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<mods:title id="417968B37D0F706F9A2B966298F597BE">Revision of the black fungus gnat species (Diptera: Sciaridae) described by W. A. Steffan from Micronesia</mods:title>
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<mods:namePart id="10F0320D6C18B7CDA2CCDC6DA0EB68DD">Broadley, Adam</mods:namePart>
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<mods:affiliation id="3270CD6062C44F6B097FC01F9525C083">Ellen Kauschke, Justus Liebig University Giessen, Institute of Zoology, Stephanstrasse 24, 35390 Giessen, Germany</mods:affiliation>
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<mods:number id="D382744A7BBAA8F467D223A141760E1D">4683</mods:number>
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<treatment id="03F2879E8158FFFBFF4DF96DAB9EF8BB" ID-DOI="http://doi.org/10.5281/zenodo.3797645" ID-GBIF-Taxon="163830093" ID-Zenodo-Dep="3797645" LSID="urn:lsid:plazi:treatment:03F2879E8158FFFBFF4DF96DAB9EF8BB" httpUri="http://treatment.plazi.org/id/03F2879E8158FFFBFF4DF96DAB9EF8BB" lastPageNumber="221" pageId="6" pageNumber="221">
<subSubSection id="C34165038158FFFBFF4DF96DAB42FB3E" box="[151,622,1129,1156]" pageId="6" pageNumber="221" type="nomenclature">
<paragraph id="8BE436888158FFFBFF4DF96DAB42FB3E" blockId="6.[151,622,1129,1191]" box="[151,622,1129,1156]" pageId="6" pageNumber="221">
<heading id="D0AC81E48158FFFBFF4DF96DAB42FB3E" bold="true" box="[151,622,1129,1156]" fontSize="11" level="1" pageId="6" pageNumber="221" reason="1">
<taxonomicName id="4C5B4D0B8158FFFBFF4DF96DAB42FB3E" ID-CoL="YRL3" authority="Steffan, 1969" authorityName="Steffan" authorityYear="1969" box="[151,622,1129,1156]" class="Insecta" family="Sciaridae" genus="Corynoptera" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="heterochela">
<emphasis id="B92FEA9A8158FFFBFF4DF96DAB42FB3E" bold="true" box="[151,622,1129,1156]" pageId="6" pageNumber="221">
<emphasis id="B92FEA9A8158FFFBFF4DF96DA8E8FB39" bold="true" box="[151,452,1129,1156]" italics="true" pageId="6" pageNumber="221">Corynoptera heterochela</emphasis>
<bibRefCitation id="EFCA4B798158FFFBFE11F96DAB42FB3E" author="Steffan, W. A." box="[459,622,1129,1155]" pageId="6" pageNumber="221" pagination="669 - 732" refId="ref10929" refString="Steffan, W. A. (1969) Insects of Micronesia. Diptera: Sciaridae. Insects of Micronesia, 12 (7), 669 - 732." type="journal article" year="1969">Steffan, 1969</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C34165038158FFFBFF4DF989A857FB1A" box="[151,379,1164,1191]" pageId="6" pageNumber="221" type="description">
<paragraph id="8BE436888158FFFBFF4DF989A857FB1A" blockId="6.[151,622,1129,1191]" box="[151,379,1164,1191]" pageId="6" pageNumber="221">
(
<figureCitation id="13602A0D8158FFFBFF7AF989A9CEFB1A" box="[160,226,1164,1191]" captionStart="FIGURE 5" captionStartId="7.[151,250,1663,1688]" captionTargetBox="[153,1181,391,1627]" captionTargetId="figure@7.[151,1197,373,1643]" captionTargetPageId="7" captionText="FIGURE 5 AB. Corynoptera heterochela Steffan, 1969. Holotype.A. Hypopygium; B. Flagellomeres 2-4 (medium turbid and with included air)." figureDoi="http://doi.org/10.5281/zenodo.3772717" httpUri="https://zenodo.org/record/3772717/files/figure.png" pageId="6" pageNumber="221">Fig. 5</figureCitation>
AB, 6 AE)
</paragraph>
</subSubSection>
<subSubSection id="C34165038158FFFBFF4DF9D6AA82FB54" box="[151,942,1232,1257]" pageId="6" pageNumber="221" type="reference_group">
<paragraph id="8BE436888158FFFBFF4DF9D6AA82FB54" blockId="6.[151,942,1232,1257]" box="[151,942,1232,1257]" pageId="6" pageNumber="221">
<treatmentCitationGroup id="AB4B11A68158FFFBFF4DF9D6AA82FB54" box="[151,942,1232,1257]" pageId="6" pageNumber="221">
