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<document id="382C439BFAB56E4C3103FADCC6D3BF9C" ID-DOI="10.11646/zootaxa.3478.1.32" ID-GBIF-Dataset="f3e4ca1e-a40b-465a-a900-7a898e472cde" ID-ISSN="1175-5326" ID-Zenodo-Dep="282268" ID-ZooBank="FC92E481-4FF7-4DD8-B7C9-9F192F373D2E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460524755954" checkinUser="plazi" docAuthor="Aarvik, Leif, Karisch, Timm &amp; Marthinsen, Gunnhild" docDate="2012" docId="03C13418FFEDFF921AE7FBC5384CF869" docLanguage="en" docName="zt03478p372.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dracontogena bjornstadi Aarvik &amp; Karisch, new species" docType="treatment" docVersion="13" lastPageNumber="353" masterDocId="FFF84C60FFE5FF9A1A70FFA73C2AFFB5" masterDocTitle="Review of the Afrotropical genus Dracontogena Diakonoff, 1970 (Lepidoptera, Tortricidae) with descriptions of eight new species" masterLastPageNumber="372" masterPageNumber="345" pageNumber="353" updateTime="1698315014682" updateUser="plazi">
<mods:mods id="8B54E91E1DED063B0BBD4C8D308BE9EB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CC39492BD867515C45B761A73A91E4D0">Review of the Afrotropical genus Dracontogena Diakonoff, 1970 (Lepidoptera, Tortricidae) with descriptions of eight new species</mods:title>
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<mods:namePart id="0064C15123DB9262C1A0922522A44928">Aarvik, Leif</mods:namePart>
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<mods:namePart id="04606354011B92273EBC38ACE85BC84D">Karisch, Timm</mods:namePart>
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<mods:namePart id="71C0F60B51FAAF641E3E58B81B0DC80A">Marthinsen, Gunnhild</mods:namePart>
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<mods:typeOfResource id="7B2DAC1CCC79EAC323BDE9AAD47C7F90">text</mods:typeOfResource>
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<mods:title id="0FEDE40FB17D87903C978524B64AAA7B">Zootaxa</mods:title>
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<mods:part id="4971D9F18166CC12D2873EE5E17C32BF">
<mods:date id="90534173BF943F1EAFFBC7557D24122D">2012</mods:date>
<mods:detail id="7879356737760E4A97B9AAEBC0AC836D" type="pubDate">
<mods:number id="9B82F90DBBC7D17E946FE09BBE6870D1">2012-09-11</mods:number>
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<mods:number id="7AE397F8D7B6CDD9DCAF61E1A7F24036">3478</mods:number>
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<mods:start id="42A4301893043100E69F3F0BEA0EE712">345</mods:start>
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<mods:identifier id="AFADEE89340BC792AEBF163C1DE3E7B3" type="DOI">10.11646/zootaxa.3478.1.32</mods:identifier>
<mods:identifier id="565F12FB2151838DE72BE814D8BAD8DC" type="GBIF-Dataset">f3e4ca1e-a40b-465a-a900-7a898e472cde</mods:identifier>
<mods:identifier id="DC62833F5040F78D7326A2EBA41EDA41" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C13418FFEDFF921AE7FBC5384CF869" ID-DOI="http://doi.org/10.5281/zenodo.5697404" ID-GBIF-Taxon="119526157" ID-Zenodo-Dep="5697404" LSID="urn:lsid:plazi:treatment:03C13418FFEDFF921AE7FBC5384CF869" httpUri="http://treatment.plazi.org/id/03C13418FFEDFF921AE7FBC5384CF869" lastPageNumber="353" pageId="8" pageNumber="353">
<subSubSection id="C372D685FFEDFF921AE7FBC53F66FBC9" box="[151,844,1122,1148]" pageId="8" pageNumber="353" type="nomenclature">
<paragraph id="8BD7850EFFEDFF921AE7FBC53F66FBC9" blockId="8.[151,844,1122,1181]" box="[151,844,1122,1148]" pageId="8" pageNumber="353">
<heading id="D09F3262FFEDFF921AE7FBC53F66FBC9" bold="true" box="[151,844,1122,1148]" fontSize="11" level="1" pageId="8" pageNumber="353" reason="1">
<emphasis id="B91C591CFFEDFF921AE7FBC53F66FBC9" bold="true" box="[151,844,1122,1148]" pageId="8" pageNumber="353">
<taxonomicName id="4C68FE8DFFEDFF921AE7FBC53E85FBC9" ID-CoL="B3QV7" authority="Aarvik &amp; Karisch" authorityName="Aarvik &amp; Karisch" box="[151,687,1122,1148]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="bjornstadi" status="sp. nov.">
<emphasis id="B91C591CFFEDFF921AE7FBC53DEFFBC9" bold="true" box="[151,453,1122,1148]" italics="true" pageId="8" pageNumber="353">Dracontogena bjornstadi</emphasis>
Aarvik &amp; Karisch
</taxonomicName>
,
<taxonomicNameLabel id="A22FE467FFEDFF9218CCFBC53F66FBC9" box="[700,844,1122,1148]" pageId="8" pageNumber="353" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AE7FB223D5EFB28" box="[151,372,1157,1181]" pageId="8" pageNumber="353" type="description">
<paragraph id="8BD7850EFFEDFF921AE7FB223D5EFB28" blockId="8.[151,844,1122,1181]" box="[151,372,1157,1181]" pageId="8" pageNumber="353">
(
<figureCitation id="1353998BFFEDFF921AEFFB223CC7FB28" box="[159,237,1157,1181]" captionStart="FIGURES 2 – 9" captionStartId="5.[151,264,1917,1940]" captionTargetBox="[151,1418,226,1851]" captionTargetId="figure@5.[151,1436,193,1896]" captionTargetPageId="5" captionText="FIGURES 2 – 9. Adults of Dracontogena. 2 – 4. D. continentalis Karisch. males. 5. D. continentalis Karisch. female. 6. D. agassizi n. sp. male. 7. D. rubondoensis n. sp. male. 8. D. bjornstadi n. sp. male. 9. D. angolensis n. sp. male. Fig 3. Shows holotype of D. hoppei Karisch. Scale 10 mm." httpUri="https://zenodo.org/record/282270/files/figure.png" pageId="8" pageNumber="353">Figs. 8</figureCitation>
,
<figureCitation id="1353998BFFEDFF921A8AFB223D33FB28" box="[250,281,1157,1181]" captionStart="FIGURES 23 – 33" captionStartId="12.[151,264,1845,1868]" captionTargetBox="[164,1432,524,1797]" captionTargetId="figure@12.[151,1436,519,1823]" captionTargetPageId="12" captionText="FIGURES 23 – 33. Tergite 8 in males of Dracontogena. 23. D. continentalis Karisch (slide L. Aarvik 2735). 24. D. agassizi n. sp. (slide T. Karisch 2235). 25. D. rubondoensis n. sp. (slide L. Aarvik 2734). 26. D. bjornstadi n. sp. (slide L. Aarvik 2733). 27. D. angolensis n. sp. (slide BMNH 28995). 28. D. bernardi Karisch. (slide L. Aarvik 2808). 29. D. solii n. sp. (slide NHMO 2010). 30. D. kingstoni n sp. (slide L. Aarvik 2010.004). 31. D. gilligani n. sp. (slide BMNH 28993). 32. D. sundi n. sp. (slide L. Aarvik 2809). 33. D. deltozyga (Meyrick) (slide L. Aarvik 2007.010). Scale 1 mm." httpUri="https://zenodo.org/record/282273/files/figure.png" pageId="8" pageNumber="353">26</figureCitation>
,
<figureCitation id="1353998BFFEDFF921B54FB223D41FB28" box="[292,363,1157,1181]" captionStart="FIGURES 34 – 40" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[153,1414,197,1906]" captionTargetId="figure@14.[151,1436,175,1914]" captionTargetPageId="14" captionText="FIGURES 34 – 40. Male genitalia in Dracontogena. 34. D. continentalis Karisch (slide L. Aarvik 2735). 35. D. agassizi n. sp. (slide T. Karisch 2235). 36. D. agassizi n. sp. phallus (slide T. Karisch 2235). 37. D. rubondoensis n. sp. (slide L. Aarvik 2734). 38. D. angolensis n. sp. (slide BMNH 28995). 39. D. bjornstadi n. sp. (slide L. Aarvik 2733). 40. D. bjornstadi n. sp. phallus (slide L. Aarvik 2733). Scale figs. 34 – 35, 37 – 39: 1 mm., scale figs. 36, 40: 0.5 mm." httpUri="https://zenodo.org/record/282274/files/figure.png" pageId="8" pageNumber="353">39, 40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AE7FB6A3E9BFABC" pageId="8" pageNumber="353" type="materials_examined">
<paragraph id="8BD7850EFFEDFF921AE7FB6A3E9BFABC" blockId="8.[151,1437,1229,2012]" pageId="8" pageNumber="353">
<emphasis id="B91C591CFFEDFF921AE7FB6A3D63FB53" bold="true" box="[151,329,1229,1254]" pageId="8" pageNumber="353">
<typeStatus id="54D33BACFFEDFF921AE7FB6A3CF8FB50" box="[151,210,1229,1253]" pageId="8" pageNumber="353">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="3B008F53FFEDFF921B24FB6A3E87FABC" ID-GBIF-Occurrence="3712346305" collectingDate="1991-02-08" collectionCode="NHMO" collectorName="A. Bjornstad" country="Tanzania" elevation="1180" location="Minziro Forest" municipality="Bukoba District" pageId="8" pageNumber="353" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D33BACFFEDFF921B24FB6A3D95FB50" box="[340,447,1229,1253]" pageId="8" pageNumber="353" type="holotype">Holotype</typeStatus>
, 3,
<collectingCountry id="F37FC59EFFEDFF921B87FB693EA0FB53" box="[503,650,1230,1254]" name="Tanzania" pageId="8" pageNumber="353">TANZANIA</collectingCountry>
:
<collectingMunicipality id="6BB31F74FFEDFF9218E9FB6A3F7EFB50" box="[665,852,1229,1253]" pageId="8" pageNumber="353">Bukoba District</collectingMunicipality>
:
<location id="8EB7D3D5FFEDFF921912FB6A383BFB50" LSID="urn:lsid:plazi:treatment:03C13418FFEDFF921AE7FBC5384CF869:8EB7D3D5FFEDFF921912FB6A383BFB50" box="[866,1041,1229,1253]" country="Tanzania" municipality="Bukoba District" name="Minziro Forest" pageId="8" pageNumber="353">Minziro Forest</location>
,
<quantity id="4C9028EBFFEDFF921E50FB6A3854FB50" box="[1056,1150,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="8" pageNumber="353" unit="m" value="1180.0">
<elevation id="0045623DFFEDFF921E50FB6A3854FB50" box="[1056,1150,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="8" pageNumber="353" unit="m" value="1180.0">1180 m</elevation>
</quantity>
.,
<date id="FFD6A3CEFFEDFF921EE2FB6A38D2FB50" box="[1170,1272,1229,1253]" pageId="8" pageNumber="353" value="1991-02-08">
<collectingDate id="EF925A26FFEDFF921EE2FB6A38D2FB50" box="[1170,1272,1229,1253]" pageId="8" pageNumber="353" value="1991-02-08">8.ii.1991</collectingDate>
</date>
,
<collectorName id="269DE0D8FFEDFF921F78FB6939B6FB50" box="[1288,1436,1229,1254]" pageId="8" pageNumber="353">A. Bjørnstad</collectorName>
24226, genitalia slide L. Aarvik 2733 (
<collectionCode id="ED791DCBFFEDFF921839FB563E8FFABC" box="[585,677,1265,1289]" collectionName="NHMO" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="8" pageNumber="353">NHMO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AB7FAB23DBAF920" pageId="8" pageNumber="353" type="description">
<paragraph id="8BD7850EFFEDFF921AB7FAB23E4EF9B0" blockId="8.[151,1437,1229,2012]" pageId="8" pageNumber="353">
Description. Adult. Male (
<figureCitation id="1353998BFFEDFF921870FAB23E60FA98" box="[512,586,1301,1325]" captionStart="FIGURES 2 – 9" captionStartId="5.[151,264,1917,1940]" captionTargetBox="[151,1418,226,1851]" captionTargetId="figure@5.[151,1436,193,1896]" captionTargetPageId="5" captionText="FIGURES 2 – 9. Adults of Dracontogena. 2 – 4. D. continentalis Karisch. males. 5. D. continentalis Karisch. female. 6. D. agassizi n. sp. male. 7. D. rubondoensis n. sp. male. 8. D. bjornstadi n. sp. male. 9. D. angolensis n. sp. male. Fig 3. Shows holotype of D. hoppei Karisch. Scale 10 mm." httpUri="https://zenodo.org/record/282270/files/figure.png" pageId="8" pageNumber="353">Fig. 8</figureCitation>
). Head: Blackish brown. Antenna dark brown. Labial palpus 1.5 times diameter of eye, blackish brown, extreme tip white. Thorax: Dark grey, white patch in front, ochreous and white posterior scale tuft; tegulae white, ochreous posteriorly. Legs dark grey, tarsi with paler rings, hind tibia with grey scale tuft. Wingspan 24.0 mm. Forewing upperside brownish black, brown suffusion dominant at dorsum at tornus; connection of white dorsal maculae interrupted; outer white macula with two small dark dots on dorsum. Hindwing dark brownish grey; basal scales modified, narrow, black; termen strongly notched before anal corner; no fringes present along termen from apex to notch.
</paragraph>
<paragraph id="8BD7850EFFEDFF921AB7F9B63D65F9C4" blockId="8.[151,1437,1229,2012]" pageId="8" pageNumber="353">
Male genitalia (
<figureCitation id="1353998BFFEDFF921B06F9B63DD4F99C" box="[374,510,1553,1577]" captionStart="FIGURES 34 – 40" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[153,1414,197,1906]" captionTargetId="figure@14.[151,1436,175,1914]" captionTargetPageId="14" captionText="FIGURES 34 – 40. Male genitalia in Dracontogena. 34. D. continentalis Karisch (slide L. Aarvik 2735). 35. D. agassizi n. sp. (slide T. Karisch 2235). 36. D. agassizi n. sp. phallus (slide T. Karisch 2235). 37. D. rubondoensis n. sp. (slide L. Aarvik 2734). 38. D. angolensis n. sp. (slide BMNH 28995). 39. D. bjornstadi n. sp. (slide L. Aarvik 2733). 40. D. bjornstadi n. sp. phallus (slide L. Aarvik 2733). Scale figs. 34 – 35, 37 – 39: 1 mm., scale figs. 36, 40: 0.5 mm." httpUri="https://zenodo.org/record/282274/files/figure.png" pageId="8" pageNumber="353">Figs. 39, 40</figureCitation>
). Valva broad, with moderate scale patch; phallus (
<figureCitation id="1353998BFFEDFF921E44F9B638AFF99C" box="[1076,1157,1553,1577]" captionStart="FIGURES 34 – 40" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[153,1414,197,1906]" captionTargetId="figure@14.[151,1436,175,1914]" captionTargetPageId="14" captionText="FIGURES 34 – 40. Male genitalia in Dracontogena. 34. D. continentalis Karisch (slide L. Aarvik 2735). 35. D. agassizi n. sp. (slide T. Karisch 2235). 36. D. agassizi n. sp. phallus (slide T. Karisch 2235). 37. D. rubondoensis n. sp. (slide L. Aarvik 2734). 38. D. angolensis n. sp. (slide BMNH 28995). 39. D. bjornstadi n. sp. (slide L. Aarvik 2733). 40. D. bjornstadi n. sp. phallus (slide L. Aarvik 2733). Scale figs. 34 – 35, 37 – 39: 1 mm., scale figs. 36, 40: 0.5 mm." httpUri="https://zenodo.org/record/282274/files/figure.png" pageId="8" pageNumber="353">Fig. 40</figureCitation>
) long, 831 μm., broadest in middle, with ca. 112 small cornuti counted from their attachment sockets. Tergite 8 (
<figureCitation id="1353998BFFEDFF921E19F9923891F9F8" box="[1129,1211,1589,1613]" captionStart="FIGURES 23 – 33" captionStartId="12.[151,264,1845,1868]" captionTargetBox="[164,1432,524,1797]" captionTargetId="figure@12.[151,1436,519,1823]" captionTargetPageId="12" captionText="FIGURES 23 – 33. Tergite 8 in males of Dracontogena. 23. D. continentalis Karisch (slide L. Aarvik 2735). 24. D. agassizi n. sp. (slide T. Karisch 2235). 25. D. rubondoensis n. sp. (slide L. Aarvik 2734). 26. D. bjornstadi n. sp. (slide L. Aarvik 2733). 27. D. angolensis n. sp. (slide BMNH 28995). 28. D. bernardi Karisch. (slide L. Aarvik 2808). 29. D. solii n. sp. (slide NHMO 2010). 30. D. kingstoni n sp. (slide L. Aarvik 2010.004). 31. D. gilligani n. sp. (slide BMNH 28993). 32. D. sundi n. sp. (slide L. Aarvik 2809). 33. D. deltozyga (Meyrick) (slide L. Aarvik 2007.010). Scale 1 mm." httpUri="https://zenodo.org/record/282273/files/figure.png" pageId="8" pageNumber="353">Fig. 26</figureCitation>
) short, with distinct lateral incisions.
</paragraph>
<paragraph id="8BD7850EFFEDFF921AB7F9DA3DBAF920" blockId="8.[151,1437,1229,2012]" box="[199,400,1661,1685]" pageId="8" pageNumber="353">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AB7F9063EE9F89D" pageId="8" pageNumber="353" type="diagnosis">
<paragraph id="8BD7850EFFEDFF921AB7F9063EE9F89D" blockId="8.[151,1437,1229,2012]" pageId="8" pageNumber="353">
Diagnosis.
<taxonomicName id="4C68FE8DFFEDFF921B3CF9053DC2F90C" box="[332,488,1697,1721]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="bjornstadi" status="sp. nov.">
<emphasis id="B91C591CFFEDFF921B3CF9053DC2F90C" box="[332,488,1697,1721]" italics="true" pageId="8" pageNumber="353">D. bjornstadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22FE467FFEDFF921B87F9053E10F90C" box="[503,570,1698,1721]" pageId="8" pageNumber="353" rank="species">n. sp.</taxonomicNameLabel>
resembles the two other species with a strongly notched hindwing,
<taxonomicName id="4C68FE8DFFEDFF921FF1F9053D02F968" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="B91C591CFFEDFF921FF1F9053D02F968" italics="true" pageId="8" pageNumber="353">D. continentalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C68FE8DFFEDFF921B2FF9613DD0F968" box="[351,506,1733,1757]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="angolensis" status="sp. nov.">
<emphasis id="B91C591CFFEDFF921B2FF9613DD0F968" box="[351,506,1733,1757]" italics="true" pageId="8" pageNumber="353">D. angolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22FE467FFEDFF921872F9613E15F968" box="[514,575,1734,1757]" pageId="8" pageNumber="353" rank="species">n. sp.</taxonomicNameLabel>
In the genitalia it differs from
<taxonomicName id="4C68FE8DFFEDFF9219EEF9613878F968" box="[926,1106,1733,1757]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="B91C591CFFEDFF9219EEF9613878F968" box="[926,1106,1733,1757]" italics="true" pageId="8" pageNumber="353">D. continentalis</emphasis>
</taxonomicName>
by the broader valva and the much higher number of cornuti, as well as the differently shaped 8th tergite. From
<taxonomicName id="4C68FE8DFFEDFF921E36F94A38C8F8B1" box="[1094,1250,1772,1796]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis id="B91C591CFFEDFF921E36F94A38C8F8B1" box="[1094,1250,1772,1796]" italics="true" pageId="8" pageNumber="353">D. angolensis</emphasis>
</taxonomicName>
it differs by the more slender phallus and more numerous cornuti.
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AB7F8933FA9F8F9" box="[199,899,1844,1868]" pageId="8" pageNumber="353" type="distribution">
<paragraph id="8BD7850EFFEDFF921AB7F8933FA9F8F9" blockId="8.[151,1437,1229,2012]" box="[199,899,1844,1868]" pageId="8" pageNumber="353">
Distribution. Known only from Minziro Forest, NW
<collectingCountry id="F37FC59EFFEDFF921966F8933F54F8F9" box="[790,894,1844,1868]" name="Tanzania" pageId="8" pageNumber="353">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AB7F8FF3DC8F80D" pageId="8" pageNumber="353" type="biology_ecology">
<paragraph id="8BD7850EFFEDFF921AB7F8FF3DC8F80D" blockId="8.[151,1437,1229,2012]" pageId="8" pageNumber="353">
Biology. The habitat is a seasonally flooded swamp forest typical of the Kagera region of northwest
<collectingCountry id="F37FC59EFFEDFF921F41F8FF39B3F8C5" box="[1329,1433,1880,1904]" name="Tanzania" pageId="8" pageNumber="353">Tanzania</collectingCountry>
. The flora of this forest represents an eastern extension of the “Guineo-Congolian regional centre of endemism” into East Africa (
<bibRefCitation id="EFF9F8FFFFEDFF921B54F8073DFCF80D" author="Bjornstad, A." box="[292,470,1952,1976]" firstAuthor="Bjornstad" journalOrPublisher="Miombo" pageId="8" pageNumber="353" pagination="12 - 13" part="7" refId="ref14287" refString="Bjornstad, A. (1991) Kagera River Catchment - Threats. Miombo, 7, 12 - 13." title="Kagera River Catchment - Threats" type="journal article" year="1991">Bjørnstad 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEDFF921AE7F863384CF869" box="[151,1126,1988,2012]" pageId="8" pageNumber="353" type="etymology">
<paragraph id="8BD7850EFFEDFF921AE7F863384CF869" blockId="8.[151,1437,1229,2012]" box="[151,1126,1988,2012]" pageId="8" pageNumber="353">
Etymology. The species is named after the collector of the
<typeStatus id="54D33BACFFEDFF921959F8633FA4F869" box="[809,910,1988,2012]" pageId="8" pageNumber="353" type="holotype">holotype</typeStatus>
, Anders Bjørnstad.
</paragraph>
</subSubSection>
</treatment>
</document>
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