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<document id="A4AD981BD0A8A6A297DE96370F92B81F" ID-CLB-Dataset="30748" ID-DOI="10.5852/ejt.2018.403" ID-GBIF-Dataset="a5cecd60-c2bc-4ba3-a9f1-ecb12bc84b08" ID-Zenodo-Dep="1173235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518629854509" checkinUser="plazi" docAuthor="Zhu, Lin, Cao, Huiluo &amp; Chen, Hongwei" docDate="2018" docId="AA2687B9FFC2102F5B96684AFAC2FB6E" docLanguage="en" docName="ejt-403_zhu_cao_chen.pdf.imf" docOrigin="European Journal of Taxonomy 403" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phortica eparmata" docType="key" docVersion="13" lastPageNumber="15" masterDocId="561FFFC1FFCF10215B2A6C1CFFB4FF9E" masterDocTitle="Phortica eparmata species complex (Diptera, Drosophilidae) from the Oriental Region, with DNA barcoding information of Chinese species" masterLastPageNumber="18" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547547276763" originalUpdateUser="yanell" pageNumber="14" updateTime="1698521456743" updateUser="ExternalLinkService">
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<mods:title id="A457220D395879E8625DA066CB016313">Phortica eparmata species complex (Diptera, Drosophilidae) from the Oriental Region, with DNA barcoding information of Chinese species</mods:title>
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<mods:namePart id="A1120454DFA4451CC50ED2090D03F42A">Zhu, Lin</mods:namePart>
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<mods:namePart id="4D622278FDF5F6292ACB9F6AEDA9FC4E">Cao, Huiluo</mods:namePart>
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<treatment id="AA2687B9FFC2102F5B96684AFAC2FB6E" ID-DOI="http://doi.org/10.5281/zenodo.5619183" ID-GBIF-Taxon="140656047" ID-Zenodo-Dep="5619183" LSID="urn:lsid:plazi:treatment:AA2687B9FFC2102F5B96684AFAC2FB6E" httpUri="http://treatment.plazi.org/id/AA2687B9FFC2102F5B96684AFAC2FB6E" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<paragraph id="223036AFFFC2102C5B96684AFBB0FBEB" blockId="13.[188,1028,1109,1141]" box="[188,1028,1109,1141]" pageId="13" pageNumber="14">
<heading id="797881C3FFC2102C5B96684AFBB0FBEB" bold="true" box="[188,1028,1109,1141]" fontSize="13" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="10FBEABDFFC2102C5B96684AFBB0FBEB" bold="true" box="[188,1028,1109,1141]" pageId="13" pageNumber="14">
Key to all species of the
<taxonomicName id="E58F4D2CFFC2102C59356849FC80FBEB" ID-CoL="8DBVM" baseAuthorityName="Okada" baseAuthorityYear="1977" box="[543,820,1109,1141]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="eparmata">Phortica eparmata</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="223036AFFFC2102C5B966891FEC6FB39" blockId="13.[188,370,1165,1191]" box="[188,370,1165,1191]" pageId="13" pageNumber="14">
<heading id="797881C3FFC2102C5B966891FEC6FB39" bold="true" box="[188,370,1165,1191]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="10FBEABDFFC2102C5B966891FEC6FB39" bold="true" box="[188,370,1165,1191]" pageId="13" pageNumber="14">Adults (males)</emphasis>
</heading>
</paragraph>
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1. Anepisternum lacking setulae ……………………………other species of the subgenus
<taxonomicName id="E58F4D2CFFC2102C5E2768A1FAC1FB46" ID-CoL="9G2PY" authorityName="Schiner" authorityYear="1862" box="[1293,1397,1213,1240]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="10FBEABDFFC2102C5E2768A1FAC1FB46" bold="true" box="[1293,1397,1213,1240]" italics="true" pageId="13" pageNumber="14">Phortica</emphasis>
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<paragraph id="223036AFFFC2102C5BEA68F8FAC2FB63" blockId="13.[188,1398,1213,1278]" box="[192,1398,1251,1278]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5BEA68F8FAC2FB63" box="[192,1398,1251,1278]" pageId="13" pageNumber="14">
Anepisternum with setulae ………………………………………………
<emphasis id="10FBEABDFFC2102C5F1668F8FAA9FB63" bold="true" box="[1084,1309,1251,1278]" pageId="13" pageNumber="14">
<taxonomicName id="E58F4D2CFFC2102C5F1668F8FB1AFB60" ID-CoL="8DBVM" baseAuthorityName="Okada" baseAuthorityYear="1977" box="[1084,1198,1252,1278]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="15" phylum="Arthropoda" rank="species" species="eparmata">
<emphasis id="10FBEABDFFC2102C5F1668F8FB1AFB60" bold="true" box="[1084,1198,1252,1278]" italics="true" pageId="13" pageNumber="14">eparmata</emphasis>
</taxonomicName>
complex
</emphasis>
…… 2
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<keyStep id="997B2E0AFFC2102C5B96693BFAC2FA48" pageId="13" pageNumber="14">
<paragraph id="223036AFFFC2102C5B96693BFAC2FA12" blockId="13.[188,1398,1319,1495]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5B96693BFAC2FA12" pageId="13" pageNumber="14">
2. Aedeagal median rod not expanded subapically (as in
<bibRefCitation id="461E4B5EFFC2102C5840693BFB85FADF" box="[874,1073,1319,1345]" pageId="13" pageNumber="14" refId="ref7704" refString="Chen H. W., Toda M. J., Lakim M. B. &amp; Mohamed M. B. 2007. The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology 55: 23 - 41." type="journal article">
Chen
<emphasis id="10FBEABDFFC2102C589A6934FC5FFADF" box="[944,1003,1319,1345]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2007
</bibRefCitation>
, fig. 29); anterior branch of aedeagal basal bridge with ca. 10 acute projections along margin (as in
<bibRefCitation id="461E4B5EFFC2102C5F6D6950FAA6FAF8" box="[1095,1298,1356,1382]" pageId="13" pageNumber="14" refId="ref7704" refString="Chen H. W., Toda M. J., Lakim M. B. &amp; Mohamed M. B. 2007. The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology 55: 23 - 41." type="journal article">
Chen
<emphasis id="10FBEABDFFC2102C5FA46951FB7FFAF8" box="[1166,1227,1356,1382]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2007
</bibRefCitation>
, fig. 29) ……………………………………………………………………
<taxonomicName id="E58F4D2CFFC2102C58CE696EFAC2FA12" ID-CoL="8DBW2" authority="Chen &amp; Toda, 2007" authorityName="Chen &amp; Toda" authorityYear="2007" box="[996,1398,1393,1420]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lanuginosa">
<emphasis id="10FBEABDFFC2102C58CE696EFB3FFA12" bold="true" box="[996,1163,1393,1420]" pageId="13" pageNumber="14">
<emphasis id="10FBEABDFFC2102C58CE696EFC42FA12" bold="true" box="[996,1014,1394,1420]" italics="true" pageId="13" pageNumber="14">P</emphasis>
.