<taxonomicName id="4C5B4D0B8158FFFBFF4DF9D6AB08FB54" ID-CoL="YRL3" authority="Steffan, 1969" authorityName="Steffan" authorityYear="1969" box="[151,548,1232,1257]" class="Insecta" family="Sciaridae" genus="Corynoptera" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="heterochela">
<emphasis id="B92FEA9A8158FFFBFF4DF9D6A8BAFB54" box="[151,406,1233,1257]" italics="true" pageId="6" pageNumber="221">Corynoptera heterochela</emphasis>
<bibRefCitation id="EFCA4B798158FFFBFE46F9D4AB08FB54" author="Steffan, W. A." box="[412,548,1232,1257]" pageId="6" pageNumber="221" pagination="669 - 732" refId="ref10929" refString="Steffan, W. A. (1969) Insects of Micronesia. Diptera: Sciaridae. Insects of Micronesia, 12 (7), 669 - 732." type="journal article" year="1969">Steffan, 1969</bibRefCitation>
</taxonomicName>
[
<treatmentCitation id="0AFA10998158FFFBFDEAF9D4ABD6FB54" author="Steffan, W. A." box="[560,762,1232,1257]" page="697" pageId="6" pageNumber="221" year="1969">
<bibRefCitation id="EFCA4B798158FFFBFDEAF9D4ABE5FB54" author="Steffan, W. A." box="[560,713,1232,1257]" pageId="6" pageNumber="221" pagination="669 - 732" refId="ref10929" refString="Steffan, W. A. (1969) Insects of Micronesia. Diptera: Sciaridae. Insects of Micronesia, 12 (7), 669 - 732." type="journal article" year="1969">Steffan (1969)</bibRefCitation>
: 697
</treatmentCitation>
699, fig. 11 aj].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34165038158FFFBFF4DF813A897FA58" pageId="6" pageNumber="221" type="materials_examined">
<paragraph id="8BE436888158FFFBFF4DF813AD23FAC7" blockId="6.[151,1437,1303,1798]" pageId="6" pageNumber="221">
<emphasis id="B92FEA9A8158FFFBFF4DF813A845FA8F" bold="true" box="[151,361,1303,1330]" pageId="6" pageNumber="221">Material studied:</emphasis>
<materialsCitation id="3B333CD58158FFFBFEA9F81CABE5FAEB" ID-GBIF-Occurrence="2609012095" collectingDate="1953-01-19" collectionCode="BPBM" collectorName="J. L. Gressitt" country="Micronesia, Caroline Islands" elevation="180" location="Mt Temwetemwensekir" pageId="6" pageNumber="221" specimenCode="BPBM 8224" specimenCount="1" typeStatus="holotype">
<emphasis id="B92FEA9A8158FFFBFEA9F81CA8CCFA8F" bold="true" box="[371,480,1304,1330]" pageId="6" pageNumber="221">
<typeStatus id="54E0882A8158FFFBFEA9F81CA8CCFA8F" box="[371,480,1304,1330]" pageId="6" pageNumber="221" type="holotype">Holotype</typeStatus>
</emphasis>
: Male.
<date id="FFE510488158FFFBFDE6F813AB84FA8F" box="[572,680,1303,1330]" pageId="6" pageNumber="221" value="1953-01-19">
<collectingDate id="EFA1E9A08158FFFBFDE6F813AB84FA8F" box="[572,680,1303,1330]" pageId="6" pageNumber="221" value="1953-01-19">19.i.1953</collectingDate>
</date>
,
<collectingCountry id="F34C76188158FFFBFD6CF813AD2DFA8C" box="[694,1025,1303,1329]" isEnumeration="true" pageId="6" pageNumber="221">Micronesia, Caroline Islands</collectingCountry>
, Ponape I.,
<location id="8E8460538158FFFBFB51F81CACB4FA8C" LSID="urn:lsid:plazi:treatment:03F2879E8158FFFBFF4DF96DAB9EF8BB:8E8460538158FFFBFB51F81CACB4FA8C" box="[1163,1432,1303,1330]" country="Micronesia, Caroline Islands" name="Mt Temwetemwensekir" pageId="6" pageNumber="221">Mt Temwetemwensekir</location>
,
<quantity id="4CA39B6D8158FFFBFF4DF838A9CDFAEB" box="[151,225,1340,1366]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="221" unit="m" value="180.0">
<elevation id="0076D1BB8158FFFBFF4DF838A9CDFAEB" box="[151,225,1340,1366]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="221" unit="m" value="180.0">180 m</elevation>
</quantity>
,
<collectingMethod id="521A4E9F8158FFFBFF34F83FA87BFAEB" box="[238,343,1339,1366]" pageId="6" pageNumber="221">light trap</collectingMethod>
, leg.
