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<document id="ADF870AE6F0C19A1F07AA2809E1A027F" ID-DOI="10.5281/zenodo.12996779" ID-Zenodo-Dep="12996779" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="luisschmitz" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1721870935690" checkinUser="luisschmitz" docAuthor="Gerry Cassis &amp; Loren Vanags" docDate="2006" docId="0380878FFFE3FFC0FF5EFE36C153FB24" docLanguage="en" docName="Denisia.19.275-398.pdf.imf" docOrigin="Denisia 19" docTitle="Heissiphara Gerry Cassis &amp; Loren Vanags 2006, nov.gen." docType="treatment" docVersion="9" lastPageNumber="367" masterDocId="FFB9FFF7FFBFFF9DFFC4FF8CC239FFED" masterDocTitle="Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)" masterLastPageNumber="398" masterPageNumber="275" pageNumber="367" updateTime="1725646846613" updateUser="ExternalLinkService">
<mods:mods id="9ACE6EF0E48AD5A0DC229556597216F3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D18E28C4B159BC14D39A456CBB251160">
<mods:title id="2DB8440AF58CC42036D619D0F8B0AF0A">Jewel Bugs of Australia (Insecta, Heteroptera, Scutelleridae)</mods:title>
</mods:titleInfo>
<mods:name id="B1215E7FFE935651543863EC64578589" type="personal">
<mods:role id="2591FC63CE2316BDA2066C57ABF3DEC3">
<mods:roleTerm id="F68391C53F6312157FFF5D513D3126E7">Author</mods:roleTerm>
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<mods:namePart id="499737C889B56F812DB117E1F1A502E8">Gerry Cassis</mods:namePart>
</mods:name>
<mods:name id="EA69B08EB89B5C90D479E5D7415CAD47" type="personal">
<mods:role id="B4DD5589509DA381BE46481B9EF76638">
<mods:roleTerm id="C7176D61C1373FB5148AD61E4C649B68">Author</mods:roleTerm>
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<mods:namePart id="EAAB2FC074294F0CCCF34048EBC1EAF1">Loren Vanags</mods:namePart>
</mods:name>
<mods:typeOfResource id="52B76EA042AAF27FBBD50CB84C014C45">text</mods:typeOfResource>
<mods:relatedItem id="030375F0D96C0B441E5E92BD75647919" type="host">
<mods:titleInfo id="F24933B31C66452D51020ECBC5D42799">
<mods:title id="404723EEEBCA8A38D0B965E801B3C676">Denisia</mods:title>
</mods:titleInfo>
<mods:part id="A641CCFBC3B1CE6A17ED54C499B7328C">
<mods:date id="9F08D228904B1E68624653F60ADD4D38">2006</mods:date>
<mods:detail id="A1C2C3A6B8B5FE74E4AE70252988719A" type="volume">
<mods:number id="30F2DC3CD2AF72BF9130B2015D5E865B">19</mods:number>
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<mods:classification id="16282E234CAFCCC47875E16A658D0DDA">journal article</mods:classification>
<mods:identifier id="AF51DA62C16CC5A477EA559F13BD27D8" type="DOI">10.5281/zenodo.12996779</mods:identifier>
<mods:identifier id="58CA94FECFF32496032BF2954D556D64" type="Zenodo-Dep">12996779</mods:identifier>
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<treatment id="0380878FFFE3FFC0FF5EFE36C153FB24" LSID="urn:lsid:plazi:treatment:0380878FFFE3FFC0FF5EFE36C153FB24" httpUri="http://treatment.plazi.org/id/0380878FFFE3FFC0FF5EFE36C153FB24" lastPageId="93" lastPageNumber="367" pageId="92" pageNumber="367">
<subSubSection id="C3336512FFE3FFC1FF5EFE36C3BBFE1F" pageId="92" pageNumber="367" type="nomenclature">
<paragraph id="8B963699FFE3FFC1FF5EFE36C3BBFE1F" blockId="92.[154,588,110,1917]" pageId="92" pageNumber="367">
<heading id="D0DE81F5FFE3FFC1FF5EFE36C3B6FE3F" bold="true" box="[154,399,442,466]" fontSize="10" level="2" pageId="92" pageNumber="367" reason="6">
<taxonomicName id="4C294D1AFFE3FFC1FF5EFE36C322FE3F" authority="Gerry Cassis &amp; Loren Vanags, 2006" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[154,283,442,466]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95DEA8BFFE3FFC1FF5EFE36C322FE3F" box="[154,283,442,466]" italics="true" pageId="92" pageNumber="367">Heissiphara</emphasis>
</taxonomicName>
<emphasis id="B95DEA8BFFE3FFC1FEE0FE36C3B6FE3F" bold="true" box="[292,399,442,466]" pageId="92" pageNumber="367">
