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<mods:title id="13C05BB35E5172B947C728C9089D3670">Distribution of haploniscids (Isopoda, Asellota, Haploniscidae) in Icelandic waters, with description of Haploniscus astraphes n. sp. from the Iceland basin and the Southeast Atlantic Ocean</mods:title>
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<paragraph id="8BD28208CF1A780CF0A6A7DA23DBFE65" blockId="20.[151,1017,465,491]" box="[151,1017,465,491]" pageId="20" pageNumber="321">
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Distribution of
<taxonomicName id="4C6DF98BCF1A780CF166A7DA222BFE65" ID-CoL="7NHRW" box="[343,521,465,491]" class="Malacostraca" family="Haploniscidae" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="family">Haploniscidae</taxonomicName>
at the Greenland-Iceland-Faeroe Ridge
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</heading>
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<paragraph id="8BD28208CF1A780CF0A6A41322C1FC89" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
Ten species of the
<taxonomicName id="4C6DF98BCF1A780CF158A41C222BFDBE" box="[361,521,535,560]" class="Malacostraca" family="Haploniscidae" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="family">Haploniscidae</taxonomicName>
were sampled during the BIOICE expeditions in the GIF area (
<tableCitation id="C6EFB7B3CF1A780CF4E7A41C250EFDBE" box="[1238,1324,535,560]" captionStart="TABLE 1" captionStartId="17.[151,242,1052,1075]" captionText="TABLE 1. List of haploniscid species occurring in the GIF area, the number of stations, number of individuals, minimum and maximum depths area (north or south of the Ridge), minimum and maximum temperature and water mass (es)." pageId="20" pageNumber="321">Table 1</tableCitation>
,
<figureCitation id="13569E8DCF1A780CF50BA41C2094FDDA" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="21.[151,250,1984,2006]" captionStartId-1="22.[151,250,1958,1980]" captionStartId-2="23.[151,250,1705,1727]" captionTargetBox-0="[398,1184,195,1955]" captionTargetBox-1="[400,1174,195,1913]" captionTargetBox-2="[285,1301,196,1683]" captionTargetId-0="figure@21.[396,1185,193,1957]" captionTargetId-1="figure@22.[398,1176,193,1915]" captionTargetId-2="figure@23.[282,1304,193,1684]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURE 10. Distribution of haploniscid species around Iceland (BIOICE localities): A, Haploniscus astraphes n. sp. (dot), H. ampliatus (triangle) and H. hamatus (square); B, H. aduncus; C, H. angustus." captionText-1="FIGURE 11. Distribution of haploniscid species around Iceland (BIOICE localities): A, H. bicuspis; B, H. foresti; C, H. spinifer." captionText-2="FIGURE 12. Distribution of haploniscid species around Iceland (BIOICE localities): A, Antennuloniscus simplex; B, Chauliodoniscus armadilloides." httpUri-0="https://zenodo.org/record/291356/files/figure.png" httpUri-1="https://zenodo.org/record/291357/files/figure.png" httpUri-2="https://zenodo.org/record/291358/files/figure.png" pageId="20" pageNumber="321">Figs 10 12</figureCitation>
), eight species of
<taxonomicName id="4C6DF98BCF1A780CF14EA4372229FDDA" box="[383,523,572,596]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9195E1ACF1A780CF14EA4372229FDDA" box="[383,523,572,596]" italics="true" pageId="20" pageNumber="321">Haploniscus</emphasis>
</taxonomicName>
and one species each of genera
<taxonomicName id="4C6DF98BCF1A780CF342A437240EFDDA" box="[883,1068,572,596]" class="Malacostraca" family="Haploniscidae" genus="Antennuloniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9195E1ACF1A780CF342A437240EFDDA" box="[883,1068,572,596]" italics="true" pageId="20" pageNumber="321">Antennuloniscus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF453A437253CFDDA" box="[1122,1310,572,596]" class="Malacostraca" family="Haploniscidae" genus="Chauliodoniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9195E1ACF1A780CF453A437253CFDDA" box="[1122,1310,572,596]" italics="true" pageId="20" pageNumber="321">Chauliodoniscus</emphasis>
</taxonomicName>
. By far the most frequent species was
<taxonomicName id="4C6DF98BCF1A780CF1E5A46B22E5FDF6" box="[468,711,608,632]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="B9195E1ACF1A780CF1E5A46B22E5FDF6" box="[468,711,608,632]" italics="true" pageId="20" pageNumber="321">Haploniscus bicuspis</emphasis>
</taxonomicName>
with 11,410 specimens sampled at 136 stations (
<tableCitation id="C6EFB7B3CF1A780CF53DA4542546FDF6" box="[1292,1380,607,632]" captionStart="TABLE 1" captionStartId="17.[151,242,1052,1075]" captionText="TABLE 1. List of haploniscid species occurring in the GIF area, the number of stations, number of individuals, minimum and maximum depths area (north or south of the Ridge), minimum and maximum temperature and water mass (es)." pageId="20" pageNumber="321">Table 1</tableCitation>
).
