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<document id="7175D0F15877136607F9A9325FC6D453" ID-CLB-Dataset="20907" ID-DOI="10.11646/zootaxa.5091.4.6" ID-GBIF-Dataset="78f4b69d-fa47-4485-923a-816d823d9686" ID-ISSN="1175-5326" ID-Zenodo-Dep="5864259" ID-ZooBank="8C277AFD-BFE9-407E-B0C8-6DCF5F1BA9FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642423292860" checkinUser="plazi" docAuthor="Osawa, Masayuki &amp; Sato, Taigi" docDate="2022" docId="03D67B44FF94E46215E36846C51BF8AB" docLanguage="en" docName="zootaxa.5091.4.6.pdf" docOrigin="Zootaxa 5091 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polyonyx deezi Osawa &amp; Sato 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="594" masterDocId="FFEF033CFF95E46515746929C17AFFDA" masterDocTitle="A distinctive new species of the genus Polyonyx Stimpson, 1858 (Crustacea Decapoda: Anomura: Porcellanidae) from Okinawa, southwestern Japan" masterLastPageNumber="597" masterPageNumber="587" pageNumber="588" updateTime="1699302582326" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8B9D08127EC087EDFD5AFD07F3BC9101" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5E72E3341042E36D5876650A0FFA7A9B">
<mods:title id="36EF838F6D37E9B750711C218037E2DE">A distinctive new species of the genus Polyonyx Stimpson, 1858 (Crustacea Decapoda: Anomura: Porcellanidae) from Okinawa, southwestern Japan</mods:title>
</mods:titleInfo>
<mods:name id="9F79A02564ADCC34B3FF695EC8220025" type="personal">
<mods:role id="8ACAE50922D784DB9216A4A6790BACF5">
<mods:roleTerm id="F1E2BEABA389B47F73A2205E10D57FDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6785EE8A441AD2490ED9911A20A1BE39">Osawa, Masayuki</mods:namePart>
<mods:nameIdentifier id="3207D7D8685E8D9E0F20F19715DAB3B6" type="ORCID">0000-0001-8600-7550</mods:nameIdentifier>
<mods:affiliation id="1260EBF3466B7EC94685AE59E4A2A771">Estuary Research Center, Shimane University, 1060 Nishikawatsu-cho, Matsue, Shimane 690 - 8504, Japan. osawam @ soc. shimane-u. ac. jp; https: // orcid. org / 0000 - 0001 - 8600 - 7550</mods:affiliation>
<mods:nameIdentifier id="FDD9FFCCD550A61F605519D1FEC5FF33" type="email">osawam@soc.shimane-u.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:name id="B5C49C998B0CC482F651C6DD76B1202A" type="personal">
<mods:role id="570E8F610E494B6FBC766CFBEA184401">
<mods:roleTerm id="B1A5144940C37DFD51766695D436C856">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75F8FBB20F5F5F79243AAF2C33EB75DB">Sato, Taigi</mods:namePart>
<mods:nameIdentifier id="0D0E108F45453794C9268454061EDBAA" type="ORCID">0000-0002-8854-7339</mods:nameIdentifier>
<mods:affiliation id="3E00D874378FC7300BE371BCF113229C">Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Nakagami, Okinawa 903 - 0213, Japan. manzyuugani @ gmail. com; https: // orcid. org / 0000 - 0002 - 8854 - 7339</mods:affiliation>
<mods:nameIdentifier id="99CA7F5D7F69C45E04ED9DCAA1AEABA6" type="email">manzyuugani@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="04C3622120A7D7117C30A169C7D26387">text</mods:typeOfResource>
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<mods:titleInfo id="9CD52CFB9D8722968D2E32998F06ECBE">
<mods:title id="012E0779D0398324E387CF1AFC5A67CF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D525D5E0F79753A67BEF98A95CDF2FF2">
<mods:date id="8834E0F7DCC24D6C99E288D32D01F043">2022</mods:date>
<mods:detail id="DE0AD1F2C48EEEEBDC27F554EE1CC747" type="pubDate">
<mods:number id="4F9D63F31C6EAD7B0EDA2C11031DA0F5">2022-01-17</mods:number>
</mods:detail>
<mods:detail id="44EEBA15ED702A27A6A72D2E8EC49D68" type="volume">
<mods:number id="B6FA406005886878AACD3525B70F47B9">5091</mods:number>
</mods:detail>
<mods:detail id="77A4AF3EBFDB397725C74E30ED6987D1" type="issue">
<mods:number id="9396624B14BE68338ED50D655FA7093F">4</mods:number>
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<mods:classification id="C08AB4464E649664F22748A9E2A4CE1B">journal article</mods:classification>
<mods:identifier id="068C65626BD41DE199F518A51402B3B6" type="CLB-Dataset">20907</mods:identifier>
<mods:identifier id="CFC94FFD8F0BEFC46A1B2BBB9C06E856" type="DOI">10.11646/zootaxa.5091.4.6</mods:identifier>
<mods:identifier id="EED4231D98481FCA608D63B492141612" type="GBIF-Dataset">78f4b69d-fa47-4485-923a-816d823d9686</mods:identifier>
<mods:identifier id="361ED3045133031EC79DEB0D20EC728F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6B2FD679ED36B9CC68B01E4AD62DBF58" type="ZooBank">8C277AFD-BFE9-407E-B0C8-6DCF5F1BA9FC</mods:identifier>
</mods:mods>
<treatment id="03D67B44FF94E46215E36846C51BF8AB" ID-DOI="http://doi.org/10.5281/zenodo.5915191" ID-GBIF-Taxon="192716476" ID-Zenodo-Dep="5915191" LSID="urn:lsid:plazi:treatment:03D67B44FF94E46215E36846C51BF8AB" httpUri="http://treatment.plazi.org/id/03D67B44FF94E46215E36846C51BF8AB" lastPageId="7" lastPageNumber="594" pageId="1" pageNumber="588">
<subSubSection id="C36599D9FF94E46415E36846C3ACFE77" pageId="1" pageNumber="588" type="nomenclature">
<paragraph id="8BC0CA52FF94E46415E36846C0EFFE53" blockId="1.[151,726,367,465]" box="[151,405,367,394]" pageId="1" pageNumber="588">
<heading id="D0887D3EFF94E46415E36846C0EFFE53" bold="true" box="[151,405,367,394]" fontSize="11" level="1" pageId="1" pageNumber="588" reason="1">
<emphasis id="B90B1640FF94E46415E36846C0EFFE53" bold="true" box="[151,405,367,394]" pageId="1" pageNumber="588">
<taxonomicName id="4C7FB1D1FF94E46415E36846C031FE50" ID-CoL="9a64bcac-d3c4-4bd8-bc07-038e76f8ccba" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[151,331,367,394]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF94E46415E36846C031FE50" bold="true" box="[151,331,367,394]" italics="true" pageId="1" pageNumber="588">Polyonyx deezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238AB3BFF94E46414276846C0EFFE53" box="[339,405,367,393]" pageId="1" pageNumber="588" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0CA52FF94E46415E368BDC3ACFE77" blockId="1.[151,726,367,465]" box="[151,726,402,430]" pageId="1" pageNumber="588">[New Japanese name: Kawari-yadori-kanidamashi]</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF94E46415E3689EC068FE0B" box="[151,274,439,465]" pageId="1" pageNumber="588" type="description">
<paragraph id="8BC0CA52FF94E46415E3689EC068FE0B" blockId="1.[151,726,367,465]" box="[151,274,439,465]" pageId="1" pageNumber="588">
(
<figureCitation id="1344D6D7FF94E46415EB689EC070FE0B" box="[159,266,439,465]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1725,1749]" captionStartId-1="3.[151,250,1902,1926]" captionStartId-2="4.[151,250,1836,1860]" captionStartId-3="5.[151,250,1070,1094]" captionTargetBox-0="[169,1412,431,1688]" captionTargetBox-1="[198,1389,189,1871]" captionTargetBox-2="[220,1368,196,1801]" captionTargetBox-3="[165,1423,181,1046]" captionTargetId-0="figure-118@2.[151,1436,417,1702]" captionTargetId-1="figure-17@3.[180,1408,181,1878]" captionTargetId-2="figure-18@4.[208,1379,181,1812]" captionTargetId-3="figure-17@5.[165,1423,181,1046]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." captionText-1="FIGURE 2. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Left larger cheliped. A, entire, dorsal view; B, carpus and distal part of merus, perpendicular dorsal view; C, chela, perpendicular dorsal view; D, entire, ventral view; E, chela, anterior margin, ventro-anterior view; F, distal part of chela, perpendicular ventral view. For setae, only soft plumose setae drawn and shown simplified. Scale bar = 1.0 mm." captionText-2="FIGURE 3. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, smaller cheliped, merus to chela, dorsal view (only soft plumose setae on dorso-anterior surface of chela drawn); B, same, distal part of chela, perpendicular ventral view; C, left second pereopod, lateral view; D, same, merus, mesial view; E, same, dactylus and distal part of propodus, lateral view (setae not drawn); F, right second pereopod, propodus, lateral view (setae not drawn); G, left third pereopod, lateral view (granules, ridges, and setae not drawn); I, left fourth pereopod, lateral view (granules, ridges, and setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." captionText-3="FIGURE 4. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Entire animal, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.5864261" figureDoi-1="http://doi.org/10.5281/zenodo.5864265" figureDoi-2="http://doi.org/10.5281/zenodo.5864269" figureDoi-3="http://doi.org/10.5281/zenodo.5864273" httpUri-0="https://zenodo.org/record/5864261/files/figure.png" httpUri-1="https://zenodo.org/record/5864265/files/figure.png" httpUri-2="https://zenodo.org/record/5864269/files/figure.png" httpUri-3="https://zenodo.org/record/5864273/files/figure.png" pageId="1" pageNumber="588">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF94E46415E368D6C21DFDBB" pageId="1" pageNumber="588" type="materials_examined">
<paragraph id="8BC0CA52FF94E46415E368D6C21DFDBB" blockId="1.[151,1437,511,2014]" pageId="1" pageNumber="588">
<emphasis id="B90B1640FF94E46415E368D6C03FFDC3" bold="true" box="[151,325,511,537]" pageId="1" pageNumber="588">Type material.</emphasis>
<materialsCitation id="3B17C00FFF94E464143868D6C5DCFDE7" ID-GBIF-Occurrence="3459548302" collectingDate="2021-02-21" collectionCode="T" county="Sakimotobu" elevation="10" location="Island" municipality="Motobu" pageId="1" pageNumber="588" specimenCode="RUMF-ZC 7601" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="54C474F0FF94E464143868D6C0C3FDC3" box="[332,441,511,537]" pageId="1" pageNumber="588" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBD96229FF94E46414B768D6C3F8FDC3" box="[451,642,511,537]" collectionCode="RUMF-ZC" pageId="1" pageNumber="588">RUMF-ZC 7601</specimenCode>
, male (cl
<quantity id="4C8767B7FF94E464179B68D6C232FDC0" box="[751,840,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="588" unit="mm" value="2.3">2.3 mm</quantity>
, cw 3.0 mm),
<collectingCounty id="62A1B2DEFF94E464169C68D6C508FDC3" box="[1000,1138,511,537]" pageId="1" pageNumber="588">Sakimotobu</collectingCounty>
,
<collectingMunicipality id="6BA45028FF94E464110A68D6C5A0FDC3" box="[1150,1242,511,537]" pageId="1" pageNumber="588">Motobu</collectingMunicipality>
,
<collectingRegion id="49BB04B0FF94E464119168D6C430FDC3" box="[1253,1354,511,537]" country="Japan" name="Okinawa" pageId="1" pageNumber="588">Okinawa</collectingRegion>
<location id="8EA09C89FF94E464102568D6C4E2FDC3" LSID="urn:lsid:plazi:treatment:03D67B44FF94E46215E36846C51BF8AB:8EA09C89FF94E464102568D6C4E2FDC3" box="[1361,1432,511,537]" county="Sakimotobu" municipality="Motobu" name="Island" pageId="1" pageNumber="588" stateProvince="Okinawa">Island</location>
,
<quantity id="4C8767B7FF94E46415E36B0AC194FDE4" box="[151,238,547,574]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="588" unit="m" value="10.0" valueMax="12.0" valueMin="8.0">
<elevation id="00522D61FF94E46415E36B0AC194FDE4" box="[151,238,547,574]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="588" unit="m" value="10.0" valueMax="12.0" valueMin="8.0">8–12 m</elevation>
</quantity>
, coral sand, associated with
<taxonomicName id="4C7FB1D1FF94E464175D6B0AC392FDE4" box="[553,744,547,574]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="1" pageNumber="588" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="B90B1640FF94E464175D6B0AC3BAFDE7" box="[553,704,547,573]" italics="true" pageId="1" pageNumber="588">Chaetopterus</emphasis>
sp.
</taxonomicName>
,
<date id="FFC1EC92FF94E46417806B0AC2C7FDE7" box="[756,957,547,573]" pageId="1" pageNumber="588" value="2021-02-21">
<collectingDate id="EF85157AFF94E46417806B0AC2C7FDE7" box="[756,957,547,573]" pageId="1" pageNumber="588" value="2021-02-21">21 February 2021</collectingDate>
</date>
, collected by
<collectionCode id="ED6E5297FF94E46411206B0DC51CFDE4" box="[1108,1126,548,574]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="588">T</collectionCode>
. Sato
</materialsCitation>
.
<materialsCitation id="3B17C00FFF94E46411C46B0DC21EFDBB" ID-GBIF-Occurrence="3459548301" collectingDate="2021-02-21" collectionCode="RUMF-ZC" county="Sakimotobu" elevation="10" location="Island" municipality="Motobu" pageId="1" pageNumber="588" specimenCode="RUMF-ZC 7602" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<typeStatus id="54C474F0FF94E46411C46B0DC46CFDE4" box="[1200,1302,548,574]" pageId="1" pageNumber="588" type="paratype">Paratype</typeStatus>
:
<specimenCode id="DBD96229FF94E46410546B0AC1A8FDBB" collectionCode="RUMF-ZC" pageId="1" pageNumber="588">RUMF-ZC 7602</specimenCode>
,
<specimenCount id="9D7901DBFF94E46415A96B6EC052FDBB" box="[221,296,583,609]" pageId="1" pageNumber="588" type="male">1 male</specimenCount>
(cl
<quantity id="4C8767B7FF94E46414206B6EC0D6FDB8" box="[340,428,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="588" unit="mm" value="2.2">2.2 mm</quantity>
, cw
<quantity id="4C8767B7FF94E46414AB6B6EC343FDB8" box="[479,569,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="588" unit="mm" value="2.8">2.8 mm</quantity>
), same data as in
<typeStatus id="54C474F0FF94E464178A6B6EC21EFDBB" box="[766,868,583,609]" pageId="1" pageNumber="588" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF94E46415B36B42C267FB7B" pageId="1" pageNumber="588" type="diagnosis">
<paragraph id="8BC0CA52FF94E46415B36B42C267FB7B" blockId="1.[151,1437,511,2014]" pageId="1" pageNumber="588">
<emphasis id="B90B1640FF94E46415B36B42C038FD5F" bold="true" box="[199,322,619,645]" pageId="1" pageNumber="588">Diagnosis.</emphasis>
Numerous granules and short granular ridges on dorsal surfaces of carapace and chelipeds. Carapace with median branchial margins being bluntly angular and produced laterally; gastric, branchial, cardiac regions with 1, 2, 1 distinct protuberances, respectively. Rostrum produced anteriorly in dorsal view, trilobed, median lobe roundly triangular; anterior margin with row of sparse short setae. Third thoracic sternite slightly concave on anteromedian margin. Telson distal plates each distinctly longer than broad. Antennal first article approximately 0.8 length of combined second to fourth articles. Third maxilliped merus with roundly subrectangular lobe on ventrolateral margin. Chelipeds somewhat unequal in size; meri each with broad, rounded lobe distally on dorsoanterior margin; carpi and palms each with at least 1 distinct protuberance on dorsal surface; carpi comparatively short, 1.3–1.4 times as long as broad, dorso-anterior margin gently convex, without distinct row of setae; chelae with dense, soft plumose setae on dorso-anterior surface; palms inflated dorsoventrally; dactylus of larger cheliped short, 0.3–0.4 length of chela. Ambulatory legs moderately short, numerous short ridges on lateral surfaces of meri and carpi; meri wide, 1.5–1.8 times as long as high in mesial view, strongly convex on dorsal margins; ventral surfaces concave distally to receive ventral margins of carpi; propodi comparatively short, 2.8–3.1 times as long as high, ventral margins each with 1 or 2 corneous spines in addition to 2 corneous spines at terminal end; dactyli each with dorsal claw much smaller than ventral claw, ventral margins with 2 teeth each bearing small corneous spine. Male with pair of gonopods on second abdominal somite.
</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF94E46315B36D82C583F9F7" lastPageId="6" lastPageNumber="593" pageId="1" pageNumber="588" type="description">
<paragraph id="8BC0CA52FF94E46415B36D82C0E3F98B" blockId="1.[151,1437,511,2014]" pageId="1" pageNumber="588">
<emphasis id="B90B1640FF94E46415B36D82C023FB1F" bold="true" box="[199,345,1195,1221]" pageId="1" pageNumber="588">Description.</emphasis>
Carapace (
<figureCitation id="1344D6D7FF94E46414AA6D82C326FB1C" box="[478,604,1195,1222]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1A, B</figureCitation>
) hexagonal in general outline, 1.2–1.3 times as broad as long, broadest on median branchial margins. Dorsal surface slightly convex, covered with numerous granules and short granular ridges and sparse short setae; 5 transverse protuberances present (1 on mesogastric region, 2 mesially on branchial region, and 1 cardiac region, respectively). Protogastric ridges well demarcated, elevated. Cervical grooves shallow. Hepatic margins convex. Branchial margins divergent on anterior half and convergent on posterior half, with row of short and moderately long plumose setae; median margin bluntly angular, produced laterally; anterior half margin carinate, thin, with shallow sulcus mesially; posterior half gently convex. Posterior margin with elevated ridge of granules. Rostrum (
<figureCitation id="1344D6D7FF94E46414006C8EC097FA1B" box="[372,493,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1A, C</figureCitation>
) moderately broad, reaching beyond anterior margin of eyes, produced anteriorly in dorsal view, ventrally bent toward anterior tip, trilobate in anterior view; median lobe triangular, with rounded apex, slightly overreaching rounded lateral lobes, with shallow median longitudinal groove on dorsal surface; anterior margin with row of sparse short setae. Orbits moderately shallow; supra-orbital margins slightly concave; lateral orbital angles rounded.
</paragraph>
<paragraph id="8BC0CA52FF94E46415B36F72C017F967" blockId="1.[151,1437,511,2014]" pageId="1" pageNumber="588">
Pterygostomial flaps (
<figureCitation id="1344D6D7FF94E46414B26F72C365F9AF" box="[454,543,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1B</figureCitation>
) entire; surface with short ridges on dorsal half and long undulate ridge on longitudinal midline; scattered plumose setae, numerous ventrally; dorso-anterior surface somewhat concave, rounded anteriorly.
</paragraph>
<paragraph id="8BC0CA52FF94E46415B36FEEC43BF894" blockId="1.[151,1437,511,2014]" pageId="1" pageNumber="588">
Third thoracic sternite (
<figureCitation id="1344D6D7FF94E46414A26FEEC354F93B" box="[470,558,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1D</figureCitation>
) 3.5 times as broad as long measured on midline; surface somewhat convex medially, with few granules anterolaterally; anterior margin slightly concave on median part, with row of sparse setae; lateral lobes narrow, produced, each with rounded apex. Fourth thoracic sternite with anterior margin moderately concave; surface depressed medially, with 2 transverse rows of small tubercles, anterior row bearing setae.
</paragraph>
<paragraph id="8BC0CA52FF94E46415B36E7EC397F84F" blockId="1.[151,1437,511,2014]" pageId="1" pageNumber="588">
Telson (
<figureCitation id="1344D6D7FF94E46414506E7EC000F8AB" box="[292,378,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1E</figureCitation>
) slightly broader than long, composed of 7 plates; lateroproximal plates much smaller than others; distal plates each distinctly longer than broad.
</paragraph>
<paragraph id="8BC0CA52FF94E46415B36EB6C447F863" blockId="1.[151,1437,511,2014]" box="[199,1341,1951,1978]" pageId="1" pageNumber="588">
Ocular peduncles (
<figureCitation id="1344D6D7FF94E46414EE6EB6C34FF860" box="[410,565,1951,1978]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1A, G, H</figureCitation>
) moderately small, short; dorsal extension onto cornea low, rounded.
</paragraph>
<paragraph id="8BC0CA52FF94E46715B36EEAC545FF0F" blockId="1.[151,1437,511,2014]" lastBlockId="2.[151,1436,151,393]" lastPageId="2" lastPageNumber="589" pageId="1" pageNumber="588">
Basal article of antennular peduncle (
<figureCitation id="1344D6D7FF94E464171C6EEAC3C0F807" box="[616,698,1987,2013]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="1" pageNumber="588">Fig. 1F</figureCitation>
) approximately as long as broad; anterior margin slightly convex, somewhat undulate; ventral surface with row of small tubercles on anterior margin and transversely oblique row of small tubercles laterally; ventro-anterior face slightly concave, with many granules.
