,679,572,598]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFD67FDD9FCBBFDC3" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[734,843,572,598]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nubila">
<emphasis id="B90BEAC2FFABF53EFD67FDD9FCBBFDC3" box="[734,843,572,598]" italics="true" pageId="22" pageNumber="123">N. nubila</emphasis>
</taxonomicName>
; (VII) 1828 paragnaths on area III, in contrast to one in
<taxonomicName id="4C7F4D53FFABF53EFE88FDBBFE7EFDED" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[305,398,606,632]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFE88FDBBFE7EFDED" box="[305,398,606,632]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
and
<quantity id="4C879B35FFABF53EFE70FDBAFDE4FDED" box="[457,532,606,633]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.286" metricValueMin="1.27" pageId="22" pageNumber="123" unit="in" value="7.0" valueMax="9.0" valueMin="5.0">59 in</quantity>
<taxonomicName id="4C7F4D53FFABF53EFDA6FDBBFD05FDED" baseAuthorityName="Day" baseAuthorityYear="1957" box="[543,757,606,632]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFDA6FDBBFD05FDED" box="[543,757,606,632]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
; (VIII) the heterogomph falciger blade with hammerheaded terminal tooth and distinct tendon in middle and posterior parapodia, in comparison to those with blunt terminal tooth and inconspicuous tendon in
<taxonomicName id="4C7F4D53FFABF53EFCFDFD47FC6FFD29" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[836,927,674,700]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFCFDFD47FC6FFD29" box="[836,927,674,700]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFC63FD47FB6AFD29" baseAuthorityName="Fauvel" baseAuthorityYear="1932" box="[986,1178,674,700]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="talehsapensis">
<emphasis id="B90BEAC2FFABF53EFC63FD47FB6AFD29" box="[986,1178,674,700]" italics="true" pageId="22" pageNumber="123">N. talehsapensis</emphasis>
</taxonomicName>
; (IX) the presence of merged paragnaths, in contrast to their absence in
<taxonomicName id="4C7F4D53FFABF53EFC8AFD22FC60FD4A" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[819,912,709,735]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFC8AFD22FC60FD4A" box="[819,912,709,735]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFC72FD22FB51FD4A" baseAuthorityName="Day" baseAuthorityYear="1957" box="[971,1185,709,735]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFC72FD22FB51FD4A" box="[971,1185,709,735]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
; (X) the antennae well separated from each other, in comparison to those that are closely together in
<taxonomicName id="4C7F4D53FFABF53EFBC0FD0CFB23FC94" authorityName="Wu, Sun &amp; Yang" authorityYear="1981" box="[1145,1235,743,769]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B90BEAC2FFABF53EFBC0FD0CFB23FC94" box="[1145,1235,743,769]" italics="true" pageId="22" pageNumber="123">N. flava</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFAB2FD0CFEEEFCB6" baseAuthorityName="Day" baseAuthorityYear="1957" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFAB2FD0CFEEEFCB6" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
; (XI) the campanulate prostomium, in contrast to an ovoid shape in
<taxonomicName id="4C7F4D53FFABF53EFBDEFCEEFAB2FCB6" baseAuthorityName="Day" baseAuthorityYear="1957" box="[1127,1346,777,803]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFBDEFCEEFAB2FCB6" box="[1127,1346,777,803]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D53FFABF53EFF04FCC8FEDAFCD0" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[189,298,811,837]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nubila">
<emphasis id="B90BEAC2FFABF53EFF04FCC8FEDAFCD0" box="[189,298,811,837]" italics="true" pageId="22" pageNumber="123">N. nubila</emphasis>
</taxonomicName>
; (XII) the π-shaped ridge pattern of areas VIVVI, in comparison to that λ-shaped pattern in
<taxonomicName id="4C7F4D53FFABF53EFF04FCAAFE7EFCF2" baseAuthorityName="Day" baseAuthorityYear="1957" box="[189,398,845,871]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFF04FCAAFE7EFCF2" box="[189,398,845,871]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
; (XIII) the dorsum of segments with pigmentation bands, in contrast to a pigment band present only in the apodous segment in