<taxonomicName id="4C6DF98BCF1A780CF5B0A46A20FCFD12" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B9195E1ACF1A780CF5B0A46A25B7FDF6" box="[1409,1429,609,632]" italics="true" pageId="20" pageNumber="321">H</emphasis>
.
<emphasis id="B9195E1ACF1A780CF0A6A48F20FCFD12" box="[151,222,644,668]" italics="true" pageId="20" pageNumber="321">foresti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF129A48F21ACFD12" box="[280,398,644,668]" class="Malacostraca" family="Haploniscidae" genus="Antennuloniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B9195E1ACF1A780CF129A48F21ACFD12" box="[280,398,644,668]" italics="true" pageId="20" pageNumber="321">A. simplex</emphasis>
</taxonomicName>
were also fairly frequent, occurring at 19 and 12 stations, with
<date id="FFD3A4C8CF1A780CF45CA48F253AFD13" box="[1133,1304,644,669]" pageId="20" pageNumber="321" value="1375" valueMax="1816">1375 and 1816</date>
specimens, respectively.
<taxonomicName id="4C6DF98BCF1A780CF11FA4A32201FD4E" box="[302,547,680,704]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="aduncus">
<emphasis id="B9195E1ACF1A780CF11FA4A32201FD4E" box="[302,547,680,704]" italics="true" pageId="20" pageNumber="321">Haploniscus aduncus</emphasis>
</taxonomicName>
occurred at ten stations, while
<taxonomicName id="4C6DF98BCF1A780CF3ABA4A324DFFD4E" box="[922,1277,680,704]" class="Malacostraca" family="Haploniscidae" genus="Chauliodoniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="armadilloides">
<emphasis id="B9195E1ACF1A780CF3ABA4A324DFFD4E" box="[922,1277,680,704]" italics="true" pageId="20" pageNumber="321">Chauliodoniscus armadilloides</emphasis>
</taxonomicName>
was reported from 7 stations. The remaining species were found at one to four stations only, i.e.
<taxonomicName id="4C6DF98BCF1A780CF45DA4C7254DFD6A" box="[1132,1391,716,740]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="astraphes">
<emphasis id="B9195E1ACF1A780CF45DA4C7254DFD6A" box="[1132,1391,716,740]" italics="true" pageId="20" pageNumber="321">Haploniscus astraphes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6DF98BCF1A780CF5B0A4C62127FC86" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="ampliatus">
<emphasis id="B9195E1ACF1A780CF5B0A4C62127FC86" italics="true" pageId="20" pageNumber="321">H. ampliatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6DF98BCF1A780CF122A4FA21BBFC86" box="[275,409,753,776]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B9195E1ACF1A780CF122A4FA21BBFC86" box="[275,409,753,776]" italics="true" pageId="20" pageNumber="321">H. angustus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6DF98BCF1A780CF196A4FA220AFC86" box="[423,552,752,776]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B9195E1ACF1A780CF196A4FA220AFC86" box="[423,552,752,776]" italics="true" pageId="20" pageNumber="321">H. hamatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C6DF98BCF1A780CF254A4FA22FEFC86" box="[613,732,752,776]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="spinifer">
<emphasis id="B9195E1ACF1A780CF254A4FA22FEFC86" box="[613,732,752,776]" italics="true" pageId="20" pageNumber="321">H. spinifer</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD28208CF1A780CF0F6A51F2342FCFB" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
The only species occurring north of the Greenland-Iceland-Faeroe Ridge were
<taxonomicName id="4C6DF98BCF1A780CF465A51F2567FCA2" box="[1108,1349,788,812]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="B9195E1ACF1A780CF465A51F2567FCA2" box="[1108,1349,788,812]" italics="true" pageId="20" pageNumber="321">Haploniscus bicuspis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF5B0A51E20D9FCDE" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B9195E1ACF1A780CF5B0A51E20D9FCDE" italics="true" pageId="20" pageNumber="321">H. angustus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6DF98BCF1A780CF13CA53321CCFCDE" box="[269,494,824,848]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="ingolfi">
<emphasis id="B9195E1ACF1A780CF13CA53321CCFCDE" box="[269,494,824,848]" italics="true" pageId="20" pageNumber="321">Haploniscus ingolfi</emphasis>
</taxonomicName>
is the only other haploniscid species previously reported from the Nordic Seas (
<bibRefCitation id="EFFCFFF9CF1A780CF0AFA5572100FCFB" author="Wolff" box="[158,290,860,885]" pageId="20" pageNumber="321" refString="Wolff, T. (1962) The systematics and biology of bathyal and abyssal Isopoda Asellota. Galathea Report, 6, 1 - 320." type="journal article" year="1962">Wolff 1962</bibRefCitation>
;
<bibRefCitation id="EFFCFFF9CF1A780CF11FA55721D2FCFA" author="Svavarsson" box="[302,496,860,884]" pageId="20" pageNumber="321" refString="Svavarsson, J. (1988) Bathyal and abyssal Asellota (Crustacea, Isopoda) from the Norwegian, Greenland, and North Polar Seas. Sarsia, 73, 83 - 106." type="journal article" year="1988">Svavarsson 1988</bibRefCitation>
), but was not found is this study.