</paragraph>
<paragraph id="8BC0CA52FF97E46715B369F6C570FE53" blockId="2.[151,1436,151,393]" pageId="2" pageNumber="589">
Antennal peduncle (
<figureCitation id="1344D6D7FF97E46714DF69F6C331FF23" box="[427,587,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="2" pageNumber="589">Fig. 1A, G, H</figureCitation>
) slender. First article broad, largest, approximately 0.8 length of combined second to fourth articles, produced forward in lateral view, broadly in contact with lower orbital margin; surface shallowly concave, with numerous granules and blunt ridge along ventral margin; anterior margin tapering, terminating in rounded apex. Second and third articles roundly rectangular, with granules and short granular ridges marginally; third article elongate. Fourth article short, rounded, nearly smooth.
</paragraph>
<caption id="DF009ADAFF97E46715E36F94C5C0F877" ID-DOI="http://doi.org/10.5281/zenodo.5864261" ID-Zenodo-Dep="5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="2" pageNumber="589" startId="2.[151,250,1725,1749]" targetBox="[169,1412,431,1688]" targetPageId="2">
<paragraph id="8BC0CA52FF97E46715E36F94C5C0F877" blockId="2.[151,1436,1725,1966]" pageId="2" pageNumber="589">
<emphasis id="B90B1640FF97E46715E36F94C06EF90F" bold="true" box="[151,276,1725,1749]" pageId="2" pageNumber="589">FIGURE 1.</emphasis>
<taxonomicName id="4C7FB1D1FF97E467146F6F94C0CFF90F" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[283,437,1725,1749]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF97E467146F6F94C0CFF90F" box="[283,437,1725,1749]" italics="true" pageId="2" pageNumber="589">Polyonyx deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF97E46714C86F97C08CF90C" bold="true" box="[444,502,1726,1750]" pageId="2" pageNumber="589">
<taxonomicNameLabel id="A238AB3BFF97E46714C86F97C08CF90C" box="[444,502,1726,1750]" pageId="2" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF009ADAFF96E46615E36E47C3B0F828" ID-DOI="http://doi.org/10.5281/zenodo.5864265" ID-Zenodo-Dep="5864265" httpUri="https://zenodo.org/record/5864265/files/figure.png" pageId="3" pageNumber="590" startId="3.[151,250,1902,1926]" targetBox="[198,1389,189,1871]" targetPageId="3">
<paragraph id="8BC0CA52FF96E46615E36E47C3B0F828" blockId="3.[151,1436,1902,2034]" pageId="3" pageNumber="590">
<emphasis id="B90B1640FF96E46615E36E47C06EF85C" bold="true" box="[151,276,1902,1926]" pageId="3" pageNumber="590">FIGURE 2.</emphasis>
<taxonomicName id="4C7FB1D1FF96E466146F6E47C0CEF85C" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[283,436,1902,1926]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF96E466146F6E47C0CEF85C" box="[283,436,1902,1926]" italics="true" pageId="3" pageNumber="590">Polyonyx deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF96E46614CF6E47C08EF85C" bold="true" box="[443,500,1902,1926]" pageId="3" pageNumber="590">
<taxonomicNameLabel id="A238AB3BFF96E46614CF6E47C08EF85C" box="[443,500,1902,1926]" pageId="3" pageNumber="590" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Left larger cheliped. A, entire, dorsal view; B, carpus and distal part of merus, perpendicular dorsal view; C, chela, perpendicular dorsal view; D, entire, ventral view; E, chela, anterior margin, ventro-anterior view; F, distal part of chela, perpendicular ventral view. For setae, only soft plumose setae drawn and shown simplified. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF009ADAFF91E46115E36E05C3BBF822" ID-DOI="http://doi.org/10.5281/zenodo.5864269" ID-Zenodo-Dep="5864269" httpUri="https://zenodo.org/record/5864269/files/figure.png" pageId="4" pageNumber="591" startId="4.[151,250,1836,1860]" targetBox="[220,1368,196,1801]" targetPageId="4">
<paragraph id="8BC0CA52FF91E46115E36E05C3BBF822" blockId="4.[151,1437,1836,2040]" pageId="4" pageNumber="591">
<emphasis id="B90B1640FF91E46115E36E05C06EF89E" bold="true" box="[151,276,1836,1860]" pageId="4" pageNumber="591">FIGURE 3.</emphasis>
<taxonomicName id="4C7FB1D1FF91E46114686E05C0CFF89E" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[284,437,1836,1860]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF91E46114686E05C0CFF89E" box="[284,437,1836,1860]" italics="true" pageId="4" pageNumber="591">Polyonyx deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF91E46114C96E05C08DF89E" bold="true" box="[445,503,1836,1860]" pageId="4" pageNumber="591">
<taxonomicNameLabel id="A238AB3BFF91E46114C96E05C08DF89E" box="[445,503,1836,1860]" pageId="4" pageNumber="591" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, smaller cheliped, merus to chela, dorsal view (only soft plumose setae on dorso-anterior surface of chela drawn); B, same, distal part of chela, perpendicular ventral view; C, left second pereopod, lateral view; D, same, merus, mesial view; E, same, dactylus and distal part of propodus, lateral view (setae not drawn); F, right second pereopod, propodus, lateral view (setae not drawn); G, left third pereopod, lateral view (granules, ridges, and setae not drawn); I, left fourth pereopod, lateral view (granules, ridges, and setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF009ADAFF90E46015E36D07C456FB9C" ID-DOI="http://doi.org/10.5281/zenodo.5864273" ID-Zenodo-Dep="5864273" box="[151,1324,1070,1094]" httpUri="https://zenodo.org/record/5864273/files/figure.png" pageId="5" pageNumber="592" startId="5.[151,250,1070,1094]" targetBox="[165,1423,181,1046]" targetPageId="5">
<paragraph id="8BC0CA52FF90E46015E36D07C456FB9C" blockId="5.[151,1324,1070,1094]" box="[151,1324,1070,1094]" pageId="5" pageNumber="592">
<emphasis id="B90B1640FF90E46015E36D07C069FB9C" bold="true" box="[151,275,1070,1094]" pageId="5" pageNumber="592">FIGURE 4.</emphasis>
<taxonomicName id="4C7FB1D1FF90E460146D6D07C0C8FB9C" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[281,434,1070,1094]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="592" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF90E460146D6D07C0C8FB9C" box="[281,434,1070,1094]" italics="true" pageId="5" pageNumber="592">Polyonyx deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF90E46014CC6D07C088FB9C" bold="true" box="[440,498,1070,1094]" pageId="5" pageNumber="592">
<taxonomicNameLabel id="A238AB3BFF90E46014CC6D07C088FB9C" box="[440,498,1070,1094]" pageId="5" pageNumber="592" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Entire animal, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC0CA52FF90E46015B36D5DC3E3FA09" blockId="5.[151,1437,1140,2031]" pageId="5" pageNumber="592">
Third maxilliped (
<figureCitation id="1344D6D7FF90E46014E76D5DC083FB55" box="[403,505,1140,1167]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[169,1412,431,1688]" captionTargetId="figure-118@2.[151,1436,417,1702]" captionTargetPageId="2" captionText="FIGURE 1. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, carapace and ocular and antennal peduncles, dorsal view (granules and setae drawn only on left side); B, carapace, pterygostomial flap, and ocular and antennal peduncles, left lateral view; C, rostrum, dorso-anterior view; D, third and fourth thoracic sternites, ventral view (setae drawn only on right side); E, telson, external view; F, left basal article of antennular peduncle, ventral view; G, left ocular and antennal peduncles and anterolateral part of carapace, dorsal view; H, left ocular peduncle and first article of antennal peduncle, lateroanterior view; I, left third maxilliped, coxa to merus and exopod, lateral view (for setae, only soft plumose setae drawn); J, same, merus to dactylus, lateral view (setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864261" httpUri="https://zenodo.org/record/5864261/files/figure.png" pageId="5" pageNumber="592">Fig. 1I, J</figureCitation>
) with coxa bearing blunt projection on ventrodistal margin; distomedian projection with blunt transverse ridge, not articulated. Basis articulating with ischium, subtriangular. Ischium broad, with ventral margin strongly convex; lateral surface with sparse granules and weak longitudinal ridge along dorsal margin; dorsodistal projection blunt; long, soft plumose setae on ventrodistal surface. Merus with laminate, roundly subrectangular lobe on ventrolateral margin; lateral surface with scattered granules and short ridges. Carpus with small subtriangular projection on median part of ventral margin and with row of short blunt ridges on dorsal surface. Propodus moderately elongate, with row of short blunt ridges on dorsal surface. Dactylus short, subtriangular, nearly smooth. Merus to dactylus with long plumose setae on ventral margins. Exopod with proximal article small, rounded; distal article laminate, robust, reaching distal two-thirds of merus, constricted on median part, with row of short ridges dorsally; distal flagellum present.
