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<document id="96E64E7DE8AAB151B40FA8396FA24CC9" ID-DOI="10.5281/zenodo.199324" ID-GBIF-Dataset="b2b6dac5-02be-402e-8af9-558020d958e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="199324" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460333904422" checkinUser="plazi" docAuthor="Winkelmann, Kai, Buckeridge, John S., Costa, Ana Cristina, Dionísio, Maria Ana Manso, Medeiros, André, Cachão, Mário &amp; Ávila, Sérgio P." docDate="2010" docId="03C1FA67FF89FFD4CCD4FC6D96B75935" docLanguage="en" docName="zt02680p044.pdf" docOrigin="Zootaxa 2680" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Zullobalanus santamariaensis Buckeridge &amp; Winkelmann, sp. nov." docType="treatment" docVersion="10" lastPageNumber="39" masterDocId="FFF8821FFF8AFFD2CC43FFE7955B5D49" masterDocTitle="Zullobalanus santamariaensis sp. nov., a new late Miocene barnacle species of the family Archaeobalanidae (Cirripedia: Thoracica), from the Azores" masterLastPageNumber="44" masterPageNumber="33" pageNumber="36" updateTime="1698259254858" updateUser="plazi">
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<mods:title id="2A37EDF59140003C3434FF7EB5DF7039">Zullobalanus santamariaensis sp. nov., a new late Miocene barnacle species of the family Archaeobalanidae (Cirripedia: Thoracica), from the Azores</mods:title>
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<mods:namePart id="8D478CF01858B110FD55792BE13B6777">Winkelmann, Kai</mods:namePart>
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<mods:namePart id="BE37E3391E14F90DD1861CE89438B4C2">Buckeridge, John S.</mods:namePart>
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<mods:namePart id="1AC06BD43A89FE8A07AAF46490204DE3">Costa, Ana Cristina</mods:namePart>
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<mods:namePart id="F09D6D57EC33AA83FEFEFC121F640A39">Dionísio, Maria Ana Manso</mods:namePart>
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<mods:namePart id="EEA17E00FF9D880247D996620DE5D63C">Medeiros, André</mods:namePart>
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<mods:namePart id="2AD378C4140FFC7DB15F7FEE8E40D002">Cachão, Mário</mods:namePart>
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<mods:namePart id="3D866360BD365938BC40C4AE8C13CCBC">Ávila, Sérgio P.</mods:namePart>
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<mods:date id="3FC8EA5C83FBCCC0B77B3C2EE22C35E1">2010</mods:date>
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<treatment id="03C1FA67FF89FFD4CCD4FC6D96B75935" ID-DOI="http://doi.org/10.5281/zenodo.5697432" ID-GBIF-Taxon="119404878" ID-Zenodo-Dep="5697432" LSID="urn:lsid:zoobank.org:act:B2C679D0-8F13-414F-8673-D5F64A94AB90" httpUri="http://treatment.plazi.org/id/03C1FA67FF89FFD4CCD4FC6D96B75935" lastPageId="6" lastPageNumber="39" pageId="3" pageNumber="36">
<subSubSection id="C37218FAFF89FFD1CCD4FC6D946D5E8E" pageId="3" pageNumber="36" type="nomenclature">
<paragraph id="8BD74B71FF89FFD1CCD4FC6D96935EED" blockId="3.[151,968,906,967]" box="[151,968,906,932]" pageId="3" pageNumber="36">
<heading id="D09FFC1DFF89FFD1CCD4FC6D96935EED" bold="true" box="[151,968,906,932]" fontSize="11" level="1" pageId="3" pageNumber="36" reason="1">
<emphasis id="B91C9763FF89FFD1CCD4FC6D96935EED" bold="true" box="[151,968,906,932]" pageId="3" pageNumber="36">
<taxonomicName id="4C6830F2FF89FFD1CCD4FC6D963F5EED" authority="Buckeridge &amp; Winkelmann" authorityName="Buckeridge &amp; Winkelmann" box="[151,868,906,932]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF89FFD1CCD4FC6D975F5EED" bold="true" box="[151,516,906,932]" italics="true" pageId="3" pageNumber="36">Zullobalanus santamariaensis</emphasis>
Buckeridge &amp; Winkelmann
</taxonomicName>
<taxonomicNameLabel id="A22F2A18FF89FFD1CF28FC6D96935EED" box="[875,968,906,932]" pageId="3" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD74B71FF89FFD1CCD4FC4A946D5E8E" blockId="3.[151,968,906,967]" box="[151,310,941,967]" pageId="3" pageNumber="36">
(
