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<document id="EC6D59D445AA409171B42A52D7BA1862" ID-DOI="10.11646/zootaxa.5293.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7961475" ID-ZooBank="8B6480D9-3A65-4319-BC4C-089F7AC6946F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1684826307319" checkinUser="plazi" docAuthor="Telnov, Dmitry &amp; Gusakov, Alexey A." docDate="2023" docId="03D63A302535FFC010EFFEA2FE59FDBA" docLanguage="en" docName="zootaxa.5293.3.7.pdf" docOrigin="Zootaxa 5293 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomosomus subtestaceus Motschulsky 1855" docType="treatment" docVersion="3" lastPageNumber="555" masterDocId="FFEF42482538FFCE1078FFE9FFABFF87" masterDocTitle="Rediscovery of Tomosomus Motschulsky, 1855 (Coleoptera: Anthicidae) with redescription of the genus, a new genus-rank synonym, and a key to species" masterLastPageNumber="556" masterPageNumber="541" pageNumber="554" updateTime="1684827086626" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CED4823BDCDB5165113B6B8A8F3D167F">Rediscovery of Tomosomus Motschulsky, 1855 (Coleoptera: Anthicidae) with redescription of the genus, a new genus-rank synonym, and a key to species</mods:title>
</mods:titleInfo>
<mods:name id="73DA2701C255A7D7D7A2CFD6F98F8CDF" type="personal">
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<mods:roleTerm id="D75BEFC0CA29A51EE9A22994BFC5D8B2">Author</mods:roleTerm>
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<mods:namePart id="E931FAEDCDF8FE81D099DCC766190AF2">Telnov, Dmitry</mods:namePart>
<mods:affiliation id="FF33541C72361FF60568BABB40392310">Department of Life Sciences, Natural History Museum, SW 7 5 BD, London, United Kingdom &amp; Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401, &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004, Rīga, Latvia</mods:affiliation>
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</mods:role>
<mods:namePart id="F14B8EF0C21FE5FB5F601A88AF698F10">Gusakov, Alexey A.</mods:namePart>
<mods:nameIdentifier id="1DDD84C6B722E83E2A561FA3A332BBB2" type="ORCID">0000-0001-7709-7515</mods:nameIdentifier>
<mods:affiliation id="6EFA355658081B510B0E4AF70CEA392A">Zoological Museum, Moscow Lomonosov State University, 2 Bol’shaya Nikitskaya Str., 125009, Moscow, Russia https: // orcid. org / 0000 - 0001 - 7709 - 7515</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D25742A48A97BE1EA1F4277EECAF4B26">text</mods:typeOfResource>
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<mods:title id="E17A8B971767EEEAE580E69A3C2F6A2F">Zootaxa</mods:title>
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<mods:part id="123BC69EBDB7498B303ECBDFE23A80EC">
<mods:date id="E94919627C8DA508B1B43D0F4B90DE88">2023</mods:date>
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<mods:number id="3ED028E38B780E7906C28C5175E0F288">2023-05-23</mods:number>
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<mods:number id="306CFCA4C63D151181DD846D11F6E67E">5293</mods:number>
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<mods:classification id="C4240C7F5E222806F1DE76C8174C23EA">journal article</mods:classification>
<mods:identifier id="C1FAEF15D91376B9984EB55F8EDAF7B5" type="DOI">10.11646/zootaxa.5293.3.6</mods:identifier>
<mods:identifier id="2B393DB1A0A7F7CD7130709071834CBA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4B70D093F39A834997F29C2694B70130" type="Zenodo-Dep">7961475</mods:identifier>
<mods:identifier id="3779E054D26C385E9CA7E0C45CD4FD7A" type="ZooBank">8B6480D9-3A65-4319-BC4C-089F7AC6946F</mods:identifier>
</mods:mods>
<treatment id="03D63A302535FFC010EFFEA2FE59FDBA" LSID="urn:lsid:plazi:treatment:03D63A302535FFC010EFFEA2FE59FDBA" httpUri="http://treatment.plazi.org/id/03D63A302535FFC010EFFEA2FE59FDBA" lastPageId="14" lastPageNumber="555" pageId="13" pageNumber="554">
<subSubSection id="C365D8AD2535FFC310EFFEA2FC9CFEE3" box="[151,823,330,357]" pageId="13" pageNumber="554" type="nomenclature">
<paragraph id="8BC08B262535FFC310EFFEA2FC9CFEE3" blockId="13.[151,823,330,357]" box="[151,823,330,357]" pageId="13" pageNumber="554">
<heading id="D0883C4A2535FFC310EFFEA2FC9CFEE3" bold="true" box="[151,823,330,357]" fontSize="11" level="1" pageId="13" pageNumber="554" reason="1">
<emphasis id="B90B57342535FFC310EFFEA2FC9CFEE3" bold="true" box="[151,823,330,357]" pageId="13" pageNumber="554">