</paragraph>
<paragraph id="8BC0CA52FF90E46315B36CF5C092FE53" blockId="5.[151,1437,1140,2031]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="593" pageId="5" pageNumber="592">
Chelipeds (first pereopods;
<figureCitation id="1344D6D7FF90E46017706CF5C3FFFA2D" box="[516,645,1500,1527]" captionStart="FIGURE 2" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[198,1389,189,1871]" captionTargetId="figure-17@3.[180,1408,181,1878]" captionTargetPageId="3" captionText="FIGURE 2. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Left larger cheliped. A, entire, dorsal view; B, carpus and distal part of merus, perpendicular dorsal view; C, chela, perpendicular dorsal view; D, entire, ventral view; E, chela, anterior margin, ventro-anterior view; F, distal part of chela, perpendicular ventral view. For setae, only soft plumose setae drawn and shown simplified. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864265" httpUri="https://zenodo.org/record/5864265/files/figure.png" pageId="5" pageNumber="592">Figs. 2A–F</figureCitation>
,
<figureCitation id="1344D6D7FF90E46017E66CF4C3A2FA2D" box="[658,728,1501,1527]" captionStart="FIGURE 3" captionStartId="4.[151,250,1836,1860]" captionTargetBox="[220,1368,196,1801]" captionTargetId="figure-18@4.[208,1379,181,1812]" captionTargetPageId="4" captionText="FIGURE 3. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, smaller cheliped, merus to chela, dorsal view (only soft plumose setae on dorso-anterior surface of chela drawn); B, same, distal part of chela, perpendicular ventral view; C, left second pereopod, lateral view; D, same, merus, mesial view; E, same, dactylus and distal part of propodus, lateral view (setae not drawn); F, right second pereopod, propodus, lateral view (setae not drawn); G, left third pereopod, lateral view (granules, ridges, and setae not drawn); I, left fourth pereopod, lateral view (granules, ridges, and setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864269" httpUri="https://zenodo.org/record/5864269/files/figure.png" pageId="5" pageNumber="592">3A, B</figureCitation>
) somewhat unequal in size, left thicker than right; dorsal and ventral surfaces of merus to dactylus covered with numerous granules and short granular ridges and with sparse very short setae. Larger cheliped (
<figureCitation id="1344D6D7FF90E46017656F0DC3FCF9E5" box="[529,646,1572,1599]" captionStart="FIGURE 2" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[198,1389,189,1871]" captionTargetId="figure-17@3.[180,1408,181,1878]" captionTargetPageId="3" captionText="FIGURE 2. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Left larger cheliped. A, entire, dorsal view; B, carpus and distal part of merus, perpendicular dorsal view; C, chela, perpendicular dorsal view; D, entire, ventral view; E, chela, anterior margin, ventro-anterior view; F, distal part of chela, perpendicular ventral view. For setae, only soft plumose setae drawn and shown simplified. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864265" httpUri="https://zenodo.org/record/5864265/files/figure.png" pageId="5" pageNumber="592">Fig. 2A–F</figureCitation>
) with ischium slightly crenulated but without spines on ventro-anterior margin; scattered, soft plumose setae on dorsal surface and anterior margin. Merus with blunt transverse crest submedially on dorsal surface; dorso-anterior margin with broad, rounded lobe distally; ventro-anterior margin crenulate, anterodistal corner with small tubercle; sparse, soft plumose setae on dorsoproximal part and ventroanterior distal corner. Carpus 1.3–1.4 times as long as broad, broadest on median part; dorsal surface slightly convex, with transverse protuberance on median part; dorso-anterior margin with elevated lobe, anterior margin of lobe gently convex, slightly crenulated, bearing few short setae, proximal corner of lobe roundly angled; dorsodistal margin posteriorly with roundly produced lobe bearing tuft of plumose setae; posterior margin rounded; ventro-anterior margin concave on median or distal one third part; ventrodistal margin with row of plumose setae. Chela 1.8–1.9 times as long as carpus, relatively narrow, 3.0–3.2 times as long as broad, lying on anterior side; anterior margin delimited by row of small, blunt and subacute denticles; dorso-anterior surface with dense, short and long, soft plumose setae. Palm somewhat inflated; dorsal surface convex, with 2 elongate protuberances on midline, proximal protuberance much lower than distal protuberance or obsolete; dorsoposterior margin with weak, longitudinal blunt ridge; ventral surface convex. Fixed finger with weakly curved, blunt distal claw; ventral surface with tuft of short plumose setae on base of fixed finger; cutting edge with row of blunt teeth, median tooth larger. Dactylus 0.3–0.4 length of chela, opening at oblique angle, with curved, blunt distal claw; dorsal surface with longitudinal ridge composed of small blunt and subacute denticles posteriorly on midline, dorso-anterior surface proximally with soft plumose setae; posterior margin rounded, with short blunt carina proximally; cutting edge with 2 large blunt teeth each on subproximal and subdistal parts and small blunt teeth; ventral surface proximally with sparse, short plumose setae adjacent to cutting edge.
</paragraph>
<paragraph id="8BC0CA52FF93E46315B368BAC03FFDC3" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="593">
Smaller cheliped (
<figureCitation id="1344D6D7FF93E46314EE68BAC362FE77" box="[410,536,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1836,1860]" captionTargetBox="[220,1368,196,1801]" captionTargetId="figure-18@4.[208,1379,181,1812]" captionTargetPageId="4" captionText="FIGURE 3. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, smaller cheliped, merus to chela, dorsal view (only soft plumose setae on dorso-anterior surface of chela drawn); B, same, distal part of chela, perpendicular ventral view; C, left second pereopod, lateral view; D, same, merus, mesial view; E, same, dactylus and distal part of propodus, lateral view (setae not drawn); F, right second pereopod, propodus, lateral view (setae not drawn); G, left third pereopod, lateral view (granules, ridges, and setae not drawn); I, left fourth pereopod, lateral view (granules, ridges, and setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864269" httpUri="https://zenodo.org/record/5864269/files/figure.png" pageId="6" pageNumber="593">Fig. 3A, B</figureCitation>
) generally similar to larger cheliped except for: chela proportionally broader, 2.8–3.1 times as long as broad; distal claws of fixed finger and dactylus narrower; cutting edges of fixed finger and dactylus each with entire row of much smaller blunt and subacute teeth; dactylus proportionally longer, 0.4–0.5 length of chela.
</paragraph>
<paragraph id="8BC0CA52FF93E46315B36B0AC3FFFBA7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="593">
Ambulatory legs (second to fourth pereopods, P2–P4;
<figureCitation id="1344D6D7FF93E463163C6B0AC2BEFDE7" box="[840,964,547,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1836,1860]" captionTargetBox="[220,1368,196,1801]" captionTargetId="figure-18@4.[208,1379,181,1812]" captionTargetPageId="4" captionText="FIGURE 3. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. A, smaller cheliped, merus to chela, dorsal view (only soft plumose setae on dorso-anterior surface of chela drawn); B, same, distal part of chela, perpendicular ventral view; C, left second pereopod, lateral view; D, same, merus, mesial view; E, same, dactylus and distal part of propodus, lateral view (setae not drawn); F, right second pereopod, propodus, lateral view (setae not drawn); G, left third pereopod, lateral view (granules, ridges, and setae not drawn); I, left fourth pereopod, lateral view (granules, ridges, and setae not drawn). Plumose setae shown simplified. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5864269" httpUri="https://zenodo.org/record/5864269/files/figure.png" pageId="6" pageNumber="593">Fig. 3C–H</figureCitation>
) moderately short, compressed laterally, decreasing in size posteriorly (P2 largest); P2 merus 1.1 and 1.3–1.4 times respective P3 and P4 meri lengths; P2 propodus 1.1 and 1.2–1.3 times respective P3 and P4 propodi lengths; dorsal and ventral margins with long, soft plumose setae on meri to propodi; latero-distal margins of propodi and dorsal and ventral margins of dactyli with sparse stiff setae. Ischia without weakly calcified parts on each mesial surface. Meri 1.8 (P2), 1.6 (P3), and 1.5 (P4) times as long as high in mesial view, highest on each submedian or median part; dorsal margin strongly convex, bluntly cristate; lateral surface with numerous transverse granular ridges bearing sparse short setae; mesial surfaces covered with granules and granular ridges, weakly calcified on dorsoproximal part; ventrodistal margins of lateral and mesial surfaces rounded; ventral surface concave distally to receive ventral margin of carpus, ventromesial margin thin, gently convex. Carpi moderately elongate; lateral surface with irregular rows of short ridges bearing sparse short setae; dorsodistal and ventrodistal margins rounded. Propodi 3.0–3.1 (P2), 2.8 (P3 and P4) times as long as high, broadest on each subproximal part; lateral surface with short transverse ridges bearing sparse short setae; dorsal margin weak crenulated; ventral margin slightly concave on median part, with 1 or 2 corneous spines, subdistal spine always present, subproximal spine present or absent; ventrodistal margin with 2 corneous spines subequal in size. Dactyli 0.4–0.5 times as long as propodi, each terminating in curved, sharply pointed, bifurcate claws, dorsal claw much smaller than ventral claw; ventral margin with 2 low teeth, distal tooth larger than proximal tooth, each tooth with small corneous spine.
</paragraph>
<paragraph id="8BC0CA52FF93E46315B36DAEC41FFB7B" blockId="6.[151,1437,151,2013]" box="[199,1381,1159,1185]" pageId="6" pageNumber="593">Fifth pereopod slender, chelate; propodus with numerous short simple setae and 2 subdistal hooked setae.</paragraph>
<paragraph id="8BC0CA52FF93E46315B36D82C3BCFB33" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="593">Male with pair of gonopods on second abdominal somite; endopod elongate, oval, somewhat twisted medially, rounded on distal margin; exopod small, rounded.</paragraph>
<paragraph id="8BC0CA52FF93E46315B36DDBC583F9F7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="593">
<emphasis id="B90B1640FF93E46315B36DDBC0E4FAD6" bold="true" box="[199,414,1266,1292]" pageId="6" pageNumber="593">Coloration in life.</emphasis>
Body and pereopods white in ground color. Carapace with numerous but scattered, small brown spots; some dark brown spots on anterior lateral margins and posterior margin. Basal article of antennular peduncle white; penultimate and ultimate peduncular articles and flagella translucent white. Antennal peduncle white, flagella translucent, pale brown. Third maxilliped also white in general. Merus and carpus of each cheliped with pale brown tint and with some brown blotches and small dark brown spots on dorsal surface; palm with 3 or 4, small dark brown blotches on proximal half of dorsal surface; dactylus with large dark brown blotch on proximal half posteriorly. Ambulatory legs with dark brown markings; meri each with blotch dorsally on midlength in P2 and P3 (size of blotch much smaller in P3) or no blotch (P4); carpi each with transverse blotch on half of dorsal surface; propodi each with ring (P2 and P3) or dorsal blotch (P4) on midlength. All setae whitish. See
<figureCitation id="1344D6D7FF93E46311C46F3AC58FF9F7" box="[1200,1269,1555,1581]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1094]" captionTargetBox="[165,1423,181,1046]" captionTargetId="figure-17@5.[165,1423,181,1046]" captionTargetPageId="5" captionText="FIGURE 4. Polyonyx deezi n. sp., holotype, (cl 2.3 mm, cw 3.0 mm), RUMF-ZC 7601. Entire animal, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5864273" httpUri="https://zenodo.org/record/5864273/files/figure.png" pageId="6" pageNumber="593">Fig. 4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF93E46315B36F1FC55AF8F3" pageId="6" pageNumber="593" type="distribution">
<paragraph id="8BC0CA52FF93E46315B36F1FC4E3F98B" blockId="6.[151,1437,151,2013]" box="[199,1433,1590,1617]" pageId="6" pageNumber="593">
<emphasis id="B90B1640FF93E46315B36F1FC01BF98A" bold="true" box="[199,353,1590,1616]" pageId="6" pageNumber="593">Distribution.</emphasis>
So far only known from the
<typeStatus id="54C474F0FF93E46317D16F1EC3AFF98B" box="[677,725,1591,1617]" pageId="6" pageNumber="593">type</typeStatus>
locality,
<collectingRegion id="49BB04B0FF93E46316486F1EC2DBF98B" box="[828,929,1591,1617]" country="Japan" name="Okinawa" pageId="6" pageNumber="593">Okinawa</collectingRegion>
Island, Ryukyu Islands, southwestern
<collectingCountry id="F3688AC2FF93E46310256F1EC4EEF98B" box="[1361,1428,1591,1617]" name="Japan" pageId="6" pageNumber="593">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BC0CA52FF93E46315B36F73C55AF8F3" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="593">
<emphasis id="B90B1640FF93E46315B36F73C050F9AE" bold="true" box="[199,298,1626,1652]" pageId="6" pageNumber="593">Habitat.</emphasis>
The
<specimenCount id="9D7901DBFF93E46314146F72C0C5F9AF" box="[352,447,1627,1653]" pageId="6" pageNumber="593" type="generic" typeStatus="type">two type</specimenCount>
specimens were extracted from sand bottom at a depth of
<quantity id="4C8767B7FF93E463114E6F72C5EAF9AF" box="[1082,1168,1627,1653]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="6" pageNumber="593" unit="m" value="10.0" valueMax="12.0" valueMin="8.0">8–12 m</quantity>
, coral reef environment, using a commercial suction pump (yabby pump). They were found inside tubes of an unidentified species of the polychaete genus
<taxonomicName id="4C7FB1D1FF93E463141C6F8AC3E4F967" authority="Cuvier, 1830" authorityName="Cuvier" authorityYear="1830" box="[360,670,1699,1725]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="6" pageNumber="593" phylum="Annelida" rank="genus">
<emphasis id="B90B1640FF93E463141C6F8AC085F967" box="[360,511,1699,1725]" italics="true" pageId="6" pageNumber="593">Chaetopterus</emphasis>
<bibRefCitation id="EFEEB7A3FF93E463177D6F8AC3E4F967" author="Cuvier, G." box="[521,670,1699,1725]" pageId="6" pageNumber="593" refId="ref7785" refString="Cuvier, G. (1830) Le Regne Animal Distribue d'apres son Organisation, pour Servir de Base a l'Histoire Naturelle des Animaux et d'Introduction a l'Anatomie Comparee. Tome 3. 2 nd Edition. Deterville et Crochard, Paris, 504 pp." type="book" year="1830">Cuvier, 1830</bibRefCitation>
</taxonomicName>
(family
<taxonomicName id="4C7FB1D1FF93E46316726F8AC2CFF964" box="[774,949,1699,1726]" class="Polychaeta" family="Chaetopteridae" kingdom="Animalia" pageId="6" pageNumber="593" phylum="Annelida" rank="family">Chaetopteridae</taxonomicName>
). The worm specimens are not preserved. Although the tubes obtained were broken and incomplete, they were externally covered with sand and small shell fragment and their bare surfaces have a stiff parchment-like appearance. The largest widths of the tubes were approximately
<quantity id="4C8767B7FF93E46314346E26C0C6F8F3" box="[320,444,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="6" pageNumber="593" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
. No females of the present new species were obtained.