<collectorName id="26AE535E8158FFFBFE4EF838AB31FAE8" box="[404,541,1339,1366]" pageId="6" pageNumber="221">J.L. Gressitt</collectorName>
(
<specimenCode id="DBFD9EF38158FFFBFDF7F838ABEDFAEB" box="[557,705,1340,1366]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="221" type="Museum">BPBM 8224</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B333CD58158FFFBFD0EF838AC0BFAEB" ID-GBIF-Occurrence="2609012103" box="[724,1319,1339,1366]" collectingDate="1953-01-19" collectionCode="BPBM" collectorName="J. L. Gressitt" country="Micronesia, Caroline Islands" elevation="180" location="Mt Temwetemwensekir" pageId="6" pageNumber="221" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B92FEA9A8158FFFBFD0EF838AA60FAEB" bold="true" box="[724,844,1340,1366]" pageId="6" pageNumber="221">
<typeStatus id="54E0882A8158FFFBFD0EF838AA60FAEB" box="[724,844,1340,1366]" pageId="6" pageNumber="221" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D5DFD018158FFFBFC86F838AA9AFAE8" box="[860,950,1339,1366]" pageId="6" pageNumber="221" type="male">3 males</specimenCount>
, same data as holotype (
<collectionCode id="ED4AAE4D8158FFFBFB11F838AC33FAEB" box="[1227,1311,1340,1366]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="6" pageNumber="221" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B333CD58158FFFBFAE9F838AD20FAC7" ID-GBIF-Occurrence="2609012094" collectingDate="1953-01-12" collectionCode="PWMP" collectorName="J. L. Gressitt" country="Ponape Island" location="Ponape Island" pageId="6" pageNumber="221" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D5DFD018158FFFBFAE9F838ACAEFAE8" box="[1331,1410,1339,1366]" pageId="6" pageNumber="221" type="male">1 male</specimenCount>
,
<specimenCount id="9D5DFD018158FFFBFA54F838A9CAFAC4" pageId="6" pageNumber="221" type="female">1 female</specimenCount>
,
<date id="FFE510488158FFFBFF2BF85BA872FAC7" box="[241,350,1375,1402]" pageId="6" pageNumber="221" value="1953-01-12">
<collectingDate id="EFA1E9A08158FFFBFF2BF85BA872FAC7" box="[241,350,1375,1402]" pageId="6" pageNumber="221" value="1953-01-12">12.i.1953</collectingDate>
</date>
,
<collectingCountry id="F34C76188158FFFBFEB3F864AB26FAC4" box="[361,522,1375,1402]" name="Ponape Island" pageId="6" pageNumber="221">Ponape Island</collectingCountry>
,
<collectingMethod id="521A4E9F8158FFFBFDCFF85BAB51FAC7" box="[533,637,1375,1402]" pageId="6" pageNumber="221">light trap</collectingMethod>
, leg.
<collectorName id="26AE535E8158FFFBFD63F864AA6DFAC4" box="[697,833,1375,1402]" pageId="6" pageNumber="221">J.L. Gressitt</collectorName>
(male in
<collectionCode id="ED4AAE4D8158FFFBFC76F864AD28FAC7" box="[940,1028,1376,1402]" pageId="6" pageNumber="221">PWMP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE436888158FFFBFF1DF887A897FA58" blockId="6.[151,1437,1303,1798]" pageId="6" pageNumber="221">
Conservation status: Body and head separated, both strongly deformed by dehydration of the Hoyers medium, air inclusion and turbidities. The
<typeStatus id="54E0882A8158FFFBFDCDF8ACABAEFA7F" box="[535,642,1448,1474]" pageId="6" pageNumber="221" type="paratype">paratypes</typeStatus>
are not so strongly deformed and allow a better interpretation of the morphological characters.
</paragraph>
</subSubSection>
<subSubSection id="C34165038158FFFBFF1DF8EBAB3BF95C" pageId="6" pageNumber="221" type="discussion">
<paragraph id="8BE436888158FFFBFF1DF8EBAB3BF95C" blockId="6.[151,1437,1303,1798]" pageId="6" pageNumber="221">
<emphasis id="B92FEA9A8158FFFBFF1DF8EBA864F9B7" bold="true" box="[199,328,1519,1546]" pageId="6" pageNumber="221">Comments</emphasis>
. The morphological details are difficult to interpret. We agree with Steffan that the species belongs to the genus
<taxonomicName id="4C5B4D0B8158FFFBFEF6FB10A897F993" authorityName="Winnertz" authorityYear="1867" box="[300,443,1556,1582]" class="Insecta" family="Sciaridae" genus="Corynoptera" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9A8158FFFBFEF6FB10A897F993" box="[300,443,1556,1582]" italics="true" pageId="6" pageNumber="221">Corynoptera</emphasis>
</taxonomicName>
by one bristle on the basal segment of the 3-segmented palpus, rather long bristles on the scutum, the wing venation and the shape and armature of the gonostylus with 4 strong apical spines, two more dorsally located. The teeth of the claws are not so typical for the genus and are similar to
<taxonomicName id="4C5B4D0B8158FFFBFBA3FB58AC24F9CB" authorityName="Winnertz" authorityYear="1867" box="[1145,1288,1628,1654]" class="Insecta" family="Sciaridae" genus="Corynoptera" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEA9A8158FFFBFBA3FB58AC24F9CB" box="[1145,1288,1628,1654]" italics="true" pageId="6" pageNumber="221">Corynoptera</emphasis>
</taxonomicName>
species from
<collectingCountry id="F34C76188158FFFBFF4DFB84A801F924" box="[151,301,1663,1690]" name="New Zealand" pageId="6" pageNumber="221">New Zealand</collectingCountry>
(
<bibRefCitation id="EFCA4B798158FFFBFEE6FB7BAB70F927" author="Mohrig, W. &amp; Jaschhof, M." box="[316,604,1663,1690]" pageId="6" pageNumber="221" pagination="1 - 110" refId="ref9988" refString="Mohrig, W. &amp; Jaschhof, M. (1999) Sciarid flies (Diptera, Sciaridae) of New Zealand. Studia dipterologica, 7 (Supplement), 1 - 110." type="book chapter" year="1999">Mohrig &amp; Jaschhof 1999</bibRefCitation>
). It could belong to the
<taxonomicName id="4C5B4D0B8158FFFBFCB2FB84AAC1F927" authorityName="Tuomikoski" authorityYear="1960" box="[872,1005,1664,1690]" class="Insecta" family="Sciaridae" genus="Corynoptera" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B92FEA9A8158FFFBFCB2FB84AAC1F927" box="[872,1005,1664,1690]" italics="true" pageId="6" pageNumber="221">C. spinifera</emphasis>
</taxonomicName>
group according to
<bibRefCitation id="EFCA4B798158FFFBFB0AFB7BA9C9F903" author="Menzel, F. &amp; Mohrig, W." pageId="6" pageNumber="221" pagination="1 - 761" refId="ref9585" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica, 6 (Supplement), 1 - 761." type="journal article" year="2000">Menzel &amp; Mohrig (2000)</bibRefCitation>
. Nevertheless, the taxonomic position of this species is not certain and another interpretation, different to the one given here, might be possible.
</paragraph>
</subSubSection>
<subSubSection id="C34165038158FFFBFF1DFBEFAB9EF8BB" box="[199,690,1771,1798]" pageId="6" pageNumber="221" type="distribution">
<paragraph id="8BE436888158FFFBFF1DFBEFAB9EF8BB" blockId="6.[151,1437,1303,1798]" box="[199,690,1771,1798]" pageId="6" pageNumber="221">
<emphasis id="B92FEA9A8158FFFBFF1DFBEFA84DF8BB" bold="true" box="[199,353,1771,1798]" pageId="6" pageNumber="221">Distribution.</emphasis>
<collectingCountry id="F34C76188158FFFBFEBDFBEFAB32F8B8" box="[359,542,1771,1797]" name="Micronesia" pageId="6" pageNumber="221">Caroline Islands</collectingCountry>
(
<collectingRegion id="499FF86A8158FFFBFDF7FBEFABA7F8B8" box="[557,651,1771,1797]" country="Micronesia" name="Pohnpei" pageId="6" pageNumber="221">Pohnpei</collectingRegion>
I.).
</paragraph>
</subSubSection>
</treatment>
</document>