<taxonomicName id="4C7F4D53FFABF53EFD37FC94FC93FC1C" baseAuthorityName="Day" baseAuthorityYear="1957" box="[654,867,879,905]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="subSpecies" species="indica" subSpecies="brunnea">
<emphasis id="B90BEAC2FFABF53EFD37FC94FC93FC1C" box="[654,867,879,905]" italics="true" pageId="22" pageNumber="123">N. indica brunnea</emphasis>
</taxonomicName>
; (XIV) the paragnaths present on both ridges and furrows of areas VIIVIII of the pharynx, in contrast to those only present on ridges in
<taxonomicName id="4C7F4D53FFABF53EFAB8FC76FA9EFC3E" baseAuthorityName="Savigny" baseAuthorityYear="1822" box="[1281,1390,913,939]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nubila">
<emphasis id="B90BEAC2FFABF53EFAB8FC76FA9EFC3E" box="[1281,1390,913,939]" italics="true" pageId="22" pageNumber="123">N. nubila</emphasis>
</taxonomicName>
; and (XV) the aciculae mostly black, in comparison to those rather pale colour in
<taxonomicName id="4C7F4D53FFABF53EFBCBFC50FAC2FC5B" baseAuthorityName="Fauvel" baseAuthorityYear="1932" box="[1138,1330,948,974]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="talehsapensis">
<emphasis id="B90BEAC2FFABF53EFBCBFC50FAC2FC5B" box="[1138,1330,948,974]" italics="true" pageId="22" pageNumber="123">N. talehsapensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC036D0FFABF53EFF04FC10FA93FBE1" blockId="22.[189,1400,1012,1141]" pageId="22" pageNumber="123">
<taxonomicName id="4C7F4D53FFABF53EFF04FC10FE7DFB9B" authorityName="Kinberg" authorityYear="1865" box="[189,397,1012,1038]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFABF53EFF04FC10FE7DFB9B" box="[189,397,1012,1038]" italics="true" pageId="22" pageNumber="123">Neanthes helenae</emphasis>
</taxonomicName>
was described without illustrations by
<bibRefCitation id="EFEE4B21FFABF53EFCE7FC10FBE5FB9A" author="Kinberg J. G. H." box="[862,1045,1012,1039]" pageId="22" pageNumber="123" pagination="167 - 179" refId="ref21148" refString="Kinberg J. G. H. 1865. Annulata nova. Nereidum dispositio nova. Ofversigt af Kongelige Vetenskaps- Akademiens Forhandlingar 22: 167 - 179. Available from https: // www. biodiversitylibrary. org / page / 32339443 [accessed 1 Jul. 2021]." type="journal article" year="1865">Kinberg (1865)</bibRefCitation>
using an unknown number of specimens collected near St Helena Island. The type material was not addressed in the literature, and most of its morphology remained unknown until this study. The
<specimenCount id="9D79FD59FFABF53EFC16FBDDFBB9FBC6" box="[943,1097,1081,1107]" pageId="22" pageNumber="123" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
are in poor condition, but the relevant diagnostic features are still visible, and thus are useful to define the species morphology.
</paragraph>
<paragraph id="8BC036D0FFABF53EFF04FB7FFCA1FA53" blockId="22.[187,1400,1179,1479]" pageId="22" pageNumber="123">
<bibRefCitation id="EFEE4B21FFABF53EFF04FB7FFEB4FB20" author="Day J. H." box="[189,324,1179,1205]" pageId="22" pageNumber="123" pagination="434 - 451" refId="ref19757" refString="Day J. H. 1949. On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnean Society of London, Zoology 41 (281): 434 - 451. https: // doi. org / 10.1111 / j. 1096 - 3642.1940. tb 02416. x" type="journal article" year="1949">Day (1949)</bibRefCitation>
described
<taxonomicName id="4C7F4D53FFABF53EFE7FFB78FD0FFB20" authorityName="Day" authorityYear="1949" box="[454,767,1179,1205]" class="Polychaeta" family="Nereididae" genus="Nereis" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nanciae" subGenus="Neanthes">
<emphasis id="B90BEAC2FFABF53EFE7FFB78FDE2FB20" box="[454,530,1180,1205]" italics="true" pageId="22" pageNumber="123">Nereis</emphasis>
(
<emphasis id="B90BEAC2FFABF53EFD9AFB7FFD60FB20" box="[547,656,1179,1205]" italics="true" pageId="22" pageNumber="123">Neanthes</emphasis>
)
<emphasis id="B90BEAC2FFABF53EFD1BFB78FD0FFB20" box="[674,767,1180,1205]" italics="true" pageId="22" pageNumber="123">nanciae</emphasis>
</taxonomicName>
from St Helena Island with several atoke specimens, but he did not justify the proposal as a new species. Later,
<bibRefCitation id="EFEE4B21FFABF53EFC3CFB59FBB3FB4D" author="Hartman O." box="[901,1091,1213,1240]" pageId="22" pageNumber="123" pagination="1 - 353" refId="ref20606" refString="Hartman O. 1959. Catalogue of the polychaetous annelids of the world, 1. Occasional Papers of the Allan Hancock Foundation 23: 1 - 353." type="journal article" year="1959">Hartman (1959)</bibRefCitation>