<figureCitation id="135357F4FF89FFD1CCDCFC4A94765E8E" box="[159,301,941,967]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,256,1813,1837]" captionStartId-1="5.[151,256,1868,1892]" captionTargetBox-0="[199,1424,468,1764]" captionTargetBox-1="[444,1152,194,1845]" captionTargetId-0="figure@4.[151,1436,418,1792]" captionTargetId-1="figure@5.[430,1152,194,1845]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal); samples DBUA-F (see. Table 1). I: Zullobalanus santamariaensis sp. nov. lateral and apical views. Letters at apical view; C = carina, CL = carinolatus, L = latus, R = rostrum. II: C - R; left interior, right exterior view. III: wall from below. Scale: I = 10 mm, II = 1 mm, III = 200 µm." captionText-1="FIGURE 3. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal), from samples DBUA-F (see Table 1); drawings based on SEM pictures. A: moveable scutum exterior (enlarged area showing detail from the lower part of the exterior of the scutum, accentuating fine apico-basal striae), B: moveable scutum interior, C: moveable tergum exterior, D: moveable tergum interior." httpUri-0="https://zenodo.org/record/199326/files/figure.png" httpUri-1="https://zenodo.org/record/199327/files/figure.png" pageId="3" pageNumber="36">Figures 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF89FFD1CCD4FC1091125915" pageId="3" pageNumber="36" type="diagnosis">
<paragraph id="8BD74B71FF89FFD1CCD4FC1091125915" blockId="3.[151,1437,1015,2012]" pageId="3" pageNumber="36">
<emphasis id="B91C9763FF89FFD1CCD4FC1094445958" bold="true" box="[151,287,1015,1041]" pageId="3" pageNumber="36">Diagnosis.</emphasis>
<emphasis id="B91C9763FF89FFD1CD6BFC1F94965958" box="[296,461,1016,1041]" italics="true" pageId="3" pageNumber="36">Zullobalanus</emphasis>
with shell lacking external apico-basal ribbing; tergum only weakly reflexed apically, but possessing extensive, well-formed crests for depressor muscle attachment; scutum with moderately produced articular ridge and moderate to weak adductor muscle scar.
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF89FFD1CC85FB8096F059CB" box="[198,939,1127,1154]" pageId="3" pageNumber="36" type="distribution">
<paragraph id="8BD74B71FF89FFD1CC85FB8096F059CB" blockId="3.[151,1437,1015,2012]" box="[198,939,1127,1154]" pageId="3" pageNumber="36">
<emphasis id="B91C9763FF89FFD1CC85FB80949559C8" bold="true" box="[198,462,1127,1153]" pageId="3" pageNumber="36">Distribution and age.</emphasis>
Azores; Late Miocene Early Pliocene.
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF89FFD1CC85FB6A906C5B55" pageId="3" pageNumber="36" type="materials_examined">
<paragraph id="8BD74B71FF89FFD1CC85FB6A906C5B55" blockId="3.[151,1437,1015,2012]" pageId="3" pageNumber="36">
<emphasis id="B91C9763FF89FFD1CC85FB6A942159EE" bold="true" box="[198,378,1165,1191]" pageId="3" pageNumber="36">
<typeStatus id="54D3F5D3FF89FFD1CC85FB6A945A59EE" box="[198,257,1165,1191]" pageId="3" pageNumber="36">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54D3F5D3FF89FFD1CDC0FB6A94AB59EE" box="[387,496,1165,1191]" pageId="3" pageNumber="36" type="holotype">Holotype</typeStatus>
containing 2 sample compositions: DBUA-F 536.1, complete shell (
<quantity id="4C90E694FF89FFD1C957FB6A902559EE" box="[1300,1406,1165,1191]" metricMagnitude="-3" metricUnit="m" metricValue="3.23" pageId="3" pageNumber="36" unit="mm" value="3.23">3.23 mm</quantity>
in height and
<quantity id="4C90E694FF89FFD1CD5DFB5594D25985" box="[286,393,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="3.39" pageId="3" pageNumber="36" unit="mm" value="3.39">3.39 mm</quantity>
in basal diameter), carina (
<quantity id="4C90E694FF89FFD1CE97FB55961B5985" box="[724,832,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="5.44" pageId="3" pageNumber="36" unit="mm" value="5.44">5.44 mm</quantity>
in height), carinolatus, (
<quantity id="4C90E694FF89FFD1C829FB55918C5985" box="[1130,1239,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="5.36" pageId="3" pageNumber="36" unit="mm" value="5.36">5.36 mm</quantity>
in height), latus (
<quantity id="4C90E694FF89FFD1CCDCFB3F945359BB" box="[159,264,1240,1266]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="3" pageNumber="36" unit="mm" value="1.84">1.84 mm</quantity>
in height), rostrum (
<quantity id="4C90E694FF89FFD1CDB9FB3F973F59BB" box="[506,612,1240,1266]" metricMagnitude="-3" metricUnit="m" metricValue="2.59" pageId="3" pageNumber="36" unit="mm" value="2.59">2.59 mm</quantity>
in height), scutum (
<quantity id="4C90E694FF89FFD1CF0CFB3F96E259BB" box="[847,953,1240,1266]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="36" unit="mm" value="2.5">2.50 mm</quantity>
in articular margin), tergum (
<quantity id="4C90E694FF89FFD1C957FB3F902659BB" box="[1300,1405,1240,1266]" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="3" pageNumber="36" unit="mm" value="3.44">3.44 mm</quantity>
in articular margin); DBUA-F 536.2, complete shell (
<quantity id="4C90E694FF89FFD1CEADFB1A9603585E" box="[750,856,1277,1303]" metricMagnitude="-3" metricUnit="m" metricValue="6.31" pageId="3" pageNumber="36" unit="mm" value="6.31">6.31 mm</quantity>
in height and
<quantity id="4C90E694FF89FFD1C841FB1A9130585E" box="[1026,1131,1277,1303]" metricMagnitude="-3" metricUnit="m" metricValue="6.63" pageId="3" pageNumber="36" unit="mm" value="6.63">6.63 mm</quantity>
in basal diameter), carina (
<quantity id="4C90E694FF89FFD1CCDCFAC594525875" box="[159,265,1314,1340]" metricMagnitude="-3" metricUnit="m" metricValue="4.89" pageId="3" pageNumber="36" unit="mm" value="4.89">4.89 mm</quantity>
in height), carinolatus, (
<quantity id="4C90E694FF89FFD1CE66FAC597D45875" box="[549,655,1314,1340]" metricMagnitude="-3" metricUnit="m" metricValue="4.79" pageId="3" pageNumber="36" unit="mm" value="4.79">4.79 mm</quantity>
in height), latus (
<quantity id="4C90E694FF89FFD1CF19FAC5969F5875" box="[858,964,1314,1340]" metricMagnitude="-3" metricUnit="m" metricValue="3.36" pageId="3" pageNumber="36" unit="mm" value="3.36">3.36 mm</quantity>
in height), rostrum (
<quantity id="4C90E694FF89FFD1C8F0FAC590465875" box="[1203,1309,1314,1340]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="36" unit="mm" value="2.3">2.30 mm</quantity>
in height), scutum (
<quantity id="4C90E694FF89FFD1CCB8FAAF943D582B" box="[251,358,1352,1378]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="3" pageNumber="36" unit="mm" value="2.43">2.43 mm</quantity>
in articular margin), tergum (
<quantity id="4C90E694FF89FFD1CE87FAAF9674582B" box="[708,815,1352,1378]" metricMagnitude="-3" metricUnit="m" metricValue="3.61" pageId="3" pageNumber="36" unit="mm" value="3.61">3.61 mm</quantity>
in articular margin); Santa Maria, Malbusca outcrop (
<geoCoordinate id="EE5C2DB6FF89FFD1CCDCFA8A946258CE" box="[159,313,1389,1415]" direction="west" orientation="longitude" pageId="3" pageNumber="36" precision="1" value="-25.068623">25°4'7.04&quot;W</geoCoordinate>
,
<geoCoordinate id="EE5C2DB6FF89FFD1CD06FA8A94A858CE" box="[325,499,1389,1415]" direction="north" orientation="latitude" pageId="3" pageNumber="36" precision="1" value="36.929314">36°55'45.53&quot;N</geoCoordinate>
[
<quantity id="4C90E694FF89FFD1CE41FA8A971A58CE" box="[514,577,1389,1415]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="36" unit="m" value="30.0">30 m</quantity>
asl]); S.P. Ávila &amp; K. Winkelmann,
<date id="FFD66DB1FF89FFD1CFB2FA8A91E658CE" box="[1009,1213,1389,1415]" pageId="3" pageNumber="36" value="2008-06-22" valueMax="2008-06-29" valueMin="2008-06-22">2229 June 2008</date>
.
<typeStatus id="54D3F5D3FF89FFD1C88AFA8A906058CE" box="[1225,1339,1389,1415]" pageId="3" pageNumber="36" type="paratype">Paratypes</typeStatus>
all with same locality, collectors and date as the
<typeStatus id="54D3F5D3FF89FFD1CE2CFA75978158E5" box="[623,730,1426,1452]" pageId="3" pageNumber="36" type="holotype">holotype</typeStatus>
: Natural History Museum Berlin: MB.A 1733, MB.A 1734; Natural History Museum Vienna:
<collectionCode id="ED79D3B4FF89FFD1CE60FA5F97D8589B" box="[547,643,1464,1490]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="36">NHMW</collectionCode>
2010/0088/0001,
<collectionCode id="ED79D3B4FF89FFD1CF1BFA5F96E3589B" box="[856,952,1464,1490]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="36">NHMW</collectionCode>
2010/0088/0002; Royal Belgian Institute of Natural Sciences Brussels:
<collectionCode id="ED79D3B4FF89FFD1CDB4FA3A971558BE" box="[503,590,1501,1527]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="3" pageNumber="36">IRSNB</collectionCode>
7255ag,
<collectionCode id="ED79D3B4FF89FFD1CE8AFA3A964458BE" box="[713,799,1501,1527]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="3" pageNumber="36">IRSNB</collectionCode>
7256ag; Natural History Museum London: NHM IC 549 NHM IC 562; Muséum National dHistoire Naturelle Paris:
<collectionCode id="ED79D3B4FF89FFD1CFE1F9E596A75B55" LSID="urn:lsid:biocol.org:col:34988" box="[930,1020,1538,1564]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="36">MNHN</collectionCode>
A32053,
<collectionCode id="ED79D3B4FF89FFD1C833F9E591915B55" LSID="urn:lsid:biocol.org:col:34988" box="[1136,1226,1538,1564]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="36">MNHN</collectionCode>
A32054.
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF89FFD4CC85F9C097DF5F4B" lastPageId="6" lastPageNumber="39" pageId="3" pageNumber="36" type="description">
<paragraph id="8BD74B71FF89FFD1CC85F9C094865A6B" blockId="3.[151,1437,1015,2012]" pageId="3" pageNumber="36">
<emphasis id="B91C9763FF89FFD1CC85F9C094045B08" bold="true" box="[198,351,1575,1601]" pageId="3" pageNumber="36">Description.</emphasis>
Shell conic, height approximating basal diameter; exterior rough, longitudinal ribbing weak to absent, lateral growth lines weak; orifice rhomboidal (
<figureCitation id="135357F4FF89FFD1CF77F9AA96215B2E" box="[820,890,1613,1639]" captionStart="FIGURE 2" captionStartId="4.[151,256,1813,1837]" captionTargetBox="[199,1424,468,1764]" captionTargetId="figure@4.[151,1436,418,1792]" captionTargetPageId="4" captionText="FIGURE 2. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal); samples DBUA-F (see. Table 1). I: Zullobalanus santamariaensis sp. nov. lateral and apical views. Letters at apical view; C = carina, CL = carinolatus, L = latus, R = rostrum. II: C - R; left interior, right exterior view. III: wall from below. Scale: I = 10 mm, II = 1 mm, III = 200 µm." httpUri="https://zenodo.org/record/199326/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
A, B); interior with strong ribbing, with minor bifurcation of some ribs near basis; rib cross-section showing simple arborescent interlaminate structures (
<figureCitation id="135357F4FF89FFD1C92EF99595FC5BFB" captionStart="FIGURE 2" captionStartId="4.[151,256,1813,1837]" captionTargetBox="[199,1424,468,1764]" captionTargetId="figure@4.[151,1436,418,1792]" captionTargetPageId="4" captionText="FIGURE 2. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal); samples DBUA-F (see. Table 1). I: Zullobalanus santamariaensis sp. nov. lateral and apical views. Letters at apical view; C = carina, CL = carinolatus, L = latus, R = rostrum. II: C - R; left interior, right exterior view. III: wall from below. Scale: I = 10 mm, II = 1 mm, III = 200 µm." httpUri="https://zenodo.org/record/199326/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
B, C); sheath smooth, well developed, pendant, occupying up to 50% of height of inner surface; basis firmly interlocking with parietal ribs; radii moderately narrow, with oblique summits, growth lines parallel to basis crossed by fine growth striae parallel to alar abutment (
<figureCitation id="135357F4FF89FFD1CF03F90596D05BB5" box="[832,907,1762,1788]" captionStart="FIGURE 2" captionStartId="4.[151,256,1813,1837]" captionTargetBox="[199,1424,468,1764]" captionTargetId="figure@4.[151,1436,418,1792]" captionTargetPageId="4" captionText="FIGURE 2. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal); samples DBUA-F (see. Table 1). I: Zullobalanus santamariaensis sp. nov. lateral and apical views. Letters at apical view; C = carina, CL = carinolatus, L = latus, R = rostrum. II: C - R; left interior, right exterior view. III: wall from below. Scale: I = 10 mm, II = 1 mm, III = 200 µm." httpUri="https://zenodo.org/record/199326/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
B); alae with marginal extensions, summits almost horizontal (
<figureCitation id="135357F4FF89FFD1CD30F8EF94E15A6B" box="[371,442,1800,1826]" captionStart="FIGURE 2" captionStartId="4.[151,256,1813,1837]" captionTargetBox="[199,1424,468,1764]" captionTargetId="figure@4.[151,1436,418,1792]" captionTargetPageId="4" captionText="FIGURE 2. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal); samples DBUA-F (see. Table 1). I: Zullobalanus santamariaensis sp. nov. lateral and apical views. Letters at apical view; C = carina, CL = carinolatus, L = latus, R = rostrum. II: C - R; left interior, right exterior view. III: wall from below. Scale: I = 10 mm, II = 1 mm, III = 200 µm." httpUri="https://zenodo.org/record/199326/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="8BD74B71FF89FFD6CC86F8CA96155C25" blockId="3.[151,1437,1015,2012]" lastBlockId="4.[151,1437,152,364]" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36">
Scutum (
<figureCitation id="135357F4FF89FFD1CD6DF8CA94285A0E" box="[302,371,1837,1863]" captionStart="FIGURE 3" captionStartId="5.[151,256,1868,1892]" captionTargetBox="[444,1152,194,1845]" captionTargetId="figure@5.[430,1152,194,1845]" captionTargetPageId="5" captionText="FIGURE 3. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal), from samples DBUA-F (see Table 1); drawings based on SEM pictures. A: moveable scutum exterior (enlarged area showing detail from the lower part of the exterior of the scutum, accentuating fine apico-basal striae), B: moveable scutum interior, C: moveable tergum exterior, D: moveable tergum interior." httpUri="https://zenodo.org/record/199327/files/figure.png" pageId="3" pageNumber="36">Fig. 3</figureCitation>
A, B) very weakly reflexed apically; exterior with strong transverse growth ribs crossed by weak apico-basal striae; interior with articular ridge moderately produced, slightly pendant basally; adductor ridge weak, broadly rounded; adductor muscle scar moderate to weak, pit for lateral depressor muscles well formed lacking any obvious muscle attachment crests. Tergum (
<figureCitation id="135357F4FF89FFD1CFD2F87A968C5AFE" box="[913,983,1949,1975]" captionStart="FIGURE 3" captionStartId="5.[151,256,1868,1892]" captionTargetBox="[444,1152,194,1845]" captionTargetId="figure@5.[430,1152,194,1845]" captionTargetPageId="5" captionText="FIGURE 3. Zullobalanus santamariaensis sp. nov. All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal), from samples DBUA-F (see Table 1); drawings based on SEM pictures. A: moveable scutum exterior (enlarged area showing detail from the lower part of the exterior of the scutum, accentuating fine apico-basal striae), B: moveable scutum interior, C: moveable tergum exterior, D: moveable tergum interior." httpUri="https://zenodo.org/record/199327/files/figure.png" pageId="3" pageNumber="36">Fig. 3</figureCitation>
C, D) moderately elongate, spur wellformed, removed from basi-scutal angle by own width and extending beyond same angle by approximately its own width, furrow well-formed distally, externally with well spaced growth lines, in some areas (especially basally), fine apico-basal striae present; interior with longitudinal furrow on articular margin wide, open, shallow, smooth except for weak growth striae; articular ridge well-formed, concave towards articular margin; crests for depressor muscles well developed basally, extending as a zone of raised papillae nearer the apex; articular ridge gently curved to extend slightly beyond a broadly concave basal margin; apex with approximately 34 strong transverse growth lines in adults.
</paragraph>
<caption id="DF171BF9FF8EFFD6CCD4F8F2912C5AC4" httpUri="https://zenodo.org/record/199326/files/figure.png" pageId="4" pageNumber="37" targetBox="[199,1424,468,1764]" targetPageId="4">
<paragraph id="8BD74B71FF8EFFD6CCD4F8F2912C5AC4" blockId="4.[151,1437,1813,1933]" pageId="4" pageNumber="37">
<emphasis id="B91C9763FF8EFFD6CCD4F8F294405A64" bold="true" box="[151,283,1813,1837]" pageId="4" pageNumber="37">FIGURE 2.</emphasis>
<taxonomicName id="4C6830F2FF8EFFD6CD67F8F197365A64" box="[292,621,1814,1837]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF8EFFD6CD67F8F197365A64" box="[292,621,1814,1837]" italics="true" pageId="4" pageNumber="37">Zullobalanus santamariaensis</emphasis>
</taxonomicName>
<emphasis id="B91C9763FF8EFFD6CE37F8F297965A64" bold="true" box="[628,717,1813,1837]" pageId="4" pageNumber="37">
<taxonomicNameLabel id="A22F2A18FF8EFFD6CE37F8F297965A64" box="[628,717,1813,1837]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal); samples DBUA-F (see. Table 1). I:
<taxonomicName id="4C6830F2FF8EFFD6C86FF8D1902F5A04" box="[1068,1396,1846,1869]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF8EFFD6C86FF8D1902F5A04" box="[1068,1396,1846,1869]" italics="true" pageId="4" pageNumber="37">Zullobalanus santamariaensis</emphasis>
</taxonomicName>
<emphasis id="B91C9763FF8EFFD6C93FF8D2959D5A24" bold="true" pageId="4" pageNumber="37">
<taxonomicNameLabel id="A22F2A18FF8EFFD6C93FF8D2959D5A24" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lateral and apical views. Letters at apical view; C = carina, CL = carinolatus, L = latus, R = rostrum. II: C - R; left interior, right exterior view. III: wall from below. Scale: I = 10 mm, II = 1 mm, III = 200 µm.
</paragraph>
</caption>
<caption id="DF171BF9FF8FFFD7CCD4F8AB95B05AB9" httpUri="https://zenodo.org/record/199327/files/figure.png" pageId="5" pageNumber="38" targetBox="[444,1152,194,1845]" targetPageId="5">
<paragraph id="8BD74B71FF8FFFD7CCD4F8AB95B05AB9" blockId="5.[151,1437,1868,2032]" pageId="5" pageNumber="38">
<emphasis id="B91C9763FF8FFFD7CCD4F8AB94475A2D" bold="true" box="[151,284,1868,1892]" pageId="5" pageNumber="38">FIGURE 3.</emphasis>
<taxonomicName id="4C6830F2FF8FFFD7CD67F8AA97365A2D" box="[292,621,1869,1892]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF8FFFD7CD67F8AA97365A2D" box="[292,621,1869,1892]" italics="true" pageId="5" pageNumber="38">Zullobalanus santamariaensis</emphasis>
</taxonomicName>
<emphasis id="B91C9763FF8FFFD7CE37F8AB97965A2D" bold="true" box="[628,717,1868,1892]" pageId="5" pageNumber="38">
<taxonomicNameLabel id="A22F2A18FF8FFFD7CE37F8AB97965A2D" box="[628,717,1868,1892]" pageId="5" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
All specimens from lower-Miocene / early-Pliocene of Malbusca outcrop (Santa Maria Island, Azores, Portugal), from samples DBUA-F (see Table 1); drawings based on SEM pictures. A: moveable scutum exterior (enlarged area showing detail from the lower part of the exterior of the scutum, accentuating fine apico-basal striae), B: moveable scutum interior, C: moveable tergum exterior, D: moveable tergum interior.
</paragraph>
</caption>
<paragraph id="8BD74B71FF8CFFD4CC85FF7097DF5F4B" blockId="6.[151,1437,151,1148]" pageId="6" pageNumber="39">
<emphasis id="B91C9763FF8CFFD4CC85FF7097285DF8" bold="true" box="[198,627,151,177]" pageId="6" pageNumber="39">Occurence at Santa Maria Island.</emphasis>
All the Late-Miocene and Early-Pliocene outcrops studied in Santa Maria contained the endemic barnacle
<taxonomicName id="4C6830F2FF8CFFD4CE22FF5A969F5D9F" box="[609,964,189,214]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF8CFFD4CE22FF5A969F5D9F" box="[609,964,189,214]" italics="true" pageId="6" pageNumber="39">Zullobalanus santamariaensis</emphasis>
</taxonomicName>
<emphasis id="B91C9763FF8CFFD4CF8FFF5A91715D9E" bold="true" box="[972,1066,189,215]" pageId="6" pageNumber="39">
<taxonomicNameLabel id="A22F2A18FF8CFFD4CF8FFF5A91715D9E" box="[972,1066,189,215]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Most specimens were found at Malbusca and at “Pedra-que-Pica” outcrops. The beds at “Pedra-que-Pica” contained some well preserved but rare barnacles with opercula cemented to diverse marine molluscs (e.g.
<taxonomicName id="4C6830F2FF8CFFD4CFB6FEEF95805C0E" authority="Brocchi, 1814" authorityName="Brocchi" authorityYear="1814" class="Bivalvia" family="Pectinidae" genus="Gigantopecten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreoida" pageId="6" pageNumber="39" phylum="Mollusca" rank="species" species="latissimus">
<emphasis id="B91C9763FF8CFFD4CFB6FEEF907F5C68" box="[1013,1316,264,289]" italics="true" pageId="6" pageNumber="39">Gigantopecten latissimus</emphasis>
(Brocchi, 1814)
</taxonomicName>
and
<taxonomicName id="4C6830F2FF8CFFD4CD57FECA97E05C0E" authority="Mayer, 1864" authorityName="Mayer" authorityYear="1864" box="[276,699,301,327]" class="Bivalvia" family="Ostreidae" genus="Lopha" kingdom="Animalia" order="Ostreoida" pageId="6" pageNumber="39" phylum="Mollusca" rank="species" species="plicatuloides">
<emphasis id="B91C9763FF8CFFD4CD57FECA94A55C0F" box="[276,510,301,326]" italics="true" pageId="6" pageNumber="39">Lopha plicatuloides</emphasis>
(Mayer, 1864))
</taxonomicName>
; the beds at Malbusca contained a number of loose opercula in a sandy matrix. At Cré outcrop, rare single walls as well as opercula were present. In the more lithified marine sediments of Figueiral and Ponta do Castelo, barnacle fragments were only observed on eroded surfaces. At Ponta dos Frades, a few barnacle fragments were found together with well preserved micro-molluscs on eroded bottom surfaces. Nevertheless, we detected fragments of
<taxonomicName id="4C6830F2FF8CFFD4CFE5FE2590485C92" box="[934,1299,450,475]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF8CFFD4CFE5FE2590485C92" box="[934,1299,450,475]" italics="true" pageId="6" pageNumber="39">Zullobalanus santamariaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F2A18FF8CFFD4C958FE2590275C95" box="[1307,1404,450,476]" pageId="6" pageNumber="39" rank="species">
<emphasis id="B91C9763FF8CFFD4C958FE2590275C95" bold="true" box="[1307,1404,450,476]" pageId="6" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
in every sampled Miocene/Pliocene outcrop.
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF8CFFD4CC85FDEA94745FAB" pageId="6" pageNumber="39" type="discussion">
<paragraph id="8BD74B71FF8CFFD4CC85FDEA94745FAB" blockId="6.[151,1437,151,1148]" pageId="6" pageNumber="39">
<emphasis id="B91C9763FF8CFFD4CC85FDEA94A15F6E" bold="true" box="[198,506,525,551]" pageId="6" pageNumber="39">Remarks and affinities.</emphasis>
There is an extensive archaeobalanid fauna known from western Europe (
<bibRefCitation id="EFF93680FF8CFFD4CCDCFDD594655F05" author="Carriol" box="[159,318,562,588]" pageId="6" pageNumber="39" refString="Carriol, R. - P. (2008) New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France). Zootaxa, 1675, 31 - 48." type="journal article" year="2008">Carriol, 2008</bibRefCitation>
) and the eastern seaboard of North
<collectingCountry id="F37F0BE1FF8CFFD4CE9CFDD5961F5F05" box="[735,836,562,588]" name="United States of America" pageId="6" pageNumber="39">America</collectingCountry>
(
<bibRefCitation id="EFF93680FF8CFFD4CF10FDD596825F05" author="Zullo" box="[851,985,562,588]" pageId="6" pageNumber="39" refString="Zullo, V. A. (1984) New genera and species of balanoid barnacles from the Oligocene and Miocene of North Carolina. Journal of Paleontology, 58, 1312 - 1338." type="journal article" year="1984">Zullo 1984</bibRefCitation>
;
<bibRefCitation id="EFF93680FF8CFFD4CFA6FDD5919A5F05" author="Zullo" box="[997,1217,562,588]" pageId="6" pageNumber="39" refString="Zullo, V. A. &amp; Kite, L. E. (1985) Barnacles of the Jacksonian (Upper Eocene) Griffiths Landing Member, Dry Branch Formation in South Carolina and Georgia. South Carolina Geology, 28, 1 - 32." type="journal article" year="1985">Zullo &amp; Kite 1985</bibRefCitation>
;
<bibRefCitation id="EFF93680FF8CFFD4C88EFDD5958C5F3B" author="Zullo" pageId="6" pageNumber="39" refString="Zullo, V. A. &amp; Perreault, R. T. (1989) Review of Actinobalanus Moroni (Cirripedia, Archaeobalanidae), with the description of new Miocene species from Florida and Belgium. Tulane Studies in Geology and Paleontology, 22, 1 - 12." type="journal article" year="1989">Zullo &amp; Perreault 1989</bibRefCitation>
;
<bibRefCitation id="EFF93680FF8CFFD4CCA7FDBF94B95F3B" author="Zullo" box="[228,482,600,626]" pageId="6" pageNumber="39" refString="Zullo, V. A. &amp; Portell, R. W. (1991) Balanoid barnacles from the early Miocene Parachucla and Penney Farms Formations, Northern Florida. Tulane Studies in Geology and Paleontology, 24, 79 - 86." type="journal article" year="1991">Zullo &amp; Portell 1991</bibRefCitation>
). Of these, species of
<taxonomicName id="4C6830F2FF8CFFD4CEB2FDBF96C45F38" box="[753,927,600,625]" class="Maxillopoda" family="Archaeobalanidae" genus="Actinobalanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B91C9763FF8CFFD4CEB2FDBF96C45F38" box="[753,927,600,625]" italics="true" pageId="6" pageNumber="39">Actinobalanus</emphasis>
</taxonomicName>
are distinguished from this taxon by their porous bases;
<taxonomicName id="4C6830F2FF8CFFD4CD7FFD9A94AD5FDF" box="[316,502,637,662]" class="Maxillopoda" family="Archaeobalanidae" genus="Hesperibalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B91C9763FF8CFFD4CD7FFD9A94AD5FDF" box="[316,502,637,662]" italics="true" pageId="6" pageNumber="39">Hesperibalanus</emphasis>
</taxonomicName>
, the terga of which lack the well-formed nodose papillae on the internal surface and the broad spur;
<taxonomicName id="4C6830F2FF8CFFD4CDC6FD4597755FF2" box="[389,558,674,699]" class="Maxillopoda" family="Archaeobalanidae" genus="Solidobalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B91C9763FF8CFFD4CDC6FD4597755FF2" box="[389,558,674,699]" italics="true" pageId="6" pageNumber="39">Solidobalanus</emphasis>
</taxonomicName>
by the externally smooth compartments; and
<emphasis id="B91C9763FF8CFFD4C811FD4591A05FF2" box="[1106,1275,674,699]" italics="true" pageId="6" pageNumber="39">Lophobalanus</emphasis>
by their very narrow radii.
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF8CFFD4CC86FD0A9426591E" pageId="6" pageNumber="39" type="reference_group">
<paragraph id="8BD74B71FF8CFFD4CC86FD0A96715E8B" blockId="6.[151,1437,151,1148]" pageId="6" pageNumber="39">
<taxonomicName id="4C6830F2FF8CFFD4CC86FD0A976A5E4F" box="[197,561,749,774]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="santamariaensis" status="sp. nov.">
<emphasis id="B91C9763FF8CFFD4CC86FD0A976A5E4F" box="[197,561,749,774]" italics="true" pageId="6" pageNumber="39">Zullobalanus santamariaensis</emphasis>
</taxonomicName>
<emphasis id="B91C9763FF8CFFD4CE79FD0A97C15E4E" bold="true" box="[570,666,749,775]" pageId="6" pageNumber="39">
<taxonomicNameLabel id="A22F2A18FF8CFFD4CE79FD0A97C15E4E" box="[570,666,749,775]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other
<emphasis id="B91C9763FF8CFFD4CFE8FD0A91115E4F" box="[939,1098,749,774]" italics="true" pageId="6" pageNumber="39">Zullobalanus</emphasis>
species by a lack of clearly formed external ribbing on the shell. Although there are some weak “ribs” these are somewhat obscured by the generally rough surface of the shell. The weakly reflexed scutal apex is close to that seen in the Australian species
<taxonomicName id="4C6830F2FF8CFFD4CCB0FCBA963D5E3E" authority="Buckeridge, 1983" authorityName="Buckeridge" authorityYear="1983" box="[243,870,861,887]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="australiae" subSpecies="victoriae">
<emphasis id="B91C9763FF8CFFD4CCB0FCBA97275E3F" box="[243,636,861,886]" italics="true" pageId="6" pageNumber="39">Zullobalanus australiae victoriae</emphasis>
(
<bibRefCitation id="EFF93680FF8CFFD4CEC8FCBA96055E3E" author="Buckeridge" box="[651,862,861,887]" pageId="6" pageNumber="39" refString="Buckeridge, J. S. (1983) The fossil barnacles (Cirripedia: Thoracica) of New Zealand and Australia. New Zealand Geological Survey Paleontological Bulletin, 50, 1 - 151 + 14 pls." type="journal article" year="1983">Buckeridge, 1983</bibRefCitation>
)
</taxonomicName>
and in most respects it is closest to this species. In addition to the lack of external ribbing, it differs from
<taxonomicName id="4C6830F2FF8CFFD4CF79FC65911B5ED2" box="[826,1088,898,923]" class="Maxillopoda" family="Archaeobalanidae" genus="Zullobalanus" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="australiae" subSpecies="victoriae">
<emphasis id="B91C9763FF8CFFD4CF79FC65911B5ED2" box="[826,1088,898,923]" italics="true" pageId="6" pageNumber="39">Z. australiae victoriae</emphasis>
</taxonomicName>
by the presence of numerous well-formed crests for depressor muscles on the tergum.
</paragraph>
<paragraph id="8BD74B71FF8CFFD4CC86FC2A9426591E" blockId="6.[151,1437,151,1148]" pageId="6" pageNumber="39">
The scutal articular ridge is often basally pendant in larger specimens of
<emphasis id="B91C9763FF8CFFD4C8C9FC2A906E5EAF" box="[1162,1333,973,998]" italics="true" pageId="6" pageNumber="39">Zullobalanus</emphasis>
species (
<bibRefCitation id="EFF93680FF8CFFD4CCDDFC1594375945" author="Buckeridge" box="[158,364,1010,1036]" pageId="6" pageNumber="39" refString="Buckeridge, J. S. (1983) The fossil barnacles (Cirripedia: Thoracica) of New Zealand and Australia. New Zealand Geological Survey Paleontological Bulletin, 50, 1 - 151 + 14 pls." type="journal article" year="1983">Buckeridge 1983</bibRefCitation>
), and although this has not been clearly seen in the material available, we should be mindful that all larger scuta are isolated and since shell disarticulation, have been abraded, as such wearing away delicate extensions.
</paragraph>
</subSubSection>
<subSubSection id="C37218FAFF8CFFD4CC85FB8596B75935" box="[198,1004,1122,1148]" pageId="6" pageNumber="39" type="etymology">
<paragraph id="8BD74B71FF8CFFD4CC85FB8596B75935" blockId="6.[151,1437,151,1148]" box="[198,1004,1122,1148]" pageId="6" pageNumber="39">
<emphasis id="B91C9763FF8CFFD4CC85FB8594085935" bold="true" box="[198,339,1122,1148]" pageId="6" pageNumber="39">Etymology.</emphasis>
Geographic: Island of Santa Maria (noun in apposition).
</paragraph>
</subSubSection>
</treatment>
</document>