</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF93E46315B36E1AC56FF8AB" pageId="6" pageNumber="593" type="etymology">
<paragraph id="8BC0CA52FF93E46315B36E1AC56FF8AB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="593">
<emphasis id="B90B1640FF93E46315B36E1AC034F897" bold="true" box="[199,334,1843,1869]" pageId="6" pageNumber="593">Etymology.</emphasis>
The species name is derived from the Ryukyu language
<taxonomicName id="4C7FB1D1FF93E46316BC6E1AC578F897" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[968,1026,1843,1869]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="deezi">
<emphasis id="B90B1640FF93E46316BC6E1AC578F897" box="[968,1026,1843,1869]" italics="true" pageId="6" pageNumber="593">deezi</emphasis>
</taxonomicName>
which means “very”, in reference to the clearly discriminable morphology of the new species in the genus
<taxonomicName id="4C7FB1D1FF93E46316D26E7EC574F8AB" authorityName="Stimpson" authorityYear="1858" box="[934,1038,1879,1905]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1640FF93E46316D26E7EC574F8AB" box="[934,1038,1879,1905]" italics="true" pageId="6" pageNumber="593">Polyonyx</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36599D9FF93E46215B36E52C51BF8AB" lastPageId="7" lastPageNumber="594" pageId="6" pageNumber="593" type="discussion">
<paragraph id="8BC0CA52FF93E46215B36E52C391FF0F" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1437,151,1905]" lastPageId="7" lastPageNumber="594" pageId="6" pageNumber="593">
<emphasis id="B90B1640FF93E46315B36E52C041F84F" bold="true" box="[199,315,1915,1941]" pageId="6" pageNumber="593">Remarks.</emphasis>
<taxonomicName id="4C7FB1D1FF93E46314356E52C090F84F" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[321,490,1915,1941]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF93E46314356E52C090F84F" box="[321,490,1915,1941]" italics="true" pageId="6" pageNumber="593">Polyonyx deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF93E463149B6E52C357F84F" bold="true" box="[495,557,1915,1941]" pageId="6" pageNumber="593">
<taxonomicNameLabel id="A238AB3BFF93E463149B6E52C357F84F" box="[495,557,1915,1941]" pageId="6" pageNumber="593" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is immediately distinguished from all other congeners by the median branchial margins of the carapace being bluntly angular and produced laterally and the dorsal surfaces of the carapace and of carpi and palms of the chelipeds with distinct protuberances. In other congeners, the branchial margins of the carapace are evenly convex or nearly straight and subparallel, and somewhat constricted on the median part, and no distinct protuberances on the carapace and chelipeds.
</paragraph>
<paragraph id="8BC0CA52FF92E46215B369F6C260FC87" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="594">
The new species belongs to the
<taxonomicName id="4C7FB1D1FF92E462174669F6C3DDFF22" authorityName="Stimpson" authorityYear="1858" box="[562,679,223,248]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B90B1640FF92E462174669F6C3DDFF22" box="[562,679,223,248]" italics="true" pageId="7" pageNumber="594">P. sinensis</emphasis>
</taxonomicName>
group, an informal species group of the genus defined by
<bibRefCitation id="EFEEB7A3FF92E462103469F6C19EFEC7" author="Johnson, D. S." pageId="7" pageNumber="594" pagination="95 - 118" refId="ref7962" refString="Johnson, D. S. (1958) The Indo-West Pacific species of the genus Polyonyx (Crustacea, Decapoda, Porcellanidae). The Annals of Zoology, 2, 95 - 118." type="journal article" year="1958">Johnson (1958)</bibRefCitation>
, primarily characterized by having the carapace without spines on the lateral margins, the chela with various degrees of setation on the dorso-anterior surface and the ambulatory dactyli each with the dorsal claw much smaller than the ventral claw. The species group includes 18 species from the Indo-West Pacific (cf. Osawa 2018;
<bibRefCitation id="EFEEB7A3FF92E46210246862C063FE53" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">
Osawa
<emphasis id="B90B1640FF92E46215E36846C1B7FE53" box="[151,205,367,393]" italics="true" pageId="7" pageNumber="594">et al</emphasis>
. 2018
</bibRefCitation>
;
<bibRefCitation id="EFEEB7A3FF92E462145D6846C344FE53" author="Werding, B. &amp; Hiller, A." box="[297,574,367,393]" pageId="7" pageNumber="594" pagination="25 - 34" refId="ref9086" refString="Werding, B. &amp; Hiller, A. (2019) A new species of Polyonyx (Crustacea, Anomura, Porcellanidae) inhabiting polychaete-worm tubes (Annelida, Chaetopteridae) in the Indo-West Pacific. ZooKeys, 818, 25 - 34. https: // doi. org / 10.3897 / zookeys. 818.30587" type="journal article" year="2019">Werding &amp; Hiller 2019</bibRefCitation>
):
<taxonomicName id="4C7FB1D1FF92E46217216846C207FE53" authority="Osawa, 2018" authorityName="Osawa" authorityYear="2018" box="[597,893,367,393]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B90B1640FF92E46217216846C3A3FE52" box="[597,729,367,392]" italics="true" pageId="7" pageNumber="594">P. angustus</emphasis>
Osawa, 2018
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E46216F96846C5CBFE53" authority="Osawa, 2007" authorityName="Osawa" authorityYear="2007" box="[909,1201,367,393]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="boucheti">
<emphasis id="B90B1640FF92E46216F96846C576FE53" box="[909,1036,367,393]" italics="true" pageId="7" pageNumber="594">P. boucheti</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462116C6846C5CBFE53" author="Osawa, M." box="[1048,1201,367,393]" pageId="7" pageNumber="594" pagination="1 - 49" refId="ref8350" refString="Osawa, M. (2007) Porcellanidae (Crustacea: Decapoda: Anomura) from New Caledonia and the Loyalty Islands. Zootaxa, 1548 (1), 1 - 49. https: // doi. org / 10.11646 / zootaxa. 1548.1.1" type="journal article" year="2007">Osawa, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E46211B56846C1A8FE77" authority="McNeill, 1968" authorityName="McNeill" authorityYear="1968" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="haigae">
<emphasis id="B90B1640FF92E46211B56846C457FE53" box="[1217,1325,367,393]" italics="true" pageId="7" pageNumber="594">P. haigae</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462104C6846C1A8FE77" author="McNeill, F. A." pageId="7" pageNumber="594" pagination="1 - 98" refId="ref8061" refString="McNeill, F. A. (1968) Crustacea, Decapoda &amp; Stomatopoda. Great Barrier Reef Expedition 1928 - 29, Scientific Reports, 7, 1 - 98, pls. 1 - 2. [British Museum (Natural History), London]" type="journal article" year="1968">McNeill, 1968</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462159468BAC364FE77" authority="Osawa &amp; Ng, 2016" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[224,542,403,429]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="heok">
<emphasis id="B90B1640FF92E462159468BAC049FE77" box="[224,307,403,429]" italics="true" pageId="7" pageNumber="594">P. heok</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462144868BAC364FE77" author="Osawa, M. &amp; Ng, P. K. L." box="[316,542,403,429]" pageId="7" pageNumber="594" pagination="499 - 518" refId="ref8748" refString="Osawa, M. &amp; Ng, P. K. L. (2016) Revision of Polyonyx pedalis Nobili, 1906 (Crustacea: Decapoda: Anomura: Porcellanidae), with descriptions of three new species. Raffles Bulletin of Zoology, Supplement, 34, 499 - 518." type="journal article" year="2016">Osawa &amp; Ng, 2016</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462175F68BAC2D8FE77" authority="Osawa &amp; Ng, 2016" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[555,930,403,429]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="kumejima">
<emphasis id="B90B1640FF92E462175F68BAC3CCFE77" box="[555,694,403,429]" italics="true" pageId="7" pageNumber="594">P. kumejima</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46217CB68BAC2D8FE77" author="Osawa, M. &amp; Ng, P. K. L." box="[703,930,403,429]" pageId="7" pageNumber="594" pagination="499 - 518" refId="ref8748" refString="Osawa, M. &amp; Ng, P. K. L. (2016) Revision of Polyonyx pedalis Nobili, 1906 (Crustacea: Decapoda: Anomura: Porcellanidae), with descriptions of three new species. Raffles Bulletin of Zoology, Supplement, 34, 499 - 518." type="journal article" year="2016">Osawa &amp; Ng, 2016</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E46216DB68BAC592FE77" authority="Sankolli, 1965" authorityName="Sankolli" authorityYear="1965" box="[943,1256,402,429]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="loimicola">
<emphasis id="B90B1640FF92E46216DB68BAC54CFE77" box="[943,1078,403,429]" italics="true" pageId="7" pageNumber="594">P. loimicola</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462114B68BAC592FE77" author="Sankolli, K. N." box="[1087,1256,402,429]" pageId="7" pageNumber="594" pagination="285 - 291" refId="ref8850" refString="Sankolli, K. N. (1965) On a new species of commensal porcellanid crab, Polyonyx loimicola sp. nov., from India (Crustacea, Anomura, Porcellanidae). Journal of the Bombay Natural History Society, 62, 285 - 291." type="journal article" year="1965">Sankolli, 1965</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462118268BAC06DFE0B" authority="Haig, 1965" authorityName="Haig" authorityYear="1965" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="maccullochi">
<emphasis id="B90B1640FF92E462118268BAC4E6FE77" box="[1270,1436,403,429]" italics="true" pageId="7" pageNumber="594">P. maccullochi</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46215E3689EC06DFE0B" author="Haig, J." box="[151,279,438,465]" pageId="7" pageNumber="594" pagination="97 - 117" refId="ref7831" refString="Haig, J. (1965) The Porcellanidae (Crustacea, Anomura) of Western Australia with description of four new Australian species. Journal of the Royal Society of Western Australia, 48, 97 - 117." type="journal article" year="1965">Haig, 1965</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E4621450689EC350FE0B" authority="Nobili, 1906" authorityName="Nobili" authorityYear="1906" box="[292,554,439,465]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="pedalis">
<emphasis id="B90B1640FF92E4621450689EC0EBFE0B" box="[292,401,439,465]" italics="true" pageId="7" pageNumber="594">P. pedalis</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46214ED689EC350FE0B" author="Nobili, G." box="[409,554,439,465]" pageId="7" pageNumber="594" pagination="393 - 411" refId="ref8299" refString="Nobili, G. (1906) Diagnoses preliminaires de 34 especes et varietes nouvelles, et de 2 genres nouveaux de decapodes de la Mer Rouge. Bulletin du Museum National d'Histoire, Paris, 11 (1905), 393 - 411, figs. 1, 2." type="book chapter" year="1906">Nobili, 1906</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E4621742689EC545FE0B" authority="Osawa, Naruse &amp; Ng, 2018" authorityName="Osawa, Naruse &amp; Ng" authorityYear="2018" box="[566,1087,439,465]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="pilosibrachium">
