treatments-xml/data/03/D6/5A/03D65A6C634BFFBBC399FD3DFC8FF833.xml

<document id="5DF4985A361D7A3D6D33933469A3EB79" ID-DOI="10.5281/zenodo.197944" ID-GBIF-Dataset="f821c77c-a895-4e82-ac0c-0daa618ba83b" ID-ISSN="1175-5326" ID-Zenodo-Dep="197944" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460282257266" checkinUser="plazi" docAuthor="Conway, Kevin W. &amp; Britz, Ralf" docDate="2010" docId="03D65A6C634BFFBBC399FD3DFC8FF833" docLanguage="en" docName="zt02616p047.pdf" docOrigin="Zootaxa 2616" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Psilorhynchus brachyrhynchus Conway &amp; Britz, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFEF2214634AFFBCC30EFF9FFFA6FF8E" masterDocTitle="Three new species of Psilorhynchus from the Ayeyarwaddy River drainage, Myanmar (Teleostei: Psilorhynchidae)" masterLastPageNumber="47" masterPageNumber="31" pageNumber="32" updateTime="1698249656017" updateUser="plazi">
<mods:mods id="52786E2C94115A219CFF33B7089BFA19" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5B8396F728AC1D007A81F84981B2316D">
<mods:title id="F03F099E6205CCB843934A5542915B3A">Three new species of Psilorhynchus from the Ayeyarwaddy River drainage, Myanmar (Teleostei: Psilorhynchidae)</mods:title>
</mods:titleInfo>
<mods:name id="892EC6E8295950EA2C26BAA613DDAF86" type="personal">
<mods:role id="8B2F7A0DDB11629C496346A424242286">
<mods:roleTerm id="E0F4BA374D32409C9D207B01CA1F4E31">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="40050D11B7E9000EB669431F2298E713">Conway, Kevin W.</mods:namePart>
</mods:name>
<mods:name id="DFAEFAB45F0D9EC8791528882D825A0C" type="personal">
<mods:role id="196B7B840ED74F5B23A0023B34FD6704">
<mods:roleTerm id="03ADC8C5D925D0E897CB97C018684AE0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="991D211203FB07293FAA88268CB1512A">Britz, Ralf</mods:namePart>
</mods:name>
<mods:typeOfResource id="67A461EDC5327151A634D97182847437">text</mods:typeOfResource>
<mods:relatedItem id="13E4700B3A71909BFF7CF118AA1108D3" type="host">
<mods:titleInfo id="36F0482607EAC9B76A0E5A917B82509E">
<mods:title id="3D633A722B76CE68A835AD1A425D34BD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F82D42F2DD03270A6D171630696A5882">
<mods:date id="80B4A95BC87D05C71EEF8A5EABD62A25">2010</mods:date>
<mods:detail id="EB1E1339889C27BC1CD1626FDFB395BC" type="volume">
<mods:number id="02E1FB840D392A4C2BBA3710C50A8503">2616</mods:number>
</mods:detail>
<mods:extent id="F6F112E48325656AABE52177A41064B5" unit="page">
<mods:start id="6C2B643A66980D9C9C9E23AE8C567E65">31</mods:start>
<mods:end id="E5CE3946A4B31AE9420D5F74B5E31439">47</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D527A37713B6341475B4EEEDDC9ED1EC">journal article</mods:classification>
<mods:identifier id="EABE3D3A14D0A7BA479AA6946D7854E8" type="DOI">10.5281/zenodo.197944</mods:identifier>
<mods:identifier id="74AD3051E5C34F479B57C41404374375" type="GBIF-Dataset">f821c77c-a895-4e82-ac0c-0daa618ba83b</mods:identifier>
<mods:identifier id="AEE8CE1503B411D7D98B920C07E0A948" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EBA20EA203DB9F66A36354E22B0515DF" type="Zenodo-Dep">197944</mods:identifier>
</mods:mods>
<treatment id="03D65A6C634BFFBBC399FD3DFC8FF833" ID-DOI="http://doi.org/10.5281/zenodo.5630197" ID-GBIF-Taxon="119396896" ID-Zenodo-Dep="5630197" LSID="urn:lsid:plazi:treatment:03D65A6C634BFFBBC399FD3DFC8FF833" httpUri="http://treatment.plazi.org/id/03D65A6C634BFFBBC399FD3DFC8FF833" lastPageId="7" lastPageNumber="38" pageId="1" pageNumber="32">
<subSubSection id="C365B8F1634BFFBDC399FD3DFD65FCA9" pageId="1" pageNumber="32" type="nomenclature">
<paragraph id="8BC0EB7A634BFFBDC399FD3DFD0EFD32" blockId="1.[151,680,674,735]" box="[151,680,674,700]" pageId="1" pageNumber="32">
<heading id="D0885C16634BFFBDC399FD3DFD0EFD32" bold="true" box="[151,680,674,700]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
<emphasis id="B90B3768634BFFBDC399FD3DFD0EFD32" bold="true" box="[151,680,674,700]" pageId="1" pageNumber="32">
<taxonomicName id="4C7F90F9634BFFBDC399FD3DFDB7FD32" ID-CoL="4PJHT" box="[151,529,674,700]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="brachyrhynchus" status="sp. nov.">
<emphasis id="B90B3768634BFFBDC399FD3DFDB7FD32" bold="true" box="[151,529,674,700]" italics="true" pageId="1" pageNumber="32">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388A13634BFFBDC117FD3DFD0EFD32" box="[537,680,674,700]" pageId="1" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0EB7A634BFFBDC399FD5AFEABFD51" blockId="1.[151,680,674,735]" box="[151,269,709,735]" pageId="1" pageNumber="32">
(
<figureCitation id="1344F7FF634BFFBDC391FD5AFEA7FD51" box="[159,257,709,735]" captionStart="FIGURE 1" captionStartId="2.[151,255,1450,1474]" captionTargetBox="[151,1436,185,1425]" captionTargetId="figure@2.[151,1436,185,1426]" captionTargetPageId="2" captionText="FIGURE 1. Psilorhynchus brachyrhynchus, NRM 40935, male, 44.9 mm SL; Myanmar: Kachin State, Putao." httpUri="https://zenodo.org/record/197945/files/figure.png" pageId="1" pageNumber="32">Figure 1</figureCitation>
)
</paragraph>
<paragraph id="8BC0EB7A634BFFBDC399FC8FFD65FCA9" blockId="1.[151,707,784,807]" box="[151,707,784,807]" pageId="1" pageNumber="32">
<taxonomicName id="4C7F90F9634BFFBDC399FC8FFE2DFCA9" box="[151,395,784,807]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634BFFBDC399FC8FFE2DFCA9" box="[151,395,784,807]" italics="true" pageId="1" pageNumber="32">Psilorhynchus balitora</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFEE968B634BFFBDC297FC8FFD11FCA9" author="Mukerji" box="[409,695,784,807]" pageId="1" pageNumber="32" refString="Mukerji, D. D. (1933) Report on Burmese fishes collected by Lt. - Col. R. W. Burton from the tributary streams of the Mali Hka River of the Myitkyina District (Upper Burma). Part I. Journal of the Bombay Natural History Society, 36, 813 - 831." type="journal article" year="1933">Mukerji 1933: plate I, 2–4</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C365B8F1634BFFBDC399FCCAFD0BFB12" pageId="1" pageNumber="32" type="materials_examined">
<paragraph id="8BC0EB7A634BFFBDC399FCCAFC3CFC32" blockId="1.[151,1437,853,1927]" pageId="1" pageNumber="32">
<emphasis id="B90B3768634BFFBDC399FCCAFEB6FCE1" bold="true" box="[151,272,853,879]" pageId="1" pageNumber="32">
<typeStatus id="54C455D8634BFFBDC399FCCAFEADFCE1" box="[151,267,853,879]" pageId="1" pageNumber="32" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED6E73BF634BFFBDC219FCCAFEFCFCE1" box="[279,346,853,879]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="32">NRM</collectionCode>
40935, male, 
<quantity id="4C87469F634BFFBDC2F0FCCAFDC0FCE1" box="[510,614,853,879]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="1" pageNumber="32" unit="mm" value="44.9">44.9 mm</quantity>
SL; 
<collectingCountry id="F368ABEA634BFFBDC190FCCAFCB3FCE1" box="[670,789,853,879]" name="Myanmar" pageId="1" pageNumber="32">Myanmar</collectingCountry>
: Kachin State, Putao, Ayeyarwaddy River drainage, Ma Kyaww Wa Chaung and its tributary Nan Hto Chaung, ca 
<quantity id="4C87469F634BFFBDC031FCE2FC2EFC19" box="[831,904,893,919]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="1" pageNumber="32" unit="mi" value="1.0">1 mile</quantity>
from 46th regiment, close to rice mill, 
<geoCoordinate id="EE4B8DBD634BFFBDC647FCE2FF79FC32" direction="north" orientation="latitude" pageId="1" pageNumber="32" precision="15" value="27.32889">27° 19' 44&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE4B8DBD634BFFBDC3E5FC3DFE2CFC32" box="[235,394,930,956]" direction="east" orientation="longitude" pageId="1" pageNumber="32" precision="15" value="97.37667">97° 22' 36&quot; E</geoCoordinate>
, 
<date id="FFC1CDBA634BFFBDC298FC3DFDEAFC32" box="[406,588,930,956]" pageId="1" pageNumber="32" value="1998-03-28">28 March 1998</date>
, S. 
<collectingCountry id="F368ABEA634BFFBDC179FC3DFD29FC32" box="[631,655,930,956]" name="Iceland" pageId="1" pageNumber="32">O</collectingCountry>
. Kullander &amp; R. Britz.
</paragraph>
<paragraph id="8BC0EB7A634BFFBDC3C8FC57FC28FBDC" blockId="1.[151,1437,853,1927]" pageId="1" pageNumber="32">
<emphasis id="B90B3768634BFFBDC3C8FC57FEEDFC6C" bold="true" box="[198,331,968,994]" pageId="1" pageNumber="32">
<typeStatus id="54C455D8634BFFBDC3C8FC57FEE0FC6C" box="[198,326,968,994]" pageId="1" pageNumber="32" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED6E73BF634BFFBDC25DFC57FE30FC6C" box="[339,406,968,994]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="32">NRM</collectionCode>
60761, 1, 
<quantity id="4C87469F634BFFBDC11AFC57FDDBFC6C" box="[532,637,968,994]" metricMagnitude="-2" metricUnit="m" metricValue="4.26" pageId="1" pageNumber="32" unit="mm" value="42.6">42.6 mm</quantity>
SL; same data as 
<typeStatus id="54C455D8634BFFBDC05DFC57FC18FC6C" box="[851,958,968,994]" pageId="1" pageNumber="32" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED6E73BF634BFFBDC0C7FC57FBAAFC6C" box="[969,1036,968,994]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="32">NRM</collectionCode>
40922, 15 (2c&amp;s), 
<quantity id="4C87469F634BFFBDC7FFFC57FA3AFC6C" box="[1265,1436,968,994]" metricMagnitude="-2" metricUnit="m" metricValue="3.965" metricValueMax="5.11" metricValueMin="2.82" pageId="1" pageNumber="32" unit="mm" value="39.65" valueMax="51.1" valueMin="28.2">28.2–51.1 mm</quantity>
SL; same locality as 
<typeStatus id="54C455D8634BFFBDC29FFC72FE5AFB89" box="[401,508,1005,1031]" pageId="1" pageNumber="32" type="holotype">holotype</typeStatus>
, 
<date id="FFC1CDBA634BFFBDC107FC72FD64FB89" box="[521,706,1005,1031]" pageId="1" pageNumber="32" value="1998-03-27">27 March 1998</date>
, S. 
<collectingCountry id="F368ABEA634BFFBDC1FEFC72FCAFFB89" box="[752,777,1005,1031]" name="Iceland" pageId="1" pageNumber="32">O</collectingCountry>
. Kullander &amp; R. Britz. 
<collectionCode id="ED6E73BF634BFFBDC72FFC72FBC3FB89" box="[1057,1125,1005,1031]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="32">NRM</collectionCode>
59658, 1 (c&amp;s), 
<quantity id="4C87469F634BFFBDC63CFC72FA3AFB89" box="[1330,1436,1005,1031]" metricMagnitude="-2" metricUnit="m" metricValue="4.09" pageId="1" pageNumber="32" unit="mm" value="40.9">40.9 mm</quantity>
SL; same locality as 
<typeStatus id="54C455D8634BFFBDC280FB8DFE5EFBA2" box="[398,504,1042,1068]" pageId="1" pageNumber="32" type="holotype">holotype</typeStatus>
, 
<date id="FFC1CDBA634BFFBDC10BFB8DFD1AFBA2" box="[517,700,1042,1068]" pageId="1" pageNumber="32" value="1998-03-27">27 March 1998</date>
, S. 
<collectingCountry id="F368ABEA634BFFBDC1E7FB8DFCA7FBA2" box="[745,769,1042,1068]" name="Iceland" pageId="1" pageNumber="32">O</collectingCountry>
. Kullander &amp; R. Britz. 
<collectionCode id="ED6E73BF634BFFBDC719FB8DFBFCFBA2" box="[1047,1114,1042,1068]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="32">NRM</collectionCode>
57222, 
<quantity id="4C87469F634BFFBDC7B2FB8DFA83FBA2" box="[1212,1317,1042,1068]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" pageId="1" pageNumber="32" unit="mm" value="32.9">32.9 mm</quantity>
SL; same locality as 
<typeStatus id="54C455D8634BFFBDC21BFBA7FED8FBDC" box="[277,382,1080,1106]" pageId="1" pageNumber="32" type="holotype">holotype</typeStatus>
, 
<date id="FFC1CDBA634BFFBDC285FBA7FDE6FBDC" box="[395,576,1080,1106]" pageId="1" pageNumber="32" value="1998-03-27">27 March 1998</date>
, S. 
<collectingCountry id="F368ABEA634BFFBDC165FBA7FD25FBDC" box="[619,643,1080,1106]" name="Iceland" pageId="1" pageNumber="32">O</collectingCountry>
. Kullander &amp; R. Britz.
</paragraph>
<paragraph id="8BC0EB7A634BFFBDC3C8FBC2FD0BFB12" blockId="1.[151,1437,853,1927]" pageId="1" pageNumber="32">
<emphasis id="B90B3768634BFFBDC3C8FBC2FDA8FBF9" bold="true" box="[198,526,1117,1143]" pageId="1" pageNumber="32">Other material examined.</emphasis>
<collectionCode id="ED6E73BF634BFFBDC119FBC2FDFCFBF9" box="[535,602,1117,1143]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="1" pageNumber="32">NRM</collectionCode>
40630, 3, 40.0– 
<quantity id="4C87469F634BFFBDC011FBC2FC2FFBF9" box="[799,905,1117,1143]" metricMagnitude="-2" metricUnit="m" metricValue="4.57" pageId="1" pageNumber="32" unit="mm" value="45.7">45.7 mm</quantity>
SL; 
<collectingCountry id="F368ABEA634BFFBDC0C9FBC2FB98FBF9" box="[967,1086,1117,1143]" name="Myanmar" pageId="1" pageNumber="32">Myanmar</collectingCountry>
: Kachin State, Putao, 
<geoCoordinate id="EE4B8DBD634BFFBDC649FBC2FF79FB12" direction="north" orientation="latitude" pageId="1" pageNumber="32" precision="15" value="27.35">27° 20' 60&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE4B8DBD634BFFBDC3E5FB1DFE2CFB12" box="[235,394,1154,1180]" direction="east" orientation="longitude" pageId="1" pageNumber="32" precision="15" value="97.4">97° 23' 60&quot; E</geoCoordinate>
, 
<date id="FFC1CDBA634BFFBDC298FB1DFDBFFB12" box="[406,537,1154,1180]" pageId="1" pageNumber="32" value="1934-04">April 1934</date>
, Fan Li San.
</paragraph>
</subSubSection>
<subSubSection id="C365B8F1634BFFBDC3C8FB37FDF6F899" pageId="1" pageNumber="32" type="diagnosis">
<paragraph id="8BC0EB7A634BFFBDC3C8FB37FDF6F899" blockId="1.[151,1437,853,1927]" pageId="1" pageNumber="32">
<emphasis id="B90B3768634BFFBDC3C8FB37FEE0FB4C" bold="true" box="[198,326,1192,1218]" pageId="1" pageNumber="32">Diagnosis.</emphasis>
A species of 
<taxonomicName id="4C7F90F9634BFFBDC2FDFB37FD3CFB4F" box="[499,666,1192,1217]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B90B3768634BFFBDC2FDFB37FD3CFB4F" box="[499,666,1192,1217]" italics="true" pageId="1" pageNumber="32">Psilorhynchus</emphasis>
</taxonomicName>
most similar to 
<taxonomicName id="4C7F90F9634BFFBDC068FB37FB94FB4F" box="[870,1074,1192,1217]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634BFFBDC068FB37FB94FB4F" box="[870,1074,1192,1217]" italics="true" pageId="1" pageNumber="32">P. amplicephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC74AFB37FB64FB4F" box="[1092,1218,1192,1217]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634BFFBDC74AFB37FB64FB4F" box="[1092,1218,1192,1217]" italics="true" pageId="1" pageNumber="32">P. balitora</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC7DAFB37FAD2FB4F" box="[1236,1396,1192,1217]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634BFFBDC7DAFB37FAD2FB4F" box="[1236,1396,1192,1217]" italics="true" pageId="1" pageNumber="32">P. breviminor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC688FB37FF40FB69" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="gokkyi">
<emphasis id="B90B3768634BFFBDC688FB37FF40FB69" italics="true" pageId="1" pageNumber="32">P. gokkyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC3F8FB51FE33FB69" box="[246,405,1230,1255]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634BFFBDC3F8FB51FE33FB69" box="[246,405,1230,1255]" italics="true" pageId="1" pageNumber="32">P. nepalensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC2A8FB51FDC1FB69" box="[422,615,1230,1255]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634BFFBDC2A8FB51FDC1FB69" box="[422,615,1230,1255]" italics="true" pageId="1" pageNumber="32">P. pavimentatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC179FB51FCAFFB69" box="[631,777,1230,1255]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634BFFBDC179FB51FCAFFB69" box="[631,777,1230,1255]" italics="true" pageId="1" pageNumber="32">P. piperatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634BFFBDC048FB51FC6AFB69" box="[838,972,1230,1255]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="rahmani">
<emphasis id="B90B3768634BFFBDC048FB51FC6AFB69" box="[838,972,1230,1255]" italics="true" pageId="1" pageNumber="32">P. rahmani</emphasis>
</taxonomicName>
in general appearance. 
<taxonomicName id="4C7F90F9634BFFBDC7E1FB51FEF5FA82" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634BFFBDC7E1FB51FEF5FA82" italics="true" pageId="1" pageNumber="32">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C7F90F9634BFFBDC154FB6CFC84FA82" box="[602,802,1267,1292]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634BFFBDC154FB6CFC84FA82" box="[602,802,1267,1292]" italics="true" pageId="1" pageNumber="32">P. amplicephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC03FFB6CFC0DFA82" box="[817,939,1267,1292]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634BFFBDC03FFB6CFC0DFA82" box="[817,939,1267,1292]" italics="true" pageId="1" pageNumber="32">P. balitora</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC0B5FB6CFBFEFA82" box="[955,1112,1267,1292]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634BFFBDC0B5FB6CFBFEFA82" box="[955,1112,1267,1292]" italics="true" pageId="1" pageNumber="32">P. breviminor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC769FB6CFB76FA82" box="[1127,1232,1267,1292]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="gokkyi">
<emphasis id="B90B3768634BFFBDC769FB6CFB76FA82" box="[1127,1232,1267,1292]" italics="true" pageId="1" pageNumber="32">P. gokkyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC7D1FB6CFADEFA82" box="[1247,1400,1267,1292]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634BFFBDC7D1FB6CFADEFA82" box="[1247,1400,1267,1292]" italics="true" pageId="1" pageNumber="32">P. nepalensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC688FB6CFE95FABF" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634BFFBDC688FB6CFE95FABF" italics="true" pageId="1" pageNumber="32">P. pavimentatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634BFFBDC262FA87FE5EFABF" box="[364,504,1304,1329]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634BFFBDC262FA87FE5EFABF" box="[364,504,1304,1329]" italics="true" pageId="1" pageNumber="32">P. piperatus</emphasis>
</taxonomicName>
by its shorter snout (snout length 43–48% HL vs. 48–56); from 
<taxonomicName id="4C7F90F9634BFFBDC7E4FA87FA33FABF" box="[1258,1429,1304,1329]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634BFFBDC7E4FA87FA33FABF" box="[1258,1429,1304,1329]" italics="true" pageId="1" pageNumber="32">amplicephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC399FAA1FEB4FAD9" box="[151,274,1342,1367]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634BFFBDC399FAA1FEB4FAD9" box="[151,274,1342,1367]" italics="true" pageId="1" pageNumber="32">P. balitora</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC22EFAA1FE1FFAD9" box="[288,441,1342,1367]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634BFFBDC22EFAA1FE1FFAD9" box="[288,441,1342,1367]" italics="true" pageId="1" pageNumber="32">P. nepalensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC2C9FAA1FD26FAD9" box="[455,640,1342,1367]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634BFFBDC2C9FAA1FD26FAD9" box="[455,640,1342,1367]" italics="true" pageId="1" pageNumber="32">P. pavimentatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634BFFBDC1B7FAA1FCE3FAD9" box="[697,837,1342,1367]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634BFFBDC1B7FAA1FCE3FAD9" box="[697,837,1342,1367]" italics="true" pageId="1" pageNumber="32">P. piperatus</emphasis>
</taxonomicName>
by having more principal rays in the upper caudalfin lobe (10 vs. 8–9); from 
<emphasis id="B90B3768634BFFBDC2D4FAFCFC89FAF2" box="[474,815,1379,1404]" italics="true" pageId="1" pageNumber="32">
<taxonomicName id="4C7F90F9634BFFBDC2D4FAFCFD01FAF2" box="[474,679,1379,1404]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="amplicephalus">P. amplicephalus</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC1BDFAFCFC89FAF2" box="[691,815,1379,1404]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="balitora">P. balitora</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C7F90F9634BFFBDC064FAFCFBA5FAF2" box="[874,1027,1379,1404]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634BFFBDC064FAFCFBA5FAF2" box="[874,1027,1379,1404]" italics="true" pageId="1" pageNumber="32">P. nepalensis</emphasis>
</taxonomicName>
by the absence (vs. presence) of a dark irregular vertical bar across the distal edge of the caudal fin; from 
<taxonomicName id="4C7F90F9634BFFBDC0FAFA17FBD4FA2F" box="[1012,1138,1416,1441]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634BFFBDC0FAFA17FBD4FA2F" box="[1012,1138,1416,1441]" italics="true" pageId="1" pageNumber="32">P. balitora</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634BFFBDC7A3FA17FAECFA2F" box="[1197,1354,1416,1441]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634BFFBDC7A3FA17FAECFA2F" box="[1197,1354,1416,1441]" italics="true" pageId="1" pageNumber="32">P. nepalensis</emphasis>
</taxonomicName>
by the absence (vs. presence) of a broad swath of melanophores across the anterior half of the dorsal fin; from 
<emphasis id="B90B3768634BFFBDC688FA31FE37FA62" italics="true" pageId="1" pageNumber="32">
<taxonomicName id="4C7F90F9634BFFBDC688FA31FEBCFA62" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="breviminor">P. breviminor</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC228FA4CFE37FA62" box="[294,401,1491,1516]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="gokkyi">P. gokkyi</taxonomicName>
</emphasis>
, 
<emphasis id="B90B3768634BFFBDC2ACFA4CFD5AFA62" box="[418,764,1491,1516]" italics="true" pageId="1" pageNumber="32">
<taxonomicName id="4C7F90F9634BFFBDC2ACFA4CFDC4FA62" box="[418,610,1491,1516]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="pavimentatus">P. pavimentatus</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC160FA4CFD5AFA62" box="[622,764,1491,1516]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="piperatus">P. piperatus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C7F90F9634BFFBDC036FA4CFC1CFA62" box="[824,954,1491,1516]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="rahmani">
<emphasis id="B90B3768634BFFBDC036FA4CFC1CFA62" box="[824,954,1491,1516]" italics="true" pageId="1" pageNumber="32">P. rahmani</emphasis>
</taxonomicName>
by the presence (vs. absence) of a large dark blotch at the base of each caudal fin lobe; and from 
<emphasis id="B90B3768634BFFBDC05FFA67FB21F99F" box="[849,1159,1528,1553]" italics="true" pageId="1" pageNumber="32">
<taxonomicName id="4C7F90F9634BFFBDC05FFA67FC53F99F" box="[849,1013,1528,1553]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="breviminor">P. breviminor</taxonomicName>
, 
<taxonomicName id="4C7F90F9634BFFBDC70CFA67FB21F99F" box="[1026,1159,1528,1553]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="piperata">P. piperata</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C7F90F9634BFFBDC7CAFA67FAEEF99F" box="[1220,1352,1528,1553]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="rahmani">
<emphasis id="B90B3768634BFFBDC7CAFA67FAEEF99F" box="[1220,1352,1528,1553]" italics="true" pageId="1" pageNumber="32">P. rahmani</emphasis>
</taxonomicName>
by the presence (vs. absence) of an irregular vertical bar across the center of the caudal fin. 
<taxonomicName id="4C7F90F9634BFFBDC7E9F981FEF5F9D2" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="32" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634BFFBDC7E9F981FEF5F9D2" italics="true" pageId="1" pageNumber="32">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
is further distinguished from the aforementioned species and all remaining congeners by the following combination of characters: 2 branchiostegal rays; ventral surface between paired fins with traingular scale-less patch; presence of a large postepiphysial fontanelle; branched dorsal-fin rays 9; unbranched pectoral-fin rays 5; caudal-fin rays 10+9; lateral-line scales 32–34; lateral sides of body marked with 5–7 indistinct round to squarish dark blotches, arranged in a longitudinal row; caudal fin with a dark, irregular vertical bar across its center.
</paragraph>
</subSubSection>
<subSubSection id="C365B8F1634BFFB9C3C8F8BDFAC6FD8A" lastPageId="5" lastPageNumber="36" pageId="1" pageNumber="32" type="description">
<paragraph id="8BC0EB7A634BFFBDC3C8F8BDFBD5F809" blockId="1.[151,1437,853,1927]" pageId="1" pageNumber="32">
<emphasis id="B90B3768634BFFBDC3C8F8BDFEF9F8B2" bold="true" box="[198,351,1826,1852]" pageId="1" pageNumber="32">Description.</emphasis>
General body shape as in 
<figureCitation id="1344F7FF634BFFBDC1ADF8BDFCAFF8B2" box="[675,777,1826,1852]" captionStart="FIGURE 1" captionStartId="2.[151,255,1450,1474]" captionTargetBox="[151,1436,185,1425]" captionTargetId="figure@2.[151,1436,185,1426]" captionTargetPageId="2" captionText="FIGURE 1. Psilorhynchus brachyrhynchus, NRM 40935, male, 44.9 mm SL; Myanmar: Kachin State, Putao." httpUri="https://zenodo.org/record/197945/files/figure.png" pageId="1" pageNumber="32">Figure 1</figureCitation>
. Morphometric and meristic data are listed in 
<tableCitation id="C6FDDEC1634BFFBDC635F8BDFA30F8B2" box="[1339,1430,1826,1852]" captionStart="TABLE 1" captionStartId="3.[151,239,304,328]" captionTargetBox="[165,1403,363,1971]" captionTargetPageId="3" captionText="TABLE 1. Selected morphometric and meristic data for Psilorhynchus brachyrhynchus holotype and 11 paratypes." httpUri="http://table.plazi.org/id/DF00BBF26349FFBFC399FEAFFAC4FEC9" pageId="1" pageNumber="32" tableUuid="DF00BBF26349FFBFC399FEAFFAC4FEC9">Table 1</tableCitation>
. Body deep, greatest depth at dorsal-fin origin. Dorsal profile arched, ventral profile straight from lower jaw to anal-fin origin, with slight dorsad inclination from anal-fin origin to caudal-fin base.
</paragraph>
<caption id="DF00BBF26348FFBEC399FA35FA88FA4F" box="[151,1326,1450,1474]" httpUri="https://zenodo.org/record/197945/files/figure.png" pageId="2" pageNumber="33" targetBox="[151,1436,185,1425]" targetPageId="2">
<paragraph id="8BC0EB7A6348FFBEC399FA35FA88FA4F" blockId="2.[151,1326,1450,1474]" box="[151,1326,1450,1474]" pageId="2" pageNumber="33">
<emphasis id="B90B37686348FFBEC399FA35FEBFFA4C" bold="true" box="[151,281,1450,1474]" pageId="2" pageNumber="33">FIGURE 1.</emphasis>
<taxonomicName id="4C7F90F96348FFBEC22EFA35FDCCFA4F" box="[288,618,1450,1473]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="33" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B37686348FFBEC22EFA35FDCCFA4F" box="[288,618,1450,1473]" italics="true" pageId="2" pageNumber="33">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
, NRM 40935, male, 44.9 mm SL; Myanmar: Kachin State, Putao.