<taxonomicNameLabel id="A26E57F0FFE3FFC1FEE0FE36C3B6FE3F" box="[292,399,442,466]" pageId="92" pageNumber="367" rank="genus">nov.gen.</taxonomicNameLabel>
</emphasis>
</heading>
(
<figureCitation id="13122A1CFFE3FFC1FF5BFE56C33BFE1F" box="[159,258,474,498]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FEC9FE56C315FE1F" box="[269,300,474,498]" captionStart="Fig" captionStartId="73.[60,96,868,887]" captionTargetBox="[60,961,106,821]" captionTargetId="figure-588@73.[60,969,105,849]" captionTargetPageId="73" captionText="Fig. 27: Distribution of Scutellerinae species (a) Calliscyta stalii (b) Cantao parentum (c) Heissiphara minuta nov.sp." figureDoi="http://doi.org/10.5281/zenodo.13714033" httpUri="https://zenodo.org/record/13714033/files/figure.png" pageId="92" pageNumber="367">27</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FEF2FE56C36CFE1F" box="[310,341,474,498]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FE9BFE56C345FE1F" box="[351,380,474,498]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">38</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC1FF5EFD88C339FDD5" pageId="92" pageNumber="367" type="type_taxon">
<paragraph id="8B963699FFE3FFC1FF5EFD88C339FDD5" blockId="92.[154,588,110,1917]" pageId="92" pageNumber="367">
Type species:
<taxonomicName id="4C294D1AFFE3FFC1FED1FD88C38FFDF4" authority="Gerry Cassis &amp; Loren Vanags, 2006" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[277,438,516,537]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="species" species="minuta" status="sp. nov.">
<emphasis id="B95DEA8BFFE3FFC1FED1FD88C38FFDF4" box="[277,438,516,537]" italics="true" pageId="92" pageNumber="367">HeissipHara minuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57F0FFE3FFC1FE79FD88C3C4FDF4" box="[445,509,516,537]" pageId="92" pageNumber="367" rank="species">nov.sp.</taxonomicNameLabel>
, original designation
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC1FF04FDDEC3E1FB45" pageId="92" pageNumber="367" type="diagnosis">
<paragraph id="8B963699FFE3FFC1FF04FDDEC3E1FB45" blockId="92.[154,588,110,1917]" pageId="92" pageNumber="367">
Diagnosis:
<taxonomicName id="4C294D1AFFE3FFC1FEE8FDDEC3AEFD84" authority="Gerry Cassis &amp; Loren Vanags, 2006" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[300,407,594,617]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95DEA8BFFE3FFC1FEE8FDDEC3AEFD84" box="[300,407,594,617]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57F0FFE3FFC1FE5AFDDEC3CBFD84" box="[414,498,594,617]" pageId="92" pageNumber="367" rank="genus">nov.gen.</taxonomicNameLabel>
is recognised by the following combination of characters: small species (
<figureCitation id="13122A1CFFE3FFC1FEABFD1AC3FEFD40" box="[367,455,662,685]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FE17FD1AC3D6FD40" box="[467,495,662,685]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); dorsum strongly convex; dorsum heavily punctate with setigerous punctures (
<figureCitation id="13122A1CFFE3FFC1FE71FD55C02BFD1D" box="[437,530,729,752]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FDE7FD55C006FD1D" box="[547,575,729,752]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); head strongly declivent and rounded (
<figureCitation id="13122A1CFFE3FFC1FDE1FD77C2FBFCD9" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38b</figureCitation>
); jugae rounded (
<figureCitation id="13122A1CFFE3FFC1FE44FC91C3E5FCD9" box="[384,476,797,820]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38b</figureCitation>
); bucculae small, arcuate (
