360 lines
37 KiB
XML
360 lines
37 KiB
XML
<document id="4AC6555DBC825467EB49D09EB6525C24" ID-DOI="10.11646/zootaxa.3734.1.6" ID-GBIF-Dataset="0d7cf01e-dc78-41ca-951b-87b0108bbb1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="223677" ID-ZooBank="5CB1F0B9-752B-4746-8865-F3587DF7074E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460695170467" checkinUser="plazi" docAuthor="Nguyen, Truong Quang, Nguyen, Khoi Vu, Devender, Robert Wayne Van, Bonkowski, Michael & Ziegler, Thomas" docDate="2013" docId="03D68789AC681623AA90FBE76393FC7E" docLanguage="en" docName="zt03734p062.pdf" docOrigin="Zootaxa 3734 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphenomorphus sheai Nguyen, Nguyen, Devender, Bonkowski & Ziegler, 2013, sp. nov." docType="treatment" docVersion="7" masterDocId="FFEFFFF1AC691626AA07FFC86108FFD3" masterDocTitle="A new species of Sphenomorphus Fitzinger, 1843 (Squamata: Sauria: Scincidae) from Vietnam" updateTime="1698353766526" updateUser="plazi">
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<mods:mods id="49DFC5CFA3F16E5DFF13B5FD3A7789F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A943E610BEE4DA03E43C533673CBAA19">
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<mods:title id="0CCD7063698B151F2135119308D86B84">A new species of Sphenomorphus Fitzinger, 1843 (Squamata: Sauria: Scincidae) from Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="F696CC9B92BB7B70027164F16FE6B84E" type="personal">
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<mods:role id="02FCAB580B33CAF8DACF34B0D3139431">
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<mods:roleTerm id="4995D0020B4EBDC6AD201BF0368DDDED">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="59B9B2CBE56612CB27214B8AB1E32351">Nguyen, Truong Quang</mods:namePart>
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</mods:name>
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<mods:name id="A6B5D8199A744D68C78B1041008749BC" type="personal">
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<mods:role id="FCCF37106DB7E9CCE207000048F9673D">
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<mods:roleTerm id="C594CD35A36CC251F5319945BD40CAB8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="773D894A4DDFE86F6FF2AFF6A299DBB0">Nguyen, Khoi Vu</mods:namePart>
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</mods:name>
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<mods:name id="880F0BE96B6541F594D75B4961892C8E" type="personal">
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<mods:role id="A3FA58839EFED18EA33D38797AB8D49A">
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<mods:roleTerm id="AFF675BA7CB9AFDACF66BE70A047E8F8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5BCB284ABA10609DDBD60EA6BE82B650">Devender, Robert Wayne Van</mods:namePart>
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</mods:name>
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<mods:name id="A04E67E791DD5DBE579827D7721A2F06" type="personal">
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<mods:role id="8D38E9D75299681B9E349F53455ADC1F">
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<mods:roleTerm id="372B21255D076C79B7830E0510F3813E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6B8760ADD082910F1EAC2E7F08528E48">Bonkowski, Michael</mods:namePart>
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</mods:name>
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<mods:name id="0D195F6BC1854E6D48ED419B4CA66BAC" type="personal">
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<mods:role id="A1D63BC51E2D4CBC7E0BAF855D0BA859">
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<mods:roleTerm id="A71E3EBC53CA1CAA357903465D6912D0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B7440C3C5FBBC96CB4984F11F916D0F2">Ziegler, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="684DA1F7352FAFAF1BABBFFB99469C6E">text</mods:typeOfResource>
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<mods:titleInfo id="4FECAEACA065ACCD35B98C71B18DB34C">
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<mods:title id="407D9B1C03CB0B082BA2DB24B45E6FD3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D13424EFBBA0F9381F38B8E35D26A6C4">
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<mods:date id="8719A3C2E597B8FEE13305FE56F267EE">2013</mods:date>
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<mods:detail id="519C9C3A857043AB9695161494EEE74B" type="volume">
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<mods:number id="C253423C58D189A7424FCC4E99666B78">3734</mods:number>
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</mods:detail>
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<mods:detail id="1AC3EE45F15595C5D766C498C6F1E0CF" type="issue">
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<mods:number id="277AE58FBF8CC897C471F294092A435E">1</mods:number>
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</mods:detail>
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</mods:part>
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<mods:classification id="CCDF58F389021DF66A92A760A5AE7401">journal volume</mods:classification>
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<mods:identifier id="7F804ACB3A85B5405B373F82B140775A" type="DOI">10.11646/zootaxa.3734.1.6</mods:identifier>
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<mods:identifier id="9ED220302753A31D8949D28C783C8F7D" type="GBIF-Dataset">0d7cf01e-dc78-41ca-951b-87b0108bbb1b</mods:identifier>
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<mods:identifier id="CEA8FE538CD269BBD39F1E4624C81B6E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F8C4B8E2A1FE52C96FCC8F7B106EA9DE" type="Zenodo-Dep">223677</mods:identifier>
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<mods:identifier id="955702A890C84CBCC5A4BAA679FD36C2" type="ZooBank">5CB1F0B9-752B-4746-8865-F3587DF7074E</mods:identifier>
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</mods:mods>
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<treatment id="03D68789AC681623AA90FBE76393FC7E" ID-DOI="http://doi.org/10.5281/zenodo.6146951" ID-GBIF-Taxon="119581734" ID-Zenodo-Dep="6146951" LSID="urn:lsid:plazi:treatment:03D68789AC681623AA90FBE76393FC7E" httpUri="http://treatment.plazi.org/id/03D68789AC681623AA90FBE76393FC7E" lastPageId="5" pageId="1">
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<subSubSection id="C3656514AC681627AA90FBE7601AFBA3" pageId="1" type="nomenclature">
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<paragraph id="8BC0369FAC681627AA90FBE76301FB99" blockId="1.[151,521,1071,1136]" box="[151,521,1071,1098]" pageId="1">
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<heading id="D08881F3AC681627AA90FBE76301FB99" bold="true" box="[151,521,1071,1098]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="B90BEA8DAC681627AA90FBE76301FB99" bold="true" box="[151,521,1071,1098]" pageId="1">
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<taxonomicName id="4C7F4D1CAC681627AA90FBE760ACFB9A" box="[151,420,1071,1097]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="species" species="sheai" status="sp. nov.">
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<emphasis id="B90BEA8DAC681627AA90FBE760ACFB9A" bold="true" box="[151,420,1071,1097]" italics="true" pageId="1">Sphenomorphus sheai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857F6AC681627ABABFBF86301FB99" box="[428,521,1072,1098]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC0369FAC681627AA90FB9F601AFBA3" blockId="1.[151,521,1071,1136]" box="[151,274,1111,1136]" pageId="1">
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(
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<figureCitation id="13442A1AAC681627AA98FB9F61E4FBA3" box="[159,236,1111,1136]" captionStart="FIGURE 1" captionStartId="2.[151,255,1926,1950]" captionTargetBox="[151,1436,1156,1902]" captionTargetId="figure@2.[151,1436,1156,1903]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal view of the holotype (IEBR A. 2013.22) of Sphenomorphus sheai sp. nov. Photo R. W. Van Devender." httpUri="https://zenodo.org/record/223678/files/figure.png" pageId="1">Figs. 1</figureCitation>
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,
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<figureCitation id="13442A1AAC681627AAFDFB906001FBA3" box="[250,265,1112,1136]" captionStart="FIGURE 2" captionStartId="3.[151,255,1574,1598]" captionTargetBox="[280,1276,205,1517]" captionTargetId="figure@3.[280,1308,194,1558]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal (A) and lateral (B) views of the head of Sphenomorphus sheai sp. nov. (holotype IEBR A. 2013.22). Photos T. Q. Nguyen." httpUri="https://zenodo.org/record/223679/files/figure.png" pageId="1">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656514AC681627AA90FB6F6439FAC0" pageId="1" type="materials_examined">
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<paragraph id="8BC0369FAC681627AA90FB6F6439FAC0" blockId="1.[151,1436,1191,2019]" pageId="1">
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<emphasis id="B90BEA8DAC681627AA90FB6F6003FB13" bold="true" box="[151,267,1191,1216]" pageId="1">
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<typeStatus id="54C4883DAC681627AA90FB6F600EFB13" box="[151,262,1191,1216]" pageId="1" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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IEBR A.2013.22 (Field number ASU 10715), adult female, collected on
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<date id="FFC1105FAC681627AE4EFB6065DAFB13" box="[1097,1234,1192,1216]" pageId="1" value="2012-06-04">4 June 2012</date>
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by K. V. Nguyen and R. W. Van Devender near Camp 1 (
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<geoCoordinate id="EE4B5058AC681627A85EFB1B63E3FB38" box="[601,747,1231,1259]" direction="north" orientation="latitude" pageId="1" precision="1" value="14.6183">14o37.098’N</geoCoordinate>
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,
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<geoCoordinate id="EE4B5058AC681627A8F1FB1B6299FB38" box="[758,913,1231,1259]" direction="east" orientation="longitude" pageId="1" precision="1" value="108.50295">108o30.177’E</geoCoordinate>
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, elevation
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<quantity id="4C879B7AAC681627AE0CFB1A6562FB39" box="[1035,1130,1234,1259]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="1" unit="m" value="1025.0">1,025 m</quantity>
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; datum WGS84) from Kon Tum Plateau in the border area between Quang Ngai and Kon Tum provinces, southern Central
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<collectingCountry id="F368760FAC681627AECEFB326425FAC0" box="[1225,1325,1274,1299]" name="Vietnam" pageId="1">Vietnam</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656514AC681627AAC0FAEA60CBF9F8" pageId="1" type="diagnosis">
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<paragraph id="8BC0369FAC681627AAC0FAEA60CBF9F8" blockId="1.[151,1436,1191,2019]" pageId="1">
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<emphasis id="B90BEA8DAC681627AAC0FAEA6049FAE8" bold="true" box="[199,321,1314,1339]" pageId="1">Diagnosis.</emphasis>
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Size small (SVL
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<quantity id="4C879B7AAC681627A81FFAEA638DFAE8" box="[536,645,1314,1339]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="1" unit="mm" value="35.2">35.2 mm</quantity>
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); prefrontals separated from each other; supraoculars four; enlarged nuchals in two pairs; lower eyelid scaly; supralabials six, first fused with nasal; primary temporals two; external ear openings absent; midbody scales in 20 rows; dorsal scales smooth, paravertebral scales 53, not widened; limbs short, pentadactyl; scales on dorsal surface of base of fourth toe in three rows; lamellae under fourth toe six; free margins of upper and lower eyelids not edged in white; dorsum and tail base bronze brown with dark dots; dorsolateral light stripe present on neck, shoulder and anterior part of dorsum; upper zone of lateral head and flank with a narrow black stripe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656514AC681624AAC0F9F26460FB9B" lastPageId="2" pageId="1" type="description">
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<paragraph id="8BC0369FAC681627AAC0F9F262DAF970" blockId="1.[151,1436,1191,2019]" pageId="1">
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<emphasis id="B90BEA8DAC681627AAC0F9F260EBF980" bold="true" box="[199,483,1594,1619]" pageId="1">
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Description of
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<typeStatus id="54C4883DAC681627AB72F9F260D6F980" box="[373,478,1594,1619]" pageId="1" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Size small (SVL
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<quantity id="4C879B7AAC681627A8AFF9F26218F980" box="[680,784,1594,1619]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="1" unit="mm" value="35.2">35.2 mm</quantity>
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), tail complete (TaL
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<quantity id="4C879B7AAC681627A9F1F9F26556F980" box="[1014,1118,1594,1619]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" pageId="1" unit="mm" value="58.5">58.5 mm</quantity>
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), head longer than wide (HL 6.0 mm, HW
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<quantity id="4C879B7AAC681627AB38F9AB6095F9A8" box="[319,413,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" unit="mm" value="4.2">4.2 mm</quantity>
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), SL
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<quantity id="4C879B7AAC681627ABE5F9AA6337F9A9" box="[482,575,1634,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" unit="mm" value="2.6">2.6 mm</quantity>
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, SFlL
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<quantity id="4C879B7AAC681627A894F9AB63F6F9A9" box="[659,766,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="1" unit="mm" value="10.8">10.8 mm</quantity>
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, AG
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<quantity id="4C879B7AAC681627A944F9AB62A6F9A9" box="[835,942,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="1" unit="mm" value="21.2">21.2 mm</quantity>
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, limbs short (FlL
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<quantity id="4C879B7AAC681627AE8DF9AA65EFF9A9" box="[1162,1255,1634,1659]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" unit="mm" value="5.1">5.1 mm</quantity>
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, HlL
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<quantity id="4C879B7AAC681627AF37F9AA6498F9A8" box="[1328,1424,1634,1659]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="1" unit="mm" value="7.5">7.5 mm</quantity>
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); proportions: TaL/SVL 1.66, HL/SVL 0.17, FlL/SVL 0.14, HlL/SVL 0.21.
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</paragraph>
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<paragraph id="8BC0369FAC681627AAC0F97B6537F868" blockId="1.[151,1436,1191,2019]" pageId="1">Snout obtuse, rounded anteriorly; rostral wider than high, distinctly visible from above; supranasals absent; frontonasal wider than long, in contact with rostral, nasals, anterior loreals, and prefrontals; prefrontals separated from each other; frontal narrowing posteriorly, longer than the distance to the tip of snout, in contact with frontonasal, prefrontals, first and second supraoculars, and frontoparietals; frontoparietals in contact with each other anteriorly, and bordered by frontal, three posterior supraoculars, parietals, and interparietal; interparietal lozenge-shaped with a small transparent spot in posterior angle; parietals in contact posteriorly, posterolateral border surrounded by three scales on each side; enlarged nuchal scales in two pairs.</paragraph>
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<paragraph id="8BC0369FAC681624AAC0F80262A5FEAB" blockId="1.[151,1436,1191,2019]" lastBlockId="2.[151,1436,151,1096]" lastPageId="2" pageId="1">Nostril in center of nasal; postnasals absent; loreals two, anterior larger than posterior; preocular single; presubocular single; supraciliaries seven, first largest, first to third in contact with first supraocular; supraoculars four, second widest, fourth supraocular followed by a small scale; postocular single; postsuboculars three, lower one in contact with fifth supralabial; primary temporals two, lower one in contact with sixth supralabial; secondary temporals two, upper one very large, in contact with parietal, overlapped by lower one; lower eyelid moveable, scaly, separated from supralabials by one row of small scales; supralabials six, anterior first fused with nasal, fifth and sixth enlarged, fourth below the eye; external ear openings absent.</paragraph>