<taxonomicName id="4C7FF0A52535FFC310EFFEA2FD1EFEE3" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[151,693,330,357]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="554" phylum="Arthropoda" rank="species" species="subtestaceus">
<emphasis id="B90B57342535FFC310EFFEA2FE6CFEE2" bold="true" box="[151,455,330,357]" italics="true" pageId="13" pageNumber="554">Tomosomus subtestaceus</emphasis>
<bibRefCitation id="EFEEF6D72535FFC311B5FEA2FD1EFEE3" author="Motschulsky, V." box="[461,693,330,357]" pageId="13" pageNumber="554" pagination="8 - 25" refId="ref6524" refString="Motschulsky, V. (1855) Voyages, Lettre de M. de Motschulsky a M. Menetries. № 2. Etudes entomologiques, [1855], 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
</taxonomicName>
(
<figureCitation id="134497A32535FFC312BCFEA3FC85FEE3" box="[708,814,330,356]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1731,1755]" captionStartId-2="5.[151,250,1943,1967]" captionStartId-3="6.[151,250,1927,1951]" captionTargetBox-0="[173,1407,187,1914]" captionTargetBox-1="[157,1430,187,1701]" captionTargetBox-2="[189,1420,201,1903]" captionTargetBox-3="[156,1426,190,1898]" captionTargetId-0="figure-27@2.[173,1414,181,1919]" captionTargetId-1="figure-17@3.[151,1436,181,1708]" captionTargetId-2="figure-17@5.[167,1420,181,1919]" captionTargetId-3="figure-27@6.[151,1436,181,1903]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Tomosomus subtestaceus Motschulsky, 1855. A—Paralectotypes ♀♀, habitus, laterodorsal view; B—Lectotype ♁ (upper specimen) and paralectotypes ♁♁, habitus, dorsal view; C—Labels [not to scale]." captionText-1="FIGURE 2. Tomosomus subtestaceus Motschulsky, 1855. A—Lectotype ♁, habitus, dorsal view; B—ditto, lateral view; C— Apex of elytra, tergite VII and aedeagus, dorsal view [not to scale]. Images courtesy Kirill V. Makarov (Moscow, Russia)." captionText-2="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." captionText-3="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi-0="http://doi.org/10.5281/zenodo.7961479" figureDoi-1="http://doi.org/10.5281/zenodo.7961481" figureDoi-2="http://doi.org/10.5281/zenodo.7961485" figureDoi-3="http://doi.org/10.5281/zenodo.7961487" httpUri-0="https://zenodo.org/record/7961479/files/figure.png" httpUri-1="https://zenodo.org/record/7961481/files/figure.png" httpUri-2="https://zenodo.org/record/7961485/files/figure.png" httpUri-3="https://zenodo.org/record/7961487/files/figure.png" pageId="13" pageNumber="554">Figs 1–4</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365D8AD2535FFC310EFFE79FCD9FE20" box="[151,882,399,423]" pageId="13" pageNumber="554" type="reference_group">
<paragraph id="8BC08B262535FFC310EFFE79FCD9FE20" blockId="13.[151,882,399,423]" box="[151,882,399,423]" pageId="13" pageNumber="554">
<treatmentCitationGroup id="AB6FAC082535FFC310EFFE79FCD9FE20" box="[151,882,399,423]" pageId="13" pageNumber="554">
<emphasis id="B90B57342535FFC310EFFE79FE00FE20" box="[151,427,399,423]" italics="true" pageId="13" pageNumber="554">
<taxonomicName id="4C7FF0A52535FFC310EFFE79FE3BFE20" authorityName="Motschulsky" authorityYear="1855" box="[151,400,399,423]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="554" phylum="Arthropoda" rank="species" species="subtestaceus">Tomosomus subtestaceus</taxonomicName>
—
</emphasis>
<treatmentCitation id="0ADEAD372535FFC311D3FE66FD35FE20" author="Motschulsky, V." box="[427,670,399,423]" page="18" pageId="13" pageNumber="554" year="1855">
<bibRefCitation id="EFEEF6D72535FFC311D3FE66FD35FE20" author="Motschulsky, V." box="[427,670,399,423]" pageId="13" pageNumber="554" pagination="8 - 25" refId="ref6524" refString="Motschulsky, V. (1855) Voyages, Lettre de M. de Motschulsky a M. Menetries. № 2. Etudes entomologiques, [1855], 8 - 25." type="journal article" year="1855">Motschulsky (1855: 18)</bibRefCitation>
</treatmentCitation>
original description.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C365D8AD2535FFC310EFFE3CFE8BFC97" pageId="13" pageNumber="554" type="materials_examined">
<paragraph id="8BC08B262535FFC310EFFE3CFACDFD93" blockId="13.[151,1437,469,2008]" pageId="13" pageNumber="554">
<emphasis id="B90B57342535FFC310EFFE3CFE6DFE68" bold="true" box="[151,454,469,495]" pageId="13" pageNumber="554">Type material examined.</emphasis>