<figureCitation id="13122A1CFFE3FFC1FEF4FCB3C3BDFCBB" box="[304,388,831,854]" captionStart="Fig" captionStartId="82.[1293,1329,107,126]" captionTargetBox="[151,1277,106,1424]" captionTargetId="figure-280@82.[150,1278,105,1425]" captionTargetPageId="82" captionText="Fig. 31: Scanning electron micrographs of key characters of Choerocoris paganus (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) External efferent system of metathoracic glands, ventral view (e) male pygophore, ventral view (f) male pygophore, dorsal view. Abbreviations: C = clypeus; Ea = evaporative area; Pe = peritreme; Pe(k) = keel of anterior margin of proepisternum; P(r) = right paramere; Pr = proctiger. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714049" httpUri="https://zenodo.org/record/13714049/files/figure.png" pageId="92" pageNumber="367">Fig. 31c</figureCitation>
); anterior margin of scutellum with submedial subreniform polished calli (
<figureCitation id="13122A1CFFE3FFC1FECAFC0EC35BFC74" box="[270,354,898,921]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Fig. 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FEAAFC0EC3B3FC74" box="[366,394,898,921]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); lateral margins of pronotum subcarinate; external efferent system of metathoracic glands moderately developed, with raised obovate peritreme (
<figureCitation id="13122A1CFFE3FFC1FDE1FC64C2F8FBCC" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38c</figureCitation>
); evaporative areas extending to mesepimeron (
<figureCitation id="13122A1CFFE3FFC1FEF5FBA0C3B2FBAE" box="[305,395,1068,1091]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38c</figureCitation>
); spermathecal fecundation canal very short (
<figureCitation id="13122A1CFFE3FFC1FE6AFBC1C03AFB89" box="[430,515,1101,1124]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38d</figureCitation>
); reservoir oval, not heavily sclerotized; and, spermathecal pump large (
<figureCitation id="13122A1CFFE3FFC1FEB2FB1DC3F5FB45" box="[374,460,1169,1192]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC1FF04FB4EC108FE0A" pageId="92" pageNumber="367" type="description">
<paragraph id="8B963699FFE3FFC1FF04FB4EC626FE5B" blockId="92.[154,588,110,1917]" lastBlockId="92.[622,1057,110,1917]" pageId="92" pageNumber="367">
Description: Small species (
<figureCitation id="13122A1CFFE3FFC1FE2CFB4EC07EFB34" box="[488,583,1218,1241]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FF5EFB68C28CFB16" box="[154,181,1252,1275]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); ovoid; dorsum strongly convex (
<figureCitation id="13122A1CFFE3FFC1FDE7FB68C2F9FAF0" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FF0EFA8AC2DCFAF0" box="[202,229,1286,1309]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); body evenly punctate, with setigerous punctures (
<figureCitation id="13122A1CFFE3FFC1FEF0FAABC3B7FAD3" box="[308,398,1319,1342]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FE58FAABC381FAD3" box="[412,440,1319,1342]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
). Head: transverse (
<figureCitation id="13122A1CFFE3FFC1FF18FAC5C30CFA8D" box="[220,309,1353,1376]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Figs 38a</figureCitation>
), strongly rounded and declivent (
<figureCitation id="13122A1CFFE3FFC1FF2BFAE7C371FA6F" box="[239,328,1387,1410]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38b</figureCitation>
); lateral margins of jugae rounded (
<figureCitation id="13122A1CFFE3FFC1FEC4FA01C364FA49" box="[256,349,1421,1444]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38b</figureCitation>
). Antennae: AIII and AIV bicompressed. Pronotum: subtrapezoidal, strongly convex (
<figureCitation id="13122A1CFFE3FFC1FE56FA5DC3D2FA05" box="[402,491,1489,1512]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FE3DFA5DC02DFA05" box="[505,532,1489,1512]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); anterolateral margins elongate, rectilinear, subcarinate; posterolateral margins short, rounded; posterior margin truncate (
<figureCitation id="13122A1CFFE3FFC1FDE7F9BAC2F9F982" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FF09F9D4C2D0F982" box="[205,233,1624,1647]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
). Scutellum: broad, U-shaped, base of forewings and abdominal connexiva III-VII visible (
<figureCitation id="13122A1CFFE3FFC1FED1F917C356F95F" box="[277,367,1691,1714]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FEB9F917C3A1F95F" box="[381,408,1691,1714]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); anterior margin with submedial, subreniform, polished calli (
<figureCitation id="13122A1CFFE3FFC1FF64F953C2CFF91B" box="[160,246,1759,1782]" captionStart="Fig" captionStartId="69.[60,96,1742,1761]" captionTargetBox="[82,967,109,1707]" captionTargetId="figure-404@69.[78,967,105,1724]" captionTargetPageId="69" captionText="Fig. 24: Habitus of Australian Scutellerinae species (a) Calliscyta stalii, ♀ (b) Heissiphara minuta nov.sp., ♀ (c) Choerocoris lattini nov.sp., ♀ (d) Choerocoris paganus, ♂ (e) Choerocoris grossi nov.sp., ♂ (f) Choerocoris variegatus, ♂. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714025" httpUri="https://zenodo.org/record/13714025/files/figure.png" pageId="92" pageNumber="367">Figs 24b</figureCitation>
,
<figureCitation id="13122A1CFFE3FFC1FEC5F953C325F91B" box="[257,284,1759,1782]" captionStart="Fig" captionStartId="93.[60,96,1168,1187]" captionTargetBox="[68,964,111,1151]" captionTargetId="figure-431@93.[60,968,105,1152]" captionTargetPageId="93" captionText="Fig. 37: Habitus of holotype of Heissiphara minuta nov.sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.13714067" httpUri="https://zenodo.org/record/13714067/files/figure.png" pageId="92" pageNumber="367">37</figureCitation>
); tip of forewings not exposed; propleuron large, platelike, with anterior margins of proepisternum arcuate; external efferent system of metathoracic glands moderately developed (
<figureCitation id="13122A1CFFE3FFC1FE97F8EAC39FF890" box="[339,422,1894,1917]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38c</figureCitation>
), evaporative ar-eas extensive, rugose, extending beyond peritreme and on to mesepimeron (
<figureCitation id="13122A1CFFE3FFC1FC65FF03C1CAFF4B" box="[929,1011,143,166]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38c</figureCitation>
); ostiole small; peritreme raised, obovate (
<figureCitation id="13122A1CFFE3FFC1FC3EFF3DC0AFFF07" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38c</figureCitation>
). Abdominal Venter: SIII carinate along midline. Female Terminalia: paratergites IX small, subelliptoid; gonocoxae I large, subtriangular, posterior margin weakly excavate. Spermatheca: basal sclerites present; fecundation canal very short (
<figureCitation id="13122A1CFFE3FFC1FCA7FEF1C181FE79" box="[867,952,381,404]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38d</figureCitation>
); reservoir oval (
<figureCitation id="13122A1CFFE3FFC1FD62FE13C0C3FE5B" box="[678,762,415,438]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38d</figureCitation>
); prominent pump (
<figureCitation id="13122A1CFFE3FFC1FC7AFE13C62AFE5B" box="[958,1043,415,438]" captionStart="Fig" captionStartId="94.[1301,1337,107,126]" captionTargetBox="[152,1285,106,983]" captionTargetId="figure-429@94.[152,1285,105,984]" captionTargetPageId="94" captionText="Fig. 38: Scanning electron micrographs of key characters of Heissiphara minuta nov.sp. (a) Head, dorsal view (b) Head, lateral view (c) Head and thorax, ventral view (d) Spermatheca. Abbreviations: Df = dorsal flange; Ea = evaporative area; Pe = peritreme; Pf = proximal flange; P(r) = right paramere; S(b) = spermathecal bulb. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13714069" httpUri="https://zenodo.org/record/13714069/files/figure.png" pageId="92" pageNumber="367">Fig. 38d</figureCitation>
).
</paragraph>
<paragraph id="8B963699FFE3FFC1FD50FE5CC108FE0A" blockId="92.[622,1057,110,1917]" box="[660,817,464,487]" pageId="92" pageNumber="367">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC1FD50FD8DC0E9FD93" pageId="92" pageNumber="367" type="etymology">
<paragraph id="8B963699FFE3FFC1FD50FD8DC0E9FD93" blockId="92.[622,1057,110,1917]" pageId="92" pageNumber="367">Etymology: This genus is named after our colleague Dr. Ernst Heiss in recognition of his significant contribution to Heteropterology.</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC1FDABFD14C626FD3C" pageId="92" pageNumber="367" type="taxon_list">
<paragraph id="8B963699FFE3FFC1FDABFD14C626FD3C" blockId="92.[622,1057,110,1917]" pageId="92" pageNumber="367">
<emphasis id="B95DEA8BFFE3FFC1FDABFD14C12CFD42" bold="true" box="[623,789,664,687]" pageId="92" pageNumber="367">Included species</emphasis>
:
<taxonomicName id="4C294D1AFFE3FFC1FDABFD36C0E8FD3C" authority="Gerry Cassis &amp; Loren Vanags, 2006" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[623,721,698,721]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="species" species="minuta" status="nov.sp.">
<emphasis id="B95DEA8BFFE3FFC1FDABFD36C0E8FD3C" box="[623,721,698,721]" italics="true" pageId="92" pageNumber="367">H. minuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57F0FFE3FFC1FD1CFD36C124FD3C" box="[728,797,698,721]" pageId="92" pageNumber="367" rank="species">nov.sp.</taxonomicNameLabel>
Australia
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC1FD50FD76C1D2FCB8" pageId="92" pageNumber="367" type="distribution">
<paragraph id="8B963699FFE3FFC1FD50FD76C1D2FCB8" blockId="92.[622,1057,110,1917]" pageId="92" pageNumber="367">
Diversity and distribution:
<taxonomicName id="4C294D1AFFE3FFC1FC5BFD76C633FCFC" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[927,1034,762,785]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FC5BFD76C633FCFC" box="[927,1034,762,785]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
is a monotypic genus, known from a single locality in temperate Western Australia.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FFE3FFC0FD50FCE3C153FB24" lastPageId="93" lastPageNumber="368" pageId="92" pageNumber="367" type="discussion">
<paragraph id="8B963699FFE3FFC1FD50FCE3C618FB99" blockId="92.[622,1057,110,1917]" pageId="92" pageNumber="367">
Remarks:
<taxonomicName id="4C294D1AFFE3FFC1FCC5FCE3C155FC6B" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[769,876,879,902]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FCC5FCE3C155FC6B" box="[769,876,879,902]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
superficially resembles a number of Australian pentatomid species, with an enlarged scutellum, such as
<emphasis id="B95DEA8BFFE3FFC1FDAAFC59C0F2FC01" box="[622,715,981,1004]" italics="true" pageId="92" pageNumber="367">
<taxonomicName id="4C294D1AFFE3FFC1FDAAFC59C0FEFC01" authorityName="Distant" authorityYear="1911" box="[622,711,981,1004]" class="Insecta" family="Pentatomidae" genus="Kapunda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">Kapunda</taxonomicName>
.
</emphasis>
However,
<taxonomicName id="4C294D1AFFE3FFC1FCF2FC59C198FC01" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[822,929,981,1004]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FCF2FC59C198FC01" box="[822,929,981,1004]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C294D1AFFE3FFC1FD76FC7BC10BFBE3" authorityName="Leach" authorityYear="1815" box="[690,818,1015,1038]" class="Insecta" family="Scutelleridae" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="family">Scutelleridae</taxonomicName>
based on the following characters: greatly enlarged scutellum, spermatheca with basal sclerites, and the spermathecal reservoir without a sclerotized rod.
</paragraph>
<paragraph id="8B963699FFE3FFC1FD50FB02C19CF94F" blockId="92.[622,1057,110,1917]" pageId="92" pageNumber="367">
The position of
<taxonomicName id="4C294D1AFFE3FFC1FC84FB02C191FB48" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[832,936,1166,1189]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FC84FB02C191FB48" box="[832,936,1166,1189]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
within the scutellerid hierarchy is difficult to determine, as males are unknown. It is conclusively excluded from the
<taxonomicName id="4C294D1AFFE3FFC1FCDCFB78C1BAFAE6" box="[792,899,1268,1291]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="300" phylum="Arthropoda" rank="subFamily" subFamily="Elvisurinae">Elvisurinae</taxonomicName>
in that it does not possess keel-like thoracic sterna. It is unlikely to be a member of the
<taxonomicName id="4C294D1AFFE3FFC1FC4DFAB4C63EFAA2" box="[905,1031,1336,1359]" pageId="92" pageNumber="380" rank="subFamily" subFamily="Tectocorinae">Tectocorinae</taxonomicName>
as its female terminalia and spermatheca are consistent with those of the scutellerines.
<taxonomicName id="4C294D1AFFE3FFC1FDAAFA12C0EFFA58" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[622,726,1438,1461]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FDAAFA12C0EFFA58" box="[622,726,1438,1461]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
shares some similarites with
<taxonomicName id="4C294D1AFFE3FFC1FDAAFA4CC084FA3A" authorityName="Hirtula BERGROTH" authorityYear="1904" box="[622,701,1472,1495]" class="Insecta" family="Scutelleridae" genus="Morbora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FDAAFA4CC084FA3A" box="[622,701,1472,1495]" italics="true" pageId="92" pageNumber="367">Morbora</emphasis>
</taxonomicName>
, in that both the body has setigerous punctures and the metathoracic peritreme is obovate. However,
<taxonomicName id="4C294D1AFFE3FFC1FCFFF98FC19AF9F7" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[827,931,1539,1562]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FCFFF98FC19AF9F7" box="[827,931,1539,1562]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
also shares these characters with
<taxonomicName id="4C294D1AFFE3FFC1FC84F9A9C192F9D1" box="[832,939,1573,1596]" class="Insecta" family="Scutelleridae" genus="Choerocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FC84F9A9C192F9D1" box="[832,939,1573,1596]" italics="true" pageId="92" pageNumber="367">CHoerocoris</emphasis>
</taxonomicName>
, and on the basis of the similar spermathecae, the former genus is putatively assigned to the
<taxonomicName id="4C294D1AFFE3FFC1FC7FF9E5C0AAF94F" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="384" phylum="Arthropoda" rank="subFamily" subFamily="Scutellerinae">Scutelleri-nae</taxonomicName>
, despite its dull colouration.
</paragraph>
<paragraph id="8B963699FFE3FFC0FD50F930C153FB24" blockId="92.[622,1057,110,1917]" lastBlockId="93.[532,966,1168,1917]" lastPageId="93" lastPageNumber="368" pageId="92" pageNumber="367">
<taxonomicName id="4C294D1AFFE3FFC1FD50F930C0C6F93E" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[660,767,1724,1747]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FD50F930C0C6F93E" box="[660,767,1724,1747]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C294D1AFFE3FFC1FC54F930C1C4F93E" box="[912,1021,1724,1747]" class="Insecta" family="Scutelleridae" genus="Choerocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FC54F930C1C4F93E" box="[912,1021,1724,1747]" italics="true" pageId="92" pageNumber="367">CHoerocoris</emphasis>
</taxonomicName>
in the shape of the head, which is more round-ed and the lateral jugal margins are not explanate. In addition,
<taxonomicName id="4C294D1AFFE3FFC1FC86F8AEC194F8D4" authorityName="Gerry Cassis &amp; Loren Vanags" authorityYear="2006" box="[834,941,1826,1849]" class="Insecta" family="Scutelleridae" genus="Heissiphara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEA8BFFE3FFC1FC86F8AEC194F8D4" box="[834,941,1826,1849]" italics="true" pageId="92" pageNumber="367">HeissipHara</emphasis>
</taxonomicName>
has a more densely punctate body, and is significantly smaller, the anterior margin of the scutellum has submedial calli, and the spermathecal reservoir is not heavily sclerotised.
</paragraph>
</subSubSection>
</treatment>
</document>
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