</paragraph>
</caption>
<paragraph id="8BC0EB7A6348FFBFC3C8F98BFC74FF72" blockId="2.[151,1437,1556,2030]" lastBlockId="3.[151,1436,152,252]" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33">Head small, eye large. Mouth inferior, snout short, rounded, its ventral surface bordered by a deep longitudinal groove on each side. Rostral cap and upper lip fused, separated by a shallow groove. Lower jaw covered by a thick squarish ‘cushion’ that can be folded backwards, the ‘cushion’ composed of two adnate tissue layers: deeper layer (lower lip), smooth, not continuous with upper lip around corner of mouth; superficial layer densely papilliated, thick, continuous with skin of isthmus, connected with rostral cap by narrow strip of skin around corner of mouth, extended posteriorly, broadened as a ridged, heavily papillated skin fold at posterolateral-most corner of mouth. Upper lip with numerous, horizontally arranged rows of unculi. Pre-epiphysial fontanelle large, rectangular, extending from ethmoid region to epiphysial bar. Postepiphysial fontanelle large, irregularly shaped, extending from epiphysial bar to occipital region. Cranial fontanelle separated by a narrow strut of frontal bone directly dorsal to the epiphysial bar. Five infraorbital bones (IO1-5); IO1-3 platelike; IO4-5 reduced in width, composed of sensory-canal ossification only. Gill membranes joined to isthmus. Fifth ceratobranchial with four needle-like pharyngeal teeth, arranged in a single row. Branchiostegal rays 2; anteriormost branchiostegal articulating with medial face of anterior ceratohyal absent. Anterior swimbladder chamber surrounded by a thick peritoneal tunic, partially enclosed in a bony capsule formed anteriorly by lateral process of 2nd vertebral centrum, laterally by outer arm of os suspensorium. Posterior swimbladder chamber greatly reduced in size.</paragraph>
<caption id="DF00BBF26349FFBFC399FEAFFAC4FEC9" ID-Table-UUID="DF00BBF26349FFBFC399FEAFFAC4FEC9" box="[151,1378,304,328]" httpUri="http://table.plazi.org/id/DF00BBF26349FFBFC399FEAFFAC4FEC9" pageId="3" pageNumber="34" targetBox="[165,1403,363,1971]" targetIsTable="true" targetPageId="3">
<paragraph id="8BC0EB7A6349FFBFC399FEAFFAC4FEC9" blockId="3.[151,1378,304,328]" box="[151,1378,304,328]" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC399FEAFFEAEFEC6" bold="true" box="[151,264,304,328]" pageId="3" pageNumber="34">TABLE 1.</emphasis>
Selected morphometric and meristic data for 
<taxonomicName id="4C7F90F96349FFBFC1FCFEAEFB9DFEC6" box="[754,1083,305,328]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B37686349FFBFC1FCFEAEFB9DFEC6" box="[754,1083,305,328]" italics="true" pageId="3" pageNumber="34">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
holotype and 11 paratypes.
</paragraph>
</caption>
<paragraph id="8BC0EB7A6349FFBFC03EFEF4FB89F83D" pageId="3" pageNumber="34">
<table id="F97F19DA63490043C3ABFEF4FADDF83D" box="[165,1403,363,1971]" gridcols="5" gridrows="38" pageId="3" pageNumber="34">
<tr id="354FE93863490043C3ABFEF4FADDFE0D" box="[165,1403,363,387]" gridrow="0" pageId="3" pageNumber="34" rowspan-0="1">
<th id="769E804463490043C03EFEF4FC3EFE0D" box="[816,920,363,387]" gridcol="1" gridrow="0" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC03EFEF4FC3EFE0D" bold="true" box="[816,920,363,387]" pageId="3" pageNumber="34">Holotype</emphasis>
</th>
<th id="769E804463490043C0EAFEF4FBFBFE0D" box="[996,1117,363,387]" gridcol="2" gridrow="0" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC0F2FEF4FBE3FE0D" bold="true" box="[1020,1093,363,387]" pageId="3" pageNumber="34">Range</emphasis>
</th>
<th id="769E804463490043C7ADFEF4FB42FE0D" box="[1187,1252,363,387]" gridcol="3" gridrow="0" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC7ADFEF4FB42FE0D" bold="true" box="[1187,1252,363,387]" pageId="3" pageNumber="34">Mean</emphasis>
</th>
<th id="769E804463490043C62DFEF4FADDFE0D" box="[1315,1403,363,387]" gridcol="4" gridrow="0" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC62DFEF4FADDFE0D" bold="true" box="[1315,1403,363,387]" pageId="3" pageNumber="34">St.Dev.</emphasis>
</th>
</tr>
<tr id="354FE93863490043C3ABFE0BFADDFE25" box="[165,1403,404,427]" gridrow="1" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABFE0BFDF5FE25" box="[165,595,404,427]" gridcol="0" gridrow="1" pageId="3" pageNumber="34">Standard length (mm)</th>
<td id="769E804463490043C03EFE0BFC3EFE25" box="[816,920,404,427]" gridcol="1" gridrow="1" pageId="3" pageNumber="34">44.9</td>
<td id="769E804463490043C0EAFE0BFBFBFE25" box="[996,1117,404,427]" gridcol="2" gridrow="1" pageId="3" pageNumber="34">28.2–51.1</td>
</tr>
<tr id="354FE93863490043C3ABFE4CFADDFE65" box="[165,1403,467,491]" gridrow="2" pageId="3" pageNumber="34" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABFE4CFDF5FE65" box="[165,595,467,491]" gridcol="0" gridrow="2" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC3ABFE4CFE31FE65" bold="true" box="[165,407,467,491]" pageId="3" pageNumber="34">% of standard length</emphasis>
</th>
</tr>
<tr id="354FE93863490043C3ABFE63FADDFD9D" box="[165,1403,508,531]" gridrow="3" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFE63FDF5FD9D" box="[165,595,508,531]" gridcol="0" gridrow="3" pageId="3" pageNumber="34">Body depth</th>
<td id="769E804463490043C03EFE63FC3EFD9D" box="[816,920,508,531]" gridcol="1" gridrow="3" pageId="3" pageNumber="34">19.4</td>
<td id="769E804463490043C0EAFE63FBFBFD9D" box="[996,1117,508,531]" gridcol="2" gridrow="3" pageId="3" pageNumber="34">18.6–21.9</td>
<td id="769E804463490043C7ADFE63FB42FD9D" box="[1187,1252,508,531]" gridcol="3" gridrow="3" pageId="3" pageNumber="34">20.0</td>
<td id="769E804463490043C62DFE63FADDFD9D" box="[1315,1403,508,531]" gridcol="4" gridrow="3" pageId="3" pageNumber="34">1.1</td>
</tr>
<tr id="354FE93863490043C3ABFDBBFADDFDB5" box="[165,1403,548,571]" gridrow="4" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFDBBFDF5FDB5" box="[165,595,548,571]" gridcol="0" gridrow="4" pageId="3" pageNumber="34">Head length</th>
<td id="769E804463490043C03EFDBBFC3EFDB5" box="[816,920,548,571]" gridcol="1" gridrow="4" pageId="3" pageNumber="34">20.9</td>
<td id="769E804463490043C0EAFDBBFBFBFDB5" box="[996,1117,548,571]" gridcol="2" gridrow="4" pageId="3" pageNumber="34">19.6–23.2</td>
<td id="769E804463490043C7ADFDBBFB42FDB5" box="[1187,1252,548,571]" gridcol="3" gridrow="4" pageId="3" pageNumber="34">21.0</td>
<td id="769E804463490043C62DFDBBFADDFDB5" box="[1315,1403,548,571]" gridcol="4" gridrow="4" pageId="3" pageNumber="34">1.0</td>
</tr>
<tr id="354FE93863490043C3ABFDD3FADDFDED" box="[165,1403,588,611]" gridrow="5" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFDD3FDF5FDED" box="[165,595,588,611]" gridcol="0" gridrow="5" pageId="3" pageNumber="34">Pre-dorsal length</th>
<td id="769E804463490043C03EFDD3FC3EFDED" box="[816,920,588,611]" gridcol="1" gridrow="5" pageId="3" pageNumber="34">48.5</td>
<td id="769E804463490043C0EAFDD3FBFBFDED" box="[996,1117,588,611]" gridcol="2" gridrow="5" pageId="3" pageNumber="34">47.7–53.6</td>
<td id="769E804463490043C7ADFDD3FB42FDED" box="[1187,1252,588,611]" gridcol="3" gridrow="5" pageId="3" pageNumber="34">50.0</td>
<td id="769E804463490043C62DFDD3FADDFDED" box="[1315,1403,588,611]" gridcol="4" gridrow="5" pageId="3" pageNumber="34">1.5</td>
</tr>
<tr id="354FE93863490043C3ABFDEBFADDFD05" box="[165,1403,628,651]" gridrow="6" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFDEBFDF5FD05" box="[165,595,628,651]" gridcol="0" gridrow="6" pageId="3" pageNumber="34">Pre-pectoral length</th>
<td id="769E804463490043C03EFDEBFC3EFD05" box="[816,920,628,651]" gridcol="1" gridrow="6" pageId="3" pageNumber="34">19.6</td>
<td id="769E804463490043C0EAFDEBFBFBFD05" box="[996,1117,628,651]" gridcol="2" gridrow="6" pageId="3" pageNumber="34">17.0–20.9</td>
<td id="769E804463490043C7ADFDEBFB42FD05" box="[1187,1252,628,651]" gridcol="3" gridrow="6" pageId="3" pageNumber="34">19.1</td>
<td id="769E804463490043C62DFDEBFADDFD05" box="[1315,1403,628,651]" gridcol="4" gridrow="6" pageId="3" pageNumber="34">1.2</td>
</tr>
<tr id="354FE93863490043C3ABFD03FADDFD3D" box="[165,1403,668,691]" gridrow="7" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFD03FDF5FD3D" box="[165,595,668,691]" gridcol="0" gridrow="7" pageId="3" pageNumber="34">Pre-pelvic length</th>
<td id="769E804463490043C03EFD03FC3EFD3D" box="[816,920,668,691]" gridcol="1" gridrow="7" pageId="3" pageNumber="34">48.7</td>
<td id="769E804463490043C0EAFD03FBFBFD3D" box="[996,1117,668,691]" gridcol="2" gridrow="7" pageId="3" pageNumber="34">48.2–52.4</td>
<td id="769E804463490043C7ADFD03FB42FD3D" box="[1187,1252,668,691]" gridcol="3" gridrow="7" pageId="3" pageNumber="34">49.8</td>
<td id="769E804463490043C62DFD03FADDFD3D" box="[1315,1403,668,691]" gridcol="4" gridrow="7" pageId="3" pageNumber="34">1.3</td>
</tr>
<tr id="354FE93863490043C3ABFD5BFADDFD55" box="[165,1403,708,731]" gridrow="8" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFD5BFDF5FD55" box="[165,595,708,731]" gridcol="0" gridrow="8" pageId="3" pageNumber="34">Pre-anal length</th>
<td id="769E804463490043C03EFD5BFC3EFD55" box="[816,920,708,731]" gridcol="1" gridrow="8" pageId="3" pageNumber="34">79.9</td>
<td id="769E804463490043C0EAFD5BFBFBFD55" box="[996,1117,708,731]" gridcol="2" gridrow="8" pageId="3" pageNumber="34">78.2–82.4</td>
<td id="769E804463490043C7ADFD5BFB42FD55" box="[1187,1252,708,731]" gridcol="3" gridrow="8" pageId="3" pageNumber="34">79.6</td>
<td id="769E804463490043C62DFD5BFADDFD55" box="[1315,1403,708,731]" gridcol="4" gridrow="8" pageId="3" pageNumber="34">1.8</td>
</tr>
<tr id="354FE93863490043C3ABFD73FADDFC8D" box="[165,1403,748,771]" gridrow="9" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFD73FDF5FC8D" box="[165,595,748,771]" gridcol="0" gridrow="9" pageId="3" pageNumber="34">Snout to anus length</th>
<td id="769E804463490043C03EFD73FC3EFC8D" box="[816,920,748,771]" gridcol="1" gridrow="9" pageId="3" pageNumber="34">57.2</td>
<td id="769E804463490043C0EAFD73FBFBFC8D" box="[996,1117,748,771]" gridcol="2" gridrow="9" pageId="3" pageNumber="34">57.2–60.0</td>
<td id="769E804463490043C7ADFD73FB42FC8D" box="[1187,1252,748,771]" gridcol="3" gridrow="9" pageId="3" pageNumber="34">58.7</td>
<td id="769E804463490043C62DFD73FADDFC8D" box="[1315,1403,748,771]" gridcol="4" gridrow="9" pageId="3" pageNumber="34">1.1</td>
</tr>
<tr id="354FE93863490043C3ABFC8BFADDFCA5" box="[165,1403,788,811]" gridrow="10" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFC8BFDF5FCA5" box="[165,595,788,811]" gridcol="0" gridrow="10" pageId="3" pageNumber="34">Anus to anal fin length</th>
<td id="769E804463490043C03EFC8BFC3EFCA5" box="[816,920,788,811]" gridcol="1" gridrow="10" pageId="3" pageNumber="34">21.6</td>
<td id="769E804463490043C0EAFC8BFBFBFCA5" box="[996,1117,788,811]" gridcol="2" gridrow="10" pageId="3" pageNumber="34">18.5–22.3</td>
<td id="769E804463490043C7ADFC8BFB42FCA5" box="[1187,1252,788,811]" gridcol="3" gridrow="10" pageId="3" pageNumber="34">20.3</td>
<td id="769E804463490043C62DFC8BFADDFCA5" box="[1315,1403,788,811]" gridcol="4" gridrow="10" pageId="3" pageNumber="34">1.1</td>
</tr>
<tr id="354FE93863490043C3ABFCA3FADDFCDD" box="[165,1403,828,851]" gridrow="11" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFCA3FDF5FCDD" box="[165,595,828,851]" gridcol="0" gridrow="11" pageId="3" pageNumber="34">Caudal-peduncle length</th>
