254 lines
31 KiB
XML
254 lines
31 KiB
XML
<document id="AC0FE6D0B7175CF7514E195B465348BC" ID-CLB-Dataset="27715" ID-DOI="10.11646/zootaxa.4541.1.1" ID-GBIF-Dataset="9c3fecf1-4e11-4c79-a908-e0a5abbaf3ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617460" ID-ZooBank="B11E734C-218B-418C-84E6-719AB3C58AFF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927824231" checkinUser="plazi" docAuthor="Arachchige, Gayashan M., Jayakody, Sevvandi, Mooi, Rich & Kroh, Andreas" docDate="2019" docId="038087B4FFB88960FF02FB1570089B0C" docLanguage="en" docName="zootaxa.4541.1.1.pdf" docOrigin="Zootaxa 4541 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Echinolampas alexandri de Loriol 1876" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="FFB9FFCCFFAA8975FF95FFB47314996C" masterDocTitle="Taxonomy and distribution of irregular echinoids (Echinoidea: Irregularia) from Sri Lanka" masterLastPageNumber="100" masterPageNumber="1" pageNumber="19" updateTime="1698715306146" updateUser="ExternalLinkService">
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<mods:titleInfo id="9FC23EC26982648C876F7E7A89B5F30D">
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<mods:title id="7E4D041969091DF2CB5A373851B75042">Taxonomy and distribution of irregular echinoids (Echinoidea: Irregularia) from Sri Lanka</mods:title>
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<mods:namePart id="51B7A461C36F69915081FE6271CB1415">Arachchige, Gayashan M.</mods:namePart>
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<mods:namePart id="E2E69A9B7EF786B54A25C790F9D27968">Jayakody, Sevvandi</mods:namePart>
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<mods:namePart id="639FFBCC4B5082906951A2C71AAF15CE">Mooi, Rich</mods:namePart>
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<mods:namePart id="6C9C18D557A1BA8C7214530776B66AFF">Kroh, Andreas</mods:namePart>
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<mods:title id="D0D2771B1CDC94D37A9DDA9B8DA35425">Zootaxa</mods:title>
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<mods:date id="ACAD5B3E1BE3B57BD75EAE0EE6DAE104">2019</mods:date>
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<mods:number id="B202F755E5C069F42D9A133F3CBDD0AE">2019-01-03</mods:number>
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<mods:number id="D115D78097A17091E1ED6715C9979159">4541</mods:number>
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<mods:identifier id="BE6FEDDB64876D454690D71CC8FE80DB" type="DOI">10.11646/zootaxa.4541.1.1</mods:identifier>
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<treatment id="038087B4FFB88960FF02FB1570089B0C" ID-DOI="http://doi.org/10.5281/zenodo.5935428" ID-GBIF-Taxon="154529635" ID-Zenodo-Dep="5935428" LSID="urn:lsid:plazi:treatment:038087B4FFB88960FF02FB1570089B0C" httpUri="http://treatment.plazi.org/id/038087B4FFB88960FF02FB1570089B0C" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19">
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<subSubSection id="C3336529FFB88967FF02FB1571979DD7" box="[151,643,1185,1211]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8B9636A2FFB88967FF02FB1571979DD7" blockId="18.[151,643,1185,1245]" box="[151,643,1185,1211]" pageId="18" pageNumber="19">
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<heading id="D0DE81CEFFB88967FF02FB1571979DD7" bold="true" box="[151,643,1185,1211]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<taxonomicName id="4C294D21FFB88967FF02FB1571979DD7" authority="de Loriol, 1876" authorityName="de Loriol" authorityYear="1876" box="[151,643,1185,1211]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="18" pageNumber="19" phylum="Echinodermata" rank="species" species="alexandri">
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<emphasis id="B95DEAB0FFB88967FF02FB1571979DD7" bold="true" box="[151,643,1185,1211]" pageId="18" pageNumber="19">
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<emphasis id="B95DEAB0FFB88967FF02FB1572D49DD7" bold="true" box="[151,448,1185,1211]" italics="true" pageId="18" pageNumber="19">Echinolampas alexandri</emphasis>
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de Loriol, 1876
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFB88967FF02FB7076789C2C" pageId="18" pageNumber="19" type="description">
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<paragraph id="8B9636A2FFB88967FF02FB7072C19DB1" blockId="18.[151,643,1185,1245]" box="[151,469,1220,1245]" pageId="18" pageNumber="19">
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<figureCitation id="13122A27FFB88967FF02FB7072049DB0" box="[151,272,1220,1245]" captionStart="FIGURE 13" captionStartId="19.[151,250,1420,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@19.[151,1436,193,1398]" captionTargetPageId="19" captionText="FIGURE 13. Echinolampas alexandri (WUSL/EI/1): A, aboral view; B, oral view; C, lateral view." httpUri="https://zenodo.org/record/2617484/files/figure.png" pageId="18" pageNumber="19">Figures 13</figureCitation>
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,
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<figureCitation id="13122A27FFB88967FE89FB70727B9DB0" box="[284,367,1220,1244]" captionStart="FIGURE 14" captionStartId="20.[151,250,1676,1698]" captionTargetBox="[154,1439,284,1569]" captionTargetId="figure@20.[154,1439,284,1569]" captionTargetPageId="20" captionText="FIGURE 14. Comparison of pore-pairs (A and B) and peristome (C and D) between Echinolampas alexandri (A and C, WUSL/EI/1) and Echinolampas ovata (B and D, WUSL/EI/3)." httpUri="https://zenodo.org/record/2617486/files/figure.png" pageId="18" pageNumber="19">14A, C</figureCitation>
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,
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<figureCitation id="13122A27FFB88967FEEEFB7072B99DB1" box="[379,429,1220,1245]" captionStart="FIGURE 15" captionStartId="21.[151,250,1582,1604]" captionTargetBox="[154,1439,763,1486]" captionTargetId="figure@21.[154,1439,763,1486]" captionTargetPageId="21" captionText="FIGURE 15. Comparison of interambulacrum 5 between Echinolampas alexandri (A, WUSL/EI/1) and Echinolampas ovata (B, WUSL/EI/3)." httpUri="https://zenodo.org/record/2617488/files/figure.png" pageId="18" pageNumber="19">15A</figureCitation>
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,
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<figureCitation id="13122A27FFB88967FE2CFB7072C19DB1" box="[441,469,1220,1245]" captionStart="FIGURE 16" captionStartId="22.[151,250,1782,1804]" captionTargetBox="[220,1391,215,1741]" captionTargetId="figure@22.[196,1401,196,1755]" captionTargetPageId="22" captionText="FIGURE 16. Distribution map of Echinolampas alexandri recorded in this study." httpUri="https://zenodo.org/record/2617490/files/figure.png" pageId="18" pageNumber="19">16</figureCitation>
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</paragraph>
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<paragraph id="8B9636A2FFB88967FF02FAB870129C4E" blockId="18.[151,1388,1292,1344]" box="[151,774,1292,1314]" pageId="18" pageNumber="19">
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1876
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<taxonomicName id="4C294D21FFB88967FF45FAB8713B9C4E" authority="de Loriol" authorityName="de Loriol" authorityYear="1876" box="[208,559,1292,1314]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="18" pageNumber="19" phylum="Echinodermata" rank="species" species="alexandri">
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<emphasis id="B95DEAB0FFB88967FF45FAB872DC9C4E" box="[208,456,1292,1314]" italics="true" pageId="18" pageNumber="19">Echinolampas alexandri</emphasis>
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de Loriol
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</taxonomicName>
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: p. 4; pl. 1: figs. 1–3.
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</paragraph>
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<paragraph id="8B9636A2FFB88967FF02FA9E76789C2C" blockId="18.[151,1388,1292,1344]" box="[151,1388,1322,1344]" pageId="18" pageNumber="19">
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1948a
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<taxonomicName id="4C294D21FFB88967FF4EFA9E71549C2C" authority="Alexandri de Loriol." authorityName="Alexandri de Loriol." box="[219,576,1322,1344]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="18" pageNumber="19" phylum="Echinodermata" rank="genus">
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<emphasis id="B95DEAB0FFB88967FF4EFA9E72C29C2C" box="[219,470,1322,1344]" italics="true" pageId="18" pageNumber="19">Echinolampas Alexandri</emphasis>
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de Loriol.
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</taxonomicName>
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—Mortensen: p. 282–286; pl. 2: figs. 20–24; pl. 3: figs. 1–3; pl. 14: figs. 1, 4, 7.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFB88967FF02FAD970F09CA2" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="8B9636A2FFB88967FF02FAD970F09CA2" blockId="18.[151,1436,1389,1990]" pageId="18" pageNumber="19">
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<emphasis id="B95DEAB0FFB88967FF02FAD972719CEA" bold="true" box="[151,357,1389,1414]" pageId="18" pageNumber="19">Material studied.</emphasis>
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Ten denuded tests: WUSL/EI/1, from Mount-Lavinia, WUSL/EI/2 and EI/110, from Negombo 1, WUSL/EI/107 and EI/108, from Baththalangunduwa 1, WUSL/EI/109, from Silavathurai, WUSL/EI/104, EI/ 105 and EI/106, from Kirinda, WUSL/EI/111 from Hambantota,
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<collectingCountry id="F33E7632FFB88967FCFAFA0170F59CA2" box="[879,993,1461,1486]" name="Sri Lanka" pageId="18" pageNumber="19">Sri Lanka</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3336529FFB88966FF52FA6D764F9E04" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
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<paragraph id="8B9636A2FFB88967FF52FA6D77F59F7A" blockId="18.[151,1436,1389,1990]" pageId="18" pageNumber="19">
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<emphasis id="B95DEAB0FFB88967FF52FA6D724D9C9E" bold="true" box="[199,345,1497,1522]" pageId="18" pageNumber="19">Description.</emphasis>
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<emphasis id="B95DEAB0FFB88967FECAFA6E71129C9E" box="[351,518,1497,1522]" italics="true" pageId="18" pageNumber="19">Size and shape</emphasis>
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—Test moderately large, 44.3–63.0 mm TL; broad, almost circular in outline, TW 90–99% TL; distinctly inflated, TH 58–66% TL; posterior margin rounded to indistinctly pointed.
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</paragraph>
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<paragraph id="8B9636A2FFB88967FF52F99573F79F31" blockId="18.[151,1436,1389,1990]" pageId="18" pageNumber="19">
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<emphasis id="B95DEAB0FFB88967FF52F99572779F55" box="[199,355,1569,1594]" italics="true" pageId="18" pageNumber="19">Apical system</emphasis>
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—Monobasal with four gonopores; situated anterior of centre 34–39% TL from anterior margin of test.
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</paragraph>
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<paragraph id="8B9636A2FFB88967FF52F9DD76499ECE" blockId="18.[151,1436,1389,1990]" pageId="18" pageNumber="19">
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<emphasis id="B95DEAB0FFB88967FF52F9DD72579FEE" box="[199,323,1641,1666]" italics="true" pageId="18" pageNumber="19">Ambulacra</emphasis>
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—Petals with unequal poriferous zones; poriferous zone IIIa of petal III, usually 2–5 pore pairs longer than IIIb; poriferous zones in anterior paired petals of very different length, posterior poriferous zones (IIa, IVb) in each petal 7 to 20 pore pairs longer; difference in posterior paired petals often not as pronounced, posterior poriferous zones (Ia, Vb) in each paired petal 4 to 9 pore pairs shorter than in anterior ones; poriferous zones parallel in anterior and posterior paired petals but converging distally in anterior paired petals; transversely elongated outer pores and circular inner pores in respiratory pore pairs; petal III shortest; posterior paired petals extending c.74%, anterior paired petals 73% and petal
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<date id="FF971062FFB88967FC93F8F670539E36" box="[774,839,1857,1882]" pageId="18" pageNumber="19" value="1960-03">III 60</date>
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% of corresponding test radius; at their widest point, petals bear 4 to 6 tubercles across interporiferous zone; phyllodes typically consisting of inner series with up to 3 pores per column, outer series with up to 9; phyllodes contain medial double row of up to 10 sphaeridial pits.
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</paragraph>
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<paragraph id="8B9636A2FFB88967FF52F81970839EAA" blockId="18.[151,1436,1389,1990]" box="[199,919,1965,1990]" pageId="18" pageNumber="19">
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<emphasis id="B95DEAB0FFB88967FF52F81972639EAA" box="[199,375,1965,1990]" italics="true" pageId="18" pageNumber="19">Interambulacra</emphasis>
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—Bourrelets indistinct; oral naked zones absent.
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</paragraph>
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<caption id="DF56662AFFB98966FF02FA3877979CCE" box="[151,1155,1420,1442]" httpUri="https://zenodo.org/record/2617484/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1420,1442]" targetBox="[151,1436,193,1398]" targetPageId="19">
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<paragraph id="8B9636A2FFB98966FF02FA3877979CCE" blockId="19.[151,1155,1420,1442]" box="[151,1155,1420,1442]" pageId="19" pageNumber="20">
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<emphasis id="B95DEAB0FFB98966FF02FA38720B9CCD" bold="true" box="[151,287,1420,1442]" pageId="19" pageNumber="20">FIGURE 13.</emphasis>
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<taxonomicName id="4C294D21FFB98966FEB0FA38710A9CCE" authorityName="de Loriol" authorityYear="1876" box="[293,542,1420,1442]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="alexandri">
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<emphasis id="B95DEAB0FFB98966FEB0FA38710A9CCE" box="[293,542,1420,1442]" italics="true" pageId="19" pageNumber="20">Echinolampas alexandri</emphasis>
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</taxonomicName>
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(WUSL/EI/1): A, aboral view; B, oral view; C, lateral view.
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</paragraph>
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</caption>
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<paragraph id="8B9636A2FFB98966FF52FA5D769F9F4F" blockId="19.[151,1436,1511,1896]" pageId="19" pageNumber="20">
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<emphasis id="B95DEAB0FFB98966FF52FA5D722F9F6C" box="[199,315,1513,1536]" italics="true" pageId="19" pageNumber="20">Peristome</emphasis>
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—Small, oval, transversely elongated; length 9–10% TL and width 12–17% TL; deeply sunken with steep-sided infundibulum; located anterior of centre, approximately 37% TL from anterior margin of test.
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</paragraph>
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<paragraph id="8B9636A2FFB98966FF52F98570879F07" blockId="19.[151,1436,1511,1896]" pageId="19" pageNumber="20">
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<emphasis id="B95DEAB0FFB98966FF52F98572209F24" box="[199,308,1585,1608]" italics="true" pageId="19" pageNumber="20">Periproct</emphasis>
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—Inframarginal, slightly larger than peristome, length 8–13% and width 13–19% TL; situated closer to posterior margin, 2–3% TL from posterior margin of test.
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</paragraph>
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<paragraph id="8B9636A2FFB98966FF52F9C3771C9FB4" blockId="19.[151,1436,1511,1896]" pageId="19" pageNumber="20">
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<emphasis id="B95DEAB0FFB98966FF52F9C372BF9FFC" bold="true" box="[199,427,1655,1680]" pageId="19" pageNumber="20">Geographic range.</emphasis>
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Indo-West Pacific, from Islands of West Indian Ocean (
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<bibRefCitation id="EFB84B53FFB98966FBA9F9C377A79FFC" author="Bell, F. J." box="[1084,1203,1655,1680]" pageId="19" pageNumber="20" pagination="12 - 22" refId="ref33960" refString="Bell, F. J. (1909) Report on the Echinoderma (other than holothurians) collected by Mr. Stanley Gardiner in the western parts of the Indian Ocean. Transactions of the Linnaean Society of London, Zoology, 13, 12 - 22. https: // doi. org / 10.1111 / j. 1096 - 3642.1909. tb 00406. x" type="journal article" year="1909">Bell 1909</bibRefCitation>
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;
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<bibRefCitation id="EFB84B53FFB98966FB55F9CC769B9FFC" author="Mortensen, T." box="[1216,1423,1655,1680]" pageId="19" pageNumber="20" pagination="67 - 72" refId="ref35463" refString="Mortensen, T. (1948 d) New Echinoidea (Cassiduloida, Clypeasteroida): Preliminary notice. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i KObenhavn, 111, 67 - 72." type="journal article" year="1948">Mortensen 1948d</bibRefCitation>
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),
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<collectingCountry id="F33E7632FFB98966FF02F92F72129FD8" box="[151,262,1691,1716]" name="Mauritius" pageId="19" pageNumber="20">Mauritius</collectingCountry>
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(
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<bibRefCitation id="EFB84B53FFB98966FE83F92F72BD9FD8" author="Clark, H. L." box="[278,425,1691,1716]" pageId="19" pageNumber="20" refId="ref34415" refString="Clark, H. L. (1925 a) A Catalogue of the Recent Sea-Urchins (Echinoidea) in the Collection of the British Museum (Natural History). Oxford University Press, London, 250 pp." type="book" year="1925">Clark 1925a</bibRefCitation>
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),
|
||
<collectingCountry id="F33E7632FFB98966FE2AF92F71069FD8" box="[447,530,1691,1716]" name="Sri Lanka" pageId="19" pageNumber="20">Ceylon</collectingCountry>
|
||
area (
|
||
<bibRefCitation id="EFB84B53FFB98966FDCFF92F71F59FD8" author="Clark, H. L." box="[602,737,1691,1716]" pageId="19" pageNumber="20" pagination="81 - 283" refId="ref34331" refString="Clark, H. L. (1917) Hawaiian and other Pacific Echini, Echinoneidae, Nucleolitidae, Urechinidae, Echinocorythidae, Calymnidae, Pourtalesiidae, Palaestomatidae, Aeropsidae, Palaeopneustidae, Hemiasteridae, and Spatangidae. Memoirs of the Museum of Comparative Zoology at Harvard College, 46, 81 - 283." type="journal article" year="1917">Clark 1917</bibRefCitation>
|
||
), Bay of Bengal (
|
||
<bibRefCitation id="EFB84B53FFB98966FC38F92F775B9FD8" author="Koehler, R." box="[941,1103,1691,1716]" pageId="19" pageNumber="20" refId="ref34949" refString="Koehler, R. (1922) Echinoderma of the Indian Museum, Part 9, An account of the Echinoidea. II. Clypeastrides et Cassidulides. Trustees of the Indian Museum, Calcutta, 161 pp." type="book" year="1922">Koehler 1922</bibRefCitation>
|
||
) and East Indies (
|
||
<bibRefCitation id="EFB84B53FFB98966FAB5F92873CA9FB4" author="Mortensen, T." pageId="19" pageNumber="20" refId="ref35354" refString="Mortensen, T. (1948 a) A Monograph of the Echinoidea IV. 1. Holectypoida, Cassiduloida. C. A. Reitzel, Copenhagen, 371 pp." type="book" year="1948">Mortensen 1948a</bibRefCitation>
|
||
) to Coral Sea (
|
||
<bibRefCitation id="EFB84B53FFB98966FE1FF90B71209FB4" author="Miskelly, A." box="[394,564,1727,1752]" pageId="19" pageNumber="20" refId="ref35198" refString="Miskelly, A. (2002) Sea urchins of Australia and the Indo-Pacific. Capricornica Publications, Lindfield, 179 pp." type="book" year="2002">Miskelly 2002</bibRefCitation>
|
||
) and South Pacific Islands (Clark 1952).
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFB98966FF52F957701D9F90" blockId="19.[151,1436,1511,1896]" box="[199,777,1763,1788]" pageId="19" pageNumber="20">
|
||
<emphasis id="B95DEAB0FFB98966FF52F95772B89F90" bold="true" box="[199,428,1763,1788]" pageId="19" pageNumber="20">Bathymetric range</emphasis>
|
||
.
|
||
<quantity id="4CD19B47FFB98966FE2EF95771349F97" box="[443,544,1763,1788]" metricMagnitude="2" metricUnit="m" metricValue="1.8649999999999998" metricValueMax="3.65" metricValueMin="0.08" pageId="19" pageNumber="20" unit="m" value="186.5" valueMax="365.0" valueMin="8.0">8–365 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFB84B53FFB98966FDBBF95071E89F90" author="Mortensen, T." box="[558,764,1764,1788]" pageId="19" pageNumber="20" refId="ref35354" refString="Mortensen, T. (1948 a) A Monograph of the Echinoidea IV. 1. Holectypoida, Cassiduloida. C. A. Reitzel, Copenhagen, 371 pp." type="book" year="1948">Mortensen 1948a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFB98966FF52F8B3764F9E04" blockId="19.[151,1436,1511,1896]" pageId="19" pageNumber="20">
|
||
<emphasis id="B95DEAB0FFB98966FF52F8B3716F9E4C" bold="true" box="[199,635,1799,1824]" pageId="19" pageNumber="20">
|
||
Observed occurrence in
|
||
<collectingCountry id="F33E7632FFB98966FE62F8B371629E4C" box="[503,630,1799,1824]" name="Sri Lanka" pageId="19" pageNumber="20">Sri Lanka</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
This species typically occurs in sandy areas between patch reefs on the western, north-western, and southern coasts of Sri Lanka, at a depth range of
|
||
<quantity id="4CD19B47FFB98966FBA8F89877809E2F" box="[1085,1172,1836,1860]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.8" metricValueMin="0.9" pageId="19" pageNumber="20" unit="m" value="18.5" valueMax="28.0" valueMin="9.0">9–28 m</quantity>
|
||
(
|
||
<figureCitation id="13122A27FFB98966FB31F89F77EE9E28" box="[1188,1274,1835,1860]" captionStart="FIGURE 16" captionStartId="22.[151,250,1782,1804]" captionTargetBox="[220,1391,215,1741]" captionTargetId="figure@22.[196,1401,196,1755]" captionTargetPageId="22" captionText="FIGURE 16. Distribution map of Echinolampas alexandri recorded in this study." httpUri="https://zenodo.org/record/2617490/files/figure.png" pageId="19" pageNumber="20">Fig. 16</figureCitation>
|
||
). This species was first reported from Sri Lanka by
|
||
<bibRefCitation id="EFB84B53FFB98966FDD4F8FB71C39E04" author="Clark, H. L." box="[577,727,1871,1896]" pageId="19" pageNumber="20" pagination="83 - 102" refId="ref34305" refString="Clark, H. L. (1915) The echinoderms of Ceylon other than holothurians. Spolia Zeylanica, 10, 83 - 102." type="journal article" year="1915">Clark (1915)</bibRefCitation>
|
||
, but has not been recorded again until the present study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF56662AFFBE8961FF02F93870009FAC" httpUri="https://zenodo.org/record/2617486/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1676,1698]" targetBox="[154,1439,284,1569]" targetPageId="20">
|
||
<paragraph id="8B9636A2FFBE8961FF02F93870009FAC" blockId="20.[151,1435,1676,1728]" pageId="20" pageNumber="21">
|
||
<emphasis id="B95DEAB0FFBE8961FF02F93872379FCD" bold="true" box="[151,291,1676,1698]" pageId="20" pageNumber="21">FIGURE 14.</emphasis>
|
||
Comparison of pore-pairs (A and B) and peristome (C and D) between
|
||
<taxonomicName id="4C294D21FFBE8961FBBEF93876339FCE" authorityName="de Loriol" authorityYear="1876" box="[1067,1319,1676,1698]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="alexandri">
|
||
<emphasis id="B95DEAB0FFBE8961FBBEF93876339FCE" box="[1067,1319,1676,1698]" italics="true" pageId="20" pageNumber="21">Echinolampas alexandri</emphasis>
|
||
</taxonomicName>
|
||
(A and C, WUSL/EI/1) and
|
||
<taxonomicName id="4C294D21FFBE8961FEDEF91E710E9FAC" baseAuthorityName="Leske" baseAuthorityYear="1778" box="[331,538,1706,1728]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="ovata">
|
||
<emphasis id="B95DEAB0FFBE8961FEDEF91E710E9FAC" box="[331,538,1706,1728]" italics="true" pageId="20" pageNumber="21">Echinolampas ovata</emphasis>
|
||
</taxonomicName>
|
||
(B and D, WUSL/EI/3).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3336529FFBF8960FF52FF2370089B0C" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="8B9636A2FFBF8960FF52FF23763E9B51" blockId="21.[151,1437,151,608]" pageId="21" pageNumber="22">
|
||
<emphasis id="B95DEAB0FFBF8960FF52FF23722999DC" bold="true" box="[199,317,151,176]" pageId="21" pageNumber="22">Remarks.</emphasis>
|
||
Based on general shape, two groups of
|
||
<taxonomicName id="4C294D21FFBF8960FC81FF2370A399DC" box="[788,951,151,176]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B95DEAB0FFBF8960FC81FF2370A399DC" box="[788,951,151,176]" italics="true" pageId="21" pageNumber="22">Echinolampas</emphasis>
|
||
</taxonomicName>
|
||
could be distinguished in
|
||
<collectingCountry id="F33E7632FFBF8960FB7BFF23767799DC" box="[1262,1379,151,176]" name="Sri Lanka" pageId="21" pageNumber="22">Sri Lanka</collectingCountry>
|
||
. The first group is characterized by a more circular outline, elongated outer pores in the petals, parallel poriferous zones in the posterior petals, and indistinct bourrelets (
|
||
<figureCitation id="13122A27FFBF8960FCB4FF6B706C9994" box="[801,888,223,248]" captionStart="FIGURE 14" captionStartId="20.[151,250,1676,1698]" captionTargetBox="[154,1439,284,1569]" captionTargetId="figure@20.[154,1439,284,1569]" captionTargetPageId="20" captionText="FIGURE 14. Comparison of pore-pairs (A and B) and peristome (C and D) between Echinolampas alexandri (A and C, WUSL/EI/1) and Echinolampas ovata (B and D, WUSL/EI/3)." httpUri="https://zenodo.org/record/2617486/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
|
||
, A and C). In the second group, the outline is more ovoid, the outer pores are circular, the posterior petal pore zones converge and the bourrelets are distinct (
|
||
<figureCitation id="13122A27FFBF8960FF0AFE9373E1982C" box="[159,245,295,320]" captionStart="FIGURE 14" captionStartId="20.[151,250,1676,1698]" captionTargetBox="[154,1439,284,1569]" captionTargetId="figure@20.[154,1439,284,1569]" captionTargetPageId="20" captionText="FIGURE 14. Comparison of pore-pairs (A and B) and peristome (C and D) between Echinolampas alexandri (A and C, WUSL/EI/1) and Echinolampas ovata (B and D, WUSL/EI/3)." httpUri="https://zenodo.org/record/2617486/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
|
||
, B and D). In addition, the first group does not have conspicuous oral naked zones (
|
||
<figureCitation id="13122A27FFBF8960FB5EFE937636982C" box="[1227,1314,295,320]" captionStart="FIGURE 15" captionStartId="21.[151,250,1582,1604]" captionTargetBox="[154,1439,763,1486]" captionTargetId="figure@21.[154,1439,763,1486]" captionTargetPageId="21" captionText="FIGURE 15. Comparison of interambulacrum 5 between Echinolampas alexandri (A, WUSL/EI/1) and Echinolampas ovata (B, WUSL/EI/3)." httpUri="https://zenodo.org/record/2617488/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
|
||
, A), while the second group does (
|
||
<figureCitation id="13122A27FFBF8960FE39FEF871179809" box="[428,515,332,357]" captionStart="FIGURE 15" captionStartId="21.[151,250,1582,1604]" captionTargetBox="[154,1439,763,1486]" captionTargetId="figure@21.[154,1439,763,1486]" captionTargetPageId="21" captionText="FIGURE 15. Comparison of interambulacrum 5 between Echinolampas alexandri (A, WUSL/EI/1) and Echinolampas ovata (B, WUSL/EI/3)." httpUri="https://zenodo.org/record/2617488/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
|
||
, B). The first group best matches descriptions of
|
||
<taxonomicName id="4C294D21FFBF8960FBD3FEF976589809" authority="alexandri" box="[1094,1356,332,357]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="subSpecies" species="alexandri" subSpecies="alexandri">
|
||
<emphasis id="B95DEAB0FFBF8960FBD3FEF976589809" box="[1094,1356,332,357]" italics="true" pageId="21" pageNumber="22">E. alexandri alexandri</emphasis>
|
||
</taxonomicName>
|
||
, being closer to the type material of de Loriol (1876) than to later material from the Siboga Expedition described by
|
||
<bibRefCitation id="EFB84B53FFBF8960FF02FE21726598C0" author="Mortensen, T." box="[151,369,404,428]" pageId="21" pageNumber="22" refId="ref35354" refString="Mortensen, T. (1948 a) A Monograph of the Echinoidea IV. 1. Holectypoida, Cassiduloida. C. A. Reitzel, Copenhagen, 371 pp." type="book" year="1948">Mortensen (1948a)</bibRefCitation>
|
||
under the name
|
||
<taxonomicName id="4C294D21FFBF8960FDA4FE21700B98C1" authorityName="Mortensen" authorityYear="1948" box="[561,799,404,429]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="subSpecies" species="alexandri" subSpecies="sibogae">
|
||
<emphasis id="B95DEAB0FFBF8960FDA4FE21700B98C1" box="[561,799,404,429]" italics="true" pageId="21" pageNumber="22">E. alexandri sibogae</emphasis>
|
||
</taxonomicName>
|
||
. In comparison to the
|
||
<typeStatus id="54928800FFBF8960FBB6FE21779398C1" box="[1059,1159,405,429]" pageId="21" pageNumber="22" type="syntype">syntypes</typeStatus>
|
||
of de Loriol (illustrated in
|
||
<bibRefCitation id="EFB84B53FFBF8960FF22FE0C729198BC" author="Mortensen, T." box="[183,389,440,464]" pageId="21" pageNumber="22" refId="ref35354" refString="Mortensen, T. (1948 a) A Monograph of the Echinoidea IV. 1. Holectypoida, Cassiduloida. C. A. Reitzel, Copenhagen, 371 pp." type="book" year="1948">Mortensen 1948a</bibRefCitation>
|
||
), the present material has a distinctly higher test, but this feature appears to be variable.
|
||
<bibRefCitation id="EFB84B53FFBF8960FF02FE6872959899" author="Mooi, R." box="[151,385,476,501]" pageId="21" pageNumber="22" pagination="63 - 85" refId="ref35221" refString="Mooi, R. (1990) Living cassiduloids (Echinodermata: Echinoidea): A key and annotated list. Proceedings of the Biological Society of Washington, 1990, 63 - 85." type="journal article" year="1990">Mooi (1990: fig. 6b)</bibRefCitation>
|
||
illustrated a prominent naked median zone in
|
||
<taxonomicName id="4C294D21FFBF8960FC00FE6977369899" authorityName="de Loriol" authorityYear="1876" box="[917,1058,476,501]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="species" species="alexandri">
|
||
<emphasis id="B95DEAB0FFBF8960FC00FE6977369899" box="[917,1058,476,501]" italics="true" pageId="21" pageNumber="22">E. alexandri</emphasis>
|
||
</taxonomicName>
|
||
, which is also true for one of the
|
||
<typeStatus id="54928800FFBF8960FF02FDB473EB9B74" box="[151,255,512,536]" pageId="21" pageNumber="22" type="syntype">syntypes</typeStatus>
|
||
. This feature, while showing little variation in the present material, seems to display intraspecific variation as well, and can thus only be used in conjunction with the additional characters listed above.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFBF8960FF52FDF370089B0C" blockId="21.[151,1437,151,608]" box="[199,796,583,608]" pageId="21" pageNumber="22">
|
||
The second group best matches
|
||
<taxonomicName id="4C294D21FFBF8960FDA0FDFD71819B0C" baseAuthorityName="Leske" baseAuthorityYear="1778" box="[565,661,585,608]" class="Echinoidea" family="Echinolampadidae" genus="Echinolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="21" pageNumber="22" phylum="Echinodermata" rank="species" species="ovata">
|
||
<emphasis id="B95DEAB0FFBF8960FDA0FDFD71819B0C" box="[565,661,585,608]" italics="true" pageId="21" pageNumber="22">E. ovata</emphasis>
|
||
</taxonomicName>
|
||
, see below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |