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<document ID-DOI="http://doi.org/10.5281/zenodo.6518559" ID-Zenodo-Dep="6518559" approvalRequired="103" approvalRequired_for_document="1" approvalRequired_for_illustrations="7" approvalRequired_for_textStreams="95" checkinTime="1651683212973" checkinUser="carolina" docAuthor="Jameson, Tom J. M., Streicher, Jeffrey W., Manuelli, Luigi, Head, Jason J. &amp; Smith, Eric N." docDate="2022" docId="03F287F8FFAEFFADC856C37BF805BC1E" docLanguage="en" docName="HerpetolMonogr.36.1.1-48.pdf" docOrigin="Herpetological Monographs 36 (1)" docSource="http://dx.doi.org/10.1655/0733-1347-36.1.1" docTitle="Craugastor Cope 1862" docType="key" docVersion="4" lastPageId="40" lastPageNumber="40" masterDocId="FFCBFF80FF88FF85CB1DC67AFB11B944" masterDocTitle="Miniaturization in Direct-Developing Frogs from Mexico with the Description of Six New Species" masterLastPageNumber="48" masterPageNumber="1" pageId="38" pageNumber="38" updateTime="1651685064813" updateUser="carolina">
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<mods:title>Miniaturization in Direct-Developing Frogs from Mexico with the Description of Six New Species</mods:title>
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<mods:namePart>Jameson, Tom J. M.</mods:namePart>
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<mods:namePart>Streicher, Jeffrey W.</mods:namePart>
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<mods:namePart>Manuelli, Luigi</mods:namePart>
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<mods:namePart>Head, Jason J.</mods:namePart>
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<mods:namePart>Smith, Eric N.</mods:namePart>
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<mods:title>Herpetological Monographs</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-04-04</mods:number>
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<paragraph blockId="38.[843,1466,1280,1304]" box="[843,1466,1280,1304]" pageId="38" pageNumber="38">
DICHOTOMOUS KEY TO THE
<emphasis box="[1139,1387,1281,1304]" italics="true" pageId="38" pageNumber="38">
<taxonomicName authorityName="Cope" authorityYear="1862" box="[1139,1269,1281,1304]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="38" pageNumber="38" phylum="Chordata" rank="genus">CRAUGASTOR</taxonomicName>
MEXICANUS
</emphasis>
SERIES
</paragraph>
</subSubSection>
<subSubSection lastPageId="40" lastPageNumber="40" pageId="38" pageNumber="38" type="key">
<key lastPageId="40" lastPageNumber="40" pageId="38" pageNumber="38">
<keyStep pageId="38" pageNumber="38">
<paragraph blockId="38.[820,1490,1326,2001]" pageId="38" pageNumber="38">
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1a. Large or medium-sized adults (females&gt;
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SVL, males&gt;
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SVL); axillary gland present or absent; supratympanic fold usually developed (except for moderately to poorly developed in
<taxonomicName baseAuthorityName="Taylor" baseAuthorityYear="1942" box="[1273,1415,1415,1438]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="38" pageNumber="38" phylum="Chordata" rank="species" species="montanus">
<emphasis box="[1273,1415,1415,1438]" italics="true" pageId="38" pageNumber="38">C. montanus</emphasis>
</taxonomicName>
); both vomerine odontophores and posterolateral projection of frontoparietal present (except for some
<taxonomicName baseAuthorityName="Taylor" baseAuthorityYear="1942" box="[1344,1490,1474,1498]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="38" pageNumber="38" phylum="Chordata" rank="species" species="montanus">
<emphasis box="[1344,1490,1474,1498]" italics="true" pageId="38" pageNumber="38">C. montanus</emphasis>
</taxonomicName>
with no odontophores [but with frontoparietal projection], and some
<taxonomicName baseAuthorityName="Brocchi" baseAuthorityYear="1877" box="[1075,1226,1533,1557]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="38" pageNumber="38" phylum="Chordata" rank="species" species="mexicanus">
<emphasis box="[1075,1226,1533,1557]" italics="true" pageId="38" pageNumber="38">C. mexicanus</emphasis>
</taxonomicName>
with no frontoparietal projection [but with vomerine odontophores]) ........... 2
</keyLead>
</paragraph>
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1b. Small sized adults (females &lt;
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SVL, males &lt;
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SVL); axillary gland always absent; supratympanic fold absent or poorly developed; with no vomerine odontophores (except in
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" box="[1161,1351,1681,1705]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="38" pageNumber="38" phylum="Chordata" rank="species" species="candelariensis">
<emphasis box="[1161,1351,1681,1705]" italics="true" pageId="38" pageNumber="38">C. candelariensis</emphasis>
</taxonomicName>
) ................ 5
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</paragraph>
</keyStep>
<paragraph blockId="38.[820,1490,1326,2001]" pageId="38" pageNumber="38">
<keyLead pageId="38" pageNumber="38">
<keyStep pageId="38" pageNumber="38">
2a. Heelnotreachingnostrilwhen legadpressedtobody; two or three metacarpal tubercles (including thenar; outer smallest, middle largest); outer/inner metatarsaltubercle ratio 0.400.55; testis usually unpigmented (sometimes lightly pigmented); ovaries unpigmented; fromthe highlands of Guerrero and central Oaxaca (
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elevation) .............
<taxonomicName authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1900" box="[1105,1490,1888,1912]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="38" pageNumber="38" phylum="Chordata" rank="species" species="omiltemanus">
<emphasis box="[1105,1372,1888,1912]" italics="true" pageId="38" pageNumber="38">Craugastor omiltemanus</emphasis>
(Günther)
</taxonomicName>
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</paragraph>
<paragraph blockId="38.[820,1490,1326,2001]" lastBlockId="39.[142,813,188,2001]" lastPageId="39" lastPageNumber="39" pageId="38" pageNumber="38">
<keyLead lastPageId="39" lastPageNumber="39" pageId="38" pageNumber="38">
2b. Heel reaching tip of snout or beyond when leg adpressed to body; always three metacarpal tubercles (including thenar); outer/inner metatarsaltubercle
<keyStep pageId="39" pageNumber="39">ratio variable (0.360.75); testis strongly pigmented; ovaries always pigmented (strongly or lightly) ............ 3</keyStep>
</keyLead>
</paragraph>
<subSection pageId="38" pageNumber="38" type="multiple">
<paragraph blockId="38.[27,876,2060,2097]" pageId="38" pageNumber="38">
Downloaded From: https://bioone.org/journals/Herpetological-Monographs on
<date box="[670,778,2061,2078]" pageId="38" pageNumber="38" value="2022-05-04">04 May 2022</date>
Terms of Use: https://bioone.org/terms-of-use Access provided by American Museum of Natural History
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<keyStep pageId="39" pageNumber="39">
<paragraph blockId="39.[142,813,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
3a. Medium-sized adult females (&lt;
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SVL); crus 53 59% SVL; with or without vomerine odontophores; supratympanic fold moderately to poorly developed; labium always barred; middle metacarpal tubercle largest and outer very small; outer tarsal heel tubercles
<emphasis box="[203,223,393,419]" italics="true" pageId="39" pageNumber="39">Ĺ</emphasis>
2 (close to heel); ovaries pigmented; highlands east of the Isthmus of Tehuantepec (from the Sierra Madre del Sur of eastern Chiapas and western Guatemala) ...................................
<taxonomicName authority="(Taylor)" baseAuthorityName="Taylor" baseAuthorityYear="1942" box="[474,812,481,505]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="montanus">
<emphasis box="[474,715,482,505]" italics="true" pageId="39" pageNumber="39">Craugastor montanus</emphasis>
(Taylor)
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</paragraph>
<paragraph blockId="39.[142,813,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
3b. Large-sized adult females (&gt;
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SVL); crus 53 73% SVL; always with vomerine odontophores; supratympanic fold well-developed; labium variable in pattern; middle metacarpal tubercle about same size as thenar, outer smaller; outer tarsal heel tubercles 0 5; ovaries pigmented or not; highlands west of the Isthmus of Tehuantepec (from Oaxaca and Guerrero to Puebla and Hidalgo, including the central highlands of Veracruz) ......................................................... 4
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<paragraph blockId="39.[142,813,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
4a. Crus&gt;62% SVL (6273%); heel always beyond tip of snout when leg adpressed to body; outer/inner metatarsal tubercle ratio (0.36[0]0.58); ovaries unpigment- ed or only slightly pigmented; from the Sierra Madre del Sur of Guerrero west of Chilpancingo (
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elevation) ....................
<taxonomicName authority="(Taylor)" baseAuthorityName="Taylor" baseAuthorityYear="1941" box="[501,812,921,945]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="saltator">
<emphasis box="[501,716,921,945]" italics="true" pageId="39" pageNumber="39">Craugastor saltator</emphasis>
(Taylor)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="39.[142,813,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
4b. Crus &lt;69% SVL (5369%); heel at snout tip or sometimes beyond when leg adpressed to body; outer/inner metatarsaltubercle ratio (0.500.75); ovaries pigmented or not; from Guerrero east of Chilpancingo to Oaxaca and to the north in Puebla, Hidalgo and the adjacent Veracruz highlands (
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.2575" metricValueMax="3.0149999999999997" metricValueMin="1.5" pageId="39" pageNumber="39" unit="m" value="2257.5" valueMax="3015.0" valueMin="1500.0">1500 3015 m</quantity>
elevation) ......
<taxonomicName authority="(Brocchi)" baseAuthorityName="Brocchi" baseAuthorityYear="1877" box="[452,812,1127,1151]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="mexicanus">
<emphasis box="[452,699,1127,1150]" italics="true" pageId="39" pageNumber="39">Craugastormexicanus</emphasis>
(Brocchi)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="39" pageNumber="39">
<paragraph blockId="39.[142,813,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
5a. Vomerine odontophores present; outer two fingers and toes lanceolate to mucronate; body flank unicolor pale; no raised eyelid tubercles; gular region not pigmented or only lightly and uniformly peppered; face never barred; postrictal tubercles 2; metacarpal tubercles about equal in size; outer tarsal ridge with 38 tiny and pointed tubercles on slightly raised fringe; posterolateral projection of frontoparietal absent; from the Sierra Madre del Sur of Oaxaca, mountains above Puerto Escondido and Puerto Angel (
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elevation)...............
<taxonomicName authority="Jameson et al." authorityName="Jameson" box="[356,812,1449,1473]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="candelariensis">
<emphasis box="[356,645,1449,1473]" italics="true" pageId="39" pageNumber="39">Craugastor candelariensis</emphasis>
Jameson et al.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="39.[142,813,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
5b. Vomerine odontophores absent; fingers and toes usually round or at most, slightly lanceolate; body flank darker anteriorly (toward axilla, except uniformly pale in some
<taxonomicName box="[323,450,1567,1591]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="cueyatl">
<emphasis box="[323,450,1567,1591]" italics="true" pageId="39" pageNumber="39">C. cueyatl</emphasis>
</taxonomicName>
); usually with raised eyelid tubercles; gular region blotched or heavily peppered (rarely lightly peppered in
<taxonomicName baseAuthorityName="Taylor" baseAuthorityYear="1936" box="[512,658,1626,1649]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis box="[512,658,1626,1649]" italics="true" pageId="39" pageNumber="39">C. pygmaeus</emphasis>
</taxonomicName>
); face barred or not; postrictal tubercles 1 or 2; outer metacarpal tubercle usually longer and/or wider than inner (except about equal in size in
<taxonomicName box="[469,585,1713,1737]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="cueyatl">
<emphasis box="[469,585,1713,1737]" italics="true" pageId="39" pageNumber="39">C. cueyatl</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Taylor" baseAuthorityYear="1936" box="[649,797,1714,1737]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis box="[649,797,1714,1737]" italics="true" pageId="39" pageNumber="39">C. pygmaeus</emphasis>
</taxonomicName>
); outer tarsal ridge with no raised fringe and variable number of tubercles (07), flat and round (sometimes slightly pointed); posterolateral projection of frontoparietal present or absent ................................................. 6
</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="39.[142,813,188,2001]" lastBlockId="39.[844,1514,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
<keyStep pageId="39" pageNumber="39">6a. Face barred, one or two prominent bars below eye; no distinctive canthal stripe; one distinctive dark and rectangular blotch under lower jaw and at level of canthus (prominent in ventral view); gular region with pale spotting; finger tips not expanded; outer tarsal</keyStep>
</keyLead>
<keyLead pageId="39" pageNumber="39">
<keyStep pageId="39" pageNumber="39">
ridge with 04 extremely small, flat and round tubercles; posterolateral projection of frontoparietal present; from the Sierra Madre del Sur of Oaxaca in the mountains above Puerto Escondido (
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elevation) ............
<taxonomicName authority="Jameson et al." authorityName="Jameson" box="[1099,1514,306,330]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="polaclavus">
<emphasis box="[1099,1347,306,330]" italics="true" pageId="39" pageNumber="39">Craugastor polaclavus</emphasis>
Jameson et al.
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph blockId="39.[844,1514,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
<keyStep pageId="39" pageNumber="39">6b. Face barred or not, bars below eye variable; canthal stripe present or absent; no distinctive rectangular blotches under lower jaw and at level of canthus (not prominent in ventral view); gular region variable but, usually spotted or peppered; finger tips expanded or not; outer tarsal ridge with 07 extremely small, flat, and round to slightly pointed tubercles; posterolateral projection of frontoparietal present or absent; ........... 7</keyStep>
</keyLead>
</paragraph>
<keyStep pageId="39" pageNumber="39">
<paragraph blockId="39.[844,1514,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
7a. Finger and toe tips round and conspicuously expanded; face barred, never with a canthal stripe; postrictal tubercles 1 (sometimes look like two fused); dorsal pattern two-toned or striped, and with a pale middorsal stripe; metacarpal tubercles about equal in size; outer tarsal ridge smooth and lacking tubercles; posterolateral projection of frontoparietal absent; from the lowlands of southern Mexico, from Puebla and Veracruz on the Gulf versant and southern Guerrero on the Pacific versant to Escuintla, Chiapas and possibly adjacent Guatemala (
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elevation)................................
<taxonomicName authority="(Taylor)" baseAuthorityName="Taylor" baseAuthorityYear="1936" box="[1174,1513,892,916]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis box="[1174,1417,893,916]" italics="true" pageId="39" pageNumber="39">Craugastor pygmaeus</emphasis>
(Taylor)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="39.[844,1514,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">
7b. Finger tips round or lanceolate, not expanded or only slightly; toe tips slightly expanded to expanded, sometimes slightly lanceolated; face barred or with a pale supralabial stripe, canthal stripe present or absent (some
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" box="[987,1128,1039,1063]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="bitonium">
<emphasis box="[987,1128,1039,1063]" italics="true" pageId="39" pageNumber="39">C. bitonium</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Taylor" baseAuthorityYear="1936" box="[1199,1381,1039,1063]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="hobartsmithi">
<emphasis box="[1199,1381,1039,1063]" italics="true" pageId="39" pageNumber="39">C. hobartsmithi</emphasis>
</taxonomicName>
); postrictal tubercles 1 or 2; dorsal pattern variable; outer tarsal ridge smooth or tuberculated (
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" box="[1257,1427,1097,1121]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="portilloensis">
<emphasis box="[1257,1427,1097,1121]" italics="true" pageId="39" pageNumber="39">C. portilloensis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="rubinus">
<emphasis italics="true" pageId="39" pageNumber="39">C. rubinus</emphasis>
</taxonomicName>
) or always tuberculated; posterolateral projection of frontoparietal present or absent....................... 8
</keyLead>
</paragraph>
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<keyStep pageId="39" pageNumber="39">
<paragraph blockId="39.[844,1514,188,2001]" pageId="39" pageNumber="39">
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8a. Always with a canthal stripe and softly barred lips; dorsum uniformly dark; postrictal tubercles always 1, on each side; body flank uniformly pale or slightly darker anteriorly; gular region with evenly distributed fine melanocytes (evenly peppered); metacarpal tubercles about equal in size; finger tips round and not expanded; outer tarsal ridge with 37 rounded tubercles; posterolateral projection of frontoparietal present; from the slopes facing the Balsas River depression in the states of Mexico and Morelos (
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elevation) .................
<taxonomicName authority="Jameson et al." authorityName="Jameson" box="[1137,1514,1479,1503]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="39" pageNumber="39" phylum="Chordata" rank="species" species="cueyatl">
<emphasis box="[1137,1347,1479,1503]" italics="true" pageId="39" pageNumber="39">Craugastor cueyatl</emphasis>
Jameson et al.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="39.[844,1514,188,2001]" pageId="39" pageNumber="39">
<keyLead pageId="39" pageNumber="39">8b. Canthal stripe present or absent, and with soft or prominent bars on lips or with a supralabial pale stripe; dorsum uniformly pale, two-toned, or striped, never uniformly colored and darker than flanks; postrictal tubercles 1 or 2; body flank always darker anteriorly; gular region variable but usually melanocyte distribution forming light spotting (particularly under lower jaws and mid-gular area); outer metacarpal tubercle slightly larger than inner; finger tips not or just barely expanded, round or slightly lanceolated; outer tarsal ridge with 07 tubercles; posterolateral projection of frontoparietal present or absent ................................... 9</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="39.[844,1514,188,2001]" lastBlockId="40.[118,788,188,1370]" lastPageId="40" lastPageNumber="40" pageId="39" pageNumber="39">
<keyLead lastPageId="40" lastPageNumber="40" pageId="39" pageNumber="39">
<keyStep lastPageId="40" lastPageNumber="40" pageId="39" pageNumber="39">
9a. Canthal stripe present or absent, and with bars on lips or a supralabial pale stripe; dorsum usually two-toned or striped; gular region with small melanocyte distribution forming a few small light spots under lower jaws and mid-gular area; finger tips slightly expanded, round or slightly lanceolated (
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" box="[440,572,188,212]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="bitonium">
<emphasis box="[440,572,188,212]" italics="true" pageId="40" pageNumber="40">C. bitonium</emphasis>
</taxonomicName>
); outer tarsal ridge with 16 tubercles, round to slightly pointed; posterolateral projection of frontoparietal absent ................. 10
</keyStep>
</keyLead>
</paragraph>
<subSection pageId="39" pageNumber="39" type="multiple">
<paragraph blockId="39.[27,876,2060,2097]" pageId="39" pageNumber="39">
Downloaded From: https://bioone.org/journals/Herpetological-Monographs on
<date box="[670,778,2061,2078]" pageId="39" pageNumber="39" value="2022-05-04">04 May 2022</date>
Terms of Use: https://bioone.org/terms-of-use Access provided by American Museum of Natural History
</paragraph>
</subSection>
<paragraph blockId="40.[118,788,188,1370]" pageId="40" pageNumber="40">
<keyLead pageId="40" pageNumber="40">
<keyStep pageId="40" pageNumber="40">
9b. Canthal stripe always present, and with bar(s) on lips; dorsum uniformly pale (as compared with flanks,
<specimenCount pageId="40" pageNumber="40" type="generic">one specimen</specimenCount>
of
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" box="[327,447,337,361]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="rubinus">
<emphasis box="[327,447,337,361]" italics="true" pageId="40" pageNumber="40">C. rubinus</emphasis>
</taxonomicName>
with traces of ill-defined and light blotching); gular region with melanocyte distribution forming prominent light spots under lower jaws and mid-gular area; finger tips usually not expanded (somewhat expanded in
<specimenCount box="[455,611,455,479]" pageId="40" pageNumber="40" type="generic">one specimen</specimenCount>
of
<taxonomicName authorityName="Jameson &amp; Streicher &amp; Manuelli &amp; Head &amp; Smith" authorityYear="2022" box="[652,772,455,479]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="rubinus">
<emphasis box="[652,772,455,479]" italics="true" pageId="40" pageNumber="40">C. rubinus</emphasis>
</taxonomicName>
), round, never lanceolated; outer tarsal ridge with 04 tubercles, extremely small, flat, and round; posterolateral projection of frontoparietal present.................... 11
</keyStep>
</keyLead>
</paragraph>
<keyStep pageId="40" pageNumber="40">
<paragraph blockId="40.[118,788,188,1370]" pageId="40" pageNumber="40">
<keyLead pageId="40" pageNumber="40">
10a. Postrictal tubercles 1, on each side; finger and toe tips slightly lanceolated; finger lengths III&gt; IV
<emphasis box="[680,700,602,627]" italics="true" pageId="40" pageNumber="40">±</emphasis>
II&gt; I; toe lengths always IV&gt; III&gt; V&gt; II&gt; I; outer/inner metatarsal tubercle ratio 0.400.75; not less than four outer tarsal tubercles, 46; from the Sierra Madre del Sur de Guerrero, south of Yextla (
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elevation).......................
<taxonomicName authority="Jameson et al." authorityName="Jameson" box="[388,788,752,776]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="bitonium">
<emphasis box="[388,621,752,776]" italics="true" pageId="40" pageNumber="40">Craugastor bitonium</emphasis>
Jameson et al.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="40.[118,788,188,1370]" pageId="40" pageNumber="40">
<keyLead pageId="40" pageNumber="40">
10b. Postrictal tubercles 12, on each side; finger and toe tips rounded, not lanceolated; finger lengths always III&gt; IV&gt; II&gt; I; toelengths IV&gt; III
<emphasis box="[508,528,840,865]" italics="true" pageId="40" pageNumber="40">±</emphasis>
V&gt; II&gt; I; outer/inner metatarsal tubercle ratio 0.620.67; outer tarsal tubercles 16; from near Uruapan, Michoacán (
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elevation) ...................
<taxonomicName authority="(Taylor)" baseAuthorityName="Taylor" baseAuthorityYear="1936" box="[421,787,930,954]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="hobartsmithi">
<emphasis box="[421,692,930,954]" italics="true" pageId="40" pageNumber="40">Craugastor hobartsmithi</emphasis>
(Taylor)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="40" pageNumber="40">
<paragraph blockId="40.[118,788,188,1370]" pageId="40" pageNumber="40">
<keyLead pageId="40" pageNumber="40">
11a. Eyelid raised tubercles present or absent; postrictal tubercles 12, on each side; toe tips round and slightly expanded; finger lengths III&gt; IV&gt; II&gt; I; toe lengths IV&gt; III
<emphasis box="[281,301,1048,1073]" italics="true" pageId="40" pageNumber="40">±</emphasis>
V&gt; II&gt; I; outer/inner metatarsaltubercle ratio 0.671.25; from the Sierra Madre del Sur of Oaxaca at Portillo del Rayo (
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elevation)..................
<taxonomicName authority="Jameson et al." authorityName="Jameson" box="[354,788,1138,1162]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="portilloensis">
<emphasis box="[354,620,1138,1162]" italics="true" pageId="40" pageNumber="40">Craugastor portilloensis</emphasis>
Jameson et al.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="40.[118,788,188,1370]" pageId="40" pageNumber="40">
<keyLead pageId="40" pageNumber="40">
11b. Eyelid raised tubercles present; postrictal tubercles 1, on each side; toe tips slightly lanceolated and barely expanded; finger lengths III&gt; IV
<emphasis box="[558,578,1226,1251]" italics="true" pageId="40" pageNumber="40">¼</emphasis>
II&gt; I; toe lengths always IV&gt; III&gt; V&gt; II&gt; I; outer/innermetatarsal tubercle ratio 0.500.55; from the Pacific slopes of
<collectingRegion box="[179,250,1316,1340]" country="Mexico" name="Jalisco" pageId="40" pageNumber="40">Jalisco</collectingRegion>
west of Talpa de Allende (
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elevation).........................
<taxonomicName authority="Jameson et al." authorityName="Jameson" box="[402,788,1346,1370]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="40" pageNumber="40" phylum="Chordata" rank="species" species="rubinus">
<emphasis box="[402,620,1346,1370]" italics="true" pageId="40" pageNumber="40">Craugastor rubinus</emphasis>
Jameson et al.
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>