<materialsCitation id="3B17817B2535FFC311A8FE3FFACDFD93" collectionCode="MZUM" country="Panama" location="Type" municipality="Obispo" pageId="13" pageNumber="554" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54C435842535FFC311A8FE3FFDEBFE77" box="[464,576,470,496]" pageId="13" pageNumber="554">Lectotype</typeStatus>
♁ [designated herewith]
<collectionCode id="ED6E13E32535FFC31313FE3FFC69FE77" box="[875,962,470,496]" pageId="13" pageNumber="554">MZUM</collectionCode>
(
<figureCitation id="134497A32535FFC313ACFE3CFB96FE68" box="[980,1085,469,495]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1731,1755]" captionTargetBox-0="[173,1407,187,1914]" captionTargetBox-1="[157,1430,187,1701]" captionTargetId-0="figure-27@2.[173,1414,181,1919]" captionTargetId-1="figure-17@3.[151,1436,181,1708]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Tomosomus subtestaceus Motschulsky, 1855. A—Paralectotypes ♀♀, habitus, laterodorsal view; B—Lectotype ♁ (upper specimen) and paralectotypes ♁♁, habitus, dorsal view; C—Labels [not to scale]." captionText-1="FIGURE 2. Tomosomus subtestaceus Motschulsky, 1855. A—Lectotype ♁, habitus, dorsal view; B—ditto, lateral view; C— Apex of elytra, tergite VII and aedeagus, dorsal view [not to scale]. Images courtesy Kirill V. Makarov (Moscow, Russia)." figureDoi-0="http://doi.org/10.5281/zenodo.7961479" figureDoi-1="http://doi.org/10.5281/zenodo.7961481" httpUri-0="https://zenodo.org/record/7961479/files/figure.png" httpUri-1="https://zenodo.org/record/7961481/files/figure.png" pageId="13" pageNumber="554">Figs 1–2</figureCitation>
): ly [handwritten, label green, circular] //
<location id="8EA0DDFD2535FFC3116BFE13FEE0FD93" LSID="urn:lsid:plazi:treatment:03D63A302535FFC010EFFEA2FE59FDBA:8EA0DDFD2535FFC3116BFE13FEE0FD93" box="[275,331,506,532]" country="Panama" municipality="Obispo" name="Type" pageId="13" pageNumber="554">Type</location>
[handwritten] //
<taxonomicName id="4C7FF0A52535FFC3127FFE13FCD5FD94" authority="Motsch" authorityName="Motschulsky" authorityYear="1855" box="[519,894,505,532]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="554" phylum="Arthropoda" rank="species" species="subtestaceus">Tomosomus subtestaceus Motsch</taxonomicName>
<collectingCountry id="F368CBB62535FFC313FDFE13FC75FD93" box="[901,990,506,532]" name="Panama" pageId="13" pageNumber="554">Panama</collectingCountry>
<collectingMunicipality id="6BA4115C2535FFC3139CFE10FB9EFD94" box="[996,1077,505,531]" pageId="13" pageNumber="554">Obispo</collectingMunicipality>
[handwritten, label green].
</materialsCitation>
</paragraph>
<paragraph id="8BC08B262535FFC310EFFDF4FBA6FDB0" blockId="13.[151,1437,469,2008]" box="[151,1037,541,570]" pageId="13" pageNumber="554">
<materialsCitation id="3B17817B2535FFC310EFFDF4FBA2FDB0" box="[151,1033,541,570]" collectionCode="MZUM" pageId="13" pageNumber="554" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<typeStatus id="54C435842535FFC310EFFDF4FE91FDB0" box="[151,314,541,567]" pageId="13" pageNumber="554">Paralectotypes</typeStatus>
2♁ &amp;
<specimenCount id="9D7940AF2535FFC311F0FDF4FE00FDB0" box="[392,427,541,567]" pageId="13" pageNumber="554" type="female">2♀</specimenCount>
<collectionCode id="ED6E13E32535FFC311CAFDF7FDA2FDBF" box="[434,521,542,568]" pageId="13" pageNumber="554">MZUM</collectionCode>
(
<figureCitation id="134497A32535FFC31261FDF4FDF0FDB0" box="[537,603,541,567]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[173,1407,187,1914]" captionTargetId="figure-27@2.[173,1414,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Tomosomus subtestaceus Motschulsky, 1855. A—Paralectotypes ♀♀, habitus, laterodorsal view; B—Lectotype ♁ (upper specimen) and paralectotypes ♁♁, habitus, dorsal view; C—Labels [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961479" httpUri="https://zenodo.org/record/7961479/files/figure.png" pageId="13" pageNumber="554">Fig. 1</figureCitation>
): same labels as
<typeStatus id="54C435842535FFC31362FDF4FC2AFDB0" box="[794,897,541,567]" pageId="13" pageNumber="554">lectotype</typeStatus>
[not sexed]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC08B262535FFC310BFFDA8FC85FDDC" blockId="13.[151,1437,469,2008]" box="[199,814,577,604]" pageId="13" pageNumber="554">
[All
<typeStatus id="54C435842535FFC31083FDABFE80FDDB" box="[251,299,578,604]" pageId="13" pageNumber="554">type</typeStatus>
specimens are in reasonably good condition].