</paragraph>
</subSubSection>
<subSubSection id="C377D183CF1A780CF0F6A58A2177FAC6" pageId="20" pageNumber="321" type="reference_group">
<paragraph id="8BD28208CF1A780CF0F6A58A2271FB1B" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
<materialsCitation id="3B058855CF1A780CF0F6A58A22E0FC6E" country="Iceland" elevation="840" location="Ridge" municipality="Ridge" pageId="20" pageNumber="321" specimenCount="1">
<taxonomicName id="4C6DF98BCF1A780CF0F6A58A2165FC16" ID-CoL="3JMYF" box="[199,327,896,920]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="B9195E1ACF1A780CF0F6A58A2165FC16" box="[199,327,896,920]" italics="true" pageId="20" pageNumber="321">H. bicuspis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF1B1A58A222AFC16" ID-CoL="6LCRT" box="[384,520,897,920]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B9195E1ACF1A780CF1B1A58A222AFC16" box="[384,520,897,920]" italics="true" pageId="20" pageNumber="321">H. angustus</emphasis>
</taxonomicName>
were the only haploniscid species occurring at depths shallower than the saddle depth at
<elevation id="0040653BCF1A780CF0C8A5AF2162FC32" box="[249,320,932,956]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="20" pageNumber="321" unit="m" value="840.0">
<quantity id="4C952FEDCF1A780CF0C8A5AF2162FC32" box="[249,320,932,956]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="20" pageNumber="321" unit="m" value="840.0">840 m</quantity>
</elevation>
on the
<location id="8EB2D4D3CF1A780CF1A4A5AF21FAFC33" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF1A4A5AF21FAFC33" box="[405,472,932,957]" country="Iceland" municipality="Ridge" name="Ridge" pageId="20" pageNumber="321">Ridge</location>
and seem to be able to cross the shallow sills of the Greenland-Iceland-Faeroe
<collectingMunicipality id="6BB61872CF1A780CF568A5AF25BEFC33" box="[1369,1436,932,957]" pageId="20" pageNumber="321">Ridge</collectingMunicipality>
and thus colonize the waters all around
<collectingCountry id="F37AC298CF1A780CF25DA5CC22E0FC6E" box="[620,706,967,992]" name="Iceland" pageId="20" pageNumber="321">Iceland</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B058855CF1A780CF2E1A5C32271FB1B" collectionCode="MNAW, ISOW, NSDW" collectorName="C. Accordingly &amp; Modified North Atlantic Water" country="Iceland" location="North Atlantic Water" municipality="Iceland" pageId="20" pageNumber="321" specimenCount="1">
<taxonomicName id="4C6DF98BCF1A780CF2E1A5C323E0FC6E" ID-CoL="3JMYF" box="[720,962,968,992]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="B9195E1ACF1A780CF2E1A5C323E0FC6E" box="[720,962,968,992]" italics="true" pageId="20" pageNumber="321">Haploniscus bicuspis</emphasis>
</taxonomicName>
has been reported to be adapted to cold conditions around 0°C (
<bibRefCitation id="EFFCFFF9CF1A780CF198A5E72261FB8B" author="Hansen" box="[425,579,1004,1029]" pageId="20" pageNumber="321" refString="Hansen, H. J. (1916) Crustacea Malacostraca. The order Isopoda. The Danish Ingolf Expedition, 3 (5), 1 - 262." type="journal article" year="1916">Hansen 1916</bibRefCitation>
;
<bibRefCitation id="EFFCFFF9CF1A780CF263A5E722F5FB8B" author="Wolff" box="[594,727,1004,1029]" pageId="20" pageNumber="321" refString="Wolff, T. (1962) The systematics and biology of bathyal and abyssal Isopoda Asellota. Galathea Report, 6, 1 - 320." type="journal article" year="1962">Wolff 1962</bibRefCitation>
), and was here reported in many instances from temperatures below zero (as low as -0.86°C), but even at temperatures as high as 7.11°
<collectorName id="2698E7DECF1A780CF3E3A21B245FFBA6" box="[978,1149,1039,1064]" pageId="20" pageNumber="321">C. Accordingly</collectorName>
, the species was found in a variety of water masses, such as the warm
<collectorName id="2698E7DECF1A780CF299A23F242CFBC2" box="[680,1038,1076,1101]" pageId="20" pageNumber="321">
Modified
<location id="8EB2D4D3CF1A780CF32AA23F242CFBC2" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF32AA23F242CFBC2" box="[795,1038,1076,1101]" country="Iceland" municipality="Iceland" name="North Atlantic Water" pageId="20" pageNumber="321">North Atlantic Water</location>
</collectorName>
(
<collectionCode id="ED7C1ACDCF1A780CF411A23F24A2FBC2" box="[1056,1152,1076,1100]" pageId="20" pageNumber="321">MNAW</collectionCode>
), the deep Labrador Sea Water (LSW) and the
<collectingCountry id="F37AC298CF1A780CF1A0A25C21C6FBFE" box="[401,484,1111,1136]" name="Iceland" pageId="20" pageNumber="321">Iceland</collectingCountry>
Sea Overflow Water (
<collectionCode id="ED7C1ACDCF1A780CF2D3A253230FFBFE" box="[738,813,1112,1136]" pageId="20" pageNumber="321">ISOW</collectionCode>
) south of
<collectingMunicipality id="6BB61872CF1A780CF391A25C23D7FBFE" box="[928,1013,1111,1136]" pageId="20" pageNumber="321">Iceland</collectingMunicipality>
, and even in the cold Norwegian Sea Deep Water (
<collectionCode id="ED7C1ACDCF1A780CF11BA27721A2FB1A" box="[298,384,1148,1172]" pageId="20" pageNumber="321">NSDW</collectionCode>
) in the deep north.
</materialsCitation>
</paragraph>
<paragraph id="8BD28208CF1A780CF0F6A2942177FAC6" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
The remaining species are confined to areas south of the Greenland-Iceland-Faeroe Ridge (
<figureCitation id="13569E8DCF1A780CF4D1A2942540FB36" box="[1248,1378,1183,1208]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="21.[151,250,1984,2006]" captionStartId-1="22.[151,250,1958,1980]" captionStartId-2="23.[151,250,1705,1727]" captionTargetBox-0="[398,1184,195,1955]" captionTargetBox-1="[400,1174,195,1913]" captionTargetBox-2="[285,1301,196,1683]" captionTargetId-0="figure@21.[396,1185,193,1957]" captionTargetId-1="figure@22.[398,1176,193,1915]" captionTargetId-2="figure@23.[282,1304,193,1684]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURE 10. Distribution of haploniscid species around Iceland (BIOICE localities): A, Haploniscus astraphes n. sp. (dot), H. ampliatus (triangle) and H. hamatus (square); B, H. aduncus; C, H. angustus." captionText-1="FIGURE 11. Distribution of haploniscid species around Iceland (BIOICE localities): A, H. bicuspis; B, H. foresti; C, H. spinifer." captionText-2="FIGURE 12. Distribution of haploniscid species around Iceland (BIOICE localities): A, Antennuloniscus simplex; B, Chauliodoniscus armadilloides." httpUri-0="https://zenodo.org/record/291356/files/figure.png" httpUri-1="https://zenodo.org/record/291357/files/figure.png" httpUri-2="https://zenodo.org/record/291358/files/figure.png" pageId="20" pageNumber="321">
Figs 10
<emphasis id="B9195E1ACF1A780CF506A2AA2567FB36" box="[1335,1349,1185,1208]" italics="true" pageId="20" pageNumber="321"></emphasis>
12
</figureCitation>
) and also to depths greater than
<quantity id="4C952FEDCF1A780CF1E2A2CF220FFB52" box="[467,557,1220,1244]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="321" unit="m" value="1200.0">1200 m</quantity>
, holding higher (&gt;2°C) bottom temperatures (
<tableCitation id="C6EFB7B3CF1A780CF471A2CF24B5FB52" box="[1088,1175,1220,1245]" captionStart="TABLE 1" captionStartId="17.[151,242,1052,1075]" captionText="TABLE 1. List of haploniscid species occurring in the GIF area, the number of stations, number of individuals, minimum and maximum depths area (north or south of the Ridge), minimum and maximum temperature and water mass (es)." pageId="20" pageNumber="321">Table 1</tableCitation>
).
<taxonomicName id="4C6DF98BCF1A780CF483A2CE2566FB52" ID-CoL="3JMYD" box="[1202,1348,1220,1244]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="astraphes">
<emphasis id="B9195E1ACF1A780CF483A2CE2566FB52" box="[1202,1348,1220,1244]" italics="true" pageId="20" pageNumber="321">H. astraphes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF5B0A2CE2127FA8E" ID-CoL="3JMY8" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="ampliatus">
<emphasis id="B9195E1ACF1A780CF5B0A2CE2127FA8E" italics="true" pageId="20" pageNumber="321">H. ampliatus</emphasis>
</taxonomicName>
occurred only in the LSW, while
<taxonomicName id="4C6DF98BCF1A780CF2BDA2E22332FA8E" ID-CoL="3JMZ4" box="[652,784,1256,1280]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B9195E1ACF1A780CF2BDA2E22332FA8E" box="[652,784,1256,1280]" italics="true" pageId="20" pageNumber="321">H. hamatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF37BA2E223E6FA8E" ID-CoL="3JN2G" box="[842,964,1256,1280]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="spinifer">
<emphasis id="B9195E1ACF1A780CF37BA2E223E6FA8E" box="[842,964,1256,1280]" italics="true" pageId="20" pageNumber="321">H. spinifer</emphasis>
</taxonomicName>
occurred only in the ISOW.
<taxonomicName id="4C6DF98BCF1A780CF521A2E320D6FAAA" ID-CoL="3JMY7" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="aduncus">
<emphasis id="B9195E1ACF1A780CF521A2E320D6FAAA" italics="true" pageId="20" pageNumber="321">Haploniscus aduncus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6DF98BCF1A780CF139A3062155FAAA" ID-CoL="6LD3T" box="[264,375,1292,1316]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B9195E1ACF1A780CF139A3062155FAAA" box="[264,375,1292,1316]" italics="true" pageId="20" pageNumber="321">H. foresti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6DF98BCF1A780CF1BDA3072285FAAA" ID-CoL="66ZC2" box="[396,679,1292,1316]" class="Malacostraca" family="Haploniscidae" genus="Antennuloniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B9195E1ACF1A780CF1BDA3072285FAAA" box="[396,679,1292,1316]" italics="true" pageId="20" pageNumber="321">Antennuloniscus simplex</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF98BCF1A780CF2DAA3072472FAAA" ID-CoL="TQP4" box="[747,1104,1292,1316]" class="Malacostraca" family="Haploniscidae" genus="Chauliodoniscus" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="armadilloides">
<emphasis id="B9195E1ACF1A780CF2DAA3072472FAAA" box="[747,1104,1292,1316]" italics="true" pageId="20" pageNumber="321">Chauliodoniscus armadilloides</emphasis>
</taxonomicName>
occurred, however, in both ISOW and LSW.
</paragraph>
</subSubSection>
<subSubSection id="C377D183CF1A780FF0F6A35F2303F801" lastPageId="23" lastPageNumber="324" pageId="20" pageNumber="321" type="description">
<paragraph id="8BD28208CF1A780CF0F6A35F2482F9CB" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
<materialsCitation id="3B058855CF1A780CF0F6A35F2482F9CB" country="Denmark" county="Faeroe Islands" elevation="620" location="The" municipality="The" pageId="20" pageNumber="321" specimenCount="1">
<location id="8EB2D4D3CF1A780CF0F6A35F20D1FAE3" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF0F6A35F20D1FAE3" box="[199,243,1364,1389]" country="Denmark" county="Faeroe Islands" municipality="The" name="The" pageId="20" pageNumber="321">The</location>
Greenland-Iceland-Faeroe
<location id="8EB2D4D3CF1A780CF20AA35F225CFAE3" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF20AA35F225CFAE3" box="[571,638,1364,1389]" country="Denmark" county="Faeroe Islands" municipality="The" name="Ridge" pageId="20" pageNumber="321">Ridge</location>
is presumably one of the most pronounced barriers of species distribution in the world oceans.
<location id="8EB2D4D3CF1A780CF211A37C2270FA1E" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF211A37C2270FA1E" box="[544,594,1399,1424]" country="Denmark" county="Faeroe Islands" municipality="The" name="This" pageId="20" pageNumber="321">This</location>
is the only extensive, shallow ridge crossing the
<location id="8EB2D4D3CF1A780CF497A37C2571FA1E" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF497A37C2571FA1E" box="[1190,1363,1399,1424]" country="Denmark" county="Faeroe Islands" municipality="The" name="Atlantic Ocean" pageId="20" pageNumber="321">Atlantic Ocean</location>
in an easterly-westerly direction, separating the
<location id="8EB2D4D3CF1A780CF249A3972323FA3A" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF249A3972323FA3A" box="[632,769,1436,1461]" country="Denmark" county="Faeroe Islands" municipality="The" name="Nordic Seas" pageId="20" pageNumber="321">Nordic Seas</location>
and the
<location id="8EB2D4D3CF1A780CF354A39723D9FA3A" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF354A39723D9FA3A" box="[869,1019,1436,1461]" country="Denmark" county="Faeroe Islands" municipality="The" name="Arctic Ocean" pageId="20" pageNumber="321">Arctic Ocean</location>
from the
<location id="8EB2D4D3CF1A780CF45CA3972533FA3B" LSID="urn:lsid:plazi:treatment:03C4331ECF1A780FF0A6A7DA2303F801:8EB2D4D3CF1A780CF45CA3972533FA3B" box="[1133,1297,1436,1461]" country="Denmark" county="Faeroe Islands" municipality="The" name="North Atlantic" pageId="20" pageNumber="321">North Atlantic</location>
proper.
<collectingMunicipality id="6BB61872CF1A780CF540A39725BFFA3B" box="[1393,1437,1436,1461]" pageId="20" pageNumber="321">The</collectingMunicipality>
deepest connections between these oceans are the
<collectingCountry id="F37AC298CF1A780CF2F7A3B4230CFA56" box="[710,814,1471,1496]" name="Denmark" pageId="20" pageNumber="321">Denmark</collectingCountry>
Strait (~
<elevation id="0040653BCF1A780CF3BCA3B423F5FA56" box="[909,983,1471,1496]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="20" pageNumber="321" unit="m" value="620.0">
<quantity id="4C952FEDCF1A780CF3BCA3B423F5FA56" box="[909,983,1471,1496]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="20" pageNumber="321" unit="m" value="620.0">620 m</quantity>
</elevation>
) in the Greenland-Iceland Ridge and the Faroe Bank Channel (~
<elevation id="0040653BCF1A780CF194A3EF21D1FA72" box="[421,499,1508,1532]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="20" pageNumber="321" unit="m" value="840.0">
<quantity id="4C952FEDCF1A780CF194A3EF21D1FA72" box="[421,499,1508,1532]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="20" pageNumber="321" unit="m" value="840.0">840 m</quantity>
</elevation>
) southwest of the
<collectingCounty id="62B3FA84CF1A780CF2FFA3EE2356FA73" box="[718,884,1508,1533]" pageId="20" pageNumber="321">Faeroe Islands</collectingCounty>
(
<bibRefCitation id="EFFCFFF9CF1A780CF3B7A3EF2490FA72" author="Hansen" box="[902,1202,1508,1533]" pageId="20" pageNumber="321" refString="Hansen, B. &amp; Osterhus, S. (2000) North Atlantic-Nordic Seas exchanges. Progress in Oceanography, 45, 109 - 208. https: // doi. org / 10.1016 / S 0079 - 6611 (99) 00052 - X" type="journal article" year="2000">Hansen &amp; Østerhus 2000</bibRefCitation>
). North of the ridge bottom temperatures are low, being nearly -0.9°C in the deepest parts. The cold overflow water passing the ridge mixes with warmer water masses resulting in higher bottom temperatures south of the ridge.
</materialsCitation>
</paragraph>
<paragraph id="8BD28208CF1A780CF0F6A05A234DF8EA" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
Most commonly benthic species living in the area have their distributional limits at the ridge, e.g. desmosomatid and nannoniscoid asellote isopods (
<bibRefCitation id="EFFCFFF9CF1A780CF2EDA07F2422F902" author="Brix" box="[732,1024,1652,1677]" pageId="20" pageNumber="321" refString="Brix, S. &amp; Svavarsson, J. (2010) Distribution and diversity of desmosomatid and nannoniscid isopods (Crustacea) on the Greenland-Iceland-Faeroe Ridge. Polar Biology, 33, 525 - 530." type="journal article" year="2010">Brix &amp; Svavarsson 2010</bibRefCitation>
); anthuridean isopods (
<bibRefCitation id="EFFCFFF9CF1A780CF53DA07F217BF93E" author="Negoescu" pageId="20" pageNumber="321" refString="Negoescu, I. &amp; Svavarsson, J. (1997) Anthurideans (Crustacea, Isopoda) from the North Atlantic and the Arctic Ocean. Sarsia, 82, 159 - 202." type="journal article" year="1997">Negoescu &amp; Svavarsson 1997</bibRefCitation>
), valviferan isopod (
<bibRefCitation id="EFFCFFF9CF1A780CF20EA09C23ABF93E" author="Stransky" box="[575,905,1687,1712]" pageId="20" pageNumber="321" refString="Stransky, B. &amp; Svavarsson, J. (2006) Astacilla boreaphilis sp. nov. (Crustacea: Isopoda: Valvifera) from shallow and deep North Atlantic waters. Zootaxa, 1259, 1 - 23." type="journal article" year="2006">Stransky &amp; Svavarsson 2006</bibRefCitation>
), amphipods (
<bibRefCitation id="EFFCFFF9CF1A780CF417A09C24D4F93E" author="Weisshappel" box="[1062,1270,1687,1712]" pageId="20" pageNumber="321" refString="Weisshappel, J. B. (2001) Distribution and diversity of the hyperbenthic amphipod family Calliopiidae in the different seas around the Greenland-Iceland-Faeroe-Ridge. Sarsia, 86, 143 - 151." type="journal article" year="2001">Weisshappel 2001</bibRefCitation>
) and pectinoid bivalve molluscs (
<bibRefCitation id="EFFCFFF9CF1A780CF156A0B7226FF95A" author="Dijkstra" box="[359,589,1724,1749]" pageId="20" pageNumber="321" refString="Dijkstra, H. H., Waren, A. &amp; Gudmundsson, G. (2009) Pectinoidea (Mollusca: Bivalvia) from Iceland. Marine Biology Research, 5, 207 - 243." type="journal article" year="2009">
Dijkstra
<emphasis id="B9195E1ACF1A780CF1FDA0B62222F95A" box="[460,512,1724,1748]" italics="true" pageId="20" pageNumber="321">et al</emphasis>
. 2009
</bibRefCitation>
). It is still poorly known how the ridge and the different associated water masses shape the distribution of the species in question; whether this is by the strong thermal gradient, by the presence of water masses of subzero temperatures being difficult to adapt to, or simply by the physical presence of the ridge itself.
<bibRefCitation id="EFFCFFF9CF1A780CF177A1232277F8CE" box="[326,597,1832,1856]" pageId="20" pageNumber="321" refString="Svavarsson, J., Stromberg, J. - O. &amp; Brattegard, T. (1993) The deep-sea asellote (Isopoda, Crustacea) fauna of the Northern Seas: species composition, distributional patterns and origin. Journal of Biogeography, 20, 537 - 555. https: // doi. org / 10.2307 / 2845725" type="journal article">
Svavarsson
<emphasis id="B9195E1ACF1A780CF1FDA12221DEF8CE" box="[460,508,1832,1856]" italics="true" pageId="20" pageNumber="321">et al</emphasis>
. (1993)
</bibRefCitation>
concluded that the diversity of the Arctic deep-water fauna was low due to the isolating effects of the ridge and the associated water masses.
</paragraph>
<paragraph id="8BD28208CF1A780CF0F6A1642125F87B" blockId="20.[151,1437,535,2037]" pageId="20" pageNumber="321">
Although haploniscid isopods are known throughout the world oceans, the Nordic Seas and the Arctic Ocean are poor of haploniscid species (
<bibRefCitation id="EFFCFFF9CF1A780CF237A19F232FF822" author="Svavarsson" box="[518,781,1940,1964]" pageId="20" pageNumber="321" refString="Svavarsson, J., Stromberg, J. - O. &amp; Brattegard, T. (1993) The deep-sea asellote (Isopoda, Crustacea) fauna of the Northern Seas: species composition, distributional patterns and origin. Journal of Biogeography, 20, 537 - 555. https: // doi. org / 10.2307 / 2845725" type="journal article" year="1993">
Svavarsson
<emphasis id="B9195E1ACF1A780CF2BEA19E22E0F822" box="[655,706,1940,1964]" italics="true" pageId="20" pageNumber="321">et al</emphasis>
. 1993
</bibRefCitation>
). Apparently, the physical presence of the ridge limits the distribution of most of the haploniscid species towards the north, due to the deep occurrence of most of the species seen here.
</paragraph>
<caption id="DF12D280CF1B780DF0A6A1CB2384F87B" httpUri="https://zenodo.org/record/291356/files/figure.png" pageId="21" pageNumber="322" targetBox="[398,1184,195,1955]" targetPageId="21">
<paragraph id="8BD28208CF1B780DF0A6A1CB2384F87B" blockId="21.[151,1437,1984,2037]" pageId="21" pageNumber="322">
<emphasis id="B9195E1ACF1B780DF0A6A1CB213DF85B" bold="true" box="[151,287,1984,2006]" pageId="21" pageNumber="322">FIGURE 10.</emphasis>
Distribution of haploniscid species around Iceland (BIOICE localities): A,
<taxonomicName id="4C6DF98BCF1B780DF42BA1CB2523F858" box="[1050,1281,1984,2006]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="astraphes" status="sp. nov.">
<emphasis id="B9195E1ACF1B780DF42BA1CB2523F858" box="[1050,1281,1984,2006]" italics="true" pageId="21" pageNumber="322">Haploniscus astraphes</emphasis>
</taxonomicName>
<emphasis id="B9195E1ACF1B780DF536A1CA251DF858" bold="true" box="[1287,1343,1985,2006]" pageId="21" pageNumber="322">
<taxonomicNameLabel id="A22AE361CF1B780DF536A1CA251DF858" box="[1287,1343,1985,2006]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(dot),
<taxonomicName id="4C6DF98BCF1B780DF5B2A1C920D9F87B" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="ampliatus">
<emphasis id="B9195E1ACF1B780DF5B2A1C920D9F87B" italics="true" pageId="21" pageNumber="322">H. ampliatus</emphasis>
</taxonomicName>
(triangle) and
<taxonomicName id="4C6DF98BCF1B780DF1A1A1EB2227F87B" box="[400,517,2015,2037]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B9195E1ACF1B780DF1A1A1EB2227F87B" box="[400,517,2015,2037]" italics="true" pageId="21" pageNumber="322">H. hamatus</emphasis>
</taxonomicName>
(square); B,
<taxonomicName id="4C6DF98BCF1B780DF2B8A1EB22DEF87B" box="[649,764,2015,2037]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="aduncus">
<emphasis id="B9195E1ACF1B780DF2B8A1EB22DEF87B" box="[649,764,2015,2037]" italics="true" pageId="21" pageNumber="322">H. aduncus</emphasis>
</taxonomicName>
; C,
<taxonomicName id="4C6DF98BCF1B780DF316A1EB2382F87B" box="[807,928,2016,2037]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B9195E1ACF1B780DF316A1EB2382F87B" box="[807,928,2016,2037]" italics="true" pageId="21" pageNumber="322">H. angustus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF12D280CF18780EF0A6A1AD20CBF854" httpUri="https://zenodo.org/record/291357/files/figure.png" pageId="22" pageNumber="323" targetBox="[400,1174,195,1913]" targetPageId="22">
<paragraph id="8BD28208CF18780EF0A6A1AD20CBF854" blockId="22.[151,1437,1958,2010]" pageId="22" pageNumber="323">
<emphasis id="B9195E1ACF18780EF0A6A1AD2100F835" bold="true" box="[151,290,1958,1980]" pageId="22" pageNumber="323">FIGURE 11.</emphasis>
Distribution of haploniscid species around Iceland (BIOICE localities): A,
<taxonomicName id="4C6DF98BCF18780EF477A1AC249EF832" box="[1094,1212,1958,1980]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="B9195E1ACF18780EF477A1AC249EF832" box="[1094,1212,1958,1980]" italics="true" pageId="22" pageNumber="323">H. bicuspis</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C6DF98BCF18780EF4DFA1AC2573F832" box="[1262,1361,1958,1980]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="B9195E1ACF18780EF4DFA1AC2573F832" box="[1262,1361,1958,1980]" italics="true" pageId="22" pageNumber="323">H. foresti</emphasis>
</taxonomicName>
; C,
<taxonomicName id="4C6DF98BCF18780EF5B2A1AC20C1F854" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="spinifer">
<emphasis id="B9195E1ACF18780EF5B2A1AC20C1F854" italics="true" pageId="22" pageNumber="323">H. spinifer</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF12D280CF19780FF0A6A0A221F8F950" httpUri="https://zenodo.org/record/291358/files/figure.png" pageId="23" pageNumber="324" targetBox="[285,1301,196,1683]" targetPageId="23">
<paragraph id="8BD28208CF19780FF0A6A0A221F8F950" blockId="23.[151,1435,1705,1758]" pageId="23" pageNumber="324">
<emphasis id="B9195E1ACF19780FF0A6A0A22105F930" bold="true" box="[151,295,1705,1727]" pageId="23" pageNumber="324">FIGURE 12.</emphasis>
Distribution of haploniscid species around Iceland (BIOICE localities): A,
<taxonomicName id="4C6DF98BCF19780FF45CA0A2254DF931" box="[1133,1391,1705,1727]" class="Malacostraca" family="Haploniscidae" genus="Antennuloniscus" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B9195E1ACF19780FF45CA0A2254DF931" box="[1133,1391,1705,1727]" italics="true" pageId="23" pageNumber="324">Antennuloniscus simplex</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C6DF98BCF19780FF0A6A0C321F1F950" box="[151,467,1736,1758]" class="Malacostraca" family="Haploniscidae" genus="Chauliodoniscus" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="armadilloides">
<emphasis id="B9195E1ACF19780FF0A6A0C321F1F950" box="[151,467,1736,1758]" italics="true" pageId="23" pageNumber="324">Chauliodoniscus armadilloides</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD28208CF19780FF0F6A1002303F801" blockId="23.[151,1437,1802,1935]" pageId="23" pageNumber="324">
Most of the haploniscids listed here have been found in the northernmost part of the Atlantic Ocean.
<taxonomicName id="4C6DF98BCF19780FF0A6A12521B8F8C8" box="[151,410,1838,1862]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="astraphes" status="sp. nov.">
<emphasis id="B9195E1ACF19780FF0A6A12521B8F8C8" box="[151,410,1838,1862]" italics="true" pageId="23" pageNumber="324">Haploniscus astraphes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22AE361CF19780FF195A12521C4F8C8" box="[420,486,1838,1863]" pageId="23" pageNumber="324" rank="species">
<emphasis id="B9195E1ACF19780FF195A12521C2F8C9" bold="true" box="[420,480,1838,1863]" pageId="23" pageNumber="324">n. sp</emphasis>
.
</taxonomicNameLabel>
, however, shows an extensive distribution, from the south side of the Greenland-
<collectingCountry id="F37AC298CF19780FF0A6A15920C8F8E5" box="[151,234,1874,1899]" name="Iceland" pageId="23" pageNumber="324">Iceland</collectingCountry>
Ridge to the
<collectingCountry id="F37AC298CF19780FF1B2A15921F6F8E5" box="[387,468,1874,1899]" name="Guinea-Bissau" pageId="23" pageNumber="324">Guinea</collectingCountry>
Basin, South Atlantic and the
<collectingCountry id="F37AC298CF19780FF31FA15923A3F8E5" box="[814,897,1874,1899]" name="Angola" pageId="23" pageNumber="324">Angola</collectingCountry>
Basin. Whether these really belong to the same species remains to be evaluated using molecular methods.
</paragraph>
</subSubSection>
</treatment>
</document>