<emphasis id="B90B1640FF92E4621742689EC387FE0B" box="[566,765,439,465]" italics="true" pageId="7" pageNumber="594">P. pilosibrachium</emphasis>
<bibRefCitation id="EFEEB7A3FF92E4621670689EC545FE0B" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." box="[772,1087,439,465]" pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">Osawa, Naruse &amp; Ng, 2018</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E4621138689EC4E2FE0B" authority="Osawa &amp; Ng, 2016" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[1100,1432,439,465]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="B90B1640FF92E4621138689EC5C9FE0B" box="[1100,1203,439,465]" italics="true" pageId="7" pageNumber="594">P. planus</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46211CE689EC4E2FE0B" author="Osawa, M. &amp; Ng, P. K. L." box="[1210,1432,439,465]" pageId="7" pageNumber="594" pagination="499 - 518" refId="ref8748" refString="Osawa, M. &amp; Ng, P. K. L. (2016) Revision of Polyonyx pedalis Nobili, 1906 (Crustacea: Decapoda: Anomura: Porcellanidae), with descriptions of three new species. Raffles Bulletin of Zoology, Supplement, 34, 499 - 518." type="journal article" year="2016">Osawa &amp; Ng, 2016</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E46215E368F2C3F2FE2F" authority="Osawa, Naruse &amp; Ng, 2018" authorityName="Osawa, Naruse &amp; Ng" authorityYear="2018" box="[151,648,475,501]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="sasekumari">
<emphasis id="B90B1640FF92E46215E368F2C04CFE2F" box="[151,310,475,501]" italics="true" pageId="7" pageNumber="594">P. sasekumari</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462143568F2C3F2FE2F" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." box="[321,648,475,501]" pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">Osawa, Naruse &amp; Ng, 2018</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E46217EC68F2C2ABFE2F" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[664,977,474,501]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B90B1640FF92E46217EC68F2C26AFE2E" box="[664,784,475,500]" italics="true" pageId="7" pageNumber="594">P. sinensis</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462166F68F2C2ABFE2F" author="Stimpson, W." box="[795,977,474,501]" pageId="7" pageNumber="594" pagination="225 - 252" refId="ref8899" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VII. Crustacea Anomura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1858, 225 - 252. [preprint (December 1858)] https: // doi. org / 10.5962 / bhl. title. 51447" type="book chapter" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462169568F2C402FE2F" authority="Werding &amp; Hiller, 2019" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[993,1400,475,501]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E462169568F2C52CFE2F" box="[993,1110,475,501]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462111468F2C402FE2F" author="Werding, B. &amp; Hiller, A." box="[1120,1400,475,501]" pageId="7" pageNumber="594" pagination="25 - 34" refId="ref9086" refString="Werding, B. &amp; Hiller, A. (2019) A new species of Polyonyx (Crustacea, Anomura, Porcellanidae) inhabiting polychaete-worm tubes (Annelida, Chaetopteridae) in the Indo-West Pacific. ZooKeys, 818, 25 - 34. https: // doi. org / 10.3897 / zookeys. 818.30587" type="journal article" year="2019">Werding &amp; Hiller, 2019</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E46210FC68F2C0A0FDC3" authority="(Haswell, 1882)" baseAuthorityName="Haswell" baseAuthorityYear="1882" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="B90B1640FF92E46210FC68F2C062FDC2" italics="true" pageId="7" pageNumber="594">P. transversus</emphasis>
(
<bibRefCitation id="EFEEB7A3FF92E462145D68D6C0A8FDC3" author="Haswell, W. A." box="[297,466,511,537]" pageId="7" pageNumber="594" pagination="750 - 763" refId="ref7871" refString="Haswell, W. A. (1882) Description of some new species of Australian Decapoda. Proceedings of the Linnean Society of New South Wales, 6, 750 - 763. https: // doi. org / 10.5962 / bhl. part. 11897" type="journal article" year="1882">Haswell, 1882</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462149C68D6C39EFDC3" authority="Werding, 2001" authorityName="Werding" authorityYear="2001" box="[488,740,511,537]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="thai">
<emphasis id="B90B1640FF92E462149C68D6C34BFDC3" box="[488,561,511,537]" italics="true" pageId="7" pageNumber="594">P. thai</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462174D68D6C39EFDC3" author="Werding, B." box="[569,740,511,537]" pageId="7" pageNumber="594" pagination="109 - 119" refId="ref9031" refString="Werding, B. (2001) Description of two new species of Polyonyx Stimpson, 1858 from the Indo-West Pacific, with a key to the species of the Polyonyx sinensis group (Crustacea: Decapoda: Porcellanidae). Proceedings of the Biological Society of Washington, 114, 109 - 119." type="journal article" year="2001">Werding, 2001</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462178568D6C56DFDC3" authority="Werding, 2001" authorityName="Werding" authorityYear="2001" box="[753,1047,511,537]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="tulearis">
<emphasis id="B90B1640FF92E462178568D6C21FFDC3" box="[753,869,511,537]" italics="true" pageId="7" pageNumber="594">P. tulearis</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462161968D6C56DFDC3" author="Werding, B." box="[877,1047,511,537]" pageId="7" pageNumber="594" pagination="109 - 119" refId="ref9031" refString="Werding, B. (2001) Description of two new species of Polyonyx Stimpson, 1858 from the Indo-West Pacific, with a key to the species of the Polyonyx sinensis group (Crustacea: Decapoda: Porcellanidae). Proceedings of the Biological Society of Washington, 114, 109 - 119." type="journal article" year="2001">Werding, 2001</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C7FB1D1FF92E462115168D6C433FDC3" authority="Miyake, 1943" authorityName="Miyake" authorityYear="1943" box="[1061,1353,511,537]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="utinomii">
<emphasis id="B90B1640FF92E462115168D6C5E5FDC2" box="[1061,1183,511,536]" italics="true" pageId="7" pageNumber="594">P. utinomii</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46211DC68D6C433FDC3" author="Miyake, S." box="[1192,1353,511,537]" pageId="7" pageNumber="594" pagination="49 - 158" refId="ref8109" refString="Miyake, S. (1943) Studies on the crab-shaped Anomura of Nippon and adjacent waters. Journal of the Department of Agriculture, Kyushu Imperial University, 7, 49 - 158." type="journal article" year="1943">Miyake, 1943</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C7FB1D1FF92E46210FC68D6C364FDE7" authority="Ng &amp; Sasekumar, 1993" authorityName="Ng &amp; Sasekumar" authorityYear="1993" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="vermicola">
<emphasis id="B90B1640FF92E46210FC68D6C07DFDE7" italics="true" pageId="7" pageNumber="594">P. vermicola</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46214646B0AC364FDE7" author="Ng, P. K. L. &amp; Sasekumar, A." box="[272,542,547,573]" pageId="7" pageNumber="594" pagination="466 - 472" refId="ref8203" refString="Ng, P. K. L. &amp; Sasekumar, A. (1993) A new species of Polyonyx Stimpson, 1858, of the P. sinensis group (Crustacea: Anomura: Porcellanidae) commensal with a chaetopterid worm from Peninsular Malaysia. Zoologische Mededelingen, 67, 466 - 472." type="journal article" year="1993">Ng &amp; Sasekumar, 1993</bibRefCitation>
</taxonomicName>
. Among the species group,
<taxonomicName id="4C7FB1D1FF92E46216286B0AC2C9FDE7" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[860,947,547,573]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46216286B0AC2C9FDE7" box="[860,947,547,573]" italics="true" pageId="7" pageNumber="594">P. deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E46216CF6B0AC286FDE7" bold="true" box="[955,1020,547,573]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E46216CF6B0AC286FDE7" box="[955,1020,547,573]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
may be closest to
<taxonomicName id="4C7FB1D1FF92E46211A06B0AC432FDE7" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[1236,1352,547,573]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E46211A06B0AC432FDE7" box="[1236,1352,547,573]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
known from
<collectingCountry id="F3688AC2FF92E46215A36B6EC04CFDBB" box="[215,310,583,609]" name="Vietnam" pageId="7" pageNumber="594">Vietnam</collectingCountry>
in the comparatively short carpi of the chelipeds and broad meri of the ambulatory legs in their proportions. Nevertheless, the new species differs from
<taxonomicName id="4C7FB1D1FF92E462166F6B42C2F5FD5F" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[795,911,619,645]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E462166F6B42C2F5FD5F" box="[795,911,619,645]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
in the dorsal surface of the carapace covered with granules instead of shallow transversal striae, the anterior margin of the median lobe of the third thoracic sternite being concave instead of convex, the carpi and palms of the chelipeds and meri of the ambulatory legs with numerous granules and granular ridges on the dorsal or lateral surfaces, and the dorso-anterior marginal lobe of the carpus of each cheliped being more weakly developed, in addition to the distinction mentioned above. Coloration is also different between
<taxonomicName id="4C7FB1D1FF92E46214E26A36C091FCE3" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[406,491,799,825]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46214E26A36C091FCE3" box="[406,491,799,825]" italics="true" pageId="7" pageNumber="594">P. deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E46214856A36C34AFCE3" bold="true" box="[497,560,799,825]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E46214856A36C34AFCE3" box="[497,560,799,825]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C7FB1D1FF92E46217126A36C3ADFCE3" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[614,727,799,825]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E46217126A36C3ADFCE3" box="[614,727,799,825]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
; the dorsal surfaces of the carapace and chelipeds are generally white in the new species, rather than brown in
<taxonomicName id="4C7FB1D1FF92E46217D66A6AC269FC87" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[674,787,835,861]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E46217D66A6AC269FC87" box="[674,787,835,861]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0CA52FF92E46215B36A4EC3D4F98B" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="594">
On the Vietnamese coast,
<bibRefCitation id="EFEEB7A3FF92E462149C6A4EC3A2FC5B" author="Britayev, T. A. &amp; Mekhova, E. &amp; Deart, Y. &amp; Martin, D." box="[488,728,871,897]" pageId="7" pageNumber="594" pagination="2930" refId="ref7721" refString="Britayev, T. A., Mekhova, E., Deart, Y. &amp; Martin, D. (2017) Do syntopic host species harbour similar symbiotic communities? The case of Chaetopterus spp. (Annelida: Chaetopteridae). PeerJ, 5, e 2930. https: // doi. org / 10.7717 / peerj. 2930" type="journal article" year="2017">
Britayev
<emphasis id="B90B1640FF92E462173B6A4EC3FDFC5B" box="[591,647,871,897]" italics="true" pageId="7" pageNumber="594">et al.</emphasis>
(2017)
</bibRefCitation>
reported that
<taxonomicName id="4C7FB1D1FF92E46216066A4EC299FC5B" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[882,995,871,897]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E46216066A4EC299FC5B" box="[882,995,871,897]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
(as an undescribed species at that time) occurred in heterosexual pairs in the tube of a possibly undescribed species of
<taxonomicName id="4C7FB1D1FF92E462117F6AA2C5D8FC7F" authorityName="Cuvier" authorityYear="1830" box="[1035,1186,907,933]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="7" pageNumber="594" phylum="Annelida" rank="genus">
<emphasis id="B90B1640FF92E462117F6AA2C5D8FC7F" box="[1035,1186,907,933]" italics="true" pageId="7" pageNumber="594">Chaetopterus</emphasis>
</taxonomicName>
and frequently shared the polychaete tube with a heterosexual pair of significantly larger congeneric porcellanid
<taxonomicName id="4C7FB1D1FF92E462110B6A86C5B5FC13" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[1151,1231,943,969]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="heok">
<emphasis id="B90B1640FF92E462110B6A86C5B5FC13" box="[1151,1231,943,969]" italics="true" pageId="7" pageNumber="594">P. heok</emphasis>
</taxonomicName>
and the tergipedid nudibranch
<taxonomicName id="4C7FB1D1FF92E462146F6AFAC0DEFC37" box="[283,420,979,1005]" class="Gastropoda" family="Trinchesiidae" genus="Phestilla" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="594" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B90B1640FF92E462146F6AFAC005FC37" box="[283,383,979,1005]" italics="true" pageId="7" pageNumber="594">Phestilla</emphasis>
sp.
</taxonomicName>
or rarely with larger pinnotherid crab
<taxonomicName id="4C7FB1D1FF92E46216326AFAC2C3FC37" box="[838,953,979,1005]" class="Malacostraca" family="Pinnotheridae" genus="Tetrias" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90B1640FF92E46216326AFAC2E9FC36" box="[838,915,979,1004]" italics="true" pageId="7" pageNumber="594">Tetrias</emphasis>
sp.
</taxonomicName>
instead of
<taxonomicName id="4C7FB1D1FF92E46211446AFAC505FC37" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[1072,1151,979,1005]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="heok">
<emphasis id="B90B1640FF92E46211446AFAC505FC37" box="[1072,1151,979,1005]" italics="true" pageId="7" pageNumber="594">P. heok</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFEEB7A3FF92E46211FE6AFAC4E6FC37" author="Werding, B. &amp; Hiller, A." box="[1162,1436,979,1005]" pageId="7" pageNumber="594" pagination="25 - 34" refId="ref9086" refString="Werding, B. &amp; Hiller, A. (2019) A new species of Polyonyx (Crustacea, Anomura, Porcellanidae) inhabiting polychaete-worm tubes (Annelida, Chaetopteridae) in the Indo-West Pacific. ZooKeys, 818, 25 - 34. https: // doi. org / 10.3897 / zookeys. 818.30587" type="journal article" year="2019">Werding &amp; Hiller (2019)</bibRefCitation>
remarked that the extremely broadened carpi of the chelipeds and meri of the ambulatory legs of
<taxonomicName id="4C7FB1D1FF92E46211896ADEC40BFBCB" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[1277,1393,1015,1041]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E46211896ADEC40BFBCB" box="[1277,1393,1015,1041]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
are distinctive characters within the genus
<taxonomicName id="4C7FB1D1FF92E46217396D32C3CFFBEF" authorityName="Stimpson" authorityYear="1858" box="[589,693,1051,1077]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1640FF92E46217396D32C3CFFBEF" box="[589,693,1051,1077]" italics="true" pageId="7" pageNumber="594">Polyonyx</emphasis>
</taxonomicName>
, and these characters are most likely adaptations for living tightly attached to the walls of the worm tube without being perceived as an obstacle for the larger crabs inhabiting the same tube.
<taxonomicName id="4C7FB1D1FF92E46214606D4AC0C7FBA7" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[276,445,1123,1149]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46214606D4AC0C7FBA7" box="[276,445,1123,1149]" italics="true" pageId="7" pageNumber="594">Polyonyx deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E46214B66D4AC37AFBA7" bold="true" box="[450,512,1123,1149]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E46214B66D4AC37AFBA7" box="[450,512,1123,1149]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has similarly broadened carpi of the chelipeds and meri of the ambulatory legs, but no females nor other symbiont animals were obtained from the same tubes of host polychaetes. The
<taxonomicName id="4C7FB1D1FF92E46210716DAEC4E6FB7B" authorityName="Cuvier" authorityYear="1830" box="[1285,1436,1159,1185]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="7" pageNumber="594" phylum="Annelida" rank="genus">
<emphasis id="B90B1640FF92E46210716DAEC4E6FB7B" box="[1285,1436,1159,1185]" italics="true" pageId="7" pageNumber="594">Chaetopterus</emphasis>
</taxonomicName>
worms as candidates of the hosts of
<taxonomicName id="4C7FB1D1FF92E46217526D82C301FB1F" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[550,635,1195,1221]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46217526D82C301FB1F" box="[550,635,1195,1221]" italics="true" pageId="7" pageNumber="594">P. deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E46217F46D82C3C4FB1F" bold="true" box="[640,702,1195,1221]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E46217F46D82C3C4FB1F" box="[640,702,1195,1221]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
have not been preserved, but they are likely relatively small-sized species considering from the size of the broken tubes collected (
<quantity id="4C8767B7FF92E462161E6DE6C29FFB33" box="[874,997,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="594" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
in the largest widths of the tubes), and the space occupied by
<taxonomicName id="4C7FB1D1FF92E46214E96DDAC08EFAD7" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[413,500,1267,1293]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46214E96DDAC08EFAD7" box="[413,500,1267,1293]" italics="true" pageId="7" pageNumber="594">P. deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E46214886DDAC347FAD7" bold="true" box="[508,573,1267,1293]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E46214886DDAC347FAD7" box="[508,573,1267,1293]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
inside the tube can be very narrow. It is difficult at present to persuasively explain the morphological significance of the broadened chelipeds and ambulatory legs in
<taxonomicName id="4C7FB1D1FF92E46211F96C3EC599FAEB" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[1165,1251,1303,1329]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46211F96C3EC599FAEB" box="[1165,1251,1303,1329]" italics="true" pageId="7" pageNumber="594">P. deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E462119D6C3EC452FAEB" bold="true" box="[1257,1320,1303,1329]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E462119D6C3EC452FAEB" box="[1257,1320,1303,1329]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
as well as the presence of distinct protuberances on the dorsal surfaces of carapace and chelipeds. In Japanese waters including the Ryukyu Islands, unlike observations of the Vietnamese material by
<bibRefCitation id="EFEEB7A3FF92E46216B76C76C5C0FAA3" author="Britayev, T. A. &amp; Mekhova, E. &amp; Deart, Y. &amp; Martin, D." box="[963,1210,1375,1401]" pageId="7" pageNumber="594" pagination="2930" refId="ref7721" refString="Britayev, T. A., Mekhova, E., Deart, Y. &amp; Martin, D. (2017) Do syntopic host species harbour similar symbiotic communities? The case of Chaetopterus spp. (Annelida: Chaetopteridae). PeerJ, 5, e 2930. https: // doi. org / 10.7717 / peerj. 2930" type="journal article" year="2017">
Britayev
<emphasis id="B90B1640FF92E462115F6C76C51FFAA3" box="[1067,1125,1375,1401]" italics="true" pageId="7" pageNumber="594">et al.</emphasis>
(2017)
</bibRefCitation>
,
<taxonomicName id="4C7FB1D1FF92E46211B36C76C463FAA3" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[1223,1305,1375,1401]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="heok">
<emphasis id="B90B1640FF92E46211B36C76C463FAA3" box="[1223,1305,1375,1401]" italics="true" pageId="7" pageNumber="594">P. heok</emphasis>
</taxonomicName>
is found in heterosexual pairs or singly in parchment tubes of
<taxonomicName id="4C7FB1D1FF92E46217B06CAAC2A9FA47" authority="Nishi, 2001" authorityName="Nishi" authorityYear="2001" box="[708,979,1411,1437]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="7" pageNumber="594" phylum="Annelida" rank="species" species="pacificus">
<emphasis id="B90B1640FF92E46217B06CAAC233FA47" box="[708,841,1411,1437]" italics="true" pageId="7" pageNumber="594">C. pacificus</emphasis>
<bibRefCitation id="EFEEB7A3FF92E462163B6CAAC2A9FA47" author="Nishi, E." box="[847,979,1411,1437]" pageId="7" pageNumber="594" pagination="1 - 26" refId="ref8259" refString="Nishi, E. (2001) Partial revision of Japanese Chaetopterus (Chaetopteridae, Polychaeta), including description of three new species from southern Pacific side of central Japan. Actinia, 14, 1 - 26." type="journal article" year="2001">Nishi, 2001</bibRefCitation>
</taxonomicName>
, which usually adhere to the underside of rocks partially buried in sand or gravel bottom of embayed coasts (cf.