<td id="769E804463490043C03EFCA3FC3EFCDD" box="[816,920,828,851]" gridcol="1" gridrow="11" pageId="3" pageNumber="34">15.1</td>
<td id="769E804463490043C0EAFCA3FBFBFCDD" box="[996,1117,828,851]" gridcol="2" gridrow="11" pageId="3" pageNumber="34">12.2–15.2</td>
<td id="769E804463490043C7ADFCA3FB42FCDD" box="[1187,1252,828,851]" gridcol="3" gridrow="11" pageId="3" pageNumber="34">14.4</td>
<td id="769E804463490043C62DFCA3FADDFCDD" box="[1315,1403,828,851]" gridcol="4" gridrow="11" pageId="3" pageNumber="34">1.2</td>
</tr>
<tr id="354FE93863490043C3ABFCFBFADDFCF5" box="[165,1403,868,891]" gridrow="12" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFCFBFDF5FCF5" box="[165,595,868,891]" gridcol="0" gridrow="12" pageId="3" pageNumber="34">Caudal-peduncle depth</th>
<td id="769E804463490043C03EFCFBFC3EFCF5" box="[816,920,868,891]" gridcol="1" gridrow="12" pageId="3" pageNumber="34">9.1</td>
<td id="769E804463490043C0EAFCFBFBFBFCF5" box="[996,1117,868,891]" gridcol="2" gridrow="12" pageId="3" pageNumber="34">8.0–9.5</td>
<td id="769E804463490043C7ADFCFBFB42FCF5" box="[1187,1252,868,891]" gridcol="3" gridrow="12" pageId="3" pageNumber="34">8.7</td>
<td id="769E804463490043C62DFCFBFADDFCF5" box="[1315,1403,868,891]" gridcol="4" gridrow="12" pageId="3" pageNumber="34">0.5</td>
</tr>
<tr id="354FE93863490043C3ABFC13FADDFC2D" box="[165,1403,908,931]" gridrow="13" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFC13FDF5FC2D" box="[165,595,908,931]" gridcol="0" gridrow="13" pageId="3" pageNumber="34">Caudal-peduncle width</th>
<td id="769E804463490043C03EFC13FC3EFC2D" box="[816,920,908,931]" gridcol="1" gridrow="13" pageId="3" pageNumber="34">5.3</td>
<td id="769E804463490043C0EAFC13FBFBFC2D" box="[996,1117,908,931]" gridcol="2" gridrow="13" pageId="3" pageNumber="34">4.2–5.4</td>
<td id="769E804463490043C7ADFC13FB42FC2D" box="[1187,1252,908,931]" gridcol="3" gridrow="13" pageId="3" pageNumber="34">4.8</td>
<td id="769E804463490043C62DFC13FADDFC2D" box="[1315,1403,908,931]" gridcol="4" gridrow="13" pageId="3" pageNumber="34">0.4</td>
</tr>
<tr id="354FE93863490043C3ABFC2BFADDFC45" box="[165,1403,948,971]" gridrow="14" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFC2BFDF5FC45" box="[165,595,948,971]" gridcol="0" gridrow="14" pageId="3" pageNumber="34">Pectoral-fin length</th>
<td id="769E804463490043C03EFC2BFC3EFC45" box="[816,920,948,971]" gridcol="1" gridrow="14" pageId="3" pageNumber="34">27.2</td>
<td id="769E804463490043C0EAFC2BFBFBFC45" box="[996,1117,948,971]" gridcol="2" gridrow="14" pageId="3" pageNumber="34">24.5–27.3</td>
<td id="769E804463490043C7ADFC2BFB42FC45" box="[1187,1252,948,971]" gridcol="3" gridrow="14" pageId="3" pageNumber="34">25.9</td>
<td id="769E804463490043C62DFC2BFADDFC45" box="[1315,1403,948,971]" gridcol="4" gridrow="14" pageId="3" pageNumber="34">0.9</td>
</tr>
<tr id="354FE93863490043C3ABFC43FADDFC7D" box="[165,1403,988,1011]" gridrow="15" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFC43FDF5FC7D" box="[165,595,988,1011]" gridcol="0" gridrow="15" pageId="3" pageNumber="34">Pelvic-fin length</th>
<td id="769E804463490043C03EFC43FC3EFC7D" box="[816,920,988,1011]" gridcol="1" gridrow="15" pageId="3" pageNumber="34">20</td>
<td id="769E804463490043C0EAFC43FBFBFC7D" box="[996,1117,988,1011]" gridcol="2" gridrow="15" pageId="3" pageNumber="34">17.8–20.8</td>
<td id="769E804463490043C7ADFC43FB42FC7D" box="[1187,1252,988,1011]" gridcol="3" gridrow="15" pageId="3" pageNumber="34">19.9</td>
<td id="769E804463490043C62DFC43FADDFC7D" box="[1315,1403,988,1011]" gridcol="4" gridrow="15" pageId="3" pageNumber="34">0.8</td>
</tr>
<tr id="354FE93863490043C3ABFB9BFADDFB95" box="[165,1403,1028,1051]" gridrow="16" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFB9BFDF5FB95" box="[165,595,1028,1051]" gridcol="0" gridrow="16" pageId="3" pageNumber="34">Length of last unbranched anal-fin ray</th>
<td id="769E804463490043C03EFB9BFC3EFB95" box="[816,920,1028,1051]" gridcol="1" gridrow="16" pageId="3" pageNumber="34">15.7</td>
<td id="769E804463490043C0EAFB9BFBFBFB95" box="[996,1117,1028,1051]" gridcol="2" gridrow="16" pageId="3" pageNumber="34">15.0–18.2</td>
<td id="769E804463490043C7ADFB9BFB42FB95" box="[1187,1252,1028,1051]" gridcol="3" gridrow="16" pageId="3" pageNumber="34">16.3</td>
<td id="769E804463490043C62DFB9BFADDFB95" box="[1315,1403,1028,1051]" gridcol="4" gridrow="16" pageId="3" pageNumber="34">1.0</td>
</tr>
<tr id="354FE93863490043C3ABFBB3FADDFBCD" box="[165,1403,1068,1091]" gridrow="17" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFBB3FDF5FBCD" box="[165,595,1068,1091]" gridcol="0" gridrow="17" pageId="3" pageNumber="34">Length of last unbranched dorsal-fin ray</th>
<td id="769E804463490043C03EFBB3FC3EFBCD" box="[816,920,1068,1091]" gridcol="1" gridrow="17" pageId="3" pageNumber="34">23.4</td>
<td id="769E804463490043C0EAFBB3FBFBFBCD" box="[996,1117,1068,1091]" gridcol="2" gridrow="17" pageId="3" pageNumber="34">18.9–23.8</td>
<td id="769E804463490043C7ADFBB3FB42FBCD" box="[1187,1252,1068,1091]" gridcol="3" gridrow="17" pageId="3" pageNumber="34">22.6</td>
<td id="769E804463490043C62DFBB3FADDFBCD" box="[1315,1403,1068,1091]" gridcol="4" gridrow="17" pageId="3" pageNumber="34">1.4</td>
</tr>
<tr id="354FE93863490043C3ABFBE4FADDFB1D" box="[165,1403,1147,1171]" gridrow="18" pageId="3" pageNumber="34" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABFBE4FDF5FB1D" box="[165,595,1147,1171]" gridcol="0" gridrow="18" pageId="3" pageNumber="34">
<emphasis id="B90B37686349FFBFC3ABFBE4FECEFB1D" bold="true" box="[165,360,1147,1171]" pageId="3" pageNumber="34">% of head length</emphasis>
</th>
</tr>
<tr id="354FE93863490043C3ABFB3BFADDFB35" box="[165,1403,1188,1211]" gridrow="19" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFB3BFDF5FB35" box="[165,595,1188,1211]" gridcol="0" gridrow="19" pageId="3" pageNumber="34">Head width</th>
<td id="769E804463490043C03EFB3BFC3EFB35" box="[816,920,1188,1211]" gridcol="1" gridrow="19" pageId="3" pageNumber="34">70.2</td>
<td id="769E804463490043C0EAFB3BFBFBFB35" box="[996,1117,1188,1211]" gridcol="2" gridrow="19" pageId="3" pageNumber="34">64.3–70.2</td>
<td id="769E804463490043C7ADFB3BFB42FB35" box="[1187,1252,1188,1211]" gridcol="3" gridrow="19" pageId="3" pageNumber="34">67.3</td>
<td id="769E804463490043C62DFB3BFADDFB35" box="[1315,1403,1188,1211]" gridcol="4" gridrow="19" pageId="3" pageNumber="34">2.6</td>
</tr>
<tr id="354FE93863490043C3ABFB53FADDFB6D" box="[165,1403,1228,1251]" gridrow="20" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFB53FDF5FB6D" box="[165,595,1228,1251]" gridcol="0" gridrow="20" pageId="3" pageNumber="34">Head depth</th>
<td id="769E804463490043C03EFB53FC3EFB6D" box="[816,920,1228,1251]" gridcol="1" gridrow="20" pageId="3" pageNumber="34">56.4</td>
<td id="769E804463490043C0EAFB53FBFBFB6D" box="[996,1117,1228,1251]" gridcol="2" gridrow="20" pageId="3" pageNumber="34">54.0–61.5</td>
<td id="769E804463490043C7ADFB53FB42FB6D" box="[1187,1252,1228,1251]" gridcol="3" gridrow="20" pageId="3" pageNumber="34">57.3</td>
<td id="769E804463490043C62DFB53FADDFB6D" box="[1315,1403,1228,1251]" gridcol="4" gridrow="20" pageId="3" pageNumber="34">2.5</td>
</tr>
<tr id="354FE93863490043C3ABFB6BFADDFA85" box="[165,1403,1268,1291]" gridrow="21" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFB6BFDF5FA85" box="[165,595,1268,1291]" gridcol="0" gridrow="21" pageId="3" pageNumber="34">Eye diameter</th>
<td id="769E804463490043C03EFB6BFC3EFA85" box="[816,920,1268,1291]" gridcol="1" gridrow="21" pageId="3" pageNumber="34">25.5</td>
<td id="769E804463490043C0EAFB6BFBFBFA85" box="[996,1117,1268,1291]" gridcol="2" gridrow="21" pageId="3" pageNumber="34">22.5–28.7</td>
<td id="769E804463490043C7ADFB6BFB42FA85" box="[1187,1252,1268,1291]" gridcol="3" gridrow="21" pageId="3" pageNumber="34">26.4</td>
<td id="769E804463490043C62DFB6BFADDFA85" box="[1315,1403,1268,1291]" gridcol="4" gridrow="21" pageId="3" pageNumber="34">1.8</td>
</tr>
<tr id="354FE93863490043C3ABFA83FADDFABD" box="[165,1403,1308,1331]" gridrow="22" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFA83FDF5FABD" box="[165,595,1308,1331]" gridcol="0" gridrow="22" pageId="3" pageNumber="34">Snout Length</th>
<td id="769E804463490043C03EFA83FC3EFABD" box="[816,920,1308,1331]" gridcol="1" gridrow="22" pageId="3" pageNumber="34">44.7</td>
<td id="769E804463490043C0EAFA83FBFBFABD" box="[996,1117,1308,1331]" gridcol="2" gridrow="22" pageId="3" pageNumber="34">43.1–48.4</td>
<td id="769E804463490043C7ADFA83FB42FABD" box="[1187,1252,1308,1331]" gridcol="3" gridrow="22" pageId="3" pageNumber="34">45.9</td>
<td id="769E804463490043C62DFA83FADDFABD" box="[1315,1403,1308,1331]" gridcol="4" gridrow="22" pageId="3" pageNumber="34">1.6</td>
</tr>
<tr id="354FE93863490043C3ABFADBFADDFAD5" box="[165,1403,1348,1371]" gridrow="23" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFADBFDF5FAD5" box="[165,595,1348,1371]" gridcol="0" gridrow="23" pageId="3" pageNumber="34">Interorbital width</th>
<td id="769E804463490043C03EFADBFC3EFAD5" box="[816,920,1348,1371]" gridcol="1" gridrow="23" pageId="3" pageNumber="34">44.7</td>
<td id="769E804463490043C0EAFADBFBFBFAD5" box="[996,1117,1348,1371]" gridcol="2" gridrow="23" pageId="3" pageNumber="34">37.7–45.0</td>
<td id="769E804463490043C7ADFADBFB42FAD5" box="[1187,1252,1348,1371]" gridcol="3" gridrow="23" pageId="3" pageNumber="34">40.9</td>
<td id="769E804463490043C62DFADBFADDFAD5" box="[1315,1403,1348,1371]" gridcol="4" gridrow="23" pageId="3" pageNumber="34">2.7</td>
</tr>
<tr id="354FE93863490043C3ABFAF3FADDFA0D" box="[165,1403,1388,1411]" gridrow="24" pageId="3" pageNumber="34">
<th id="769E804463490043C3ABFAF3FDF5FA0D" box="[165,595,1388,1411]" gridcol="0" gridrow="24" pageId="3" pageNumber="34">Mouth width</th>
<td id="769E804463490043C03EFAF3FC3EFA0D" box="[816,920,1388,1411]" gridcol="1" gridrow="24" pageId="3" pageNumber="34">30.8</td>
<td id="769E804463490043C0EAFAF3FBFBFA0D" box="[996,1117,1388,1411]" gridcol="2" gridrow="24" pageId="3" pageNumber="34">28.4–35.2</td>
<td id="769E804463490043C7ADFAF3FB42FA0D" box="[1187,1252,1388,1411]" gridcol="3" gridrow="24" pageId="3" pageNumber="34">31.7</td>
<td id="769E804463490043C62DFAF3FADDFA0D" box="[1315,1403,1388,1411]" gridcol="4" gridrow="24" pageId="3" pageNumber="34">2.0</td>
</tr>
<tr id="354FE93863490043C3ABFA23FADDFA5D" box="[165,1403,1468,1491]" gridrow="25" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABFA23FDF5FA5D" box="[165,595,1468,1491]" gridcol="0" gridrow="25" pageId="3" pageNumber="34">Dorsal-fin rays</th>
<td id="769E804463490043C03EFA23FC3EFA5D" box="[816,920,1468,1491]" gridcol="1" gridrow="25" pageId="3" pageNumber="34">iii.9</td>
<td id="769E804463490043C0EAFA23FBFBFA5D" box="[996,1117,1468,1491]" gridcol="2" gridrow="25" pageId="3" pageNumber="34">-</td>
</tr>
<tr id="354FE93863490043C3ABFA7BFADDFA75" box="[165,1403,1508,1531]" gridrow="26" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABFA7BFDF5FA75" box="[165,595,1508,1531]" gridcol="0" gridrow="26" pageId="3" pageNumber="34">Anal-fin rays</th>
<td id="769E804463490043C03EFA7BFC3EFA75" box="[816,920,1508,1531]" gridcol="1" gridrow="26" pageId="3" pageNumber="34">ii.6</td>
<td id="769E804463490043C0EAFA7BFBFBFA75" box="[996,1117,1508,1531]" gridcol="2" gridrow="26" pageId="3" pageNumber="34">-</td>
</tr>
<tr id="354FE93863490043C3ABF993FADDF9AD" box="[165,1403,1548,1571]" gridrow="27" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF993FDF5F9AD" box="[165,595,1548,1571]" gridcol="0" gridrow="27" pageId="3" pageNumber="34">Principal caudal fin-rays</th>