transferred the species to
<taxonomicName id="4C7F4D53FFABF53EFF06FB3BFEDCFB6C" authorityName="Kinberg" authorityYear="1865" box="[191,300,1247,1273]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="genus">
<emphasis id="B90BEAC2FFABF53EFF06FB3BFEDCFB6C" box="[191,300,1247,1273]" italics="true" pageId="22" pageNumber="123">Neanthes</emphasis>
</taxonomicName>
and suggested it as a possible synonym of the incompletely known
<taxonomicName id="4C7F4D53FFABF53EFB94FB04FB58FB6C" authorityName="Kinberg" authorityYear="1865" box="[1069,1192,1247,1273]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFABF53EFB94FB04FB58FB6C" box="[1069,1192,1247,1273]" italics="true" pageId="22" pageNumber="123">N. helenae</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFEE4B21FFABF53EFB0EFB3BFA86FB6C" author="Fauchald K." box="[1207,1398,1247,1273]" pageId="22" pageNumber="123" pagination="1 - 575" refId="ref20097" refString="Fauchald K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Foundation Monographs in Marine Biology 7: 1 - 575. Available from https: // repository. si. edu / handle / 10088 / 6207 [accessed 1 Jul. 2021]." type="journal article" year="1972">Fauchald (1972)</bibRefCitation>
considered
<taxonomicName id="4C7F4D53FFABF53EFEFAFAE5FDE4FA8E" baseAuthorityName="Day" baseAuthorityYear="1949" box="[323,532,1281,1307]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nanciae">
<emphasis id="B90BEAC2FFABF53EFEFAFAE5FDE4FA8E" box="[323,532,1281,1307]" italics="true" pageId="22" pageNumber="123">Neanthes nanciae</emphasis>
</taxonomicName>
as a valid species, but
<bibRefCitation id="EFEE4B21FFABF53EFCA4FAE5FC35FA8E" author="Wilson R. S." box="[797,965,1281,1307]" pageId="22" pageNumber="123" pagination="209 - 226" refId="ref22922" refString="Wilson R. S. 1984. Neanthes (Polychaeta: Nereididae) from Victoria with descriptions of two new species. Proceedings of the Royal Society of Victoria 96: 209 - 226." type="journal article" year="1984">Wilson (1984)</bibRefCitation>
agreed with Hartmans assumptions, suggesting
<taxonomicName id="4C7F4D53FFABF53EFEFDFAC0FE36FAA8" baseAuthorityName="Day" baseAuthorityYear="1949" box="[324,454,1316,1341]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nanciae">
<emphasis id="B90BEAC2FFABF53EFEFDFAC0FE36FAA8" box="[324,454,1316,1341]" italics="true" pageId="22" pageNumber="123">N. nanciae</emphasis>
</taxonomicName>
as a possible junior synonym of
<taxonomicName id="4C7F4D53FFABF53EFCE2FAC0FC29FAA8" authorityName="Kinberg" authorityYear="1865" box="[859,985,1315,1341]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFABF53EFCE2FAC0FC29FAA8" box="[859,985,1315,1341]" italics="true" pageId="22" pageNumber="123">N. helenae</emphasis>
</taxonomicName>
. After the detailed examination of both species
<typeStatus id="54C48872FFABF53EFEE6FAA2FE62FAF5" box="[351,402,1350,1376]" pageId="22" pageNumber="123">type</typeStatus>
material,
<taxonomicName id="4C7F4D53FFABF53EFDBCFAA2FD73FACA" authorityName="Kinberg" authorityYear="1865" box="[517,643,1349,1375]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="helenae">
<emphasis id="B90BEAC2FFABF53EFDBCFAA2FD73FACA" box="[517,643,1349,1375]" italics="true" pageId="22" pageNumber="123">N. helenae</emphasis>
</taxonomicName>
is here regarded as the senior synonym of
<taxonomicName id="4C7F4D53FFABF53EFBC3FAA2FB0AFACA" baseAuthorityName="Day" baseAuthorityYear="1949" box="[1146,1274,1350,1375]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="species" species="nanciae">
<emphasis id="B90BEAC2FFABF53EFBC3FAA2FB0AFACA" box="[1146,1274,1350,1375]" italics="true" pageId="22" pageNumber="123">N. nanciae</emphasis>
</taxonomicName>
as a result of the overlap in unique and diagnostic features among other species in
<taxonomicName id="4C7F4D53FFABF53EFBB2FA83FB88FA14" authorityName="Kinberg" authorityYear="1865" box="[1035,1144,1383,1409]" class="Polychaeta" family="Nereididae" genus="Neanthes" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="123" phylum="Annelida" rank="genus">
<emphasis id="B90BEAC2FFABF53EFBB2FA83FB88FA14" box="[1035,1144,1383,1409]" italics="true" pageId="22" pageNumber="123">Neanthes</emphasis>
</taxonomicName>
, such as the presence of p-bar paragnaths on area VI, ventrolateral projections on the apodous segment and the presence of simple chaetae (fused falcigers) in supracicular fascicle.