<bibRefCitation id="EFEEB7A3FF92E46216DE6C8EC503FA1B" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." box="[938,1145,1447,1473]" pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">
Osawa
<emphasis id="B90B1640FF92E46216896C8EC554FA1B" box="[1021,1070,1447,1473]" italics="true" pageId="7" pageNumber="594">et al</emphasis>
. 2018
</bibRefCitation>
); other symbionts, at least porcellanid or brachyuran crabs, from the same host tubes were never recorded. At Iriomote Island of the Ryukyus,
<bibRefCitation id="EFEEB7A3FF92E46215E36CC6C004F9D3" author="Naruse, T. &amp; Watanabe, T. &amp; Yoshida, R." box="[151,382,1519,1545]" pageId="7" pageNumber="594" pagination="17 - 28" refId="ref8144" refString="Naruse, T., Watanabe, T. &amp; Yoshida, R. (2017) Additional information on the distribution and habitats of six hexapodid species (Crustacea: Decapoda: Brachyura), with review of their Japanese names. Fauna Ryukyuana, 35, 17 - 28. [in Japanese with English abstract]" type="journal article" year="2017">
Naruse
<emphasis id="B90B1640FF92E46215846CC6C059F9D3" box="[240,291,1519,1545]" italics="true" pageId="7" pageNumber="594">et al</emphasis>
. (2017)
</bibRefCitation>
recorded an example of co-existence of an ovigerous female of
<taxonomicName id="4C7FB1D1FF92E46211136CC6C455F9D3" authorityName="Osawa, Naruse &amp; Ng" authorityYear="2018" box="[1127,1327,1519,1545]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="pilosibrachium">
<emphasis id="B90B1640FF92E46211136CC6C455F9D3" box="[1127,1327,1519,1545]" italics="true" pageId="7" pageNumber="594">P. pilosibrachium</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C7FB1D1FF92E46210166CC6C18EF9F6" authorityName="Miyake" authorityYear="1943" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="utinomii">
<emphasis id="B90B1640FF92E46210166CC6C40DF9D2" box="[1378,1399,1519,1544]" italics="true" pageId="7" pageNumber="594">P.</emphasis>
cf.
<emphasis id="B90B1640FF92E46215E36F3AC18EF9F6" box="[151,244,1555,1580]" italics="true" pageId="7" pageNumber="594">utinomii</emphasis>
</taxonomicName>
) and an ovigerous female of hexapodid brachyuran
<taxonomicName id="4C7FB1D1FF92E462164F6F3AC58CF9F7" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[827,1270,1555,1581]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B90B1640FF92E462164F6F3AC575F9F7" box="[827,1039,1555,1581]" italics="true" pageId="7" pageNumber="594">Hexapinus simplex</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46211616F3AC58CF9F7" author="Rahayu, D. L. &amp; Ng, P. K. L." box="[1045,1270,1555,1581]" pageId="7" pageNumber="594" pagination="396 - 483" refId="ref8801" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng, 2014</bibRefCitation>
</taxonomicName>
from the same tube of
<taxonomicName id="4C7FB1D1FF92E46215996F1EC3D0F98B" authority="Marenzeller, 1879" authorityName="Marenzeller" authorityYear="1879" box="[237,682,1591,1617]" class="Polychaeta" family="Chaetopteridae" genus="Chaetopterus" kingdom="Animalia" pageId="7" pageNumber="594" phylum="Annelida" rank="species" species="cautus">
<emphasis id="B90B1640FF92E46215996F1EC0AEF98A" box="[237,468,1591,1617]" italics="true" pageId="7" pageNumber="594">Chaetopterus cautus</emphasis>
<bibRefCitation id="EFEEB7A3FF92E46214AF6F1EC3D0F98B" author="Marenzeller, E. von" box="[475,682,1591,1617]" pageId="7" pageNumber="594" pagination="109 - 154" refId="ref7998" refString="Marenzeller, E. von (1879) Sudjapanische Anneliden. I. (Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea). Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Classe, Wien, 41, 109 - 154, pls. I - VI." type="journal article" year="1879">Marenzeller, 1879</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0CA52FF92E46215B36F72C19AF8DF" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="594">
The occurrence of
<taxonomicName id="4C7FB1D1FF92E46214EC6F72C097F9AF" authority="Osawa &amp; Sato, 2022" authorityName="Osawa &amp; Sato" authorityYear="2022" box="[408,493,1627,1653]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="deezi" status="sp. nov.">
<emphasis id="B90B1640FF92E46214EC6F72C097F9AF" box="[408,493,1627,1653]" italics="true" pageId="7" pageNumber="594">P. deezi</emphasis>
</taxonomicName>
<emphasis id="B90B1640FF92E46214866F72C34AF9AF" bold="true" box="[498,560,1627,1653]" pageId="7" pageNumber="594">
<taxonomicNameLabel id="A238AB3BFF92E46214866F72C34AF9AF" box="[498,560,1627,1653]" pageId="7" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from coral reefs as a rather exposed environment may be unusual in species of the
<taxonomicName id="4C7FB1D1FF92E46215B46F56C04FF942" authorityName="Stimpson" authorityYear="1858" box="[192,309,1663,1688]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B90B1640FF92E46215B46F56C04FF942" box="[192,309,1663,1688]" italics="true" pageId="7" pageNumber="594">P. sinensis</emphasis>
</taxonomicName>
group because many species of the group have been recorded from estuaries or coastal embayments. Only
<taxonomicName id="4C7FB1D1FF92E46215AC6F8AC023F966" authorityName="Osawa" authorityYear="2018" box="[216,345,1699,1724]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B90B1640FF92E46215AC6F8AC023F966" box="[216,345,1699,1724]" italics="true" pageId="7" pageNumber="594">P. angustus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7FB1D1FF92E462141D6F8AC09CF967" authorityName="Osawa" authorityYear="2007" box="[361,486,1699,1725]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="boucheti">
<emphasis id="B90B1640FF92E462141D6F8AC09CF967" box="[361,486,1699,1725]" italics="true" pageId="7" pageNumber="594">P. boucheti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7FB1D1FF92E46214826F8AC3FAF967" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[502,640,1699,1725]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="kumejima">
<emphasis id="B90B1640FF92E46214826F8AC3FAF967" box="[502,640,1699,1725]" italics="true" pageId="7" pageNumber="594">P. kumejima</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7FB1D1FF92E46217B56F8AC253F967" authorityName="Osawa &amp; Ng" authorityYear="2016" box="[705,809,1699,1725]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="B90B1640FF92E46217B56F8AC253F967" box="[705,809,1699,1725]" italics="true" pageId="7" pageNumber="594">P. planus</emphasis>
</taxonomicName>
were obtained from substrates of coral rubble or sand, although their commensal host animals were not always recorded (
<bibRefCitation id="EFEEB7A3FF92E46216F86FEEC51DF93B" author="Osawa, M. &amp; Ng, P. K. L." box="[908,1127,1735,1761]" pageId="7" pageNumber="594" pagination="499 - 518" refId="ref8748" refString="Osawa, M. &amp; Ng, P. K. L. (2016) Revision of Polyonyx pedalis Nobili, 1906 (Crustacea: Decapoda: Anomura: Porcellanidae), with descriptions of three new species. Raffles Bulletin of Zoology, Supplement, 34, 499 - 518." type="journal article" year="2016">Osawa &amp; Ng 2016</bibRefCitation>
; Osawa 2018;
<bibRefCitation id="EFEEB7A3FF92E462107B6FEEC1AEF8DF" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">
Osawa
<emphasis id="B90B1640FF92E46210176FEEC4EFF93B" box="[1379,1429,1735,1761]" italics="true" pageId="7" pageNumber="594">et al</emphasis>
. 2018
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC0CA52FF92E46215B36E26C51BF8AB" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="594">
<bibRefCitation id="EFEEB7A3FF92E46215B36E26C0D2F8F3" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." box="[199,424,1807,1833]" pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">
Osawa
<emphasis id="B90B1640FF92E46214686E26C034F8F3" box="[284,334,1807,1833]" italics="true" pageId="7" pageNumber="594">et al</emphasis>
. (2018)
</bibRefCitation>
presented an identification key to the Indo-West Pacific species of the
<taxonomicName id="4C7FB1D1FF92E46211A76E26C4E7F8F2" authorityName="Stimpson" authorityYear="1858" box="[1235,1437,1807,1833]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B90B1640FF92E46211A76E26C4E7F8F2" box="[1235,1437,1807,1833]" italics="true" pageId="7" pageNumber="594">Polyonyx sinensis</emphasis>
</taxonomicName>
group known at the time. The identification key is amended with the addition of two species described since
<bibRefCitation id="EFEEB7A3FF92E46210246E1AC059F8AB" author="Osawa, M. &amp; Naruse, T. &amp; Ng, P. K. L." pageId="7" pageNumber="594" pagination="303 - 323" refId="ref8668" refString="Osawa, M., Naruse, T. &amp; Ng, P. K. L. (2018) New records of species of the Polyonyx sinensis group (Crustacea: Decapoda: Anomura: Porcellanidae) from Japan, the Philippines, Singapore, and Malaysia, with descriptions of two new species. Zootaxa, 4429 (2), 303 - 323. https: // doi. org / 10.11646 / zootaxa. 4429.2.6" type="journal article" year="2018">
Osawa
<emphasis id="B90B1640FF92E46215E36E7EC1B2F8AB" box="[151,200,1879,1905]" italics="true" pageId="7" pageNumber="594">et al</emphasis>
. (2018)
</bibRefCitation>
,
<taxonomicName id="4C7FB1D1FF92E462145B6E7EC0D4F8AA" authorityName="Osawa" authorityYear="2018" box="[303,430,1879,1904]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B90B1640FF92E462145B6E7EC0D4F8AA" box="[303,430,1879,1904]" italics="true" pageId="7" pageNumber="594">P. angustus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7FB1D1FF92E46214906E7EC32CF8AB" authorityName="Werding &amp; Hiller" authorityYear="2019" box="[484,598,1879,1905]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B90B1640FF92E46214906E7EC32CF8AB" box="[484,598,1879,1905]" italics="true" pageId="7" pageNumber="594">P. socialis</emphasis>
</taxonomicName>
, and the present new species, as shown below.
</paragraph>
</subSubSection>
</treatment>
</document>
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