<td id="769E804463490043C03EF993FC3EF9AD" box="[816,920,1548,1571]" gridcol="1" gridrow="27" pageId="3" pageNumber="34">10+9</td>
<td id="769E804463490043C0EAF993FBFBF9AD" box="[996,1117,1548,1571]" gridcol="2" gridrow="27" pageId="3" pageNumber="34">-</td>
</tr>
<tr id="354FE93863490043C3ABF9ABFADDF9C5" box="[165,1403,1588,1611]" gridrow="28" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF9ABFDF5F9C5" box="[165,595,1588,1611]" gridcol="0" gridrow="28" pageId="3" pageNumber="34">Pectoral-fin rays</th>
<td id="769E804463490043C03EF9ABFC3EF9C5" box="[816,920,1588,1611]" gridcol="1" gridrow="28" pageId="3" pageNumber="34">v.10</td>
<td id="769E804463490043C0EAF9ABFBFBF9C5" box="[996,1117,1588,1611]" gridcol="2" gridrow="28" pageId="3" pageNumber="34">v.9–10</td>
</tr>
<tr id="354FE93863490043C3ABF9C3FADDF9FD" box="[165,1403,1628,1651]" gridrow="29" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF9C3FDF5F9FD" box="[165,595,1628,1651]" gridcol="0" gridrow="29" pageId="3" pageNumber="34">Pelvic-fin rays</th>
<td id="769E804463490043C03EF9C3FC3EF9FD" box="[816,920,1628,1651]" gridcol="1" gridrow="29" pageId="3" pageNumber="34">ii.7</td>
<td id="769E804463490043C0EAF9C3FBFBF9FD" box="[996,1117,1628,1651]" gridcol="2" gridrow="29" pageId="3" pageNumber="34">ii.6–7</td>
</tr>
<tr id="354FE93863490043C3ABF91BFADDF915" box="[165,1403,1668,1691]" gridrow="30" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF91BFDF5F915" box="[165,595,1668,1691]" gridcol="0" gridrow="30" pageId="3" pageNumber="34">Lateral-line scales</th>
<td id="769E804463490043C03EF91BFC3EF915" box="[816,920,1668,1691]" gridcol="1" gridrow="30" pageId="3" pageNumber="34">33</td>
<td id="769E804463490043C0EAF91BFBFBF915" box="[996,1117,1668,1691]" gridcol="2" gridrow="30" pageId="3" pageNumber="34">32–34</td>
</tr>
<tr id="354FE93863490043C3ABF933FADDF94D" box="[165,1403,1708,1731]" gridrow="31" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF933FDF5F94D" box="[165,595,1708,1731]" gridcol="0" gridrow="31" pageId="3" pageNumber="34">Scales between dorsal and pelvic fins</th>
<td id="769E804463490043C03EF933FC3EF94D" box="[816,920,1708,1731]" gridcol="1" gridrow="31" pageId="3" pageNumber="34">3.5/1/2</td>
<td id="769E804463490043C0EAF933FBFBF94D" box="[996,1117,1708,1731]" gridcol="2" gridrow="31" pageId="3" pageNumber="34">3.5/1/2–2.5</td>
</tr>
<tr id="354FE93863490043C3ABF94BFADDF965" box="[165,1403,1748,1771]" gridrow="32" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF94BFDF5F965" box="[165,595,1748,1771]" gridcol="0" gridrow="32" pageId="3" pageNumber="34">Circumpeduncular-scale rows</th>
<td id="769E804463490043C03EF94BFC3EF965" box="[816,920,1748,1771]" gridcol="1" gridrow="32" pageId="3" pageNumber="34">10</td>
<td id="769E804463490043C0EAF94BFBFBF965" box="[996,1117,1748,1771]" gridcol="2" gridrow="32" pageId="3" pageNumber="34">-</td>
</tr>
<tr id="354FE93863490043C3ABF963FADDF89D" box="[165,1403,1788,1811]" gridrow="33" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF963FDF5F89D" box="[165,595,1788,1811]" gridcol="0" gridrow="33" pageId="3" pageNumber="34">Preanal-scale rows</th>
<td id="769E804463490043C03EF963FC3EF89D" box="[816,920,1788,1811]" gridcol="1" gridrow="33" pageId="3" pageNumber="34">9</td>
<td id="769E804463490043C0EAF963FBFBF89D" box="[996,1117,1788,1811]" gridcol="2" gridrow="33" pageId="3" pageNumber="34">9–10</td>
</tr>
<tr id="354FE93863490043C3ABF8BBFADDF8B5" box="[165,1403,1828,1851]" gridrow="34" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF8BBFDF5F8B5" box="[165,595,1828,1851]" gridcol="0" gridrow="34" pageId="3" pageNumber="34">Predorsal-scale rows</th>
<td id="769E804463490043C03EF8BBFC3EF8B5" box="[816,920,1828,1851]" gridcol="1" gridrow="34" pageId="3" pageNumber="34">11</td>
<td id="769E804463490043C0EAF8BBFBFBF8B5" box="[996,1117,1828,1851]" gridcol="2" gridrow="34" pageId="3" pageNumber="34">10–12</td>
</tr>
<tr id="354FE93863490043C3ABF8D3FADDF8ED" box="[165,1403,1868,1891]" gridrow="35" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF8D3FDF5F8ED" box="[165,595,1868,1891]" gridcol="0" gridrow="35" pageId="3" pageNumber="34">Abdominal vertebrae</th>
<td id="769E804463490043C03EF8D3FC3EF8ED" box="[816,920,1868,1891]" gridcol="1" gridrow="35" pageId="3" pageNumber="34">-</td>
<td id="769E804463490043C0EAF8D3FBFBF8ED" box="[996,1117,1868,1891]" gridcol="2" gridrow="35" pageId="3" pageNumber="34">17–18</td>
</tr>
<tr id="354FE93863490043C3ABF8EBFADDF805" box="[165,1403,1908,1931]" gridrow="36" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF8EBFDF5F805" box="[165,595,1908,1931]" gridcol="0" gridrow="36" pageId="3" pageNumber="34">Caudal vertebrae</th>
<td id="769E804463490043C03EF8EBFC3EF805" box="[816,920,1908,1931]" gridcol="1" gridrow="36" pageId="3" pageNumber="34">-</td>
<td id="769E804463490043C0EAF8EBFBFBF805" box="[996,1117,1908,1931]" gridcol="2" gridrow="36" pageId="3" pageNumber="34">17–18</td>
</tr>
<tr id="354FE93863490043C3ABF803FADDF83D" box="[165,1403,1948,1971]" gridrow="37" pageId="3" pageNumber="34" rowspan-3="1" rowspan-4="1">
<th id="769E804463490043C3ABF803FDF5F83D" box="[165,595,1948,1971]" gridcol="0" gridrow="37" pageId="3" pageNumber="34">Total vertebrae</th>
<td id="769E804463490043C03EF803FC3EF83D" box="[816,920,1948,1971]" gridcol="1" gridrow="37" pageId="3" pageNumber="34">-</td>
<td id="769E804463490043C0EAF803FBFBF83D" box="[996,1117,1948,1971]" gridcol="2" gridrow="37" pageId="3" pageNumber="34">35</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC0EB7A634EFFB8C3C8FF07FCF8FE39" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="35">
Dorsal-fin rays 
<date id="FFC1CDBA634EFFB8C271FF07FE16FF3C" box="[383,432,152,178]" pageId="4" pageNumber="35">iii.9</date>
, anal-fin rays 
<date id="FFC1CDBA634EFFB8C156FF07FD23FF3C" box="[600,645,152,178]" pageId="4" pageNumber="35">ii.6.</date>
Principal caudal-fin rays 10+9, dorsal procurrent rays 7(2), ventral procurrent rays 5(1) or 6(1). Pelvic-fin rays 
<date id="FFC1CDBA634EFFB8C1A5FF22FD77FF59" box="[683,721,189,215]" pageId="4" pageNumber="35">ii.6</date>
(4) or 
<date id="FFC1CDBA634EFFB8C010FF22FCE0FF59" box="[798,838,189,215]" pageId="4" pageNumber="35">ii.7</date>
(13), pectoral-fin rays 
<date id="FFC1CDBA634EFFB8C75BFF22FBDBFF59" box="[1109,1149,189,215]" pageId="4" pageNumber="35">v.9</date>
(1), 
<date id="FFC1CDBA634EFFB8C7A5FF22FB7BFF59" box="[1195,1245,189,215]" pageId="4" pageNumber="35" value="1910-05">v.10</date>
(11) or 
<date id="FFC1CDBA634EFFB8C64EFF22FAD2FF59" box="[1344,1396,189,215]" pageId="4" pageNumber="35" value="1911-05">v.11</date>
(6). Paired fins large, horizontally placed. Pectoral fin reaching vertical through dorsal-fin origin, reaching to one or three scale rows anterior to pelvic-fin origin when adpressed. Pelvic-fin origin posterior to dorsal-fin origin, insertion opposite 2nd branched dorsal-fin ray. Skin on ventral surface of unbranched pectoral- and pelvic-fin rays thickened and covered in a layer of unculi, forming adhesive pads. Dorsal fin high, tip weakly rounded, posterior margin straight to weakly concave. Anal fin short, tip pointed, posterior margin concave, reaching to vertical through caudal-fin base when adpressed.
</paragraph>
<paragraph id="8BC0EB7A634EFFB8C3C8FE5DFE37FD19" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="35">Anus positioned 2 scale rows posterior to insertion of pelvic fins. Scales large, 32(2), 33(10) or 34(5) along lateral line, plus 1(2), 2(14) or 3(1) on base of caudal fin. 3.5/1/2(8) or 3.5/1/2.5(9) transverse scale rows from dorsal-fin origin to pelvic-fin origin, 10 around caudal peduncle, 10(1), 11(13) or 12(2) predorsal scales, 9(4) or 10(12) scales between anus and anal-fin origin. Ventral surface with a narrow, triangular scaleless patch between paired fins. Total number of vertebrae 35, comprising 17+18(3) or 18+17(1) abdominal and caudal vertebrae.</paragraph>
<paragraph id="8BC0EB7A634EFFB8C3C8FD3DFEBBFBD9" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="35">Small conical tubercles with keratinised tips distributed over entire surface of snout, lateral surface of head, rostral cap and papilliferous skin folds lateral to mouth. Head tuberculation more strongly developed in males. Tubercles on dorsal surface of head arranged as a series of small, irregular longitudinal ridges. Scales on anterior half of body with similar elongate tubercles, forming a series of small parallel ridges across scale surface in both sexes, more strongly developed in males. Posterior edges of scales on dorsal and lateral body surface of males each rimmed with four or five small conical tubercles with keratinized tip. Keratinized tips most greatly developed on tubercles on anterior half of body. Strip of minute conical tubercles, multiple rows thick, present along dorsal surface of unbranched and anteriormost branched pectoral-fin rays in males. Single row of minute conical tubercles present along dorsal surface of unbranched and anteriormost branched pectoral-fin rays in females. Single row of minute conical tubercles present along dorsal surface of unbranched and anteriormost branched pelvic-fin rays and lateral surface of anteriormost dorsal-fin rays of males only.</paragraph>
<paragraph id="8BC0EB7A634EFFB8C3C8FBFDFCC4F999" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="35">
<emphasis id="B90B3768634EFFB8C3C8FBFDFEF3FBF2" bold="true" box="[198,341,1122,1148]" pageId="4" pageNumber="35">Coloration.</emphasis>
In alcohol body background light cream (
<figureCitation id="1344F7FF634EFFB8C07CFBFDFC1BFBF2" box="[882,957,1122,1148]" captionStart="FIGURE 1" captionStartId="2.[151,255,1450,1474]" captionTargetBox="[151,1436,185,1425]" captionTargetId="figure@2.[151,1436,185,1426]" captionTargetPageId="2" captionText="FIGURE 1. Psilorhynchus brachyrhynchus, NRM 40935, male, 44.9 mm SL; Myanmar: Kachin State, Putao." httpUri="https://zenodo.org/record/197945/files/figure.png" pageId="4" pageNumber="35">Fig. 1</figureCitation>
). Occiput dark brown. Dorsal surface between occiput and dorsal fin with two faint brown saddles, first saddle situated closer to occiput than to dorsal-fin origin, extending between predorsal scales 4–6, second saddle situated 1–3 scales anterior to dorsal fin origin, extending to base of 1st branched dorsal-fin ray. Dorsal saddles between occiput and dorsal fin most obvious in smaller specimens (below 
<quantity id="4C87469F634EFFB8C191FB67FD55FA9C" box="[671,755,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="35" unit="mm" value="40.0">40 mm</quantity>
SL), becoming increasingly less visible with size due to occlusion by dark markings along posterior margins of predorsal scales. A small, prominent dark brown spot situated at anteriormost point of dorsal fin. Four brown saddles along dorsal surface between mid-dorsal fin and caudal-fin base. First saddle situated between insertions of branched dorsal-fins rays 3–9, second saddle directly between dorsal-fin and anal-fin origin, third saddle directly above anterior half of anal fin, fourth saddle situated anterior to caudal-fin base. Dorsal saddles between mid-dorsal fin and caudal-fin base decreasing in size caudad, extending ventrad 2–3 scale rows on body side, connecting with round to squarish dark-brown blotches arranged in a longitudinal row on flank.