</paragraph>
</subSubSection>
<subSubSection id="C365655BFFABF53EFF04FA08FE5AF9A1" box="[189,426,1516,1588]" pageId="22" pageNumber="123" type="distribution">
<paragraph id="8BC036D0FFABF53EFF04FA08FEA6F993" blockId="22.[189,342,1516,1542]" box="[189,342,1516,1542]" pageId="22" pageNumber="123">
<heading id="D08881BCFFABF53EFF04FA08FEA6F993" bold="true" box="[189,342,1516,1542]" fontSize="11" level="3" pageId="22" pageNumber="123" reason="3">
<emphasis id="B90BEAC2FFABF53EFF04FA08FEA6F993" bold="true" box="[189,342,1516,1542]" pageId="22" pageNumber="123">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFABF53EFF04F9FEFE5AF9A1" blockId="22.[189,426,1562,1588]" box="[189,426,1562,1588]" pageId="22" pageNumber="123">
<collectingCountry id="F3687640FFABF53EFF04F9FEFE56F9A1" box="[189,422,1562,1588]" name="Saint Helena Island" pageId="22" pageNumber="123">Saint Helena Island</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C365655BFFABF53EFF04F9BEFC53F937" box="[189,931,1626,1698]" pageId="22" pageNumber="123" type="biology_ecology">
<paragraph id="8BC036D0FFABF53EFF04F9BEFED0F9E1" blockId="22.[189,288,1626,1652]" box="[189,288,1626,1652]" pageId="22" pageNumber="123">
<heading id="D08881BCFFABF53EFF04F9BEFED0F9E1" bold="true" box="[189,288,1626,1652]" fontSize="11" level="3" pageId="22" pageNumber="123" reason="3">
<emphasis id="B90BEAC2FFABF53EFF04F9BEFED0F9E1" bold="true" box="[189,288,1626,1652]" pageId="22" pageNumber="123">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFABF53EFF04F963FC53F937" blockId="22.[189,931,1671,1698]" box="[189,931,1671,1698]" pageId="22" pageNumber="123">
The upper part of the intertidal zone to
<quantity id="4C879B35FFABF53EFD33F96CFD36F937" box="[650,710,1672,1698]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="22" pageNumber="123" unit="m" value="73.0">73 m</quantity>
depth (
<bibRefCitation id="EFEE4B21FFABF53EFCA4F96CFC66F937" author="Day J. H." box="[797,918,1672,1698]" pageId="22" pageNumber="123" pagination="434 - 451" refId="ref19757" refString="Day J. H. 1949. On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnean Society of London, Zoology 41 (281): 434 - 451. https: // doi. org / 10.1111 / j. 1096 - 3642.1940. tb 02416. x" type="journal article" year="1949">Day 1949</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C365655BFFABF53EFF04F92CFEC9F89A" box="[189,361,1736,1807]" pageId="22" pageNumber="123" type="description">
<paragraph id="8BC036D0FFABF53EFF04F92CFE99F977" blockId="22.[189,361,1736,1762]" box="[189,361,1736,1762]" pageId="22" pageNumber="123">
<heading id="D08881BCFFABF53EFF04F92CFE99F977" bold="true" box="[189,361,1736,1762]" fontSize="11" level="3" pageId="22" pageNumber="123" reason="3">
<emphasis id="B90BEAC2FFABF53EFF04F92CFE99F977" bold="true" box="[189,361,1736,1762]" pageId="22" pageNumber="123">Reproduction</emphasis>
</heading>
</paragraph>
<paragraph id="8BC036D0FFABF53EFF04F911FEC9F89A" blockId="22.[189,313,1781,1807]" box="[189,313,1781,1807]" pageId="22" pageNumber="123">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>