</paragraph>
<paragraph id="8BC08B262535FFC310BFFD8CFEAAFD40" blockId="13.[151,1437,469,2008]" pageId="13" pageNumber="554">
<emphasis id="B90B57342535FFC310BFFD8CFED9FDF8" bold="true" box="[199,370,613,639]" pageId="13" pageNumber="554">New material.</emphasis>
<materialsCitation id="3B17817B2535FFC31104FD8CFDCCFD07" box="[380,615,613,640]" collectionCode="MIUP" pageId="13" pageNumber="554" specimenCount="75">
<specimenCount id="9D7940AF2535FFC31104FD8CFDB3FDF8" box="[380,536,613,640]" pageId="13" pageNumber="554" type="generic">75 specimens</specimenCount>
<collectionCode id="ED6E13E32535FFC31259FD8FFDCCFD07" box="[545,615,614,640]" pageId="13" pageNumber="554">MIUP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B17817B2535FFC3120DFD8CFCC5FD07" box="[629,878,613,640]" collectionCode="BMNH" pageId="13" pageNumber="554" specimenCount="10">
<specimenCount id="9D7940AF2535FFC3120DFD8CFCBAFDF8" box="[629,785,613,639]" pageId="13" pageNumber="554" type="generic">10 specimens</specimenCount>
<collectionCode id="ED6E13E32535FFC31362FD8FFCC5FD07" box="[794,878,614,640]" pageId="13" pageNumber="554">BMNH</collectionCode>
</materialsCitation>
&amp;
<specimenCount id="9D7940AF2535FFC313E0FD8CFB9FFDF8" box="[920,1076,613,639]" pageId="13" pageNumber="554" type="generic">30 specimens</specimenCount>
<materialsCitation id="3B17817B2535FFC31445FD8CFF56FD40" collectorName="San Lorenzo NP" country="Panama" location="Colon" pageId="13" pageNumber="554" specimenCount="1" stateProvince="Colon">
DTC (
<figureCitation id="134497A32535FFC314F0FD8CFB64FDF8" box="[1160,1231,613,639]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="13" pageNumber="554">Fig. 3</figureCitation>
): IBISCA Project
<collectingCountry id="F368CBB62535FFC310EFFD63FF5BFD23" box="[151,240,650,676]" name="Panama" pageId="13" pageNumber="554">Panama</collectingCountry>
<collectingRegion id="49BB45C42535FFC31082FD60FE95FD24" box="[250,318,649,675]" country="Panama" name="Colon" pageId="13" pageNumber="554">Colon</collectingRegion>
<collectorName id="268AEEF02535FFC31131FD60FDACFD23" box="[329,519,649,676]" pageId="13" pageNumber="554">San Lorenzo NP</collectorName>
[printed] // IBISCA 20030-05 Morphospec. Sort Juergen Schmidl COANTH 8 [printed]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC08B262535FFC310BFFD38FE8BFC97" blockId="13.[151,1437,469,2008]" pageId="13" pageNumber="554">
The
<typeStatus id="54C435842535FFC31082FD38FECAFD6C" box="[250,353,721,747]" pageId="13" pageNumber="554">lectotype</typeStatus>
designation is made in order to maintain nomenclatural stability since
<taxonomicName id="4C7FF0A52535FFC31401FD3BFA88FD6C" authorityName="Motschulsky" authorityYear="1855" box="[1145,1315,721,747]" class="Insecta" family="Anthicidae" genus="Tomosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="554" phylum="Arthropoda" rank="species" species="subtestaceus">
<emphasis id="B90B57342535FFC31401FD3BFA88FD6C" box="[1145,1315,721,747]" italics="true" pageId="13" pageNumber="554">T. subtestaceus</emphasis>
</taxonomicName>
is the type of its genus.
</paragraph>
</subSubSection>
<subSubSection id="C365D8AD2535FFC010BFFCF0FB1DFE0E" lastPageId="14" lastPageNumber="555" pageId="13" pageNumber="554" type="description">
<paragraph id="8BC08B262535FFC310BFFCF0FE1EFB1C" blockId="13.[151,1437,469,2008]" pageId="13" pageNumber="554">
<emphasis id="B90B57342535FFC310BFFCF0FEDEFCB4" bold="true" box="[199,373,793,819]" pageId="13" pageNumber="554">Redescription.</emphasis>
<typeStatus id="54C435842535FFC31105FCF3FE46FCB3" box="[381,493,794,820]" pageId="13" pageNumber="554">Lectotype</typeStatus>
♁, total body length
<quantity id="4C8726C32535FFC31299FCF0FC90FCB3" box="[737,827,793,820]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="554" unit="mm" value="2.2">2.2 mm</quantity>
; head
<quantity id="4C8726C32535FFC313FCFCF0FC42FCB3" box="[900,1001,793,820]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="13" pageNumber="554" unit="mm" value="0.42">0.42 mm</quantity>
long, across compound eyes
<quantity id="4C8726C32535FFC31540FCF0FA36FCB3" box="[1336,1437,793,820]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="554" unit="mm" value="0.33">0.33 mm</quantity>
broad, pronotum
<quantity id="4C8726C32535FFC31120FCD4FE10FCDF" box="[344,443,829,856]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="13" pageNumber="554" unit="mm" value="0.56">0.56 mm</quantity>
long, maximum width
<quantity id="4C8726C32535FFC312C6FCD4FCBDFCDF" box="[702,790,829,856]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="554" unit="mm" value="0.5">0.5 mm</quantity>
, minimum width
<quantity id="4C8726C32535FFC313A2FCD4FBEBFCDF" box="[986,1088,829,856]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="554" unit="mm" value="0.27">0.27 mm</quantity>
, elytra
<quantity id="4C8726C32535FFC314E9FCD4FB5FFCDF" box="[1169,1268,829,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="13" pageNumber="554" unit="mm" value="1.35">1.35 mm</quantity>
long,
<quantity id="4C8726C32535FFC31541FCD4FA36FCDF" box="[1337,1437,829,856]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="554" unit="mm" value="0.85">0.85 mm</quantity>
combined wide.
<typeStatus id="54C435842535FFC31137FC88FE4CFCFC" box="[335,487,865,891]" pageId="13" pageNumber="554">Paralectotype</typeStatus>
1 ♁, total body length
<quantity id="4C8726C32535FFC3129EFC88FCE7FCFB" box="[742,844,865,892]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="13" pageNumber="554" unit="mm" value="1.95">1.95 mm</quantity>
; head
<quantity id="4C8726C32535FFC313EAFC88FC5EFCFB" box="[914,1013,865,892]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="554" unit="mm" value="0.38">0.38 mm</quantity>
long, across compound eyes
<quantity id="4C8726C32535FFC31541FC88FA36FCFB" box="[1337,1437,865,892]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="554" unit="mm" value="0.27">0.27 mm</quantity>
broad, pronotum
<quantity id="4C8726C32535FFC31124FC6CFE6AFC27" box="[348,449,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="13" pageNumber="554" unit="mm" value="0.47">0.47 mm</quantity>
long, maximum width
<quantity id="4C8726C32535FFC312B5FC6CFC9EFC27" box="[717,821,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="554" unit="mm" value="0.45">0.45 mm</quantity>
, minimum width
<quantity id="4C8726C32535FFC31387FC6CFBCCFC27" box="[1023,1127,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="554" unit="mm" value="0.25">0.25 mm</quantity>
, elytra
<quantity id="4C8726C32535FFC314C4FC6CFA89FC27" box="[1212,1314,901,928]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="13" pageNumber="554" unit="mm" value="1.22">1.22 mm</quantity>
long,
<quantity id="4C8726C32535FFC31513FC6CFF68FC43" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="554" unit="mm" value="0.75">0.75 mm</quantity>
combined wide.
<typeStatus id="54C435842535FFC311FBFC40FDB0FC44" box="[387,539,937,963]" pageId="13" pageNumber="554">Paralectotype</typeStatus>
2 ♁, total body length 2.0 mm; head
<quantity id="4C8726C32535FFC313C7FC40FB89FC43" box="[959,1058,937,964]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="554" unit="mm" value="0.39">0.39 mm</quantity>
long, across compound eyes
<quantity id="4C8726C32535FFC31513FC40FF68FC6F" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="554" unit="mm" value="0.28">0.28 mm</quantity>
broad, pronotum
<quantity id="4C8726C32535FFC311E9FC24FE5CFC6F" box="[401,503,973,1000]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="554" unit="mm" value="0.49">0.49 mm</quantity>
long, maximum width
<quantity id="4C8726C32535FFC3137CFC24FCC7FC6F" box="[772,876,973,1000]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="554" unit="mm" value="0.45">0.45 mm</quantity>
, minimum width
<quantity id="4C8726C32535FFC3144FFC24FB34FC6F" box="[1079,1183,973,1000]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="554" unit="mm" value="0.27">0.27 mm</quantity>
, elytra
<quantity id="4C8726C32535FFC3148DFC24FAF0FC6F" box="[1269,1371,973,1000]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="554" unit="mm" value="1.25">1.25 mm</quantity>
long,
<quantity id="4C8726C32535FFC310EFFC18FF50FB8B" box="[151,251,1009,1036]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="13" pageNumber="554" unit="mm" value="0.77">0.77 mm</quantity>
combined wide.
<typeStatus id="54C435842535FFC311C3FC18FDF8FB8C" box="[443,595,1009,1035]" pageId="13" pageNumber="554">Paralectotype</typeStatus>
<specimenCount id="9D7940AF2535FFC31222FC18FD2FFB8B" box="[602,644,1009,1036]" pageId="13" pageNumber="554" type="female">3 ♀</specimenCount>
, total body length
<quantity id="4C8726C32535FFC3132FFC18FC04FB8B" box="[855,943,1009,1036]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="554" unit="mm" value="2.1">2.1 mm</quantity>
; head
<quantity id="4C8726C32535FFC3138EFC18FBF1FB8B" box="[1014,1114,1009,1036]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="13" pageNumber="554" unit="mm" value="0.42">0.42 mm</quantity>
long, across compound eyes
<quantity id="4C8726C32535FFC310EFFBFCFF51FBB7" box="[151,250,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="554" unit="mm" value="0.32">0.32 mm</quantity>
broad, pronotum
<quantity id="4C8726C32535FFC311C6FBFCFD8AFBB7" box="[446,545,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="554" unit="mm" value="0.52">0.52 mm</quantity>
long, maximum width
<quantity id="4C8726C32535FFC31359FBFCFC2EFBB7" box="[801,901,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="554" unit="mm" value="0.49">0.49 mm</quantity>
, minimum width
<quantity id="4C8726C32535FFC3143FFBFCFB07FBB7" box="[1095,1196,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="554" unit="mm" value="0.27">0.27 mm</quantity>
, elytra
<quantity id="4C8726C32535FFC31484FBFCFAF5FBB7" box="[1276,1374,1045,1072]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="13" pageNumber="554" unit="mm" value="1.27">1.27 mm</quantity>
long,
<quantity id="4C8726C32535FFC310EFFBD0FF51FBD3" box="[151,250,1081,1108]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="13" pageNumber="554" unit="mm" value="0.83">0.83 mm</quantity>
combined wide.
<typeStatus id="54C435842535FFC311C0FBD0FDFBFBD4" box="[440,592,1081,1107]" pageId="13" pageNumber="554">Paralectotype</typeStatus>
<specimenCount id="9D7940AF2535FFC3122FFBD0FD2BFBD3" box="[599,640,1081,1108]" pageId="13" pageNumber="554" type="female">4 ♀</specimenCount>
, total body length
<quantity id="4C8726C32535FFC31337FBD0FC1EFBD3" box="[847,949,1081,1108]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="13" pageNumber="554" unit="mm" value="2.25">2.25 mm</quantity>
; head
<quantity id="4C8726C32535FFC31382FBD0FBF6FBD3" box="[1018,1117,1081,1108]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="13" pageNumber="554" unit="mm" value="0.42">0.42 mm</quantity>
long, across compound eyes
<quantity id="4C8726C32535FFC310EFFBB4FF50FBFF" box="[151,251,1117,1144]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="554" unit="mm" value="0.33">0.33 mm</quantity>
broad, pronotum
<quantity id="4C8726C32535FFC311B9FBB4FD8FFBFF" box="[449,548,1117,1144]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="13" pageNumber="554" unit="mm" value="0.56">0.56 mm</quantity>
long, maximum width
<quantity id="4C8726C32535FFC31350FBB4FC26FBFF" box="[808,909,1117,1144]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="554" unit="mm" value="0.52">0.52 mm</quantity>
, minimum width
<quantity id="4C8726C32535FFC3142AFBB4FB1CFBFF" box="[1106,1207,1117,1144]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="554" unit="mm" value="0.28">0.28 mm</quantity>
, elytra
<quantity id="4C8726C32535FFC31570FBB4FAF6FBFF" box="[1288,1373,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="554" unit="mm" value="1.3">1.3 mm</quantity>
long,
<quantity id="4C8726C32535FFC310EFFB68FF50FB1B" box="[151,251,1153,1180]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="554" unit="mm" value="0.85">0.85 mm</quantity>
combined wide.
</paragraph>
<paragraph id="8BC08B262535FFC010BFFB4FFCDCFEC6" blockId="13.[151,1437,469,2008]" lastBlockId="14.[151,1437,151,573]" lastPageId="14" lastPageNumber="555" pageId="13" pageNumber="554">
Dorsum and venter brown, forebody somewhat paler brown. Mouthparts, antennae, palps and legs pale yellow, abdomen yellowish brown. Head subtriangular, glossy dorsally and ventrally, slightly convex in dorsal aspect. Anterolateral margin of frons slightly bulging between antennal insertion and internal anterior margin of a compound eye. Compound eye moderate, protruding beyond lateral outline of head. Tempus rounded and constricted towards subtruncate head base. Head dorsal punctures minute and inconspicuous. Head dorsal setae moderately long, sporadic. Antenna extending slightly beyond humeral area of elytron when directed posteriad. Antennomere three about same length as antennomere two, antennomeres 8–10 distinctly thickened, transverse. Terminal antennomere elongate triangular with tapered apex, asymmetrical, 2.25× as long as penultimate antennomere. Penultimate maxillary palpomere subtriangular. Terminal maxillary palpomere securiform. Pronotum dorsally and laterally glossy, slightly convex in dorsal aspect, anterior lobe wider than head across compound eyes. Front margin of anterior lobe subtruncate, dorsally and laterally with very narrow anterior rim. Anterior lobe with shallow, moderately broad, median longitudinal sulcus (this sulcus is moderately present, nearly obsolete in some specimens). Posterior lobe strongly narrower and shorter than anterior. Postmedian lateral constriction deep and broad, dorsally with a row of moderate punctures with raised, narrow, longitudinally directed intervening spaces, continues onto pronotal disc as a shallow depression (in lateral view). Pronotal punctures minute, sparse, intervening spaces significantly wider than punctures. Dorsal pronotal setae whitish, sparse, subdecumbent. Pronotal tactile setae yellowish, long, erect, sporadic, mainly aligned along with lateral margins of pronotum. Scutellar shield minute, subtriangular, rounded at posterior margin. Elytra ovoid, moderately convex in dorsal aspect, widened laterally before midlength, humerus obsolete (apterous species). Elytral punctures much stronger and larger than those on the dorsal forebody including in the lateral constriction area, becoming smaller on the posterior half of elytron. Intervening spaces glossy and glabrous, in the basal third of elytron as wide as to twice as wide as punctures. Posterior sutural angle prolonged into a short, acute denticle (
<figureCitation id="134497A32535FFC311A5F870FD97F834" box="[477,572,1945,1971]" captionStart="FIGURE 2" captionStartId="3.[151,250,1731,1755]" captionTargetBox="[157,1430,187,1701]" captionTargetId="figure-17@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Tomosomus subtestaceus Motschulsky, 1855. A—Lectotype ♁, habitus, dorsal view; B—ditto, lateral view; C— Apex of elytra, tergite VII and aedeagus, dorsal view [not to scale]. Images courtesy Kirill V. Makarov (Moscow, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7961481" httpUri="https://zenodo.org/record/7961481/files/figure.png" pageId="13" pageNumber="554">Figs 2C</figureCitation>
,
<figureCitation id="134497A32535FFC31234F870FDDAF834" box="[588,625,1945,1971]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="13" pageNumber="554">3A</figureCitation>
). Epipleuron nearly complete, moderately broad. Elytral setae whitish to yellowish, long, rather sparse, suberect to erect. All femora clavate. Protibia at posterior margin with distinct, obtusely angulate postmedian swelling (
<figureCitation id="134497A32536FFC01225FF7EFD13FF36" box="[605,696,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[189,1420,201,1903]" captionTargetId="figure-17@5.[167,1420,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama.A—Habitus, dorsal view; B—ditto, lateral view; C—ditto, pro-, meso- and metathorax, ventral view; D—Foreleg including procoxa [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961485" httpUri="https://zenodo.org/record/7961485/files/figure.png" pageId="14" pageNumber="555">Fig. 3D</figureCitation>
). Basal metatarsomere longer than combined length of remaining metatarsomeres. Tergite VII broadly rounded at posterior margin, with broad median longitudinal furrow (
<figureCitation id="134497A32536FFC0154EFF52FA3BFF52" box="[1334,1424,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="14" pageNumber="555">Fig. 4A</figureCitation>
). Morphological sternite VII broadly rounded at posterior margin (
<figureCitation id="134497A32536FFC013F3FF36FC4CFF7E" box="[907,999,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="14" pageNumber="555">Fig. 4B</figureCitation>
). Morphological sternite VIII broadly emarginate at posterior margin (
<figureCitation id="134497A32536FFC01273FEEAFDC3FE9A" box="[523,616,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="14" pageNumber="555">Fig. 4C</figureCitation>
). Morphological sternite IX as in
<figureCitation id="134497A32536FFC0138DFEEAFBFAFE9A" box="[1013,1105,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="14" pageNumber="555">Fig. 4D</figureCitation>
. Aedeagus elongate, tubular, apically rounded, slightly curved in lateral view (
<figureCitation id="134497A32536FFC012C6FECEFCB2FEC6" box="[702,793,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1731,1755]" captionTargetBox="[157,1430,187,1701]" captionTargetId="figure-17@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Tomosomus subtestaceus Motschulsky, 1855. A—Lectotype ♁, habitus, dorsal view; B—ditto, lateral view; C— Apex of elytra, tergite VII and aedeagus, dorsal view [not to scale]. Images courtesy Kirill V. Makarov (Moscow, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7961481" httpUri="https://zenodo.org/record/7961481/files/figure.png" pageId="14" pageNumber="555">Figs 2C</figureCitation>
,
<figureCitation id="134497A32536FFC0135DFECEFCC0FEC6" box="[805,875,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[156,1426,190,1898]" captionTargetId="figure-27@6.[151,1436,181,1903]" captionTargetPageId="6" captionText="FIGURE 4. Tomosomus subtestaceus Motschulsky, 1855, ♁ from San Lorenzo NP, Panama, terminalia and genitalia. A— Tergite VII, dorsal view; B—Morphological sternite VII, ventral view; C—Morphological sternite VIII, ventral view; D— Morphological sternite IX; E—Aedeagus, ventral view; F—ditto, lateral view; G—ditto, apex, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7961487" httpUri="https://zenodo.org/record/7961487/files/figure.png" pageId="14" pageNumber="555">4E–G</figureCitation>
).
</paragraph>
<paragraph id="8BC08B262536FFC010BFFEA3FB1DFE0E" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="555">
<emphasis id="B90B57342536FFC010BFFEA3FE13FEE3" bold="true" box="[199,440,330,356]" pageId="14" pageNumber="555">Sexual dimorphism.</emphasis>
Female protibia not or only slightly swollen at posterior margin. Morphological sternite VIII without apical emargination. Morphological sternite IX rod-like, without accessory arms.
</paragraph>
</subSubSection>
<subSubSection id="C365D8AD2536FFC010BFFE7AFD26FE2A" box="[199,653,403,429]" pageId="14" pageNumber="555" type="biology_ecology">
<paragraph id="8BC08B262536FFC010BFFE7AFD26FE2A" blockId="14.[151,1437,151,573]" box="[199,653,403,429]" pageId="14" pageNumber="555">
<emphasis id="B90B57342536FFC010BFFE7AFE87FE2A" bold="true" box="[199,300,403,429]" pageId="14" pageNumber="555">Ecology.</emphasis>
Species of lowland rainforests.
</paragraph>
</subSubSection>
<subSubSection id="C365D8AD2536FFC010BFFE5FFBFDFD9E" pageId="14" pageNumber="555" type="distribution">
<paragraph id="8BC08B262536FFC010BFFE5FFBFDFD9E" blockId="14.[151,1437,151,573]" pageId="14" pageNumber="555">
<emphasis id="B90B57342536FFC010BFFE5FFECAFE57" bold="true" box="[199,353,438,464]" pageId="14" pageNumber="555">Distribution.</emphasis>
<collectingCountry id="F368CBB62536FFC01112FE5EFE6DFE56" box="[362,454,439,465]" name="Panama" pageId="14" pageNumber="555">Panama</collectingCountry>
.
<typeStatus id="54C435842536FFC011ABFE5EFDA0FE56" box="[467,523,439,465]" pageId="14" pageNumber="555">Type</typeStatus>
locality ‘Obispo’, as of
<bibRefCitation id="EFEEF6D72536FFC01352FE5EFBF9FE56" author="Selander, R. B. &amp; Vaurie, P." box="[810,1106,439,465]" pageId="14" pageNumber="555" pagination="1 - 70" refId="ref6738" refString="Selander, R. B. &amp; Vaurie, P. (1962) A gazetteer to accompany the &quot; Insecta &quot; volumes of the &quot; Biologia Centrali-Americana &quot;. American Museum Novitates, 2099, 1 - 70." type="journal article" year="1962">Selander &amp; Vaurie (1962)</bibRefCitation>
and
<bibRefCitation id="EFEEF6D72536FFC014F6FE5EFAE7FE56" author="Chandler, D. S." box="[1166,1356,439,465]" pageId="14" pageNumber="555" pagination="321 - 327" refId="ref6336" refString="Chandler, D. S. (2013) Further notes on the New World Pselaphinae (Coleoptera: Staphylinidae) described by Victor Motschulsky. The Coleopterists Bulletin, 67 (3), 321 - 327. https: // doi. org / 10.1649 / 0010 - 065 X- 67.3.321" type="journal article" year="2013">Chandler (2013)</bibRefCitation>
, was a settlement ‘on the Río Obispo across the Chagres River from the site of Gamboa which is today along the transisthmus railroad’. Today the area is known as Bas Obispo (
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).
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<emphasis id="B90B57342536FFC010BFFDCAFE27FDBB" bold="true" box="[199,396,546,573]" pageId="14" pageNumber="555">Type depository.</emphasis>
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.
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