</paragraph>
<paragraph id="8BC0EB7A634EFFB8C3C8F9BDFBB7F859" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="35">Flank with 5–7 indistinct round to squarish dark-brown blotches, arranged in a longitudinal row. In specimens with 5, first situated posterodorsal to opercle opening, extending beneath lateral-line scales 1–7 or 8, second extending beneath lateral-line scales 10–15, third extending beneath lateral-line scales 17–21, fourth extending beneath lateral-line scales 24–28, fifth situated at base of caudal fin, extending beneath lateral-line scales 30/31 to 32/33/34. Second and or third blotch described above often divided into two small separate dark-brown blotches in specimens with blotch counts of 6 or 7. Scale row situated below lateral-line scale row densely speckled with dark brown melanophores, forming an indistinct, irregular line between pectoral-fin base and pelvic-fin origin, continued posteriorly as an indistinct, frequently interrupted streak, ventrad to dark-brown round to squarish blotches. Scales at pelvic-fin origin darkly pigmented in a number of specimens, forming an indistinct dark brown pre-pelvic spot. Large melanophores, deeply situated in epidermis, forming short stripe between tip of snout and anterior edge of eye; a dark brown blotch-like marking on cheek below ventral margin of eye; another at center of opercle.</paragraph>
<paragraph id="8BC0EB7A634FFFB9C3C8FF07FAC6FD8A" blockId="5.[151,1436,152,666]" pageId="5" pageNumber="36">Ventral surface largely devoid of pigment except for melanophores on anterior edge of rostral cap; a small patch of melanophores situated beneath scales at anal-fin origin; a short line of melanophores posterior to anus, running along ventral midline beneath 3rd to 5th scale post anus in row between anus and anal-fin origin. Broad swath of small melanophores at base of pectoral fin. Anteriormost pectoral- and pelvic-fin rays lightly speckled with small dark melanophores. Pelvic fin with an indistinct stripe across its center, formed by small aggregations of melanophores around first branching point of branched rays. Dorsal fin intensely marked with a black proximal stripe running along base of rays; a more distal black stripe crossing center of fin, formed by small aggregations of melanophores around first branching point of branched rays. Caudal fin with two large black blotches, one at the base of each lobe; a single irregular black vertical bar across center of fin, formed by small aggregations of melanophores around first branching point of branched rays. Anal fin immaculate.</paragraph>
</subSubSection>
<subSubSection id="C365B8F1634FFFB9C3C8FD8FFE88FD14" pageId="5" pageNumber="36" type="distribution">
<paragraph id="8BC0EB7A634FFFB9C3C8FD8FFE88FD14" blockId="5.[151,1436,152,666]" pageId="5" pageNumber="36">
<emphasis id="B90B3768634FFFB9C3C8FD8FFEC1FDA4" bold="true" box="[198,359,528,554]" pageId="5" pageNumber="36">Distribution.</emphasis>
Known to date only from the upper Ayeyarwaddy River drainage, in the vicinity of Putao in northern 
<collectingCountry id="F368ABEA634FFFB9C204FDAAFE24FDC1" box="[266,386,565,591]" name="Myanmar" pageId="5" pageNumber="36">Myanmar</collectingCountry>
(
<figureCitation id="1344F7FF634FFFB9C29AFDAAFE45FDC1" box="[404,483,565,591]" captionStart="FIGURE 2" captionStartId="5.[151,255,1990,2014]" captionTargetBox="[395,1195,757,1966]" captionTargetId="figure@5.[395,1195,757,1970]" captionTargetPageId="5" captionText="FIGURE 2. Distribution of Psilorhynchus brachyrhynchus (square), P. piperatus (star) and P. gokkyi (circle)." httpUri="https://zenodo.org/record/197946/files/figure.png" pageId="5" pageNumber="36">Fig. 2</figureCitation>
). The 
<typeStatus id="54C455D8634FFFB9C13AFDAAFDCCFDC1" box="[564,618,565,591]" pageId="5" pageNumber="36">type</typeStatus>
locality, at the confluence of the Ma Kyaww Wa Chaung and its tributary the Nan Hto Chaung (
<figureCitation id="1344F7FF634FFFB9C101FDC5FDFEFDFA" box="[527,600,602,628]" captionStart="FIGURE 3" captionStartId="6.[151,255,682,706]" captionTargetBox="[416,1172,180,667]" captionTargetId="figure@6.[416,1172,180,668]" captionTargetPageId="6" captionText="FIGURE 3. Ma Kyaww Wa Chaung and its tributary Nan Hto Chaung, Putao, Kachin State, Myanmar. Type locality of Psilorhynchus brachyrhynchus. Photograph by S. O. Kullander." httpUri="https://zenodo.org/record/197947/files/figure.png" pageId="5" pageNumber="36">Fig. 3</figureCitation>
), is described in detail by 
<bibRefCitation id="EFEE968B634FFFB9C096FDC5FB6CFDFA" author="Kullander" box="[920,1226,602,628]" pageId="5" pageNumber="36" refString="Kullander, S. O. &amp; Britz, R. (2002) Revision of the family Badidae (Teleostei: Perciformes), with description of a new genus and ten new species. Ichthyological Exploration of Freshwaters, 13, 295 - 372." type="journal article" year="2002">Kullander &amp; Britz (2002)</bibRefCitation>
and 
<bibRefCitation id="EFEE968B634FFFB9C60BFDC5FE8DFD14" author="Kullander" pageId="5" pageNumber="36" refString="Kullander, S. O. &amp; Fang, F. (2005) Two new species of Puntius from northern Myanmar (Teleostei: Cyprinidae). Copeia, 2005, 290 - 302." type="journal article" year="2005">Kullander &amp; Fang (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF00BBF2634FFFB9C399F859FA80F853" box="[151,1318,1990,2014]" httpUri="https://zenodo.org/record/197946/files/figure.png" pageId="5" pageNumber="36" targetBox="[395,1195,757,1966]" targetPageId="5">
<paragraph id="8BC0EB7A634FFFB9C399F859FA80F853" blockId="5.[151,1318,1990,2014]" box="[151,1318,1990,2014]" pageId="5" pageNumber="36">
<emphasis id="B90B3768634FFFB9C399F859FEBFF850" bold="true" box="[151,281,1990,2014]" pageId="5" pageNumber="36">FIGURE 2.</emphasis>
Distribution of 
<taxonomicName id="4C7F90F9634FFFB9C2CAF858FCA8F850" box="[452,782,1991,2014]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634FFFB9C2CAF858FCA8F850" box="[452,782,1991,2014]" italics="true" pageId="5" pageNumber="36">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
(square), 
<taxonomicName id="4C7F90F9634FFFB9C077F858FC5FF850" box="[889,1017,1991,2014]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634FFFB9C077F858FC5FF850" box="[889,1017,1991,2014]" italics="true" pageId="5" pageNumber="36">P. piperatus</emphasis>
</taxonomicName>
(star) and 
<taxonomicName id="4C7F90F9634FFFB9C765F858FB6DF850" box="[1131,1227,1991,2014]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="gokkyi">
<emphasis id="B90B3768634FFFB9C765F858FB6DF850" box="[1131,1227,1991,2014]" italics="true" pageId="5" pageNumber="36">P. gokkyi</emphasis>
</taxonomicName>
(circle).
</paragraph>
</caption>
<caption id="DF00BBF2634CFFBAC399FD35FC98FD68" httpUri="https://zenodo.org/record/197947/files/figure.png" pageId="6" pageNumber="37" targetBox="[416,1172,180,667]" targetPageId="6">
<paragraph id="8BC0EB7A634CFFBAC399FD35FC98FD68" blockId="6.[151,1436,682,742]" pageId="6" pageNumber="37">
<emphasis id="B90B3768634CFFBAC399FD35FEBFFD4C" bold="true" box="[151,281,682,706]" pageId="6" pageNumber="37">FIGURE 3.</emphasis>
Ma Kyaww Wa Chaung and its tributary Nan Hto Chaung, Putao, Kachin State, Myanmar. Type locality of 
<taxonomicName id="4C7F90F9634CFFBAC399FD50FE47FD68" box="[151,481,719,742]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC399FD50FE47FD68" box="[151,481,719,742]" italics="true" pageId="6" pageNumber="37">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
. Photograph by S. O. Kullander.
</paragraph>
</caption>
<subSubSection id="C365B8F1634CFFBAC3C8FCA2FC89FCF3" pageId="6" pageNumber="37" type="etymology">
<paragraph id="8BC0EB7A634CFFBAC3C8FCA2FC89FCF3" blockId="6.[151,1437,828,2013]" pageId="6" pageNumber="37">
<emphasis id="B90B3768634CFFBAC3C8FCA2FEF5FCD9" bold="true" box="[198,339,829,855]" pageId="6" pageNumber="37">Etymology.</emphasis>
Latinized from the Greek words brachys (βραχυς), short, and rhynchos (ρυγχος), snout, in reference to the short snout of this species. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="C365B8F1634CFFBAC3C8FC17FD31F8AC" pageId="6" pageNumber="37" type="discussion">
<paragraph id="8BC0EB7A634CFFBAC3C8FC17FD31F8AC" blockId="6.[151,1437,828,2013]" pageId="6" pageNumber="37">
<emphasis id="B90B3768634CFFBAC3C8FC17FE99FC2C" bold="true" box="[198,319,904,930]" pageId="6" pageNumber="37">Remarks.</emphasis>
<taxonomicName id="4C7F90F9634CFFBAC245FC16FD1FFC2C" box="[331,697,905,930]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC245FC16FD1FFC2C" box="[331,697,905,930]" italics="true" pageId="6" pageNumber="37">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
most closely resembles 
<taxonomicName id="4C7F90F9634CFFBAC0E9FC16FB2FFC2C" box="[999,1161,905,930]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634CFFBAC0E9FC16FB2FFC2C" box="[999,1161,905,930]" italics="true" pageId="6" pageNumber="37">P. breviminor</emphasis>
</taxonomicName>
in general appearance and in several meristics characters, including number of principal caudal fin rays (10+9), and unbranched pectoral-fin rays (5). 
<taxonomicName id="4C7F90F9634CFFBAC29CFC4CFD5BFC62" box="[402,765,979,1004]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC29CFC4CFD5BFC62" box="[402,765,979,1004]" italics="true" pageId="6" pageNumber="37">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
is easily distinguished from 
<taxonomicName id="4C7F90F9634CFFBAC75DFC4CFB57FC62" box="[1107,1265,979,1004]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634CFFBAC75DFC4CFB57FC62" box="[1107,1265,979,1004]" italics="true" pageId="6" pageNumber="37">P. breviminor</emphasis>
</taxonomicName>
by features of its caudal-fin pigment pattern (
<figureCitation id="1344F7FF634CFFBAC107FC67FDF5FB9C" box="[521,595,1016,1042]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
B), which consists of two large black blotches, one at the base of each lobe, and a single irregular vertical black bar across the center of the fin, traversing both upper and lower lobes. In 
<taxonomicName id="4C7F90F9634CFFBAC208FBDCFE01FBD2" box="[262,423,1091,1116]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634CFFBAC208FBDCFE01FBD2" box="[262,423,1091,1116]" italics="true" pageId="6" pageNumber="37">P. breviminor</emphasis>
</taxonomicName>
the caudal fin lacks such distinctive blotches or vertical barring and instead exhibits irregular black markings, which are most prominent along the outer edge of the upper and lower lobes (
<figureCitation id="1344F7FF634CFFBAC662FBF7FF01FB29" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
C). Several other species of 
<taxonomicName id="4C7F90F9634CFFBAC2FDFB11FD3CFB29" box="[499,666,1166,1191]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B90B3768634CFFBAC2FDFB11FD3CFB29" box="[499,666,1166,1191]" italics="true" pageId="6" pageNumber="37">Psilorhynchus</emphasis>
</taxonomicName>
, similar in general appearance to 
<taxonomicName id="4C7F90F9634CFFBAC724FB11FAA2FB29" box="[1066,1284,1166,1191]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC724FB11FAA2FB29" box="[1066,1284,1166,1191]" italics="true" pageId="6" pageNumber="37">P. brachyrhynchus</emphasis>
</taxonomicName>
, also exhibit black vertical barring on the caudal fin, including 
<taxonomicName id="4C7F90F9634CFFBAC1EEFB2CFC0EFB42" box="[736,936,1203,1228]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634CFFBAC1EEFB2CFC0EFB42" box="[736,936,1203,1228]" italics="true" pageId="6" pageNumber="37">P. amplicephalus</emphasis>
</taxonomicName>
(see Arunachalum 
<emphasis id="B90B3768634CFFBAC782FB2CFB66FB42" box="[1164,1216,1203,1228]" italics="true" pageId="6" pageNumber="37">et al</emphasis>
. 2007), 
<taxonomicName id="4C7F90F9634CFFBAC62FFB2CFA3DFB42" box="[1313,1435,1203,1228]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634CFFBAC62FFB2CFA3DFB42" box="[1313,1435,1203,1228]" italics="true" pageId="6" pageNumber="37">P. balitora</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634CFFBAC3C4FB47FEC0FB7F" box="[202,358,1240,1265]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634CFFBAC3C4FB47FEC0FB7F" box="[202,358,1240,1265]" italics="true" pageId="6" pageNumber="37">P. nepalensis</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFEE968B634CFFBAC2A8FB47FD4CFB7C" author="Conway" box="[422,746,1240,1266]" pageId="6" pageNumber="37" refString="Conway, K. W. &amp; Mayden, R. L. (2008 b) Two new species of Psilorhynchus (Ostariophysi: Psilorhynchidae) with the redescription of P. balitora. Ichthyological Exploration of Freshwaters, 19, 215 - 232." type="journal article" year="2008" yearSuffix="b">Conway &amp; Mayden 2008b</bibRefCitation>
). In each of these species, the proximalmost caudal bar is dumbbell-shaped, being pinched at its center (
<figureCitation id="1344F7FF634CFFBAC1C4FB62FCB2FA99" box="[714,788,1277,1303]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
A), and it is likely that the blotches at the base of the caudal fin in 
<taxonomicName id="4C7F90F9634CFFBAC23BFABCFDB7FAB2" box="[309,529,1315,1340]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC23BFABCFDB7FAB2" box="[309,529,1315,1340]" italics="true" pageId="6" pageNumber="37">P. brachyrhynchus</emphasis>
</taxonomicName>
are homologous with the proximalmost bar in 
<taxonomicName id="4C7F90F9634CFFBAC74AFABCFAA9FAB2" box="[1092,1295,1315,1340]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634CFFBAC74AFABCFAA9FAB2" box="[1092,1295,1315,1340]" italics="true" pageId="6" pageNumber="37">P. amplicephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC610FABCFA3DFAB2" box="[1310,1435,1315,1340]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634CFFBAC610FABCFA3DFAB2" box="[1310,1435,1315,1340]" italics="true" pageId="6" pageNumber="37">P. balitora</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634CFFBAC3C2FAD7FEC9FAEF" box="[204,367,1352,1377]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634CFFBAC3C2FAD7FEC9FAEF" box="[204,367,1352,1377]" italics="true" pageId="6" pageNumber="37">P. nepalensis</emphasis>
</taxonomicName>
. Similarly, the vertical bar across the center of the caudal fin of 
<taxonomicName id="4C7F90F9634CFFBAC797FAD7FAD9FAEF" box="[1177,1407,1352,1377]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC797FAD7FAD9FAEF" box="[1177,1407,1352,1377]" italics="true" pageId="6" pageNumber="37">P. brachyrhynchus</emphasis>
</taxonomicName>
is considered to be homologous with the central caudal-fin bar of 
<taxonomicName id="4C7F90F9634CFFBAC08AFAF1FBEAFA09" box="[900,1100,1390,1415]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634CFFBAC08AFAF1FBEAFA09" box="[900,1100,1390,1415]" italics="true" pageId="6" pageNumber="37">P. amplicephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC755FAF1FB71FA09" box="[1115,1239,1390,1415]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634CFFBAC755FAF1FB71FA09" box="[1115,1239,1390,1415]" italics="true" pageId="6" pageNumber="37">P. balitora</emphasis>
</taxonomicName>
(
<figureCitation id="1344F7FF634CFFBAC7E9FAF2FA88FA09" box="[1255,1326,1389,1415]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
A) and 
<taxonomicName id="4C7F90F9634CFFBAC688FAF1FEB5FA22" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634CFFBAC688FAF1FEB5FA22" italics="true" pageId="6" pageNumber="37">P. nepalensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7F90F9634CFFBAC22CFA0CFD2BFA22" box="[290,653,1427,1452]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC22CFA0CFD2BFA22" box="[290,653,1427,1452]" italics="true" pageId="6" pageNumber="37">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
lacks the distalmost caudal-fin bar present in 
<taxonomicName id="4C7F90F9634CFFBAC7A0FA0CFAD1FA22" box="[1198,1399,1427,1452]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634CFFBAC7A0FA0CFAD1FA22" box="[1198,1399,1427,1452]" italics="true" pageId="6" pageNumber="37">P. amplicephalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC688FA0CFF53FA5F" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634CFFBAC688FA0CFF53FA5F" italics="true" pageId="6" pageNumber="37">P. balitora</emphasis>
</taxonomicName>
(
<figureCitation id="1344F7FF634CFFBAC20BFA27FEEDFA5C" box="[261,331,1464,1490]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
A) and 
<taxonomicName id="4C7F90F9634CFFBAC2ADFA27FD9AFA5F" box="[419,572,1464,1489]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634CFFBAC2ADFA27FD9AFA5F" box="[419,572,1464,1489]" italics="true" pageId="6" pageNumber="37">P. nepalensis</emphasis>
</taxonomicName>
. Other species of the genus lack vertical barring in the caudal fins (e.g. 
<taxonomicName id="4C7F90F9634CFFBAC688FA27FD32FA79" authority="Nebeshwar, Bagra &amp; Das" authorityName="Nebeshwar, Bagra &amp; Das" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="arunachalensis">
<emphasis id="B90B3768634CFFBAC688FA27FEECFA79" italics="true" pageId="6" pageNumber="37">P. arunachalensis</emphasis>
(Nebeshwar, Bagra &amp; Das)
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC1AEFA41FCAFFA79" box="[672,777,1502,1527]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gokkyi">
<emphasis id="B90B3768634CFFBAC1AEFA41FCAFFA79" box="[672,777,1502,1527]" italics="true" pageId="6" pageNumber="37">P. gokkyi</emphasis>
</taxonomicName>
[
<figureCitation id="1344F7FF634CFFBAC017FA42FCC4FA79" box="[793,866,1501,1527]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
D], 
<taxonomicName id="4C7F90F9634CFFBAC080FA41FBDAFA79" authority="Rainboth" authorityName="Rainboth" box="[910,1148,1501,1527]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B90B3768634CFFBAC080FA41FBA0FA79" box="[910,1030,1502,1527]" italics="true" pageId="6" pageNumber="37">P. gracilis</emphasis>
Rainboth
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC786FA41FF5CF993" authority="Hora &amp; Mukerji" authorityName="Hora &amp; Mukerji" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="homaloptera">
<emphasis id="B90B3768634CFFBAC786FA41FA9DFA79" box="[1160,1339,1502,1527]" italics="true" pageId="6" pageNumber="37">P. homaloptera</emphasis>
Hora &amp; Mukerji
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC208F99CFEDBF992" box="[262,381,1539,1564]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="melissa">
<emphasis id="B90B3768634CFFBAC208F99CFEDBF992" box="[262,381,1539,1564]" italics="true" pageId="6" pageNumber="37">P. melissa</emphasis>
</taxonomicName>
[
<figureCitation id="1344F7FF634CFFBAC283F99CFE71F993" box="[397,471,1539,1565]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
G], 
<taxonomicName id="4C7F90F9634CFFBAC10AF99CFB9FF993" authority="Vishwanath &amp; Manojkumar" authorityName="Vishwanath &amp; Manojkumar" box="[516,1081,1539,1565]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis id="B90B3768634CFFBAC10AF99CFD45F992" box="[516,739,1539,1564]" italics="true" pageId="6" pageNumber="37">P. microphthalmus</emphasis>
Vishwanath &amp; Manojkumar
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC748F99CFAA4F992" box="[1094,1282,1539,1564]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634CFFBAC748F99CFAA4F992" box="[1094,1282,1539,1564]" italics="true" pageId="6" pageNumber="37">P. pavimentatus</emphasis>
</taxonomicName>
[
<figureCitation id="1344F7FF634CFFBAC61CF99CFAFBF993" box="[1298,1373,1539,1565]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
E], 
<taxonomicName id="4C7F90F9634CFFBAC688F99CFEAEF9CF" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634CFFBAC688F99CFEAEF9CF" italics="true" pageId="6" pageNumber="37">P. piperatus</emphasis>
</taxonomicName>
[
<figureCitation id="1344F7FF634CFFBAC219F9B7FEC4F9CC" box="[279,354,1576,1602]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
F], 
<taxonomicName id="4C7F90F9634CFFBAC285F9B7FCBBF9CC" authority="Menon &amp; Datta" authorityName="Menon &amp; Datta" box="[395,797,1576,1602]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="pseudecheneis">
<emphasis id="B90B3768634CFFBAC285F9B7FDF3F9CF" box="[395,597,1576,1601]" italics="true" pageId="6" pageNumber="37">P. pseudecheneis</emphasis>
Menon &amp; Datta
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC027F9B7FB06F9CC" authority="Conway &amp; Kottelat" authorityName="Conway &amp; Kottelat" box="[809,1184,1576,1602]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B90B3768634CFFBAC027F9B7FC0AF9CF" box="[809,940,1576,1601]" italics="true" pageId="6" pageNumber="37">P. robustus</emphasis>
Conway &amp; Kottelat
</taxonomicName>
[
<figureCitation id="1344F7FF634CFFBAC7BEF9B7FB5DF9CC" box="[1200,1275,1576,1602]" captionStart="FIGURE 4" captionStartId="7.[151,256,1336,1360]" captionTargetBox="[152,1433,542,1298]" captionTargetId="figure@7.[151,1436,530,1314]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of Psilorhynchus and in P. balitora. A. Psilorhynchus balitora, KU 29191, 45.5 mm SL. B. Psilorhynchus brachyrhynchus, NRM 40922, paratype, 45.5 mm SL. C. Psilorhynchus breviminor, ZRC 51222, holotype, 35.9 mm SL. D. Psilorhynchus gokkyi, BMNH 2010.4. 14.1, holotype, 50.7 mm SL. E. Psilorhynchus pavimentatus, UMMZ 248831, holotype, 48.7 mm SL. F. Psilorhynchus piperatus, BMNH 2010.4. 14.8, holotype, 47.7 mm SL. G. Psilorhynchus melissa, UMMZ 248829, holotype, 47.6 mm SL. H. Psilorhynchus robustus, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens." httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="6" pageNumber="37">Fig. 4</figureCitation>
H], 
<taxonomicName id="4C7F90F9634CFFBAC626F9B7FEBFF9E9" authority="Hamilton" authorityName="Hamilton" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="sucatio">
<emphasis id="B90B3768634CFFBAC626F9B7FA3AF9CF" box="[1320,1436,1576,1601]" italics="true" pageId="6" pageNumber="37">P. sucatio</emphasis>
(Hamilton)
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634CFFBAC25FF9D1FC83F9E9" authority="Arunachalam &amp; Muralidharan" authorityName="Arunachalam &amp; Muralidharan" box="[337,805,1613,1639]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="tenura">
<emphasis id="B90B3768634CFFBAC25FF9D1FE1CF9E9" box="[337,442,1614,1639]" italics="true" pageId="6" pageNumber="37">P. tenura</emphasis>
Arunachalam &amp; Muralidharan
</taxonomicName>
) or lack caudal-fin pigmentation entirely (
<taxonomicName id="4C7F90F9634CFFBAC61AF9D1FA35F9E9" box="[1300,1427,1614,1639]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="rahmani">
<emphasis id="B90B3768634CFFBAC61AF9D1FA35F9E9" box="[1300,1427,1614,1639]" italics="true" pageId="6" pageNumber="37">P. rahmani</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFEE968B634CFFBAC399F9ECFE47F903" author="Conway" box="[151,481,1651,1677]" pageId="6" pageNumber="37" refString="Conway, K. W. &amp; Mayden, R. L. (2008 b) Two new species of Psilorhynchus (Ostariophysi: Psilorhynchidae) with the redescription of P. balitora. Ichthyological Exploration of Freshwaters, 19, 215 - 232." type="journal article" year="2008" yearSuffix="b">Conway &amp; Mayden 2008b</bibRefCitation>
). 
<taxonomicName id="4C7F90F9634CFFBAC2F8F9ECFCD6F902" box="[502,880,1651,1676]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC2F8F9ECFCD6F902" box="[502,880,1651,1676]" italics="true" pageId="6" pageNumber="37">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
is further distinguished from similar-looking species, including 
<emphasis id="B90B3768634CFFBAC261F907FD66F93F" box="[367,704,1688,1713]" italics="true" pageId="6" pageNumber="37">
<taxonomicName id="4C7F90F9634CFFBAC261F907FD9FF93F" box="[367,569,1688,1713]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="amplicephalus">P. amplicephalus</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC14BF907FD66F93F" box="[581,704,1688,1713]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="balitora">P. balitora</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C7F90F9634CFFBAC1C0F907FCCCF93F" box="[718,874,1688,1713]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634CFFBAC1C0F907FCCCF93F" box="[718,874,1688,1713]" italics="true" pageId="6" pageNumber="37">P. breviminor</emphasis>
</taxonomicName>
, 
<emphasis id="B90B3768634CFFBAC076F906FC7BF93C" box="[888,989,1689,1714]" italics="true" pageId="6" pageNumber="37">P. g o k k y i</emphasis>
, 
<taxonomicName id="4C7F90F9634CFFBAC0E1F906FB2EF93C" box="[1007,1160,1689,1714]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634CFFBAC0E1F906FB2EF93C" box="[1007,1160,1689,1714]" italics="true" pageId="6" pageNumber="37">P. nepalensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634CFFBAC798F906FAE8F93C" box="[1174,1358,1689,1714]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634CFFBAC798F906FAE8F93C" box="[1174,1358,1689,1714]" italics="true" pageId="6" pageNumber="37">P. pavimentatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7F90F9634CFFBAC688F906FEA0F959" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634CFFBAC688F906FEA0F959" italics="true" pageId="6" pageNumber="37">P. piperatus</emphasis>
</taxonomicName>
by its extremely short snout, which is only 43–48% HL (vs. 48–58% in 
<taxonomicName id="4C7F90F9634CFFBAC76CF921FA8DF959" box="[1122,1323,1726,1751]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="amplicephalus">
<emphasis id="B90B3768634CFFBAC76CF921FA8DF959" box="[1122,1323,1726,1751]" italics="true" pageId="6" pageNumber="37">P. amplicephalus</emphasis>
</taxonomicName>
, 50–53% in 
<emphasis id="B90B3768634CFFBAC3B6F97CFEE0F972" box="[184,326,1763,1788]" italics="true" pageId="6" pageNumber="37">
<taxonomicName id="4C7F90F9634CFFBAC3B6F97CFEE7F972" box="[184,321,1763,1788]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="balitora">P. balitora</taxonomicName>
,
</emphasis>
51–56% in 
<taxonomicName id="4C7F90F9634CFFBAC2EEF97CFD2FF972" box="[480,649,1763,1788]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634CFFBAC2EEF97CFD2FF972" box="[480,649,1763,1788]" italics="true" pageId="6" pageNumber="37">P. breviminor</emphasis>
</taxonomicName>
, 51–57% in 
<taxonomicName id="4C7F90F9634CFFBAC025F97CFC3BF972" box="[811,925,1763,1788]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gokkyi">
<emphasis id="B90B3768634CFFBAC025F97CFC3BF972" box="[811,925,1763,1788]" italics="true" pageId="6" pageNumber="37">P. gokkyi</emphasis>
</taxonomicName>
, 48–55% in 
<taxonomicName id="4C7F90F9634CFFBAC731F97CFB45F972" box="[1087,1251,1763,1788]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="B90B3768634CFFBAC731F97CFB45F972" box="[1087,1251,1763,1788]" italics="true" pageId="6" pageNumber="37">P. nepalensis</emphasis>
</taxonomicName>
, 50–55% in 
<taxonomicName id="4C7F90F9634CFFBAC68BF97CFE95F8AC" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634CFFBAC68BF97CFE95F8AC" italics="true" pageId="6" pageNumber="37">P. pavimentatus</emphasis>
</taxonomicName>
, and 53–55% in 
<emphasis id="B90B3768634CFFBAC2F4F896FD24F8AC" box="[506,642,1801,1826]" italics="true" pageId="6" pageNumber="37">P. p i p e r a t u s</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C365B8F1634CFFBBC3C8F8B1FC8FF833" lastPageId="7" lastPageNumber="38" pageId="6" pageNumber="37" type="reference_group">
<paragraph id="8BC0EB7A634CFFBBC3C8F8B1FE59FE39" blockId="6.[151,1437,828,2013]" lastBlockId="7.[151,1437,152,439]" lastPageId="7" lastPageNumber="38" pageId="6" pageNumber="37">
<taxonomicName id="4C7F90F9634CFFBAC3C8F8B1FD90F8C9" ID-CoL="4PJHT" box="[198,566,1838,1863]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC3C8F8B1FD90F8C9" box="[198,566,1838,1863]" italics="true" pageId="6" pageNumber="37">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
is the only member of the genus (and the only member of the order 
<taxonomicName id="4C7F90F9634CFFBAC399F8CCFEE0F8E3" ID-CoL="334" box="[151,326,1875,1901]" class="Actinopterygii" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="order">Cypriniformes</taxonomicName>
; Conway 
<emphasis id="B90B3768634CFFBAC2B7F8CCFE4DF8E2" box="[441,491,1875,1900]" italics="true" pageId="6" pageNumber="37">et al</emphasis>
. 2010) known to possess two branchiostegal rays (vs. three in all other species in which this character has been examined). The two branchiostegal rays of 
<taxonomicName id="4C7F90F9634CFFBAC707F8E6FB40F81C" ID-CoL="4PJHT" box="[1033,1254,1913,1938]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634CFFBAC707F8E6FB40F81C" box="[1033,1254,1913,1938]" italics="true" pageId="6" pageNumber="37">P. brachyrhynchus</emphasis>
</taxonomicName>
each articulate with the lateral face of the hyoid bar (the first at the suture between anterior and posterior ceratohyals and the second with the posterior ceratohyal) and are considered homologous to the two posteriormost branchiostegal rays of other cypriniform fishes. In a number of 
<taxonomicName id="4C7F90F9634DFFBBC018FF07FC69FF3F" ID-CoL="6434S" box="[790,975,152,177]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B90B3768634DFFBBC018FF07FC69FF3F" box="[790,975,152,177]" italics="true" pageId="7" pageNumber="38">Psilorhynchus</emphasis>
</taxonomicName>
(viz., 
<taxonomicName id="4C7F90F9634DFFBBC729FF07FB17FF3F" ID-CoL="4PJHS" box="[1063,1201,152,177]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634DFFBBC729FF07FB17FF3F" box="[1063,1201,152,177]" italics="true" pageId="7" pageNumber="38">P. balitora</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634DFFBBC7CAFF07FAD4FF3F" ID-CoL="78KG6" box="[1220,1394,152,177]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634DFFBBC7CAFF07FAD4FF3F" box="[1220,1394,152,177]" italics="true" pageId="7" pageNumber="38">P. breviminor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634DFFBBC68BFF07FE95FF58" ID-CoL="6WKG5" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="pavimentatus">
<emphasis id="B90B3768634DFFBBC68BFF07FE95FF58" italics="true" pageId="7" pageNumber="38">P. pavimentatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7F90F9634DFFBBC24FFF22FE66FF58" ID-CoL="4PJJH" box="[321,448,189,214]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="rahmani">
<emphasis id="B90B3768634DFFBBC24FFF22FE66FF58" box="[321,448,189,214]" italics="true" pageId="7" pageNumber="38">P. rahmani</emphasis>
</taxonomicName>
and 
<emphasis id="B90B3768634DFFBBC2F7FF22FDC0FF58" box="[505,614,189,214]" italics="true" pageId="7" pageNumber="38">P. s u c a t i o</emphasis>
) the anteriomost branchiostegal ray, which articulates with the medial face of the anterior ceratohyal, is composed of the articular head only and is much shorter than the two posterior rays. In 
<taxonomicName id="4C7F90F9634DFFBBC266FE97FDA0FEAF" ID-CoL="78KG6" box="[360,518,264,289]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="breviminor">
<emphasis id="B90B3768634DFFBBC266FE97FDA0FEAF" box="[360,518,264,289]" italics="true" pageId="7" pageNumber="38">P. breviminor</emphasis>
</taxonomicName>
the anteriormost branchiostegal ray exhibits asymmetry, being present on the left side only in two out of the three c&amp;s specimens examined. Reduction of the anteriomost branchiostegal ray is clearly a derived character and may be of utility in defining a monophyletic group within 
<taxonomicName id="4C7F90F9634DFFBBC7FBFECDFE4CFE1C" ID-CoL="6434S" authority="Conway &amp; Kottelat 2010" authorityName="Conway &amp; Kottelat" authorityYear="2010" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B90B3768634DFFBBC7FBFECDFA3AFEE5" box="[1269,1436,338,363]" italics="true" pageId="7" pageNumber="38">Psilorhynchus</emphasis>
(
<bibRefCitation id="EFEE968B634DFFBBC391FEE7FE44FE1C" author="Conway" box="[159,482,376,402]" pageId="7" pageNumber="38" refString="Conway, K. W. &amp; Kottelat, M. (2010) Two new species of Torrent Minnow (Ostariophysi: Psilorhynchidae: Psilorhynchus) from Western Myanmar. Raffles Zoology Bulletin, 58, 259 - 267." type="journal article" year="2010">Conway &amp; Kottelat 2010</bibRefCitation>
)
</taxonomicName>
. The complete absence of this element in 
<taxonomicName id="4C7F90F9634DFFBBC0F4FEE7FB46FE1F" ID-CoL="4PJHT" box="[1018,1248,376,401]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634DFFBBC0F4FEE7FB46FE1F" box="[1018,1248,376,401]" italics="true" pageId="7" pageNumber="38">P. brachyrhynchus</emphasis>
</taxonomicName>
should also be considered within this context.
</paragraph>
<caption id="DF00BBF2634DFFBBC399FAA7FDE6F9AF" httpUri="https://zenodo.org/record/197948/files/figure.png" pageId="7" pageNumber="38" targetBox="[152,1433,542,1298]" targetPageId="7">
<paragraph id="8BC0EB7A634DFFBBC399FAA7FDE6F9AF" blockId="7.[151,1437,1336,1569]" pageId="7" pageNumber="38">
<emphasis id="B90B3768634DFFBBC399FAA7FEBAFADE" bold="true" box="[151,284,1336,1360]" pageId="7" pageNumber="38">FIGURE 4.</emphasis>
Schematic illustration of caudal-fin pigmentation patterns in Myanmar species of 
<taxonomicName id="4C7F90F9634DFFBBC792FAA7FA90FAC1" ID-CoL="6434S" box="[1180,1334,1336,1359]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B90B3768634DFFBBC792FAA7FA90FAC1" box="[1180,1334,1336,1359]" italics="true" pageId="7" pageNumber="38">Psilorhynchus</emphasis>
</taxonomicName>
and in 
<taxonomicName id="4C7F90F9634DFFBBC686FAA7FF4BFAFC" ID-CoL="4PJHS" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634DFFBBC686FAA7FF4BFAFC" italics="true" pageId="7" pageNumber="38">P. balitora</emphasis>
</taxonomicName>
. A. 
<taxonomicName id="4C7F90F9634DFFBBC215FAC4FDB6FAFC" ID-CoL="4PJHS" box="[283,528,1371,1394]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634DFFBBC215FAC4FDB6FAFC" box="[283,528,1371,1394]" italics="true" pageId="7" pageNumber="38">Psilorhynchus balitora</emphasis>
</taxonomicName>
, KU 29191, 45.5 mm SL. B. 
<taxonomicName id="4C7F90F9634DFFBBC043FAC4FB31FAFC" ID-CoL="4PJHT" box="[845,1175,1371,1394]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634DFFBBC043FAC4FB31FAFC" box="[845,1175,1371,1394]" italics="true" pageId="7" pageNumber="38">Psilorhynchus brachyrhynchus</emphasis>
</taxonomicName>
, NRM 40922, paratype, 45.5 mm SL. C. 
<emphasis id="B90B3768634DFFBBC25AFAE2FDDBFA1A" box="[340,637,1405,1428]" italics="true" pageId="7" pageNumber="38">
<taxonomicName id="4C7F90F9634DFFBBC25AFAE2FDDFFA1A" ID-CoL="78KG6" box="[340,633,1405,1428]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="breviminor">Psilorhynchus breviminor</taxonomicName>
,
</emphasis>
ZRC 51222, holotype, 35.9 mm SL. D. 
<taxonomicName id="4C7F90F9634DFFBBC747FAE2FA9CFA1A" ID-CoL="4PJHX" box="[1097,1338,1405,1428]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="gokkyi">
<emphasis id="B90B3768634DFFBBC747FAE2FA9CFA1A" box="[1097,1338,1405,1428]" italics="true" pageId="7" pageNumber="38">Psilorhynchus gokkyi</emphasis>
</taxonomicName>
, BMNH 2010.4.14.1, holotype, 50.7 mm SL. E. 
<emphasis id="B90B3768634DFFBBC16EFA3FFC0DFA39" box="[608,939,1440,1463]" italics="true" pageId="7" pageNumber="38">
<taxonomicName id="4C7F90F9634DFFBBC16EFA3FFC01FA39" ID-CoL="6WKG5" box="[608,935,1440,1463]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="pavimentatus">Psilorhynchus pavimentatus</taxonomicName>
,
</emphasis>
UMMZ 248831, holotype, 48.7 mm SL. F. 
<taxonomicName id="4C7F90F9634DFFBBC399FA5CFE0CFA54" ID-CoL="4PJJF" box="[151,426,1475,1498]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="piperatus">
<emphasis id="B90B3768634DFFBBC399FA5CFE0CFA54" box="[151,426,1475,1498]" italics="true" pageId="7" pageNumber="38">Psilorhynchus piperatus</emphasis>
</taxonomicName>
, BMNH 2010.4.14.8, holotype, 47.7 mm SL. G. 
<taxonomicName id="4C7F90F9634DFFBBC0D9FA5CFB73FA54" ID-CoL="4PJJ8" box="[983,1237,1475,1498]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="melissa">
<emphasis id="B90B3768634DFFBBC0D9FA5CFB73FA54" box="[983,1237,1475,1498]" italics="true" pageId="7" pageNumber="38">Psilorhynchus melissa</emphasis>
</taxonomicName>
, UMMZ 248829, holotype, 47.6 mm SL. H. 
<taxonomicName id="4C7F90F9634DFFBBC2B2FA7AFD1FFA72" ID-CoL="4PJJJ" box="[444,697,1509,1532]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B90B3768634DFFBBC2B2FA7AFD1FFA72" box="[444,697,1509,1532]" italics="true" pageId="7" pageNumber="38">Psilorhynchus robustus</emphasis>
</taxonomicName>
, ZRC 51111, holotype, 60.3 mm SL. Tips of caudal-fin membranes frayed or torn in majority of specimens.
</paragraph>
</caption>
<paragraph id="8BC0EB7A634DFFBBC3C8F9E7FC8FF833" blockId="7.[151,1437,1656,1981]" pageId="7" pageNumber="38">
<bibRefCitation id="EFEE968B634DFFBBC3C8F9E7FE7CF91C" author="Mukerji" box="[198,474,1656,1682]" pageId="7" pageNumber="38" refString="Mukerji, D. D. (1933) Report on Burmese fishes collected by Lt. - Col. R. W. Burton from the tributary streams of the Mali Hka River of the Myitkyina District (Upper Burma). Part I. Journal of the Bombay Natural History Society, 36, 813 - 831." type="journal article" year="1933">Mukerji (1933: plate I)</bibRefCitation>
illustrated a specimen of 
<taxonomicName id="4C7F90F9634DFFBBC01DF9E7FC1CF91F" ID-CoL="6434S" box="[787,954,1656,1681]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="B90B3768634DFFBBC01DF9E7FC1CF91F" box="[787,954,1656,1681]" italics="true" pageId="7" pageNumber="38">Psilorhynchus</emphasis>
</taxonomicName>
collected from a tributary stream of the 
<collectingCountry id="F368ABEA634DFFBBC399F902FF68F939" box="[151,206,1693,1719]" name="Serbia" pageId="7" pageNumber="38">Mali</collectingCountry>
Hka River, northern 
<collectingCountry id="F368ABEA634DFFBBC2C2F902FD99F939" box="[460,575,1693,1719]" name="Myanmar" pageId="7" pageNumber="38">Myanmar</collectingCountry>
(Myitkyina District) (
<figureCitation id="1344F7FF634DFFBBC049F902FC37F939" box="[839,913,1693,1719]" captionStart="FIGURE 5" captionStartId="8.[151,255,680,704]" captionTargetBox="[159,1436,187,649]" captionTargetId="figure@8.[153,1438,187,651]" captionTargetPageId="8" captionText="FIGURE 5. Psilorhynchus brachyrhynchus as illustrated in Mukerji (1933: plate I)." httpUri="https://zenodo.org/record/197949/files/figure.png" pageId="7" pageNumber="38">Fig. 5</figureCitation>
). He referred this specimen to 
<taxonomicName id="4C7F90F9634DFFBBC60CF901FAD9F939" ID-CoL="4PJHS" box="[1282,1407,1694,1719]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634DFFBBC60CF901FAD9F939" box="[1282,1407,1694,1719]" italics="true" pageId="7" pageNumber="38">P. balitora</emphasis>
</taxonomicName>
, a species restricted to the Ganges-Brahmaputra drainage of northern 
<collectingCountry id="F368ABEA634DFFBBC0B5F95CFC5EF953" box="[955,1016,1731,1757]" name="India" pageId="7" pageNumber="38">India</collectingCountry>
(West Bengal and Assam), eastern 
<collectingCountry id="F368ABEA634DFFBBC399F977FF47F88C" box="[151,225,1768,1794]" name="Nepal" pageId="7" pageNumber="38">Nepal</collectingCountry>
, and northern 
<collectingCountry id="F368ABEA634DFFBBC282F977FDB0F88C" box="[396,534,1768,1794]" name="Bangladesh" pageId="7" pageNumber="38">Bangladesh</collectingCountry>
(
<bibRefCitation id="EFEE968B634DFFBBC129F977FCCCF88C" author="Conway" box="[551,874,1768,1794]" pageId="7" pageNumber="38" refString="Conway, K. W. &amp; Mayden, R. L. (2008 b) Two new species of Psilorhynchus (Ostariophysi: Psilorhynchidae) with the redescription of P. balitora. Ichthyological Exploration of Freshwaters, 19, 215 - 232." type="journal article" year="2008" yearSuffix="b">Conway &amp; Mayden 2008b</bibRefCitation>
). Based on features of the specimen figured by 
<bibRefCitation id="EFEE968B634DFFBBC399F892FEEBF8A9" author="Mukerji" box="[151,333,1805,1831]" pageId="7" pageNumber="38" refString="Mukerji, D. D. (1933) Report on Burmese fishes collected by Lt. - Col. R. W. Burton from the tributary streams of the Mali Hka River of the Myitkyina District (Upper Burma). Part I. Journal of the Bombay Natural History Society, 36, 813 - 831." type="journal article" year="1933">Mukerji (1933)</bibRefCitation>
, particularly the absence of scales from the midventral region and the presence of a distinctive stripe across the center of the dorsal fin, 
<bibRefCitation id="EFEE968B634DFFBBC17FF8ACFC1DF8C3" author="Conway" box="[625,955,1843,1869]" pageId="7" pageNumber="38" refString="Conway, K. W. &amp; Mayden, R. L. (2008 a) Psilorhynchus breviminor, a new species of psilorhynchid from Myanmar (Ostariophysi: Psilorhynchidae). Ichthyological Exploration of Freshwaters, 19, 111 - 120." type="journal article" year="2008" yearSuffix="a">Conway &amp; Mayden (2008a)</bibRefCitation>
argued that this specimen did not belong to 
<taxonomicName id="4C7F90F9634DFFBBC3B9F8C7FE9DF8FF" ID-CoL="4PJHS" box="[183,315,1880,1905]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="balitora">
<emphasis id="B90B3768634DFFBBC3B9F8C7FE9DF8FF" box="[183,315,1880,1905]" italics="true" pageId="7" pageNumber="38">P. balitora</emphasis>
</taxonomicName>
or 
<emphasis id="B90B3768634DFFBBC26BF8C7FDB6F8FF" box="[357,528,1880,1905]" italics="true" pageId="7" pageNumber="38">
<taxonomicName id="4C7F90F9634DFFBBC26BF8C7FDADF8FF" ID-CoL="78KG6" box="[357,523,1880,1905]" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="breviminor">P. breviminor</taxonomicName>
,
</emphasis>
later suggesting (
<bibRefCitation id="EFEE968B634DFFBBC1E3F8C7FB9EF8FC" author="Conway" box="[749,1080,1880,1906]" pageId="7" pageNumber="38" refString="Conway, K. W. &amp; Mayden, R. L. (2008 b) Two new species of Psilorhynchus (Ostariophysi: Psilorhynchidae) with the redescription of P. balitora. Ichthyological Exploration of Freshwaters, 19, 215 - 232." type="journal article" year="2008" yearSuffix="b">Conway &amp; Mayden 2008b</bibRefCitation>
) that it likely represented an undescribed species. We tentatively consider the specimen illustrated by 
<bibRefCitation id="EFEE968B634DFFBBC72CF8E2FB79F819" author="Mukerji" box="[1058,1247,1917,1943]" pageId="7" pageNumber="38" refString="Mukerji, D. D. (1933) Report on Burmese fishes collected by Lt. - Col. R. W. Burton from the tributary streams of the Mali Hka River of the Myitkyina District (Upper Burma). Part I. Journal of the Bombay Natural History Society, 36, 813 - 831." type="journal article" year="1933">Mukerji (1933)</bibRefCitation>
to belong to 
<taxonomicName id="4C7F90F9634DFFBBC688F8E1FEF5F832" ID-CoL="4PJHT" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B90B3768634DFFBBC688F8E1FEF5F832" italics="true" pageId="7" pageNumber="38">P. brachyrhynchus</emphasis>
</taxonomicName>
, until more material becomes available.
</paragraph>
</subSubSection>
</treatment>
</document>