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<paragraph id="8BC0369FAC6B1624AAC0FE4F62FEFDCB" blockId="2.[151,1436,151,1096]" pageId="2">Mental wider than long, rounded anteriorly, in contact with anterior infralabial on each side and postmental; infralabials five; postmental undivided, in contact with mental, first infralabial and anterior pair of chinshields; three pairs of chinshields, anterior pair in contact with each other anteriorly, second pair separated from each other by a gular scale, and posterior pair separated from each other by three scales.</paragraph>
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<paragraph id="8BC0369FAC6B1624AAC0FDEF637BFD6B" blockId="2.[151,1436,151,1096]" pageId="2">Midbody scales in 20 rows; dorsal scales smooth, subequal to lateral and ventral scales, four scale rows between dark stripes on upper lateral zones; paravertebral scales 53, not widened; ventrals smooth, in 54 rows; precloacals four, inner scales overlapping outer ones, medial two enlarged, right scale overlapped by left scale; tail thick at base, median subcaudals widened.</paragraph>
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<paragraph id="8BC0369FAC6B1624AAC0FD0F63B9FC84" blockId="2.[151,1436,151,1096]" pageId="2">Limbs short, pentadactyl; second and fourth fingers equal in length, third longest; fore and hind limbs distinctly separated when adpressed along body wall; scales on dorsal surface of base of fourth toe in three rows, reducing to two at third or fourth scale and with terminal two scales single; subdigital lamellae smooth, numbering five under fourth finger and six under fourth toe</paragraph>
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<paragraph id="8BC0369FAC6B1624AAC0FCAF6460FB9B" blockId="2.[151,1436,151,1096]" pageId="2">Coloration in alcohol. Free margins of upper and lower eyelids not edged in white; dorsum and tail base bronze brown with fine dark dots; dorsal tail tip dark brown; upper lateral head and flank with narrow black stripe, one scale wide, running from nostril to anterior corner of eye and from behind the eye through hindlimb to tail base, interrupted by small light spots from posterior half of body; dorsolateral light stripe present on neck, shoulder and anterior part of dorsum; supralabials and infralabials with dark bars on sutures; arms and legs dark brown with light spots; chin and throat cream; venter and tail base cream; underside of tail tip with very small dark brown dots.</paragraph>
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</subSubSection>
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<caption id="DF006617AC6B1624AA90F84E600CF86E" httpUri="https://zenodo.org/record/223678/files/figure.png" pageId="2" targetBox="[151,1436,1156,1902]" targetPageId="2">
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<paragraph id="8BC0369FAC6B1624AA90F84E600CF86E" blockId="2.[151,1436,1925,1981]" pageId="2">
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<emphasis id="B90BEA8DAC6B1624AA90F84E6017F84D" bold="true" box="[151,287,1926,1950]" pageId="2">FIGURE 1.</emphasis>
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Dorsal view of the holotype (IEBR A.2013.22) of
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<taxonomicName id="4C7F4D1CAC6B1624A971F84D656EF84E" box="[886,1126,1925,1949]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="sheai" status="sp. nov.">
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<emphasis id="B90BEA8DAC6B1624A971F84D656EF84E" box="[886,1126,1925,1949]" italics="true" pageId="2">Sphenomorphus sheai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857F6AC6B1624AE73F84E65C4F84E" box="[1140,1228,1926,1949]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
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Photo R. W. Van Devender.
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</paragraph>
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</caption>
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<caption id="DF006617AC6A1625AA90F9EE6076F98E" httpUri="https://zenodo.org/record/223679/files/figure.png" pageId="3" targetBox="[280,1276,205,1517]" targetPageId="3">
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<paragraph id="8BC0369FAC6A1625AA90F9EE6076F98E" blockId="3.[151,1437,1573,1629]" pageId="3">
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<emphasis id="B90BEA8DAC6A1625AA90F9EE6011F9ED" bold="true" box="[151,281,1574,1598]" pageId="3">FIGURE 2.</emphasis>
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Dorsal (A) and lateral (B) views of the head of
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<taxonomicName id="4C7F4D1CAC6A1625A926F9ED6504F9EE" box="[801,1036,1573,1597]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="sheai" status="sp. nov.">
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<emphasis id="B90BEA8DAC6A1625A926F9ED6504F9EE" box="[801,1036,1573,1597]" italics="true" pageId="3">Sphenomorphus sheai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857F6AC6A1625AE14F9EE656DF9EE" box="[1043,1125,1574,1597]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
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(holotype IEBR A.2013.22). Photos T. Q. Nguyen.
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</paragraph>
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</caption>
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<subSubSection id="C3656514AC6A1625AAC0F96663CFF8BB" pageId="3" type="distribution">
|
||
<paragraph id="8BC0369FAC6A1625AAC0F96662B9F93C" blockId="3.[151,1436,1710,2016]" pageId="3">
|
||
<emphasis id="B90BEA8DAC6A1625AAC0F9666068F914" bold="true" box="[199,352,1710,1735]" pageId="3">Distribution.</emphasis>
|
||
<taxonomicName id="4C7F4D1CAC6A1625AB6FF9666368F914" box="[360,608,1710,1735]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="sheai" status="sp. nov.">
|
||
<emphasis id="B90BEA8DAC6A1625AB6FF9666368F914" box="[360,608,1710,1735]" italics="true" pageId="3">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F6AC6A1625A86FF97863C8F914" box="[616,704,1712,1735]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus id="54C4883DAC6A1625A9DDF9786502F91B" box="[986,1034,1712,1736]" pageId="3">type</typeStatus>
|
||
locality in the border area between Quang Ngai and Kon Tumprovince, southern Central
|
||
<collectingCountry id="F368760FAC6A1625A8F6F91E6258F93C" box="[753,848,1750,1775]" name="Vietnam" pageId="3">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="13442A1AAC6A1625A958F91E62ABF93C" box="[863,931,1750,1775]" captionStart="FIGURE 3" captionStartId="4.[151,255,1980,2004]" captionTargetBox="[222,1366,183,1965]" captionTargetId="figure@4.[222,1369,183,1968]" captionTargetPageId="4" captionText="FIGURE 3. Map showing the type locality (red circle) of Sphenomorphus sheai in the border area between Quang Ngai and Kon Tum provinces, Vietnam." httpUri="https://zenodo.org/record/223680/files/figure.png" pageId="3">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC0369FAC6A1625AAC0F93763CFF8BB" blockId="3.[151,1436,1710,2016]" pageId="3">
|
||
<emphasis id="B90BEA8DAC6A1625AAC0F9376080F8CB" bold="true" box="[199,392,1791,1816]" pageId="3">Natural history.</emphasis>
|
||
The
|
||
<typeStatus id="54C4883DAC6A1625ABCFF9376322F8CB" box="[456,554,1791,1816]" pageId="3" type="holotype">holotype</typeStatus>
|
||
was collected about two hours after dark among leaf litter on a steep slope about three meters above a small stream surrounded by secondary forest at an elevation of
|
||
<quantity id="4C879B7AAC6A1625AEA6F8EF65F2F8EC" box="[1185,1274,1831,1856]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="3" unit="m" value="1025.0">1025 m</quantity>
|
||
. The
|
||
<typeStatus id="54C4883DAC6A1625AF3DF8EF6494F893" box="[1338,1436,1831,1856]" pageId="3" type="holotype">holotype</typeStatus>
|
||
contained four small white follicles in each ovary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656514AC6A1623AAC0F8BF608DFD92" lastPageId="5" pageId="3" type="etymology">
|
||
<paragraph id="8BC0369FAC6A1625AAC0F8BF6480F80C" blockId="3.[151,1436,1710,2016]" pageId="3">
|
||
<emphasis id="B90BEA8DAC6A1625AAC0F8BF6046F843" bold="true" box="[199,334,1911,1936]" pageId="3">Etymology.</emphasis>
|
||
The new species is named in honour of Dr. Glenn Shea from the University of Sydney,
|
||
<collectingCountry id="F368760FAC6A1625AF29F8BF649FF843" box="[1326,1431,1911,1936]" name="Australia" pageId="3">Australia</collectingCountry>
|
||
, in recognition of his outstanding contributions towards the systematics of scincid lizards. For the common names we suggest Shea’s Forest Skink (English), Thằn lằn phê-nô shea (Vietnamese), and Sheas Bodenskink (
|
||
<collectingCountry id="F368760FAC6A1625AF1BF80F6475F80C" box="[1308,1405,1991,2015]" name="Germany" pageId="3">German</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF006617AC6D1622AA90F874630FF820" httpUri="https://zenodo.org/record/223680/files/figure.png" pageId="4" targetBox="[222,1366,183,1965]" targetPageId="4">
|
||
<paragraph id="8BC0369FAC6D1622AA90F874630FF820" blockId="4.[151,1436,1979,2035]" pageId="4">
|
||
<emphasis id="B90BEA8DAC6D1622AA90F8746011F807" bold="true" box="[151,281,1980,2004]" pageId="4">FIGURE 3.</emphasis>
|
||
Map showing the type locality (red circle) of
|
||
<taxonomicName id="4C7F4D1CAC6D1622A901F87362F8F800" box="[774,1008,1979,2003]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="sheai">
|
||
<emphasis id="B90BEA8DAC6D1622A901F87362F8F800" box="[774,1008,1979,2003]" italics="true" pageId="4">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
in the border area between Quang Ngai and Kon Tum provinces, Vietnam.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0369FAC6C1623AAC0FF5F608DFD92" blockId="5.[151,1436,151,941]" pageId="5">
|
||
<emphasis id="B90BEA8DAC6C1623AAC0FF5F6064FF63" bold="true" box="[199,364,151,176]" pageId="5">Comparisons.</emphasis>
|
||
Based on specimen examination (Appendix) and data from the literature (e.g., Boulenger 1887; Smith 1935; Bourret 1939, 2009; Taylor 1963; Darevsky & Orlov 2005; Nguyen
|
||
<emphasis id="B90BEA8DAC6C1623AE2CFF09656BFF0B" box="[1067,1123,191,216]" italics="true" pageId="5">et al.</emphasis>
|
||
2011; Luu
|
||
<emphasis id="B90BEA8DAC6C1623AEE1FF09641EFF0B" box="[1254,1302,191,216]" italics="true" pageId="5">et al</emphasis>
|
||
. 2013; Roy
|
||
<emphasis id="B90BEA8DAC6C1623AA90FF2161C3FED2" box="[151,203,232,257]" italics="true" pageId="5">et al</emphasis>
|
||
. 2013), we compare
|
||
<taxonomicName id="4C7F4D1CAC6C1623ABC2FF2063B7FED2" box="[453,703,232,257]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="sheai">
|
||
<emphasis id="B90BEA8DAC6C1623ABC2FF2063B7FED2" box="[453,703,232,257]" italics="true" pageId="5">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
with the following species of
|
||
<taxonomicName id="4C7F4D1CAC6C1623AE21FF2065D3FED2" box="[1062,1243,232,257]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90BEA8DAC6C1623AE21FF2065D3FED2" box="[1062,1243,232,257]" italics="true" pageId="5">Sphenomorphus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F368760FAC6C1623AF21FF206461FED2" box="[1318,1385,232,257]" name="China" pageId="5">China</collectingCountry>
|
||
and mainland Southeast Asia which have midbody scale counts fewer than 26 rows:
|
||
<taxonomicName id="4C7F4D1CAC6C1623AE7CFED86016FE82" authority="Boulenger" authorityName="Boulenger" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="B90BEA8DAC6C1623AE7CFED86493FEFA" box="[1147,1435,272,297]" italics="true" pageId="5">Sphenomorphus stellatus</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
(22–24 rows),
|
||
<taxonomicName id="4C7F4D1CAC6C1623ABD6FEF163C0FE83" authority="Bourret" authorityName="Bourret" box="[465,712,312,337]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tridigitus">
|
||
<emphasis id="B90BEA8DAC6C1623ABD6FEF1635EFE82" box="[465,598,312,337]" italics="true" pageId="5">S. tridigitus</emphasis>
|
||
(Bourret)
|
||
</taxonomicName>
|
||
(18–20 rows),
|
||
<taxonomicName id="4C7F4D1CAC6C1623A97DFEF16415FE82" authority="Darevsky & Orlov" authorityName="Darevsky & Orlov" box="[890,1309,312,337]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tetradactylus">
|
||
<emphasis id="B90BEA8DAC6C1623A97DFEF16523FE82" box="[890,1067,312,337]" italics="true" pageId="5">S. tetradactylus</emphasis>
|
||
(Darevsky & Orlov)
|
||
</taxonomicName>
|
||
(20 rows).
|
||
<taxonomicName id="4C7F4D1CAC6C1623AA90FEA86085FEAA" box="[151,397,352,377]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="sheai">
|
||
<emphasis id="B90BEA8DAC6C1623AA90FEA86085FEAA" box="[151,397,352,377]" italics="true" pageId="5">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C7F4D1CAC6C1623A8BCFEA9623CFEAA" box="[699,820,352,377]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="B90BEA8DAC6C1623A8BCFEA9623CFEAA" box="[699,820,352,377]" italics="true" pageId="5">S. stellatus</emphasis>
|
||
</taxonomicName>
|
||
by having a smaller size (SVL
|
||
<quantity id="4C879B7AAC6C1623AE94FEA865FFFEAB" box="[1171,1271,352,377]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="5" unit="mm" value="35.2">35.2 mm</quantity>
|
||
vs.
|
||
<quantity id="4C879B7AAC6C1623AF23FEA86494FEAB" box="[1316,1436,352,377]" metricMagnitude="-2" metricUnit="m" metricValue="6.85" metricValueMax="8.0" metricValueMin="5.7" pageId="5" unit="mm" value="68.5" valueMax="80.0" valueMin="57.0">57–80 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C7F4D1CAC6C1623AABFFE41603EFE72" box="[184,310,392,417]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="B90BEA8DAC6C1623AABFFE41603EFE72" box="[184,310,392,417]" italics="true" pageId="5">S. stellatus</emphasis>
|
||
</taxonomicName>
|
||
), fewer lamellae under fourth toe (six vs.
|
||
<quantity id="4C879B7AAC6C1623A935FE406292FE72" box="[818,922,392,417]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.842" metricValueMin="4.572" pageId="5" unit="in" value="20.5" valueMax="23.0" valueMin="18.0">18–23 in</quantity>
|
||
<taxonomicName id="4C7F4D1CAC6C1623A9A2FE41652BFE72" box="[933,1059,392,417]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="B90BEA8DAC6C1623A9A2FE41652BFE72" box="[933,1059,392,417]" italics="true" pageId="5">S. stellatus</emphasis>
|
||
</taxonomicName>
|
||
), fore and hind limbs distinctly separated when adpressed along body wall (vs. overlapped), and by the absence of dorsal large dark blotches (vs. presence). The new species differs from
|
||
<taxonomicName id="4C7F4D1CAC6C1623A860FE1163E2FE22" box="[615,746,472,497]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tridigitus">
|
||
<emphasis id="B90BEA8DAC6C1623A860FE1163E2FE22" box="[615,746,472,497]" italics="true" pageId="5">S. tridigitus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D1CAC6C1623A926FE1162D8FE22" box="[801,976,472,497]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tetradactylus">
|
||
<emphasis id="B90BEA8DAC6C1623A926FE1162D8FE22" box="[801,976,472,497]" italics="true" pageId="5">S. tetradactylus</emphasis>
|
||
</taxonomicName>
|
||
by having forelimbs pentydactyl (versus three and four in
|
||
<taxonomicName id="4C7F4D1CAC6C1623AB5FFDC960D3FDCA" box="[344,475,512,537]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tridigitus">
|
||
<emphasis id="B90BEA8DAC6C1623AB5FFDC960D3FDCA" box="[344,475,512,537]" italics="true" pageId="5">S. tridigitus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D1CAC6C1623A815FDC963C8FDCA" box="[530,704,512,537]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tetradactylus">
|
||
<emphasis id="B90BEA8DAC6C1623A815FDC963C8FDCA" box="[530,704,512,537]" italics="true" pageId="5">S. tetradactylus</emphasis>
|
||
</taxonomicName>
|
||
, respectively) and two primary temporals (vs. one in
|
||
<taxonomicName id="4C7F4D1CAC6C1623AF1EFDC96494FDCA" box="[1305,1436,512,537]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tridigitus">
|
||
<emphasis id="B90BEA8DAC6C1623AF1EFDC96494FDCA" box="[1305,1436,512,537]" italics="true" pageId="5">S. tridigitus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D1CAC6C1623AAC1FDE1607CFD92" box="[198,372,552,577]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="tetradactylus">
|
||
<emphasis id="B90BEA8DAC6C1623AAC1FDE1607CFD92" box="[198,372,552,577]" italics="true" pageId="5">S. tetradactylus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656514AC6C1623AAC0FD986393FC7E" pageId="5" type="description">
|
||
<paragraph id="8BC0369FAC6C1623AAC0FD986382FCCE" blockId="5.[151,1436,151,941]" pageId="5">
|
||
<taxonomicName id="4C7F4D1CAC6C1623AAC0FD9860C8FDBA" box="[199,448,592,617]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="sheai">
|
||
<emphasis id="B90BEA8DAC6C1623AAC0FD9860C8FDBA" box="[199,448,592,617]" italics="true" pageId="5">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
could be mistaken for several skink species of the genus
|
||
<taxonomicName id="4C7F4D1CAC6C1623AE58FD9965D9FDBB" box="[1119,1233,593,616]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90BEA8DAC6C1623AE58FD9965D9FDBB" box="[1119,1233,593,616]" italics="true" pageId="5">Lygosoma</emphasis>
|
||
</taxonomicName>
|
||
, which also have short limbs and midbody scale rows fewer than 26, including
|
||
<taxonomicName id="4C7F4D1CAC6C1623A988FDBE65A6FD5E" authority="Theobald" authorityName="Theobald" box="[911,1198,628,653]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="anguinum">
|
||
<emphasis id="B90BEA8DAC6C1623A988FDBE652CFD5F" box="[911,1060,629,653]" italics="true" pageId="5">L. anguinum</emphasis>
|
||
(Theobald)
|
||
</taxonomicName>
|
||
(22),
|
||
<taxonomicName id="4C7F4D1CAC6C1623AF07FDBE601AFD62" authority="Stolizcka" authorityName="Stolizcka" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="lineolatum">
|
||
<emphasis id="B90BEA8DAC6C1623AF07FDBE6494FD5E" box="[1280,1436,628,653]" italics="true" pageId="5">L. lineolatum</emphasis>
|
||
(Stolizcka)
|
||
</taxonomicName>
|
||
(22),
|
||
<taxonomicName id="4C7F4D1CAC6C1623AB5FFD51632DFD62" authority="Shreve" authorityName="Shreve" box="[344,549,664,689]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="popae">
|
||
<emphasis id="B90BEA8DAC6C1623AB5FFD5160B3FD63" box="[344,443,665,688]" italics="true" pageId="5">L. popae</emphasis>
|
||
(Shreve)
|
||
</taxonomicName>
|
||
(24),
|
||
<taxonomicName id="4C7F4D1CAC6C1623A86CFD516253FD62" authority="Gmelin" authorityName="Gmelin" box="[619,859,664,689]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="punctata">
|
||
<emphasis id="B90BEA8DAC6C1623A86CFD5163E4FD63" box="[619,748,665,688]" italics="true" pageId="5">L. punctata</emphasis>
|
||
(Gmelin)
|
||
</taxonomicName>
|
||
(24–28),
|
||
<taxonomicName id="4C7F4D1CAC6C1623A9CBFD5165ECFD62" authority="Linnaeus" authorityName="Linnaeus" box="[972,1252,664,689]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="quadrupes">
|
||
<emphasis id="B90BEA8DAC6C1623A9CBFD516569FD62" box="[972,1121,664,689]" italics="true" pageId="5">L. quadrupes</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
(24–28), and
|
||
<taxonomicName id="4C7F4D1CAC6C1623AF82FD516370FD06" authority="Geissler, Hartmann & Neang" authorityName="Geissler, Hartmann & Neang" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="veunsaiensis">
|
||
<emphasis id="B90BEA8DAC6C1623AF82FD51602CFD07" italics="true" pageId="5">L. veunsaiensis</emphasis>
|
||
Geissler, Hartmann & Neang
|
||
</taxonomicName>
|
||
(22). However,
|
||
<taxonomicName id="4C7F4D1CAC6C1623A932FD746526FD06" box="[821,1070,700,725]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="sheai">
|
||
<emphasis id="B90BEA8DAC6C1623A932FD746526FD06" box="[821,1070,700,725]" italics="true" pageId="5">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
differs from all aforementioned species by the absence of supranasals (present in
|
||
<taxonomicName id="4C7F4D1CAC6C1623A8C1FD296230FD2B" box="[710,824,737,760]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90BEA8DAC6C1623A8C1FD296230FD2B" box="[710,824,737,760]" italics="true" pageId="5">Lygosoma</emphasis>
|
||
</taxonomicName>
|
||
) and having the first supralabial fused with nasal (not fused in
|
||
<taxonomicName id="4C7F4D1CAC6C1623AAF0FCCE6053FCCE" box="[247,347,774,797]" class="Reptilia" family="Scincidae" genus="Lygosma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90BEA8DAC6C1623AAF0FCCE6053FCCE" box="[247,347,774,797]" italics="true" pageId="5">Lygosma</emphasis>
|
||
</taxonomicName>
|
||
) (see Geissler
|
||
<emphasis id="B90BEA8DAC6C1623A804FCCE633CFCCE" box="[515,564,772,797]" italics="true" pageId="5">et al</emphasis>
|
||
. 2012).
|
||
</paragraph>
|
||
<paragraph id="8BC0369FAC6C1623AAC0FCE06393FC7E" blockId="5.[151,1436,151,941]" pageId="5">
|
||
<taxonomicName id="4C7F4D1CAC6C1623AAC0FCE060B5FC92" box="[199,445,808,833]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="sheai">
|
||
<emphasis id="B90BEA8DAC6C1623AAC0FCE060B5FC92" box="[199,445,808,833]" italics="true" pageId="5">Sphenomorphus sheai</emphasis>
|
||
</taxonomicName>
|
||
also resembles
|
||
<taxonomicName id="4C7F4D1CAC6C1623A869FCE0641BFC92" authority="Nguyen, Nguyen, Bohme & Ziegler" authorityName="Nguyen, Nguyen, Bohme & Ziegler" box="[622,1299,808,833]" class="Reptilia" family="Scincidae" genus="Scincella" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="apraefrontalis">
|
||
<emphasis id="B90BEA8DAC6C1623A869FCE06273FC92" box="[622,891,808,833]" italics="true" pageId="5">Scincella apraefrontalis</emphasis>
|
||
Nguyen, Nguyen, Böhme & Ziegler
|
||
</taxonomicName>
|
||
by having a low midbody scale count, no ear openings, and first supralabial fused with nasal; however, the new species can be distinguished from latter by the presence of prefrontals and opaque window on lower eyelid which are absent in
|
||
<taxonomicName id="4C7F4D1CAC6C1623AA90FC5C639FFC7E" authority="Nguyen et al. 2010" authorityName="Nguyen et al." authorityYear="2010" box="[151,663,916,941]" class="Reptilia" family="Scincidae" genus="Scincella" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="apraefrontalis">
|
||
<emphasis id="B90BEA8DAC6C1623AA90FC5C60ABFC7E" box="[151,419,916,941]" italics="true" pageId="5">Scincella apraefrontalis</emphasis>
|
||
(Nguyen
|
||
<emphasis id="B90BEA8DAC6C1623A813FC5E6344FC7E" box="[532,588,916,941]" italics="true" pageId="5">et al.</emphasis>
|
||
2010)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |