treatments-xml/data/03/F2/87/03F287B4FF93FFDDD639FD2A7CD8FBEA.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4324229" ID-GBIF-Dataset="d5ddaf75-c39b-49b3-bbf5-7e7af2f7ad66" ID-GBIF-Taxon="168144934" ID-Zenodo-Dep="4324229" approvalRequired="5" approvalRequired_for_bibRefs="2" approvalRequired_for_document="1" approvalRequired_for_textStreams="2" checkinTime="1601372077630" checkinUser="plazi" docAuthor="Agudelo‐Zamora, Henry D., Tavera, José, Murillo, Yiskar D. &amp; Ortega‐Lara, Armando" docDate="2020" docId="03F287B4FF93FFDDD639FD2A7CD8FBEA" docLanguage="en" docName="jFishBiology.97.4.1-14.pdf" docOrigin="Journal of Fish Biology 97 (4)" docSource="http://dx.doi.org/10.1111/jfb.14527" docStyle="DocumentStyle{}" docTitle="Characidium tatama Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara 2020, new species" docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="7" masterDocId="FFCBFFCCFF97FFDAD509FFD07C36FFEF" masterDocTitle="The unknown diversity of the genus Characidium (Characiformes: Crenuchidae) in the Chocó biogeographic region, Colombian Andes: Two new species supported by morphological and molecular data" masterLastPageNumber="14" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1635496756590" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The unknown diversity of the genus Characidium (Characiformes: Crenuchidae) in the Chocó biogeographic region, Colombian Andes: Two new species supported by morphological and molecular data</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Agudelo‐Zamora, Henry D.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8571-7235</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tavera, José</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4517-9238</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Murillo, Yiskar D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortega‐Lara, Armando</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2015-7948</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Journal of Fish Biology</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-09-27</mods:number>
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<mods:detail type="volume">
<mods:number>97</mods:number>
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<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:start>1</mods:start>
<mods:end>14</mods:end>
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<mods:url>http://dx.doi.org/10.1111/jfb.14527</mods:url>
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<mods:identifier type="DOI">10.1111/jfb.14527</mods:identifier>
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<subSubSection box="[816,1304,761,787]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph blockId="4.[816,1304,761,787]" box="[816,1304,761,787]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[816,853,762,787]" pageId="4" pageNumber="5">3.2</emphasis>
| 
<heading bold="true" box="[922,1304,761,787]" fontSize="10" level="3" pageId="4" pageNumber="5" reason="0">
<emphasis bold="true" box="[922,1304,761,787]" pageId="4" pageNumber="5">
<taxonomicName authority="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[922,1153,761,787]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tatama" status="new species">
<emphasis bold="true" box="[922,1153,761,787]" italics="true" pageId="4" pageNumber="5">Characidium tatama</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1160,1304,762,787]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph blockId="4.[816,1463,835,994]" box="[816,1103,835,855]" pageId="4" pageNumber="5">
<figureCitation box="[816,903,835,855]" captionStart="FIGURE 3" captionStartId="4.[119,210,1916,1935]" captionTargetBox="[122,765,1268,1875]" captionTargetId="figure-349@4.[122,765,1268,1875]" captionTargetPageId="4" captionText="FIGURE 3 C. tatama n. sp., Holotype, IMCN 8925, 38.1 mm LS, Colombia, Chocó, San José Del Palmar. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058139" httpUri="https://zenodo.org/record/4058139/files/figure.png" pageId="4" pageNumber="5">Figures 3</figureCitation>
, 
<figureCitation box="[911,923,835,855]" captionStart="FIGURE 4" captionStartId="4.[816,907,609,628]" captionTargetBox="[819,1462,139,567]" captionTargetId="figure-380@4.[818,1463,138,568]" captionTargetPageId="4" captionText="FIGURE 4 Scale detail of C. tatama n. sp. IMCN 8924, Paratype, 36.3 mm LS, white arrows show the rounded point characteristic in the lateral line" figureDoi="http://doi.org/10.5281/zenodo.4058137" httpUri="https://zenodo.org/record/4058137/files/figure.png" pageId="4" pageNumber="5">4</figureCitation>
and 
<figureCitation box="[968,982,835,855]" captionStart="FIGURE 5" captionStartId="4.[816,907,1916,1935]" captionTargetBox="[818,1461,1393,1876]" captionTargetId="figure-414@4.[818,1461,1393,1876]" captionText="FIGURE 5 Typical habitat of C. dule, Ingará River, San José Del Palmar, Chocó, Colombia" figureDoi="http://doi.org/10.5281/zenodo.4058141" httpUri="https://zenodo.org/record/4058141/files/figure.png" pageId="4" pageNumber="5">5</figureCitation>
, and 
<tableCitation box="[1030,1098,835,855]" captionStart="TABLE 1" captionStartId="5.[124,202,139,158]" captionTargetBox="[143,1293,186,993]" captionText="TABLE 1 Morphometric data from C. tatama n. sp. (n = 7)" httpUri="http://table.plazi.org/id/DF24662AFF92FFDFD575FF5B7E85FF70" pageId="4" pageNumber="5" tableUuid="DF24662AFF92FFDFD575FF5B7E85FF70">Table 1</tableCitation>
.
</paragraph>
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<paragraph blockId="4.[816,1463,835,994]" pageId="4" pageNumber="5">urn:lsid:zoobank.org:act:4170703E80D7-4170-9212-10B5918D3E79.</paragraph>
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<paragraph blockId="4.[816,1463,835,994]" pageId="4" pageNumber="5">
<taxonomicName authorityName="J.T.Reinhardt" authorityYear="1867" box="[856,967,939,959]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis box="[856,967,939,959]" italics="true" pageId="4" pageNumber="5">Characidium</emphasis>
</taxonomicName>
sp. 2 
<emphasis bold="true" box="[1025,1033,940,959]" pageId="4" pageNumber="5">“</emphasis>
Pacifico 
<emphasis bold="true" box="[1105,1132,940,959]" pageId="4" pageNumber="5">” –</emphasis>
<bibRefCitation author="Agudelo-Zamora, H. &amp; Ortega-Lara, A. &amp; Taphorn, D." box="[1139,1407,939,959]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="5" pagination="249 - 263" part="4768" refId="ref9983" refString="Agudelo-Zamora, H., Ortega-Lara, A., &amp; Taphorn, D. (2020). Characidium chancoense, a new species of south American darter from the Rio Cauca drainage, Colombia (Characiformes: Crenuchidae). Zootaxa, 4768 (2), 249 - 263. https: // doi. org / 10.11646 / zootaxa. 4768.2.6." title="Characidium chancoense, a new species of south American darter from the Rio Cauca drainage, Colombia (Characiformes: Crenuchidae)" type="journal article" year="2020">
Agudelo-Zamora 
<emphasis box="[1300,1339,939,959]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
., 2020
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, p. 14 (material examined).
</paragraph>
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<emphasis bold="true" box="[816,853,1074,1099]" pageId="4" pageNumber="5">3.3</emphasis>
| 
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<emphasis bold="true" box="[922,1032,1074,1099]" pageId="4" pageNumber="5">
<typeStatus box="[922,1032,1074,1099]" pageId="4" pageNumber="5">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2865196308" collectingDate="2018-09-09" collectionCode="IMCN" collectorName="A. Ortega-Lara" country="Colombia" county="rio Ingara by the bridge" latitude="4.920386" location="Choco" longLatPrecision="1" longitude="-76.20596" municipality="San Jose Del Palmar" pageId="4" pageNumber="5" specimenCode="IMCN 8925" specimenCount="1" typeStatus="holotype">
<paragraph blockId="4.[816,1465,1147,1237]" pageId="4" pageNumber="5">
<specimenCode box="[816,931,1147,1167]" collectionCode="IMCN" pageId="4" pageNumber="5">IMCN 8925</specimenCode>
, 
<quantity box="[941,1023,1147,1167]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="5" unit="mm" value="38.1">38.1 mm</quantity>
<emphasis box="[1030,1040,1147,1167]" italics="true" pageId="4" pageNumber="5">L</emphasis>
<subScript attach="left" box="[1040,1049,1156,1169]" fontSize="5" pageId="4" pageNumber="5">S</subScript>
, 
<collectingCountry box="[1061,1151,1147,1167]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:03F287B4FF93FFDDD639FD2A7CD8FBEA:8E846079FF93FFDED180FBAB78F1FB60" box="[1161,1223,1147,1167]" country="Colombia" county="rio Ingara by the bridge" latitude="4.920386" longLatPrecision="1" longitude="-76.20596" municipality="San Jose Del Palmar" name="Choco" pageId="4" pageNumber="5">Chocó</location>
, 
<collectingMunicipality box="[1233,1429,1147,1167]" pageId="4" pageNumber="5">San José Del Palmar</collectingMunicipality>
, 
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, 
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04 55 
<emphasis bold="true" box="[1069,1073,1179,1192]" italics="true" pageId="4" pageNumber="5">
<superScript attach="none" box="[1069,1073,1179,1192]" fontSize="6" pageId="4" pageNumber="5">'</superScript>
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13.39 
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<superScript attach="right" box="[1128,1136,1179,1192]" fontSize="6" pageId="4" pageNumber="5">''</superScript>
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N
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, 
<geoCoordinate box="[1164,1314,1179,1202]" degrees="76" direction="west" minutes="12" orientation="longitude" pageId="4" pageNumber="5" precision="1" seconds="21.48" value="-76.20596">
76 12 
<emphasis bold="true" box="[1222,1226,1179,1192]" italics="true" pageId="4" pageNumber="5">
<superScript attach="none" box="[1222,1226,1179,1192]" fontSize="6" pageId="4" pageNumber="5">'</superScript>
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21.48 
<emphasis bold="true" box="[1281,1289,1179,1192]" italics="true" pageId="4" pageNumber="5">
<superScript attach="right" box="[1281,1289,1179,1192]" fontSize="6" pageId="4" pageNumber="5">''</superScript>
</emphasis>
W
</geoCoordinate>
, 
<collectorName box="[1323,1461,1182,1202]" pageId="4" pageNumber="5">A. Ortega-Lara</collectorName>
.
</paragraph>
<paragraph blockId="4.[816,1465,1147,1237]" box="[816,1007,1217,1237]" pageId="4" pageNumber="5">
<date box="[816,1003,1217,1237]" pageId="4" pageNumber="5" value="2018-09-09">
<collectingDate box="[816,1003,1217,1237]" pageId="4" pageNumber="5" value="2018-09-09">09 September 2018</collectingDate>
</date>
.
</paragraph>
</materialsCitation>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4058139" ID-Zenodo-Dep="4058139" httpUri="https://zenodo.org/record/4058139/files/figure.png" pageId="4" pageNumber="5" startId="4.[119,210,1916,1935]" targetBox="[122,765,1268,1875]" targetPageId="4">
<paragraph blockId="4.[119,740,1915,1965]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[119,231,1916,1935]" pageId="4" pageNumber="5">FIGURE 3</emphasis>
<taxonomicName authority="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[261,349,1915,1935]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tatama" status="n. sp.">
<emphasis box="[261,349,1915,1935]" italics="true" pageId="4" pageNumber="5">C. tatama</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[354,406,1915,1935]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
, Holotype, IMCN 8925, 38.1 mm 
<emphasis box="[716,726,1915,1935]" italics="true" pageId="4" pageNumber="5">L</emphasis>
<subScript attach="left" box="[726,735,1924,1937]" fontSize="5" pageId="4" pageNumber="5">S</subScript>
, Colombia, Chocó, San José Del Palmar. Scale bar = 1 cm
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4058141" ID-Zenodo-Dep="4058141" httpUri="https://zenodo.org/record/4058141/files/figure.png" pageId="4" pageNumber="5" startId="4.[816,907,1916,1935]" targetBox="[818,1461,1393,1876]" targetPageId="4">
<paragraph blockId="4.[816,1432,1915,1965]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[816,928,1916,1935]" pageId="4" pageNumber="5">FIGURE 5</emphasis>
Typical habitat of 
<taxonomicName authority=", Ingara River, San Jose Del Palmar" authorityName="Ingara River, San Jose Del Palmar" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="dule">
<emphasis box="[1123,1184,1915,1935]" italics="true" pageId="4" pageNumber="5">C. dule</emphasis>
, Ingará River, San José Del Palmar
</taxonomicName>
, Chocó, Colombia
</paragraph>
</caption>
<caption ID-Table-UUID="DF24662AFF92FFDFD575FF5B7E85FF70" box="[124,691,139,159]" httpUri="http://table.plazi.org/id/DF24662AFF92FFDFD575FF5B7E85FF70" pageId="5" pageNumber="6" startId="5.[124,202,139,158]" targetBox="[143,1293,186,993]" targetIsTable="true" targetPageId="5">
<paragraph blockId="5.[124,222,139,158]" box="[124,691,139,159]" lastBlockId="5.[252,691,139,159]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[124,222,139,158]" pageId="5" pageNumber="6">TABLE 1</emphasis>
Morphometric data from 
<taxonomicName authority="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[486,574,139,159]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tatama" status="n. sp.">
<emphasis box="[486,574,139,159]" italics="true" pageId="5" pageNumber="6">
C. 
<figureCitation box="[510,592,139,159]" captionStart="FIGURE 3" captionStartId="4.[119,210,1916,1935]" captionTargetBox="[122,765,1268,1875]" captionTargetId="figure-349@4.[122,765,1268,1875]" captionText="FIGURE 3 C. tatama n. sp., Holotype, IMCN 8925, 38.1 mm LS, Colombia, Chocó, San José Del Palmar. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058139" httpUri="https://zenodo.org/record/4058139/files/figure.png" pageId="5" pageNumber="6">
tatama 
<taxonomicNameLabel box="[579,629,139,159]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</figureCitation>
</emphasis>
</taxonomicName>
(
<emphasis box="[638,649,139,159]" italics="true" pageId="5" pageNumber="6">n</emphasis>
= 7)
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="6">
<table box="[143,1293,186,993]" gridcols="5" gridrows="22" pageId="5" pageNumber="6">
<tr box="[143,1293,186,205]" gridrow="0" pageId="5" pageNumber="6" rowspan-0="1">
<th box="[530,612,186,205]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">
<heading bold="true" box="[530,612,187,205]" fontSize="7" level="6" pageId="5" pageNumber="6" reason="2">Holotype</heading>
</th>
<th box="[781,870,186,205]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">
<heading bold="true" box="[781,836,187,205]" fontSize="7" level="6" pageId="5" pageNumber="6" reason="2">Range</heading>
</th>
<th box="[1038,1089,186,205]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">
<heading bold="true" box="[1038,1089,187,205]" fontSize="7" level="6" pageId="5" pageNumber="6" reason="2">
<emphasis bold="true" box="[1038,1089,187,205]" pageId="5" pageNumber="6">Mean</emphasis>
</heading>
</th>
<th box="[1257,1293,186,205]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">
<emphasis bold="true" box="[1257,1293,186,205]" pageId="5" pageNumber="6">S.D.</emphasis>
</th>
</tr>
<tr box="[143,1293,228,246]" gridrow="1" pageId="5" pageNumber="6" rowspan-4="1">
<th box="[143,363,228,246]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">TL</th>
<td box="[530,612,228,246]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">44.2</td>
<td box="[781,870,228,246]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">
33.6 
<emphasis bold="true" box="[820,831,228,246]" pageId="5" pageNumber="6">–</emphasis>
44.2
</td>
<td box="[1038,1089,228,246]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">39.5</td>
</tr>
<tr box="[143,1293,265,284]" gridrow="2" pageId="5" pageNumber="6" rowspan-4="1">
<th box="[143,363,265,284]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">SL</th>
<td box="[530,612,265,284]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">38.1</td>
<td box="[781,870,265,284]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">
27.9 
<emphasis bold="true" box="[820,831,266,284]" pageId="5" pageNumber="6">–</emphasis>
38.1
</td>
<td box="[1038,1089,265,284]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">33.2</td>
</tr>
<tr box="[143,1293,302,321]" gridrow="3" pageId="5" pageNumber="6" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th box="[143,363,302,321]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">% SL</th>
</tr>
<tr box="[143,1293,340,358]" gridrow="4" pageId="5" pageNumber="6">
<th box="[143,363,340,358]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Head length</th>
<td box="[530,612,340,358]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">21.5</td>
<td box="[781,870,340,358]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">
22.5 
<emphasis bold="true" box="[820,831,340,358]" pageId="5" pageNumber="6">–</emphasis>
25
</td>
<td box="[1038,1089,340,358]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">23.8</td>
<td box="[1257,1293,340,358]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">0.9</td>
</tr>
<tr box="[143,1293,378,396]" gridrow="5" pageId="5" pageNumber="6">
<th box="[143,363,378,396]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Pectoral distance</th>
<td box="[530,612,378,396]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">20.3</td>
<td box="[781,870,378,396]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">
20.8 
<emphasis bold="true" box="[820,831,378,396]" pageId="5" pageNumber="6">–</emphasis>
26.1
</td>
<td box="[1038,1089,378,396]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">22.8</td>
<td box="[1257,1293,378,396]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">2.0</td>
</tr>
<tr box="[143,1293,415,433]" gridrow="6" pageId="5" pageNumber="6">
<th box="[143,363,415,433]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Predorsal distance</th>
<td box="[530,612,415,433]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">45.5</td>
<td box="[781,870,415,433]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">
44.2 
<emphasis bold="true" box="[820,831,415,433]" pageId="5" pageNumber="6">–</emphasis>
50.6
</td>
<td box="[1038,1089,415,433]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">47.2</td>
<td box="[1257,1293,415,433]" gridcol="4" gridrow="6" pageId="5" pageNumber="6">2.3</td>
</tr>
<tr box="[143,1293,452,470]" gridrow="7" pageId="5" pageNumber="6">
<th box="[143,363,452,470]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Prepelvic distance</th>
<td box="[530,612,452,470]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">51.2</td>
<td box="[781,870,452,470]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">
48.3 
<emphasis bold="true" box="[820,831,452,470]" pageId="5" pageNumber="6">–</emphasis>
59.1
</td>
<td box="[1038,1089,452,470]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">53.4</td>
<td box="[1257,1293,452,470]" gridcol="4" gridrow="7" pageId="5" pageNumber="6">4.0</td>
</tr>
<tr box="[143,1293,490,508]" gridrow="8" pageId="5" pageNumber="6">
<th box="[143,363,490,508]" gridcol="0" gridrow="8" pageId="5" pageNumber="6">Preanal distance</th>
<td box="[530,612,490,508]" gridcol="1" gridrow="8" pageId="5" pageNumber="6">76.2</td>
<td box="[781,870,490,508]" gridcol="2" gridrow="8" pageId="5" pageNumber="6">
71.1 
<emphasis bold="true" box="[820,831,490,508]" pageId="5" pageNumber="6">–</emphasis>
83.1
</td>
<td box="[1038,1089,490,508]" gridcol="3" gridrow="8" pageId="5" pageNumber="6">76.6</td>
<td box="[1257,1293,490,508]" gridcol="4" gridrow="8" pageId="5" pageNumber="6">3.9</td>
</tr>
<tr box="[143,1293,527,545]" gridrow="9" pageId="5" pageNumber="6">
<th box="[143,363,527,545]" gridcol="0" gridrow="9" pageId="5" pageNumber="6">Anal-apex distance</th>
<td box="[530,612,527,545]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">84.0</td>
<td box="[781,870,527,545]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">
82.1 
<emphasis bold="true" box="[820,831,527,545]" pageId="5" pageNumber="6">–</emphasis>
90.2
</td>
<td box="[1038,1089,527,545]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">85.8</td>
<td box="[1257,1293,527,545]" gridcol="4" gridrow="9" pageId="5" pageNumber="6">3.4</td>
</tr>
<tr box="[143,1293,564,582]" gridrow="10" pageId="5" pageNumber="6">
<th box="[143,363,564,582]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">Body width</th>
<td box="[530,612,564,582]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">10.5</td>
<td box="[781,870,564,582]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">
9.4 
<emphasis bold="true" box="[820,831,564,582]" pageId="5" pageNumber="6">–</emphasis>
11.3
</td>
<td box="[1038,1089,564,582]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">10.2</td>
<td box="[1257,1293,564,582]" gridcol="4" gridrow="10" pageId="5" pageNumber="6">0.7</td>
</tr>
<tr box="[143,1293,601,620]" gridrow="11" pageId="5" pageNumber="6">
<th box="[143,363,601,620]" gridcol="0" gridrow="11" pageId="5" pageNumber="6">Body depth at DO</th>
<td box="[530,612,601,620]" gridcol="1" gridrow="11" pageId="5" pageNumber="6">17.4</td>
<td box="[781,870,601,620]" gridcol="2" gridrow="11" pageId="5" pageNumber="6">
15.2 
<emphasis bold="true" box="[820,831,602,620]" pageId="5" pageNumber="6">–</emphasis>
19.3
</td>
<td box="[1038,1089,601,620]" gridcol="3" gridrow="11" pageId="5" pageNumber="6">18.0</td>
<td box="[1257,1293,601,620]" gridcol="4" gridrow="11" pageId="5" pageNumber="6">1.6</td>
</tr>
<tr box="[143,1293,638,657]" gridrow="12" pageId="5" pageNumber="6">
<th box="[143,363,638,657]" gridcol="0" gridrow="12" pageId="5" pageNumber="6">Body depth at AO</th>
<td box="[530,612,638,657]" gridcol="1" gridrow="12" pageId="5" pageNumber="6">12.9</td>
<td box="[781,870,638,657]" gridcol="2" gridrow="12" pageId="5" pageNumber="6">
12.7 
<emphasis bold="true" box="[820,831,639,657]" pageId="5" pageNumber="6">–</emphasis>
16.6
</td>
<td box="[1038,1089,638,657]" gridcol="3" gridrow="12" pageId="5" pageNumber="6">14.4</td>
<td box="[1257,1293,638,657]" gridcol="4" gridrow="12" pageId="5" pageNumber="6">1.3</td>
</tr>
<tr box="[143,1293,676,695]" gridrow="13" pageId="5" pageNumber="6">
<th box="[143,363,676,695]" gridcol="0" gridrow="13" pageId="5" pageNumber="6">Caudal peduncle depth</th>
<td box="[530,612,676,695]" gridcol="1" gridrow="13" pageId="5" pageNumber="6">11.0</td>
<td box="[781,870,676,695]" gridcol="2" gridrow="13" pageId="5" pageNumber="6">
9.4 
<emphasis bold="true" box="[820,831,676,694]" pageId="5" pageNumber="6">–</emphasis>
11.5
</td>
<td box="[1038,1089,676,695]" gridcol="3" gridrow="13" pageId="5" pageNumber="6">10.5</td>
<td box="[1257,1293,676,695]" gridcol="4" gridrow="13" pageId="5" pageNumber="6">0.7</td>
</tr>
<tr box="[143,1293,714,732]" gridrow="14" pageId="5" pageNumber="6" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th box="[143,363,714,732]" gridcol="0" gridrow="14" pageId="5" pageNumber="6">% HL</th>
</tr>
<tr box="[143,1293,750,769]" gridrow="15" pageId="5" pageNumber="6">
<th box="[143,363,750,769]" gridcol="0" gridrow="15" pageId="5" pageNumber="6">Snout length</th>
<td box="[530,612,750,769]" gridcol="1" gridrow="15" pageId="5" pageNumber="6">23.0</td>
<td box="[781,870,750,769]" gridcol="2" gridrow="15" pageId="5" pageNumber="6">
17.7 
<emphasis bold="true" box="[820,831,751,769]" pageId="5" pageNumber="6">–</emphasis>
22.8
</td>
<td box="[1038,1089,750,769]" gridcol="3" gridrow="15" pageId="5" pageNumber="6">19.8</td>
<td box="[1257,1293,750,769]" gridcol="4" gridrow="15" pageId="5" pageNumber="6">2.1</td>
</tr>
<tr box="[143,1293,788,807]" gridrow="16" pageId="5" pageNumber="6">
<th box="[143,363,788,807]" gridcol="0" gridrow="16" pageId="5" pageNumber="6">Snout-maxillary tip</th>
<td box="[530,612,788,807]" gridcol="1" gridrow="16" pageId="5" pageNumber="6">27.5</td>
<td box="[781,870,788,807]" gridcol="2" gridrow="16" pageId="5" pageNumber="6">
21.8 
<emphasis bold="true" box="[820,831,788,806]" pageId="5" pageNumber="6">–</emphasis>
28
</td>
<td box="[1038,1089,788,807]" gridcol="3" gridrow="16" pageId="5" pageNumber="6">25.7</td>
<td box="[1257,1293,788,807]" gridcol="4" gridrow="16" pageId="5" pageNumber="6">2.3</td>
</tr>
<tr box="[143,1293,826,844]" gridrow="17" pageId="5" pageNumber="6">
<th box="[143,363,826,844]" gridcol="0" gridrow="17" pageId="5" pageNumber="6">Anterior naris-eye</th>
<td box="[530,612,826,844]" gridcol="1" gridrow="17" pageId="5" pageNumber="6">8.2</td>
<td box="[781,870,826,844]" gridcol="2" gridrow="17" pageId="5" pageNumber="6">
3.7 
<emphasis bold="true" box="[820,831,826,844]" pageId="5" pageNumber="6">–</emphasis>
10.2
</td>
<td box="[1038,1089,826,844]" gridcol="3" gridrow="17" pageId="5" pageNumber="6">6.0</td>
<td box="[1257,1293,826,844]" gridcol="4" gridrow="17" pageId="5" pageNumber="6">2.3</td>
</tr>
<tr box="[143,1293,863,881]" gridrow="18" pageId="5" pageNumber="6">
<th box="[143,363,863,881]" gridcol="0" gridrow="18" pageId="5" pageNumber="6">Posterior naris - eye</th>
<td box="[530,612,863,881]" gridcol="1" gridrow="18" pageId="5" pageNumber="6">1,7</td>
<td box="[781,870,863,881]" gridcol="2" gridrow="18" pageId="5" pageNumber="6">
1.3 
<emphasis bold="true" box="[820,831,863,881]" pageId="5" pageNumber="6">–</emphasis>
3.8
</td>
<td box="[1038,1089,863,881]" gridcol="3" gridrow="18" pageId="5" pageNumber="6">2.6</td>
<td box="[1257,1293,863,881]" gridcol="4" gridrow="18" pageId="5" pageNumber="6">1.0</td>
</tr>
<tr box="[143,1293,900,919]" gridrow="19" pageId="5" pageNumber="6">
<th box="[143,363,900,919]" gridcol="0" gridrow="19" pageId="5" pageNumber="6">Cheek</th>
<td box="[530,612,900,919]" gridcol="1" gridrow="19" pageId="5" pageNumber="6">23.4</td>
<td box="[781,870,900,919]" gridcol="2" gridrow="19" pageId="5" pageNumber="6">
18.5 
<emphasis bold="true" box="[820,831,900,918]" pageId="5" pageNumber="6">–</emphasis>
25.8
</td>
<td box="[1038,1089,900,919]" gridcol="3" gridrow="19" pageId="5" pageNumber="6">21.3</td>
<td box="[1257,1293,900,919]" gridcol="4" gridrow="19" pageId="5" pageNumber="6">2.8</td>
</tr>
<tr box="[143,1293,937,956]" gridrow="20" pageId="5" pageNumber="6">
<th box="[143,363,937,956]" gridcol="0" gridrow="20" pageId="5" pageNumber="6">Orbital diameter</th>
<td box="[530,612,937,956]" gridcol="1" gridrow="20" pageId="5" pageNumber="6">26.6</td>
<td box="[781,870,937,956]" gridcol="2" gridrow="20" pageId="5" pageNumber="6">
24.7 
<emphasis bold="true" box="[820,831,938,956]" pageId="5" pageNumber="6">–</emphasis>
31.4
</td>
<td box="[1038,1089,937,956]" gridcol="3" gridrow="20" pageId="5" pageNumber="6">28.0</td>
<td box="[1257,1293,937,956]" gridcol="4" gridrow="20" pageId="5" pageNumber="6">2.3</td>
</tr>
<tr box="[143,1293,975,993]" gridrow="21" pageId="5" pageNumber="6">
<th box="[143,363,975,993]" gridcol="0" gridrow="21" pageId="5" pageNumber="6">Interorbital distance</th>
<td box="[530,612,975,993]" gridcol="1" gridrow="21" pageId="5" pageNumber="6">10.8</td>
<td box="[781,870,975,993]" gridcol="2" gridrow="21" pageId="5" pageNumber="6">
8.7 
<emphasis bold="true" box="[820,831,975,993]" pageId="5" pageNumber="6">–</emphasis>
17.7
</td>
<td box="[1038,1089,975,993]" gridcol="3" gridrow="21" pageId="5" pageNumber="6">13.1</td>
<td box="[1257,1293,975,993]" gridcol="4" gridrow="21" pageId="5" pageNumber="6">3.3</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[124,1222,1018,1038]" box="[124,1222,1018,1038]" pageId="5" pageNumber="6">
<tableNote box="[124,1222,1018,1038]" pageId="5" pageNumber="6" targetBox="[143,1293,186,993]" targetPageId="5">Abbreviations: DO, dorsal origin; AO, anal origin; HL, head length; S.D; standard deviation; LS, standard length; TL, total length.</tableNote>
</paragraph>
<paragraph blockId="5.[124,347,1109,1134]" box="[124,347,1109,1134]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[124,161,1109,1134]" pageId="5" pageNumber="6">3.4</emphasis>
| 
<heading bold="true" box="[230,347,1109,1134]" fontSize="10" level="3" pageId="5" pageNumber="6" reason="0">
<emphasis bold="true" box="[230,347,1109,1134]" pageId="5" pageNumber="6">
<typeStatus box="[230,347,1109,1134]" pageId="5" pageNumber="6">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[124,773,1182,1514]" pageId="5" pageNumber="6">
All from 
<materialsCitation ID-GBIF-Occurrence="2865196309" collectionCode="IMCN" country="Colombia" location="Colombia" pageId="5" pageNumber="6" specimenCode="IMCN 8924, CHIFF 0011" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry box="[210,300,1182,1202]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
. 
<specimenCode box="[312,428,1182,1202]" collectionCode="IMCN" pageId="5" pageNumber="6">IMCN 8924</specimenCode>
, 1 C&amp;S 
<specimenCount box="[511,575,1182,1202]" pageId="5" pageNumber="6" type="female">female</specimenCount>
, 36.0 mm 
<emphasis box="[678,688,1182,1202]" italics="true" pageId="5" pageNumber="6">L</emphasis>
<subScript attach="left" box="[688,697,1190,1203]" fontSize="5" pageId="5" pageNumber="6">S</subScript>
; 
<specimenCode collectionCode="CHIFF" pageId="5" pageNumber="6">CHIFF 0011</specimenCode>
, 2, 
<quantity box="[217,353,1217,1237]" metricMagnitude="-2" metricUnit="m" metricValue="3.2099999999999995" metricValueMax="3.63" metricValueMin="2.79" pageId="5" pageNumber="6" unit="mm" value="32.099999999999994" valueMax="36.3" valueMin="27.9">
27.9 
<emphasis bold="true" box="[258,270,1217,1236]" pageId="5" pageNumber="6">–</emphasis>
36.3 mm
</quantity>
<emphasis box="[363,373,1217,1237]" italics="true" pageId="5" pageNumber="6">L</emphasis>
<subScript attach="left" box="[373,382,1225,1238]" fontSize="5" pageId="5" pageNumber="6">S</subScript>
, all collected with the holotype
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="2865196306" collectingDate="2018-09-09" collectionCode="IMCN, AOL" collectorName="A. Ortega-Lara." country="Colombia" county="Choco" location="rio Cruces tributary of rio Ingara" municipality="San Jose Del Palmar" pageId="5" pageNumber="6" specimenCode="IMCN 8926, 1, AOL 138" specimenCount="1" typeStatus="paratype">
<specimenCode collectionCode="IMCN" pageId="5" pageNumber="6">IMCN 8926</specimenCode>
, 
<specimenCode box="[185,197,1251,1271]" collectionCode="IMCN" pageId="5" pageNumber="6">1</specimenCode>
(
<specimenCode box="[209,296,1251,1271]" collectionCode="AOL" pageId="5" pageNumber="6">AOL 138</specimenCode>
tissue), 
<quantity box="[375,440,1251,1271]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="6" unit="mm" value="36.0">36 mm</quantity>
<emphasis box="[447,457,1251,1271]" italics="true" pageId="5" pageNumber="6">L</emphasis>
<subScript attach="left" box="[457,466,1260,1273]" fontSize="5" pageId="5" pageNumber="6">S</subScript>
, 
<collectingCountry box="[478,568,1251,1271]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
, 
<collectingCounty box="[578,640,1251,1271]" pageId="5" pageNumber="6">Chocó</collectingCounty>
, 
<collectingMunicipality pageId="5" pageNumber="6">San José Del Palmar</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03F287B4FF93FFDDD639FD2A7CD8FBEA:8E846079FF92FFDFD5DCFAD67E79FAF5" box="[213,591,1286,1306]" country="Colombia" county="Choco" municipality="San Jose Del Palmar" name="rio Cruces tributary of rio Ingara" pageId="5" pageNumber="6">río Cruces tributary of río Ingará</location>
, 
<collectorName box="[615,773,1286,1306]" pageId="5" pageNumber="6">A. Ortega-Lara.</collectorName>
<date box="[124,320,1321,1341]" pageId="5" pageNumber="6" value="2018-09-09">
<collectingDate box="[124,320,1321,1341]" pageId="5" pageNumber="6" value="2018-09-09">09 September 2018</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2865196307" collectingDate="2018-09-09" collectionCode="IMCN, AOL" collectorName="A. Ortega-Lara" country="Colombia" county="Choco" latitude="4.9152503" location="quebrada Corcobado" longLatPrecision="1" longitude="-76.21767" municipality="San Jose Del Palmar" pageId="5" pageNumber="6" specimenCode="IMCN 8927, 1, AOL 139" specimenCount="1" typeStatus="paratype">
<specimenCode box="[331,448,1321,1341]" collectionCode="IMCN" pageId="5" pageNumber="6">IMCN 8927</specimenCode>
, 
<specimenCode box="[459,471,1321,1341]" collectionCode="IMCN" pageId="5" pageNumber="6">1</specimenCode>
(
<specimenCode box="[486,575,1321,1341]" collectionCode="AOL" pageId="5" pageNumber="6">AOL 139</specimenCode>
tissue), 
<quantity box="[657,739,1321,1341]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="5" pageNumber="6" unit="mm" value="32.5">32.5 mm</quantity>
<emphasis box="[748,758,1321,1341]" italics="true" pageId="5" pageNumber="6">L</emphasis>
<subScript attach="left" box="[758,767,1329,1342]" fontSize="5" pageId="5" pageNumber="6">S</subScript>
, 
<collectingCountry box="[124,214,1355,1375]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
, 
<collectingCounty box="[234,296,1355,1375]" pageId="5" pageNumber="6">Chocó</collectingCounty>
, 
<collectingMunicipality box="[316,542,1355,1375]" pageId="5" pageNumber="6">San José Del Palmar</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03F287B4FF93FFDDD639FD2A7CD8FBEA:8E846079FF92FFDFD738FA9B7F37FAB0" box="[561,769,1355,1375]" country="Colombia" county="Choco" latitude="4.9152503" longLatPrecision="1" longitude="-76.21767" municipality="San Jose Del Palmar" name="quebrada Corcobado" pageId="5" pageNumber="6">quebrada Corcobado</location>
, 
<geoCoordinate box="[124,243,1387,1410]" degrees="4" direction="north" minutes="54" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="54.9" value="4.9152503">
4 54 
<emphasis bold="true" box="[169,173,1387,1400]" italics="true" pageId="5" pageNumber="6">
<superScript attach="none" box="[169,173,1387,1400]" fontSize="6" pageId="5" pageNumber="6">'</superScript>
</emphasis>
54.9 
<emphasis bold="true" box="[216,224,1387,1400]" italics="true" pageId="5" pageNumber="6">
<superScript attach="right" box="[216,224,1387,1400]" fontSize="6" pageId="5" pageNumber="6">''</superScript>
</emphasis>
N
</geoCoordinate>
, 
<geoCoordinate box="[260,386,1387,1410]" degrees="76" direction="west" minutes="13" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="3.6" value="-76.21767">
76 13 
<emphasis bold="true" box="[318,322,1387,1400]" italics="true" pageId="5" pageNumber="6">
<superScript attach="none" box="[318,322,1387,1400]" fontSize="6" pageId="5" pageNumber="6">'</superScript>
</emphasis>
3.6 
<emphasis bold="true" box="[352,360,1387,1400]" italics="true" pageId="5" pageNumber="6">
<superScript attach="right" box="[352,360,1387,1400]" fontSize="6" pageId="5" pageNumber="6">''</superScript>
</emphasis>
W
</geoCoordinate>
, 
<collectorName box="[402,549,1390,1410]" pageId="5" pageNumber="6">A. Ortega-Lara</collectorName>
, 
<date box="[565,769,1390,1410]" pageId="5" pageNumber="6" value="2018-09-09">
<collectingDate box="[565,769,1390,1410]" pageId="5" pageNumber="6" value="2018-09-09">09 September 2018</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2865196305" collectingDate="2010-05-16" collectionCode="CIRUV" collectorName="Diaz &amp; Serna." country="Colombia" county="Buenaventura" latitude="3.9904723" location="Buenaventura, quebrada La Brea, tributary of rio Calima" longLatPrecision="1" longitude="-76.97314" municipality="La Brea" pageId="5" pageNumber="6" specimenCode="CIRUV 010-19" specimenCount="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCode box="[124,265,1425,1445]" collectionCode="CIRUV" country="Colombia" httpUri="http://grbio.org/cool/f9et-gfx8" name="Coleccion Ictiologica de Referencia de la Universidad del Valle" pageId="5" pageNumber="6">CIRUV 010-19</specimenCode>
, 1, 
<quantity box="[298,380,1425,1445]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" pageId="5" pageNumber="6" unit="mm" value="30.5">30.5 mm</quantity>
<emphasis box="[387,397,1425,1445]" italics="true" pageId="5" pageNumber="6">L</emphasis>
<subScript attach="left" box="[397,406,1433,1446]" fontSize="5" pageId="5" pageNumber="6">S</subScript>
, 
<collectingCountry box="[417,507,1425,1445]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
, 
<collectingRegion box="[516,662,1425,1445]" country="Colombia" name="Valle del Cauca" pageId="5" pageNumber="6">Valle del Cauca</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:03F287B4FF93FFDDD639FD2A7CD8FBEA:8E846079FF92FFDFD796FA417E40FA28" country="Colombia" county="Buenaventura" latitude="3.9904723" longLatPrecision="1" longitude="-76.97314" municipality="La Brea" name="Buenaventura, quebrada La Brea, tributary of rio Calima" pageId="5" pageNumber="6" stateProvince="Valle del Cauca">Buenaventura, quebrada La Brea, tributary of río Calima</location>
, 
<geoCoordinate box="[649,769,1457,1479]" degrees="3" direction="north" minutes="59" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="25.7" value="3.9904723">
3 59 
<emphasis bold="true" box="[694,698,1457,1470]" italics="true" pageId="5" pageNumber="6">
<superScript attach="none" box="[694,698,1457,1470]" fontSize="6" pageId="5" pageNumber="6">'</superScript>
</emphasis>
25.7 
<emphasis bold="true" box="[741,749,1457,1470]" italics="true" pageId="5" pageNumber="6">
<superScript attach="right" box="[741,749,1457,1470]" fontSize="6" pageId="5" pageNumber="6">''</superScript>
</emphasis>
N
</geoCoordinate>
, 
<geoCoordinate box="[124,262,1491,1514]" degrees="76" direction="west" minutes="58" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="23.3" value="-76.97314">
76 58 
<emphasis bold="true" box="[181,185,1491,1504]" italics="true" pageId="5" pageNumber="6">
<superScript attach="none" box="[181,185,1491,1504]" fontSize="6" pageId="5" pageNumber="6">'</superScript>
</emphasis>
23.3 
<emphasis bold="true" box="[228,236,1491,1504]" italics="true" pageId="5" pageNumber="6">
<superScript attach="right" box="[228,236,1491,1504]" fontSize="6" pageId="5" pageNumber="6">''</superScript>
</emphasis>
W
</geoCoordinate>
. 
<collectorName box="[270,311,1494,1514]" pageId="5" pageNumber="6">Díaz</collectorName>
&amp; 
<collectorName box="[337,396,1494,1514]" pageId="5" pageNumber="6">Serna.</collectorName>
<date box="[400,528,1494,1514]" pageId="5" pageNumber="6" value="2010-05-16">
<collectingDate box="[400,528,1494,1514]" pageId="5" pageNumber="6" value="2010-05-16">16 May 2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph blockId="5.[124,345,1594,1619]" box="[124,345,1594,1619]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[124,161,1594,1619]" pageId="5" pageNumber="6">3.5</emphasis>
| 
<heading bold="true" box="[230,345,1594,1619]" fontSize="10" level="3" pageId="5" pageNumber="6" reason="0">
<emphasis bold="true" box="[230,345,1594,1619]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[124,773,1667,1965]" lastBlockId="5.[821,1469,1113,1722]" pageId="5" pageNumber="6">
<taxonomicName authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[124,313,1667,1687]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tatama">
<emphasis box="[124,313,1667,1687]" italics="true" pageId="5" pageNumber="6">Characidium tatama</emphasis>
</taxonomicName>
is distinguished from all 
<taxonomicName authorityName="J.T.Reinhardt" authorityYear="1867" box="[580,691,1667,1687]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis box="[580,691,1667,1687]" italics="true" pageId="5" pageNumber="6">Characidium</emphasis>
</taxonomicName>
, except 
<taxonomicName authority="Travassos 1955" authorityName="Travassos" authorityYear="1955" box="[124,353,1702,1722]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="alipioi">
<emphasis box="[124,200,1702,1722]" italics="true" pageId="5" pageNumber="6">C. alipioi</emphasis>
Travassos 1955
</taxonomicName>
, 
<taxonomicName authority="Lujan et al., 2013" authorityName="Lujan" authorityYear="2013" box="[362,611,1702,1722]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="amaila">
<emphasis box="[362,445,1702,1722]" italics="true" pageId="5" pageNumber="6">C. amaila</emphasis>
<bibRefCitation author="Lujan, N. K. &amp; Agudelo-Zamora, H. &amp; Taphorn, D. C. &amp; Booth, P. N. &amp; Fernandez, H." box="[450,611,1702,1722]" journalOrPublisher="Copeia" pageId="5" pageNumber="6" pagination="454 - 463" part="2013" refId="ref11390" refString="Lujan, N. K., Agudelo-Zamora, H., Taphorn, D. C., Booth, P. N., &amp; Lopez- Fernandez, H. (2013). Description of a new, narrowly endemic south American darter (Characiformes: Crenuchidae) from the central Guiana shield highlands of Guyana. Copeia, 2013 (3), 454 - 463. https: // doi. org / 10.1093 / molbev / msy 096." title="Description of a new, narrowly endemic south American darter (Characiformes: Crenuchidae) from the central Guiana shield highlands of Guyana" type="journal article" year="2013">
Lujan 
<emphasis box="[506,544,1702,1722]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2013
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Steindachner 1915" authorityName="Steindachner" authorityYear="1915" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="boavistae">
<emphasis box="[619,728,1702,1722]" italics="true" pageId="5" pageNumber="6">C. boavistae</emphasis>
Steindachner 1915
</taxonomicName>
, 
<taxonomicName authority="Pearson 1924" authorityName="Pearson" authorityYear="1924" box="[279,535,1737,1757]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="bolivianum">
<emphasis box="[279,399,1737,1757]" italics="true" pageId="5" pageNumber="6">C. bolivianum</emphasis>
Pearson 1924
</taxonomicName>
, 
<taxonomicName authority="Steindachner 1915" authorityName="Steindachner" authorityYear="1915" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="crandellii">
<emphasis box="[543,645,1737,1757]" italics="true" pageId="5" pageNumber="6">C. crandellii</emphasis>
Steindachner 1915
</taxonomicName>
, 
<taxonomicName authority="Steindachner 1915" authorityName="Steindachner" authorityYear="1915" box="[186,510,1771,1791]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="declivirostre">
<emphasis box="[186,319,1771,1791]" italics="true" pageId="5" pageNumber="6">C. declivirostre</emphasis>
Steindachner 1915
</taxonomicName>
, 
<taxonomicName authority="Travassos 1956" authorityName="Travassos" authorityYear="1956" box="[521,769,1771,1791]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gomesi">
<emphasis box="[521,609,1771,1791]" italics="true" pageId="5" pageNumber="6">C. gomesi</emphasis>
Travassos 1956
</taxonomicName>
, 
<taxonomicName authority="Travassos 1944" authorityName="Travassos" authorityYear="1944" box="[124,435,1806,1826]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="grajahuense">
<emphasis box="[124,262,1806,1826]" italics="true" pageId="5" pageNumber="6">C. grajahuense</emphasis>
Travassos 1944
</taxonomicName>
, 
<taxonomicName authority="Travassos 1949" authorityName="Travassos" authorityYear="1949" box="[453,769,1806,1826]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="japuhybense">
<emphasis box="[453,596,1806,1826]" italics="true" pageId="5" pageNumber="6">C. japuhybense</emphasis>
Travassos 1949
</taxonomicName>
, 
<taxonomicName authority="Zanata &amp; Camelier, 2015" authorityName="Zanata &amp; Camelier" authorityYear="2015" box="[124,504,1841,1861]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="kamakan">
<emphasis box="[124,235,1841,1861]" italics="true" pageId="5" pageNumber="6">C. kamakan</emphasis>
<bibRefCitation author="Zanata, A. M. &amp; Camelier, P." box="[249,504,1841,1861]" journalOrPublisher="Neotropical Ichthyology" pageId="5" pageNumber="6" pagination="487 - 498" part="13" refId="ref12522" refString="Zanata, A. M., &amp; Camelier, P. (2015). Two new species of Characidium Reinhardt (Characiformes: Crenuchidae) from northeastern Brazilian coastal drainages. Neotropical Ichthyology, 13 (3), 487 - 498. https: // doi. org / 10.1590 / 1982 - 0224 - 20140106." title="Two new species of Characidium Reinhardt (Characiformes: Crenuchidae) from northeastern Brazilian coastal drainages" type="journal article" year="2015">Zanata &amp; Camelier, 2015</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Travassos 1949" authorityName="Travassos" authorityYear="1949" box="[520,769,1841,1861]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="lauroi">
<emphasis box="[520,600,1841,1861]" italics="true" pageId="5" pageNumber="6">C. lauroi</emphasis>
Travassos 1949
</taxonomicName>
, 
<taxonomicName authority="Gomes 1947" authorityName="Gomes" authorityYear="1947" box="[124,414,1875,1895]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="pterostictum">
<emphasis box="[124,268,1875,1895]" italics="true" pageId="5" pageNumber="6">C. pterostictum</emphasis>
Gomes 1947
</taxonomicName>
, 
<taxonomicName authority="Steindachner 1882" authorityName="Steindachner" authorityYear="1882" box="[431,769,1875,1895]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="purpuratum">
<emphasis box="[431,568,1875,1895]" italics="true" pageId="5" pageNumber="6">C. purpuratum</emphasis>
Steindachner 1882
</taxonomicName>
, 
<taxonomicName authority="Travassos, 1946" authorityName="Travassos" authorityYear="1946" box="[124,404,1910,1930]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="timbuiense">
<emphasis box="[124,244,1910,1930]" italics="true" pageId="5" pageNumber="6">C. timbuiense</emphasis>
<bibRefCitation author="Travassos" box="[251,404,1910,1930]" journalOrPublisher="Sudeste do Brasil. (Masters thesis)" pageId="5" pageNumber="6" publicationUrl="http://portais4.ufes.br/posgrad/teses/tese_8576_Maridiesse%" refId="ref11886" refString="Travassos, 1946 (Characiformes: Crenuchidae), com descriC ~ ao de uma especie nova para o Caparao, Sudeste do Brasil. (Masters thesis). http: // portais 4. ufes. br / posgrad / teses / tese _ 8576 _ Maridiesse %" title="(Characiformes: Crenuchidae), com descriC ~ ao de uma especie nova para o Caparao" type="url" year="1946">Travassos, 1946</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Melo et al., 2016" authorityName="Melo" authorityYear="2016" box="[412,692,1910,1930]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="travassosi">
<emphasis box="[412,524,1910,1930]" italics="true" pageId="5" pageNumber="6">C. travassosi</emphasis>
<bibRefCitation author="Melo, M. R. S. &amp; Buckup, P. A. &amp; Oyakawa, O. T." box="[531,692,1910,1930]" journalOrPublisher="Neotropical Ichthyology" pageId="5" pageNumber="6" pagination="160014" part="14" refId="ref11680" refString="Melo, M. R. S., Buckup, P. A., &amp; Oyakawa, O. T. (2016). A new species of Characidium Reinhardt, 1867 (Characiformes: Crenuchidae) endemic to the Atlantic Forest in Parana state, southern Brazil. Neotropical Ichthyology, 14 (3), e 160014." title="A new species of Characidium Reinhardt, 1867 (Characiformes: Crenuchidae) endemic to the Atlantic Forest in Parana state, southern Brazil" type="journal article" year="2016">
Melo 
<emphasis box="[585,624,1910,1930]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2016
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Travassos 1967" authorityName="Travassos" authorityYear="1967" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="vidali">
<emphasis box="[701,771,1910,1930]" italics="true" pageId="5" pageNumber="6">C. vidali</emphasis>
Travassos 1967
</taxonomicName>
and the clade members of 
<taxonomicName authorityName="Reinhardt" authorityYear="1867" box="[539,653,1945,1965]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fasciatum">
<emphasis box="[539,653,1945,1965]" italics="true" pageId="5" pageNumber="6">C. fasciatum</emphasis>
</taxonomicName>
or clade C1 (
<emphasis box="[826,875,1113,1133]" italics="true" pageId="5" pageNumber="6">sensu</emphasis>
<bibRefCitation author="Buckup, P. A." box="[881,1011,1113,1133]" journalOrPublisher="Copeia" pageId="5" pageNumber="6" pagination="1066 - 1073" part="1992" refId="ref10456" refString="Buckup, P. A. (1992). Redescription of Characidium fasciatum, type species of the Characidiinae (Teleostei, Characiformes). Copeia, 1992, 1066 - 1073. https: // doi. org / 10.2307 / 1446639." title="Redescription of Characidium fasciatum, type species of the Characidiinae (Teleostei, Characiformes)" type="journal article" year="1992">Buckup, 1992</bibRefCitation>
) for having the isthmus without scales (Figure 3). Also, 
<taxonomicName authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[876,968,1147,1167]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tatama">
<emphasis box="[876,968,1147,1167]" italics="true" pageId="5" pageNumber="6">C. tatama</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName authority="Zanata" authorityName="Zanata" box="[1190,1359,1147,1167]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="helmeri">
<emphasis box="[1190,1282,1147,1167]" italics="true" pageId="5" pageNumber="6">C. helmeri</emphasis>
Zanata
</taxonomicName>
, Sarmento- Soares &amp; Martins-Pinheiro 2015, 
<taxonomicName authority="Travassos 1967" authorityName="Travassos" authorityYear="1967" box="[1158,1422,1182,1202]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="oiticicai">
<emphasis box="[1158,1256,1182,1202]" italics="true" pageId="5" pageNumber="6">C. oiticicai</emphasis>
Travassos 1967
</taxonomicName>
and 
<taxonomicName authority="Melo et al., 2016" authorityName="Melo" authorityYear="2016" box="[821,1109,1217,1237]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="travassossi">
<emphasis box="[821,942,1217,1237]" italics="true" pageId="5" pageNumber="6">C. travassossi</emphasis>
<bibRefCitation author="Melo, M. R. S. &amp; Buckup, P. A. &amp; Oyakawa, O. T." box="[948,1109,1217,1237]" journalOrPublisher="Neotropical Ichthyology" pageId="5" pageNumber="6" pagination="160014" part="14" refId="ref11680" refString="Melo, M. R. S., Buckup, P. A., &amp; Oyakawa, O. T. (2016). A new species of Characidium Reinhardt, 1867 (Characiformes: Crenuchidae) endemic to the Atlantic Forest in Parana state, southern Brazil. Neotropical Ichthyology, 14 (3), e 160014." title="A new species of Characidium Reinhardt, 1867 (Characiformes: Crenuchidae) endemic to the Atlantic Forest in Parana state, southern Brazil" type="journal article" year="2016">
Melo 
<emphasis box="[1003,1043,1217,1237]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2016
</bibRefCitation>
</taxonomicName>
by the presence of an adipose fin (
<emphasis box="[1445,1463,1217,1237]" italics="true" pageId="5" pageNumber="6">vs</emphasis>
. adipose fin absence). Additionally, 
<taxonomicName authority="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara, 2020" authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[1157,1249,1251,1271]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tatama" status="n. sp.">
<emphasis box="[1157,1249,1251,1271]" italics="true" pageId="5" pageNumber="6">
C. 
<figureCitation box="[1185,1272,1251,1271]" captionStart="FIGURE 3" captionStartId="4.[119,210,1916,1935]" captionTargetBox="[122,765,1268,1875]" captionTargetId="figure-349@4.[122,765,1268,1875]" captionText="FIGURE 3 C. tatama n. sp., Holotype, IMCN 8925, 38.1 mm LS, Colombia, Chocó, San José Del Palmar. Scale bar = 1 cm" figureDoi="http://doi.org/10.5281/zenodo.4058139" httpUri="https://zenodo.org/record/4058139/files/figure.png" pageId="5" pageNumber="6">
tatama 
<taxonomicNameLabel box="[1259,1313,1251,1271]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</figureCitation>
</emphasis>
</taxonomicName>
is distinguished among clade C1 congeners by having a rounded point in the midline of the upper scales of the lateral line (
<figureCitation box="[1166,1244,1321,1341]" captionStart="FIGURE 4" captionStartId="4.[816,907,609,628]" captionTargetBox="[819,1462,139,567]" captionTargetId="figure-380@4.[818,1463,138,568]" captionTargetPageId="4" captionText="FIGURE 4 Scale detail of C. tatama n. sp. IMCN 8924, Paratype, 36.3 mm LS, white arrows show the rounded point characteristic in the lateral line" figureDoi="http://doi.org/10.5281/zenodo.4058137" httpUri="https://zenodo.org/record/4058137/files/figure.png" pageId="5" pageNumber="6">Figure 4</figureCitation>
); it is distinguished from 
<taxonomicName box="[821,909,1355,1375]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="amaila">
<emphasis box="[821,909,1355,1375]" italics="true" pageId="5" pageNumber="6">C. amaila</emphasis>
</taxonomicName>
by having a lower number of scales on the lateral line (32 
<emphasis bold="true" box="[850,862,1391,1410]" pageId="5" pageNumber="6">–</emphasis>
33 
<emphasis box="[893,911,1390,1410]" italics="true" pageId="5" pageNumber="6">vs</emphasis>
. 34 
<emphasis bold="true" box="[947,959,1391,1410]" pageId="5" pageNumber="6">–</emphasis>
36) and by the shape of its swim bladder (wide bilobed anterior chamber and small posterior thin chamber 
<emphasis box="[1301,1319,1425,1445]" italics="true" pageId="5" pageNumber="6">vs</emphasis>
. anterior chamber as wide as the posterior, and the posterior chamber long and thin); from 
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[870,973,1494,1514]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="crandellii">
<emphasis box="[870,973,1494,1514]" italics="true" pageId="5" pageNumber="6">C. crandellii</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1018,1147,1494,1514]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="declivirostre">
<emphasis box="[1018,1147,1494,1514]" italics="true" pageId="5" pageNumber="6">C. declivirostre</emphasis>
</taxonomicName>
by having free branchiostegal rays (
<emphasis box="[826,844,1529,1549]" italics="true" pageId="5" pageNumber="6">vs</emphasis>
. joined branchiostegal rays [see 
<bibRefCitation author="Lujan, N. K. &amp; Agudelo-Zamora, H. &amp; Taphorn, D. C. &amp; Booth, P. N. &amp; Fernandez, H." box="[1160,1330,1529,1549]" journalOrPublisher="Copeia" pageId="5" pageNumber="6" pagination="454 - 463" part="2013" refId="ref11390" refString="Lujan, N. K., Agudelo-Zamora, H., Taphorn, D. C., Booth, P. N., &amp; Lopez- Fernandez, H. (2013). Description of a new, narrowly endemic south American darter (Characiformes: Crenuchidae) from the central Guiana shield highlands of Guyana. Copeia, 2013 (3), 454 - 463. https: // doi. org / 10.1093 / molbev / msy 096." title="Description of a new, narrowly endemic south American darter (Characiformes: Crenuchidae) from the central Guiana shield highlands of Guyana" type="journal article" year="2013">
Lujan 
<emphasis box="[1219,1261,1529,1549]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2013
</bibRefCitation>
, p. 455); from 
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[821,932,1563,1583]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="boavistae">
<emphasis box="[821,932,1563,1583]" italics="true" pageId="5" pageNumber="6">C. boavistae</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Pearson" authorityYear="1924" box="[944,1067,1563,1583]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="bolivianum">
<emphasis box="[944,1067,1563,1583]" italics="true" pageId="5" pageNumber="6">C. bolivianum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Travassos" authorityYear="1956" box="[1079,1167,1563,1583]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gomesi">
<emphasis box="[1079,1167,1563,1583]" italics="true" pageId="5" pageNumber="6">C. gomesi</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Travassos" authorityYear="1944" box="[1180,1315,1563,1583]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="grajahuensis">
<emphasis box="[1180,1315,1563,1583]" italics="true" pageId="5" pageNumber="6">C. grajahuensis</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Travassos" authorityYear="1949" box="[1328,1463,1563,1583]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="japuhybense">
<emphasis box="[1328,1463,1563,1583]" italics="true" pageId="5" pageNumber="6">C. japuhybense</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Zanata &amp; Camelier" authorityYear="2015" box="[821,926,1598,1618]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="kamakan">
<emphasis box="[821,926,1598,1618]" italics="true" pageId="5" pageNumber="6">C. kamakan</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Travassos" authorityYear="1949" box="[939,1014,1598,1618]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="lauroi">
<emphasis box="[939,1014,1598,1618]" italics="true" pageId="5" pageNumber="6">C. lauroi</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Steindachner" authorityYear="1882" box="[1026,1157,1598,1618]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="purpuratum">
<emphasis box="[1026,1157,1598,1618]" italics="true" pageId="5" pageNumber="6">C. purpuratum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Gomes" authorityYear="1947" box="[1169,1307,1598,1618]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="pterostictum">
<emphasis box="[1169,1307,1598,1618]" italics="true" pageId="5" pageNumber="6">C. pterostictum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Travassos" authorityYear="1955" box="[1319,1427,1598,1618]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="schubarti">
<emphasis box="[1319,1427,1598,1618]" italics="true" pageId="5" pageNumber="6">C. schubarti</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Travassos" authorityYear="1946" box="[821,939,1633,1653]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="timbuiense">
<emphasis box="[821,939,1633,1653]" italics="true" pageId="5" pageNumber="6">C. timbuiense</emphasis>
</taxonomicName>
for having the caudal and anal fins without bands on the rays (
<emphasis box="[871,889,1667,1687]" italics="true" pageId="5" pageNumber="6">vs</emphasis>
. caudal and anal fins with dark pigment or with some 
<typeStatus box="[1401,1442,1667,1687]" pageId="5" pageNumber="6">type</typeStatus>
of band) (see 
<bibRefCitation author="Lujan, N. K. &amp; Agudelo-Zamora, H. &amp; Taphorn, D. C. &amp; Booth, P. N. &amp; Fernandez, H." box="[920,1080,1702,1722]" journalOrPublisher="Copeia" pageId="5" pageNumber="6" pagination="454 - 463" part="2013" refId="ref11390" refString="Lujan, N. K., Agudelo-Zamora, H., Taphorn, D. C., Booth, P. N., &amp; Lopez- Fernandez, H. (2013). Description of a new, narrowly endemic south American darter (Characiformes: Crenuchidae) from the central Guiana shield highlands of Guyana. Copeia, 2013 (3), 454 - 463. https: // doi. org / 10.1093 / molbev / msy 096." title="Description of a new, narrowly endemic south American darter (Characiformes: Crenuchidae) from the central Guiana shield highlands of Guyana" type="journal article" year="2013">
Lujan 
<emphasis box="[975,1014,1702,1722]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2013
</bibRefCitation>
, p. 458; 
<bibRefCitation author="Zanata, A. M. &amp; Camelier, P." box="[1157,1389,1702,1722]" journalOrPublisher="Neotropical Ichthyology" pageId="5" pageNumber="6" pagination="487 - 498" part="13" refId="ref12522" refString="Zanata, A. M., &amp; Camelier, P. (2015). Two new species of Characidium Reinhardt (Characiformes: Crenuchidae) from northeastern Brazilian coastal drainages. Neotropical Ichthyology, 13 (3), 487 - 498. https: // doi. org / 10.1590 / 1982 - 0224 - 20140106." title="Two new species of Characidium Reinhardt (Characiformes: Crenuchidae) from northeastern Brazilian coastal drainages" type="journal article" year="2015">Zanata &amp; Camelier, 2015</bibRefCitation>
, p. 488).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph blockId="5.[821,1064,1802,1827]" box="[821,1064,1802,1827]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[821,858,1802,1827]" pageId="5" pageNumber="6">3.6</emphasis>
| 
<heading bold="true" box="[927,1064,1802,1827]" fontSize="10" level="3" pageId="5" pageNumber="6" reason="0">
<emphasis bold="true" box="[927,1064,1802,1827]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[821,1470,1875,1965]" lastBlockId="6.[119,769,142,855]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Morphometric data can be seen in 
<tableCitation box="[1154,1224,1875,1895]" captionStart="TABLE 1" captionStartId="5.[124,202,139,158]" captionTargetBox="[143,1293,186,993]" captionText="TABLE 1 Morphometric data from C. tatama n. sp. (n = 7)" httpUri="http://table.plazi.org/id/DF24662AFF92FFDFD575FF5B7E85FF70" pageId="5" pageNumber="6" tableUuid="DF24662AFF92FFDFD575FF5B7E85FF70">Table 1</tableCitation>
, the largest known specimen not exceeding 
<quantity box="[1022,1087,1910,1930]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="6" unit="mm" value="45.0">45 mm</quantity>
<emphasis box="[1099,1109,1910,1930]" italics="true" pageId="5" pageNumber="6">L</emphasis>
<subScript attach="left" box="[1109,1118,1919,1932]" fontSize="5" pageId="5" pageNumber="6">S</subScript>
Body slightly compressed laterally, strongly convex dorsal profile from snout to supraoccipital fontanelle, ascending slightly convex arc to dorsal-fin origin; descending through base of dorsal fin, and to first caudal-fin procurrent ray. Ventral profile concave from dentary to pelvic fin, vertex located from this point to caudal peduncle straight. Greatest body depth at, or just above, dorsal-fin origin. Snout strongly rounded tip, at level of ventral margin of orbit. Maxillary long and extending beyond anterior margin of orbit. Circular orbit larger than snout length, smaller than maxilla. Iris round.
</paragraph>
<paragraph blockId="6.[119,769,142,855]" pageId="6" pageNumber="7">Dorsal-fin rays ii8(1), 9(5), 10*(1); anal-fin rays ii6*(7); pectoral-fin rays ii6(1), 7*(2) ii8(4); pelvic-fin rays i7(6), 8*(1); principal caudal-fin rays in dorsal lobe 8(1), 9*(5), in ventral lobe 8*(7).</paragraph>
<paragraph blockId="6.[119,769,142,855]" pageId="6" pageNumber="7">Scales cycloid. Lateral line complete; pored lateral-line scales 32(2), 33*(5). Scales above lateral line 3(1), 4*(6), below 2*(7). Predorsal scales 11*(7). Circumpeduncular scales 10(1), 11*(4). Isthmus and area between pectoral fins without scales.</paragraph>
<paragraph blockId="6.[119,769,142,855]" pageId="6" pageNumber="7">
Mesethmoid bone separated from frontal, parietal fontanel absent, postcleitrum (PC) I absent, PC II and III present. Medial process of first rib with ventral concave curvature in hook shape decreasing in width to the tip of belly, dorsal upper process triangular with concave space between process and base to vertebral joint (
<figureCitation box="[544,633,766,786]" captionStart="FIGURE 8" captionStartId="7.[1107,1198,141,160]" captionTargetBox="[125,1057,139,619]" captionTargetId="figure-619@7.[124,1062,138,622]" captionText="FIGURE 8 Detail in the shape of the first rib (a)–(c) and swimbladder (b)–(d) in (a) and (b). C. tatama n. sp. (b, ventral view) IMCN 8924 (1C&amp;S) and (c) and (d) C. dule n. sp. (d, dorsal view) IMCN 8929 (1C&amp;S). The right lobule in (d) is damaged. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.4058149" httpUri="https://zenodo.org/record/4058149/files/figure.png" pageId="6" pageNumber="7">Figure 8a</figureCitation>
). Swimbladder with two chambers, anterior chamber bilobate and short, posterior chamber slim, its base wider than the tip (
<figureCitation box="[494,583,835,855]" captionStart="FIGURE 8" captionStartId="7.[1107,1198,141,160]" captionTargetBox="[125,1057,139,619]" captionTargetId="figure-619@7.[124,1062,138,622]" captionText="FIGURE 8 Detail in the shape of the first rib (a)–(c) and swimbladder (b)–(d) in (a) and (b). C. tatama n. sp. (b, ventral view) IMCN 8924 (1C&amp;S) and (c) and (d) C. dule n. sp. (d, dorsal view) IMCN 8929 (1C&amp;S). The right lobule in (d) is damaged. Scale bar = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.4058149" httpUri="https://zenodo.org/record/4058149/files/figure.png" pageId="6" pageNumber="7">Figure 8b</figureCitation>
).
</paragraph>
<paragraph blockId="6.[119,425,935,960]" box="[119,425,935,960]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[119,156,935,960]" pageId="6" pageNumber="7">3.7</emphasis>
| 
<heading bold="true" box="[226,425,935,960]" fontSize="10" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" box="[226,425,935,960]" pageId="6" pageNumber="7">Colour in alcohol</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[119,768,1009,1167]" lastBlockId="6.[816,1465,142,370]" pageId="6" pageNumber="7">
Dark colouration on head due to the high concentration of chromatophores divided longitudinally by a black lateral band that crosses the eye. Ventral region silver and divided by the lateral line. Body crossed by longitudinal band that ends in the caudal region with a dark bar on the base of the caudal fin, basicaudal spot in medial rays of the caudal fin. Dorsal region brown with the edges of the scales black, forming a pattern of chainrings, which extend beyond the longitudinal band. Pectoral, pelvic and anal fins hyaline; caudal fin in smaller specimens (&lt;
<quantity box="[1325,1389,211,231]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="28.0">28 mm</quantity>
<emphasis box="[1395,1405,211,231]" italics="true" pageId="6" pageNumber="7">L</emphasis>
<subScript attach="left" box="[1404,1412,220,233]" fontSize="5" pageId="6" pageNumber="7">S</subScript>
,) also hyaline, but larger specimens with scattered chromatophores not clustered or forming bands. Dorsal-fin base with dark black stripe. Chromatophores forming a band at the base of the dorsal fin. Unpigmented belly, except on caudal peduncle, which has a last dark transversal body band.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="biology_ecology">
<paragraph blockId="6.[816,1015,450,475]" box="[816,1015,450,475]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[816,853,450,475]" pageId="6" pageNumber="7">3.8</emphasis>
| 
<heading bold="true" box="[922,1015,450,475]" fontSize="10" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" box="[922,1015,450,475]" pageId="6" pageNumber="7">Ecology</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[816,1464,523,647]" pageId="6" pageNumber="7">
The Ingará River (
<typeStatus box="[990,1032,523,543]" pageId="6" pageNumber="7">type</typeStatus>
locality) has U-shaped banks (
<emphasis box="[1332,1381,523,543]" italics="true" pageId="6" pageNumber="7">sensu</emphasis>
<bibRefCitation author="Parsons, M. &amp; Thoms, M. &amp; Norris, R." journalOrPublisher="Canberra" pageId="6" pageNumber="7" refId="ref11940" refString="Parsons, M., Thoms, M. and Norris, R., 2002, Australian River Assessment System: Review of Physical River Assessment Methods - A Biological Perspective, Monitoring River Health Initiative Technical Report no 21, Commonwealth of Australia and University of Canberra, Canberra." title="Australian River Assessment System: Review of Physical River Assessment Methods - A Biological Perspective, Monitoring River Health Initiative Technical Report no 21, Commonwealth of Australia and University of Canberra" type="book" year="2002">
Parsons 
<emphasis box="[816,854,558,578]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
., 2002
</bibRefCitation>
) with abundant riparian coverage on both sides, and a predominance of a rocky substrate and some large ridges, which makes it a torrential river with few pools or backwaters (
<figureCitation box="[1253,1332,627,647]" captionStart="FIGURE 5" captionStartId="4.[816,907,1916,1935]" captionTargetBox="[818,1461,1393,1876]" captionTargetId="figure-414@4.[818,1461,1393,1876]" captionText="FIGURE 5 Typical habitat of C. dule, Ingará River, San José Del Palmar, Chocó, Colombia" figureDoi="http://doi.org/10.5281/zenodo.4058141" httpUri="https://zenodo.org/record/4058141/files/figure.png" pageId="6" pageNumber="7">Figure 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="distribution">
<paragraph blockId="6.[816,1063,727,752]" box="[816,1063,727,752]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[816,853,727,752]" pageId="6" pageNumber="7">3.9</emphasis>
| 
<heading bold="true" box="[922,1063,727,752]" fontSize="10" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" box="[922,1063,727,752]" pageId="6" pageNumber="7">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[816,1463,801,890]" pageId="6" pageNumber="7">
<taxonomicName authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[816,1001,801,821]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tatama">
<emphasis box="[816,1001,801,821]" italics="true" pageId="6" pageNumber="7">Characidium tatama</emphasis>
</taxonomicName>
is endemic to the 
<collectingRegion box="[1195,1255,801,821]" country="Panama" name="Embera-Wounaan" pageId="6" pageNumber="7">Chocó</collectingRegion>
biogeographic region and inhabits rivers tributaries of the upper San Juan and Calima river (Figure 1).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="etymology">
<paragraph blockId="6.[816,1064,970,995]" box="[816,1064,970,995]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[816,868,970,995]" pageId="6" pageNumber="7">3.10</emphasis>
| 
<heading bold="true" box="[938,1064,970,995]" fontSize="10" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" box="[938,1064,970,995]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[816,1463,1043,1133]" pageId="6" pageNumber="7">
A noun in apposition to the Tatamá National Natural Park created in 1987 and the place where most of the 
<typeStatus box="[1103,1142,1078,1098]" pageId="6" pageNumber="7">type</typeStatus>
specimens were collected. The word Tatamá in Embera-Chamí language means the grandfather of the rivers.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="conservation">
<paragraph blockId="6.[816,1237,1213,1238]" box="[816,1237,1213,1238]" pageId="6" pageNumber="7">
<emphasis bold="true" box="[816,868,1213,1238]" pageId="6" pageNumber="7">3.11</emphasis>
| 
<heading bold="true" box="[938,1237,1213,1238]" fontSize="10" level="3" pageId="6" pageNumber="7" reason="0">
<emphasis bold="true" box="[938,1237,1213,1238]" pageId="6" pageNumber="7">Conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[816,1464,1286,1341]" lastBlockId="7.[124,772,692,1029]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<taxonomicName authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[816,999,1286,1306]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tatama">
<emphasis box="[816,999,1286,1306]" italics="true" pageId="6" pageNumber="7">Characidium tatama</emphasis>
</taxonomicName>
is so far known from two localities in tributaries of the San Juan and Calima Rivers. The EOO estimated for the species is 5.419. 
<quantity box="[207,256,697,717]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="km" value="3.0">3 km</quantity>
<superScript attach="left" box="[256,265,692,705]" fontSize="5" pageId="7" pageNumber="8">2</superScript>
, and the AOO estimated is 
<quantity box="[545,603,697,717]" metricMagnitude="4" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="8" unit="km" value="74.0">74 km</quantity>
<superScript attach="left" box="[603,612,692,705]" fontSize="5" pageId="7" pageNumber="8">2</superScript>
(
<emphasis box="[627,676,697,717]" italics="true" pageId="7" pageNumber="8">sensu</emphasis>
<bibRefCitation author="Bachman, S. &amp; Moat, J. &amp; Hill, A. W. &amp; de la Torre, J. &amp; Scott, B." journalOrPublisher="ZooKeys" pageId="7" pageNumber="8" pagination="117 - 126" part="150" refId="ref10245" refString="Bachman, S., Moat, J., Hill, A. W., de la Torre, J., &amp; Scott, B. (2011). Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool. ZooKeys, 150, 117 - 126." title="Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool" type="journal article" year="2011">
Bachman 
<emphasis box="[124,165,731,751]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
., 2011
</bibRefCitation>
). Additional collecting efforts should be conducted in the region to better understand the geographic distribution of the species and the existent threats. From the points assessed here for the distribution of 
<taxonomicName authorityName="Agudelo‐Zamora &amp; Tavera &amp; Murillo &amp; Ortega‐Lara" authorityYear="2020" box="[216,305,835,855]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tatama">
<emphasis box="[216,305,835,855]" italics="true" pageId="7" pageNumber="8">C. tatama</emphasis>
</taxonomicName>
, the species is tentatively categorized as Vulnerable (VU) D2 under the geospatial aspect because the population is restricted to less than 
<quantity box="[331,390,905,925]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="km" value="20.0">20 km</quantity>
<superScript attach="left" box="[390,399,900,913]" fontSize="5" pageId="7" pageNumber="8">2</superScript>
and the number of localities is less than five (
<collectionCode box="[170,222,939,959]" pageId="7" pageNumber="8">IUCN</collectionCode>
Standards and Petitions Subcommittee, 2019), with accelerated deforestation inside Colombian national parks (
<bibRefCitation author="Clerici, N. &amp; Armenteras, D. &amp; Kareiva, P. &amp; Botero, R. &amp; Ramirez-Delgado, J. P. &amp; Forero-Medina, G." journalOrPublisher="Scientific Reports" pageId="7" pageNumber="8" pagination="1 - 10" part="10" refId="ref10585" refString="Clerici, N., Armenteras, D., Kareiva, P., Botero, R., Ramirez-Delgado, J. P., Forero-Medina, G., et al. (2020). Deforestation in Colombian protected areas increased duringpost-conflict periods. Scientific Reports, 10 (1), 1 - 10." title="Deforestation in Colombian protected areas increased duringpost-conflict periods" type="journal article" year="2020">
Clerici 
<emphasis box="[124,163,1009,1029]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
., 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>