<emphasis id="10FBEABDFFC2102C5F2E696DFB3FFA12" bold="true" box="[1028,1163,1393,1420]" italics="true" pageId="13" pageNumber="14">lanuginosa</emphasis>
</emphasis>
<bibRefCitation id="461E4B5EFFC2102C5FB8696DFAC2FA12" box="[1170,1398,1393,1420]" pageId="13" pageNumber="14" refId="ref7704" refString="Chen H. W., Toda M. J., Lakim M. B. &amp; Mohamed M. B. 2007. The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology 55: 23 - 41." type="journal article">Chen &amp; Toda, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="223036AFFFC2102C5BEA698BFAC2FA48" blockId="13.[188,1398,1319,1495]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5BEA698BFAC2FA48" pageId="13" pageNumber="14">
Aedeagal median rod expanded subapically (
<figureCitation id="BAB42A2AFFC2102C582B698BFC35FA2F" box="[769,897,1431,1457]" captionStart="Fig. 2" captionStartId="7.[189,232,1932,1958]" captionTargetBox="[210,1372,992,1836]" captionTargetId="figure@7.[187,1401,956,1873]" captionTargetPageId="7" captionText="Fig. 2. Phortica (Phortica) jadete sp. nov., ♂. A. Arista. B. Epandrium and cercus (lateral view). C. Surstylus (frontal view).D. Hypandrium, paramere, aedeagus and aedeagalapodeme (lateral views). E. Paramere (frontal view). F. Tip of aedeagal median rod (ventral view). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173239/files/figure.png" pageId="13" pageNumber="14">Fig. 2D, F</figureCitation>
); anterior branch of aedeagal basal bridge lacking acute projections ……………………………………………………………………………3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC2102C5B966A1CFAC2F9DE" pageId="13" pageNumber="14">
<paragraph id="223036AFFFC2102C5B966A1CFAC2F984" blockId="13.[188,1398,1536,1600]" box="[188,1398,1536,1562]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5B966A1CFAC2F984" box="[188,1398,1536,1562]" pageId="13" pageNumber="14">3. Arista lacking ventral branch ………………………………………………………………………4</keyLead>
</paragraph>
<paragraph id="223036AFFFC2102C5BEA6A3AFAC2F9DE" blockId="13.[188,1398,1536,1600]" box="[192,1398,1573,1600]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5BEA6A3AFAC2F9DE" box="[192,1398,1573,1600]" pageId="13" pageNumber="14"> Arista with ventral branch(es) ………………………………………………………………………5</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC2102C5B966A75FAC2F96A" pageId="13" pageNumber="14">
<paragraph id="223036AFFFC2102C5B966A75FAC2F937" blockId="13.[188,1398,1641,1780]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5B966A75FAC2F937" pageId="13" pageNumber="14">
4. Paramere submedially pubescent and expanded to two strong branches (as in
<bibRefCitation id="461E4B5EFFC2102C5F8B6A75FAC6F91A" box="[1185,1394,1641,1668]" pageId="13" pageNumber="14" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">
Chen
<emphasis id="10FBEABDFFC2102C5FC06A76FA9DF91D" box="[1258,1321,1641,1667]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2005
</bibRefCitation>
, fig. 7C) ……………………………………………………………
<taxonomicName id="E58F4D2CFFC2102C58DF6A93FAC2F937" ID-CoL="77D82" authority="Chen &amp; Gao, 2005" authorityName="Chen &amp; Gao" authorityYear="2005" box="[1013,1398,1678,1705]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="1" phylum="Arthropoda" rank="species" species="latipenis">
<emphasis id="10FBEABDFFC2102C58DF6A93FB35F937" bold="true" box="[1013,1153,1678,1705]" pageId="13" pageNumber="14">
<emphasis id="10FBEABDFFC2102C58DF6A93FBB3F937" bold="true" box="[1013,1031,1679,1705]" italics="true" pageId="13" pageNumber="14">P</emphasis>
.
<emphasis id="10FBEABDFFC2102C5F306A92FB35F937" bold="true" box="[1050,1153,1678,1705]" italics="true" pageId="13" pageNumber="14">latipenis</emphasis>
</emphasis>
<bibRefCitation id="461E4B5EFFC2102C5FA66A92FAC2F937" box="[1164,1398,1678,1705]" pageId="13" pageNumber="14" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">Chen &amp; Gao, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="223036AFFFC2102C5BEA6AA8FAC2F96A" blockId="13.[188,1398,1641,1780]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5BEA6AA8FAC2F96A" pageId="13" pageNumber="14">
Paramere lacking pubescence, slightly bifurcated apically (as in
<bibRefCitation id="461E4B5EFFC2102C5F0B6AA8FAB4F950" box="[1057,1280,1715,1742]" pageId="13" pageNumber="14" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">
Chen
<emphasis id="10FBEABDFFC2102C5F456AA9FB06F950" box="[1135,1202,1716,1742]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2005
</bibRefCitation>
, fig. 8C) ……………………………………………………………………
<taxonomicName id="E58F4D2CFFC2102C58CE6AC6FAC2F96A" ID-CoL="4GS6P" authority="Chen &amp; Wen, 2005" authorityName="Chen &amp; Wen" authorityYear="2005" box="[996,1398,1753,1780]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="setitabula">
<emphasis id="10FBEABDFFC2102C58CE6AC6FBCAF96A" bold="true" box="[996,1150,1753,1780]" pageId="13" pageNumber="14">
<emphasis id="10FBEABDFFC2102C58CE6AC6FC42F96A" bold="true" box="[996,1014,1754,1780]" italics="true" pageId="13" pageNumber="14">P</emphasis>
.
<emphasis id="10FBEABDFFC2102C5F236AC5FBCAF96A" bold="true" box="[1033,1150,1753,1780]" italics="true" pageId="13" pageNumber="14">setitabula</emphasis>
</emphasis>
<bibRefCitation id="461E4B5EFFC2102C5FA36AC5FAC2F96A" box="[1161,1398,1753,1780]" pageId="13" pageNumber="14" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">Chen &amp; Wen, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC2102C5B966B01FAC2F81F" pageId="13" pageNumber="14">
<paragraph id="223036AFFFC2102C5B966B01FAC2F8C3" blockId="13.[188,1398,1821,1922]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5B966B01FAC2F8C3" pageId="13" pageNumber="14">
5. Paramere with one expanded, suberect lobe-like process submedially (
<figureCitation id="BAB42A2AFFC2102C5F1F6B01FB00F8A9" box="[1077,1204,1821,1847]" captionStart="Fig. 2" captionStartId="7.[189,232,1932,1958]" captionTargetBox="[210,1372,992,1836]" captionTargetId="figure@7.[187,1401,956,1873]" captionTargetPageId="7" captionText="Fig. 2. Phortica (Phortica) jadete sp. nov., ♂. A. Arista. B. Epandrium and cercus (lateral view). C. Surstylus (frontal view).D. Hypandrium, paramere, aedeagus and aedeagalapodeme (lateral views). E. Paramere (frontal view). F. Tip of aedeagal median rod (ventral view). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173239/files/figure.png" pageId="13" pageNumber="14">Fig. 2D, F</figureCitation>
) ……………… ……………………………………………………………………………………
<taxonomicName id="E58F4D2CFFC2102C5FBE6B5FFAB1F8C3" ID-CoL="BDCZC" authority="Zhu &amp; Cao &amp; Chen, 2018" authorityName="Zhu &amp; Cao &amp; Chen" authorityYear="2018" box="[1172,1285,1858,1885]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="jadete" status="sp. nov.">
<emphasis id="10FBEABDFFC2102C5FBE6B5FFAB1F8C3" bold="true" box="[1172,1285,1858,1885]" pageId="13" pageNumber="14">
<emphasis id="10FBEABDFFC2102C5FBE6B5FFB12F8C3" bold="true" box="[1172,1190,1859,1885]" italics="true" pageId="13" pageNumber="14">P</emphasis>
.
<emphasis id="10FBEABDFFC2102C5F976B5EFAB1F8C3" bold="true" box="[1213,1285,1858,1885]" italics="true" pageId="13" pageNumber="14">jadete</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BC857C6FFC2102C5E3E6B5FFAC2F8C3" box="[1300,1398,1859,1885]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="223036AFFFC2102C5BEA6B74FAC2F81F" blockId="13.[188,1398,1821,1922]" box="[192,1398,1895,1922]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5BEA6B74FAC2F81F" box="[192,1398,1895,1922]" pageId="13" pageNumber="14"> Paramere lacking expanded, suberect lobe-like process …………………………………………6</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC2102C5B966BB7FAC2F875" pageId="13" pageNumber="14">
<paragraph id="223036AFFFC2102C5B966BB7FAC2F858" blockId="13.[188,1398,1963,2027]" box="[188,1398,1963,1990]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5B966BB7FAC2F858" box="[188,1398,1963,1990]" pageId="13" pageNumber="14">
6. Paramere much expanded medially (as in
<bibRefCitation id="461E4B5EFFC2102C59F76BB7FC13F858" box="[733,935,1963,1990]" pageId="13" pageNumber="14" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">
Chen
<emphasis id="10FBEABDFFC2102C580E6BB0FCD4F85B" box="[804,864,1963,1989]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2005
</bibRefCitation>
, fig. 9C;
<bibRefCitation id="461E4B5EFFC2102C5F3F6BB7FB6AF85B" box="[1045,1246,1963,1989]" pageId="13" pageNumber="14" refId="ref7704" refString="Chen H. W., Toda M. J., Lakim M. B. &amp; Mohamed M. B. 2007. The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology 55: 23 - 41." type="journal article">
Chen
<emphasis id="10FBEABDFFC2102C5F766BB0FB2CF85B" box="[1116,1176,1963,1989]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2007
</bibRefCitation>
, fig. 34) …7
</keyLead>
</paragraph>
<paragraph id="223036AFFFC2102C5BEA6BCDFAC2F875" blockId="13.[188,1398,1963,2027]" box="[192,1398,2000,2027]" pageId="13" pageNumber="14">
<keyLead id="997E959AFFC2102C5BEA6BCDFAC2F875" box="[192,1398,2000,2027]" pageId="13" pageNumber="14"> Paramere not expanded medially; aedeagal median rod lacking acute projections ………………8</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC1102F5B976D10FAC3FE0C" pageId="14" pageNumber="15">
<paragraph id="223036AFFFC1102F5B976D10FAC3FED7" blockId="14.[189,1399,267,402]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5B976D10FAC3FED7" pageId="14" pageNumber="15">
7. Paramere lacking process (as in
<bibRefCitation id="461E4B5EFFC1102F595B6D17FC89FEBB" box="[625,829,267,293]" pageId="14" pageNumber="15" refId="ref7704" refString="Chen H. W., Toda M. J., Lakim M. B. &amp; Mohamed M. B. 2007. The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology 55: 23 - 41." type="journal article">
Chen
<emphasis id="10FBEABDFFC1102F59926D10FD42FEBB" box="[696,758,267,293]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2007
</bibRefCitation>
, fig. 34); aedeagal median rod with one pair of small acute projections submedially (as in
<bibRefCitation id="461E4B5EFFC1102F59CE6D33FC18FED7" box="[740,940,303,329]" pageId="14" pageNumber="15" refId="ref7704" refString="Chen H. W., Toda M. J., Lakim M. B. &amp; Mohamed M. B. 2007. The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology 55: 23 - 41." type="journal article">
Chen
<emphasis id="10FBEABDFFC1102F58006D2CFCD2FED7" box="[810,870,303,329]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2007
</bibRefCitation>
, fig. 33) …
<taxonomicName id="E58F4D2CFFC1102F5F1E6D2CFAC3FED7" ID-CoL="8DBXD" authority="Chen &amp; Toda, 2007" authorityName="Chen &amp; Toda" authorityYear="2007" box="[1076,1399,303,330]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zeta">
<emphasis id="10FBEABDFFC1102F5F1E6D2CFB31FED4" bold="true" box="[1076,1157,303,330]" pageId="14" pageNumber="15">
<emphasis id="10FBEABDFFC1102F5F1E6D2CFBF2FED4" bold="true" box="[1076,1094,304,330]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5F7C6D2CFB31FED4" bold="true" box="[1110,1157,304,330]" italics="true" pageId="14" pageNumber="15">zeta</emphasis>
</emphasis>
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="223036AFFFC1102F5BEB6D48FAC3FE0C" blockId="14.[189,1399,267,402]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5BEB6D48FAC3FE0C" pageId="14" pageNumber="15">
Paramere with one small triangular process (as in
<bibRefCitation id="461E4B5EFFC1102F58156D4FFBBDFEF0" box="[831,1033,339,366]" pageId="14" pageNumber="15" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">
Chen
<emphasis id="10FBEABDFFC1102F58AC6D48FC76FEF3" box="[902,962,339,365]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2005
</bibRefCitation>
, fig. 9C); aedeagal median rod lacking projection submedially …………………………………
<taxonomicName id="E58F4D2CFFC1102F58CD6D6BFAC3FE0C" ID-CoL="4GS73" authority="Chen &amp; Wen, 2005" authorityName="Chen &amp; Wen" authorityYear="2005" box="[999,1399,374,402]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="1" phylum="Arthropoda" rank="species" species="unipetala">
<emphasis id="10FBEABDFFC1102F58CD6D6BFBC9FE0F" bold="true" box="[999,1149,374,401]" pageId="14" pageNumber="15">
<emphasis id="10FBEABDFFC1102F58CD6D6BFC4DFE0F" bold="true" box="[999,1017,375,401]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5F216D6AFBC9FE0F" bold="true" box="[1035,1149,374,401]" italics="true" pageId="14" pageNumber="15">unipetala</emphasis>
</emphasis>
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</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC1102F5B976DA5FAC2FE69" pageId="14" pageNumber="15">
<paragraph id="223036AFFFC1102F5B976DA5FAC2FE4D" blockId="14.[189,1398,441,503]" box="[189,1398,441,468]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5B976DA5FAC2FE4D" box="[189,1398,441,468]" pageId="14" pageNumber="15">
8. Posterior branch of paramere slender, neither process nor projection (
<figureCitation id="BAB42A2AFFC1102F5F686DA5FB16FE4D" box="[1090,1186,441,467]" captionStart="Fig. 3" captionStartId="8.[188,231,1966,1992]" captionTargetBox="[216,1397,1105,1897]" captionTargetId="figure@8.[188,1397,1071,1926]" captionTargetPageId="8" captionText="Fig. 3. Phortica (Phortica) kava sp. nov., ♂. A. Arista. B. Epandrium and cercus. C. Surstylus. D. Hypandrium, paramere, aedeagus and aedeagalapodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173241/files/figure.png" pageId="14" pageNumber="15">Fig. 3D</figureCitation>
) ………………9
</keyLead>
</paragraph>
<paragraph id="223036AFFFC1102F5BEB6DC1FAC2FE69" blockId="14.[189,1398,441,503]" box="[193,1398,477,503]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5BEB6DC1FAC2FE69" box="[193,1398,477,503]" pageId="14" pageNumber="15"> Posterior branch of paramere thick, with process or projection ………………………………10</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC1102F5B976E03FAC2FD3B" pageId="14" pageNumber="15">
<paragraph id="223036AFFFC1102F5B976E03FAC3FDC3" blockId="14.[189,1399,541,677]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5B976E03FAC3FDC3" pageId="14" pageNumber="15">
9. Paramere mostly yellow, bifurcated from distal ¼, the anterior branch with three pointed teeth apically (
<figureCitation id="BAB42A2AFFC1102F5A736E5FFE0DFDC3" box="[345,441,579,605]" captionStart="Fig. 3" captionStartId="8.[188,231,1966,1992]" captionTargetBox="[216,1397,1105,1897]" captionTargetId="figure@8.[188,1397,1071,1926]" captionTargetPageId="8" captionText="Fig. 3. Phortica (Phortica) kava sp. nov., ♂. A. Arista. B. Epandrium and cercus. C. Surstylus. D. Hypandrium, paramere, aedeagus and aedeagalapodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173241/files/figure.png" pageId="14" pageNumber="15">Fig. 3D</figureCitation>
); pleura mostly dark brown to black ……………………………
<taxonomicName id="E58F4D2CFFC1102F5F9D6E5FFAA6FDC3" ID-CoL="BDCZD" authority="Zhu &amp; Cao &amp; Chen, 2018" authorityName="Zhu &amp; Cao &amp; Chen" authorityYear="2018" box="[1207,1298,578,605]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kava" status="sp. nov.">
<emphasis id="10FBEABDFFC1102F5F9D6E5FFAA6FDC3" bold="true" box="[1207,1298,578,605]" pageId="14" pageNumber="15">
<emphasis id="10FBEABDFFC1102F5F9D6E5FFB7DFDC3" bold="true" box="[1207,1225,579,605]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5FF36E5EFAA6FDC3" bold="true" box="[1241,1298,578,605]" italics="true" pageId="14" pageNumber="15">kava</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BC857C6FFC1102F5E316E5FFAC3FDC3" box="[1307,1399,579,605]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="223036AFFFC1102F5BEB6E7BFAC2FD3B" blockId="14.[189,1399,541,677]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5BEB6E7BFAC2FD3B" pageId="14" pageNumber="15">
Paramere nearly black, bifurcated submedially, the anterior branch lacking teeth (as in
<bibRefCitation id="461E4B5EFFC1102F5FF76E7BFE93FD3B" pageId="14" pageNumber="15" refId="ref8440" refString="Toda M. J. &amp; Peng T. X. 1992. Some species of the subfamily Steganinae (Diptera, Drosophilidae) from Guangdong Province, southern China. Annales de la Societe entomologique de France (N. S.) 28: 201 - 213." type="journal article">Toda &amp; Peng 1992</bibRefCitation>
, fig. 16); pleura mostly orange to orange brown …………
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<emphasis id="10FBEABDFFC1102F58CC6E97FBC4FD3B" bold="true" box="[998,1136,650,677]" pageId="14" pageNumber="15">
<emphasis id="10FBEABDFFC1102F58CC6E97FC4CFD3B" bold="true" box="[998,1016,651,677]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5F236E96FBC4FD3B" bold="true" box="[1033,1136,650,677]" italics="true" pageId="14" pageNumber="15">bipartita</emphasis>
</emphasis>
(
<bibRefCitation id="461E4B5EFFC1102F5FAB6E96FADAFD3B" box="[1153,1390,650,677]" pageId="14" pageNumber="15" refId="ref8440" refString="Toda M. J. &amp; Peng T. X. 1992. Some species of the subfamily Steganinae (Diptera, Drosophilidae) from Guangdong Province, southern China. Annales de la Societe entomologique de France (N. S.) 28: 201 - 213." type="journal article">Toda &amp; Peng, 1992</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="997B2E0AFFC1102F5B976ED1FAC3FC95" pageId="14" pageNumber="15">
<paragraph id="223036AFFFC1102F5B976ED1FAC3FD79" blockId="14.[189,1399,716,779]" box="[189,1399,716,743]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5B976ED1FAC3FD79" box="[189,1399,716,743]" pageId="14" pageNumber="15">
10. Paramere strongly sclerotized distally or apically (
<figureCitation id="BAB42A2AFFC1102F58656ED0FC07FD79" box="[847,947,716,743]" captionStart="Fig.4" captionStartId="9.[189,232,1967,1993]" captionTargetBox="[202,1378,1236,1912]" captionTargetId="figure@9.[189,1398,1211,1932]" captionTargetPageId="9" captionText="Fig.4.Phortica(Phortica)mengdasp.nov.,♂.A.Epandrium andcercus.B. Surstylus.C. Hypandrium, paramere, aedeagus and aedeagal apodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173243/files/figure.png" pageId="14" pageNumber="15">Figs 4C</figureCitation>
,
<figureCitation id="BAB42A2AFFC1102F58E96ED0FC53FD79" box="[963,999,716,743]" captionStart="Fig. 6" captionStartId="11.[189,232,1965,1991]" captionTargetBox="[228,1383,1185,1917]" captionTargetId="figure@11.[189,1398,1147,1925]" captionTargetPageId="11" captionText="Fig. 6. Phortica (Phortica) yena sp. nov., ♂. A. Epandrium and cercus; B. Surstylus; C. Hypandrium, paramere, aedeagus and aedeagal apodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173247/files/figure.png" pageId="14" pageNumber="15">6C</figureCitation>
) ………………………………11
</keyLead>
</paragraph>
<paragraph id="223036AFFFC1102F5BEB6EEDFAC3FC95" blockId="14.[189,1399,716,779]" box="[193,1399,752,779]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5BEB6EEDFAC3FC95" box="[193,1399,752,779]" pageId="14" pageNumber="15"> Paramere not black ………………………………………………………………………………12</keyLead>
</paragraph>
</keyStep>
<paragraph id="223036AFFFC1102F5B976F2FFAC3FC27" blockId="14.[189,1399,818,953]" pageId="14" pageNumber="15">
<keyStep id="997B2E0AFFC1102F5B976F2FFAC3FC27" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5B976F2FFAC3FC27" pageId="14" pageNumber="15">
11. Paramere deeply bifurcated distally, anterior branch truncate, with three teeth apically (
<figureCitation id="BAB42A2AFFC1102F5E3A6F2EFADAFCD3" box="[1296,1390,818,845]" captionStart="Fig.4" captionStartId="9.[189,232,1967,1993]" captionTargetBox="[202,1378,1236,1912]" captionTargetId="figure@9.[189,1398,1211,1932]" captionTargetPageId="9" captionText="Fig.4.Phortica(Phortica)mengdasp.nov.,♂.A.Epandrium andcercus.B. Surstylus.C. Hypandrium, paramere, aedeagus and aedeagal apodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173243/files/figure.png" pageId="14" pageNumber="15">Fig. 4C</figureCitation>
) …………………………………………………………………………………..
<taxonomicName id="E58F4D2CFFC1102F5FAC6F4BFAB8FCEF" ID-CoL="BDCZH" authority="Zhu &amp; Cao &amp; Chen, 2018" authorityName="Zhu &amp; Cao &amp; Chen" authorityYear="2018" box="[1158,1292,854,881]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mengda" status="sp. nov.">
<emphasis id="10FBEABDFFC1102F5FAC6F4BFAB8FCEF" bold="true" box="[1158,1292,854,881]" pageId="14" pageNumber="15">
<emphasis id="10FBEABDFFC1102F5FAC6F4BFB2CFCEF" bold="true" box="[1158,1176,855,881]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5F866F4AFAB8FCEF" bold="true" box="[1196,1292,854,881]" italics="true" pageId="14" pageNumber="15">mengda</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BC857C6FFC1102F5E326F4BFAC3FCEF" box="[1304,1399,855,881]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
Paramere not bifurcated distally, anterior branch pointed, lacking teeth apically (
<figureCitation id="BAB42A2AFFC1102F5FE26F66FA93FC0B" box="[1224,1319,890,917]" captionStart="Fig. 6" captionStartId="11.[189,232,1965,1991]" captionTargetBox="[228,1383,1185,1917]" captionTargetId="figure@11.[189,1398,1147,1925]" captionTargetPageId="11" captionText="Fig. 6. Phortica (Phortica) yena sp. nov., ♂. A. Epandrium and cercus; B. Surstylus; C. Hypandrium, paramere, aedeagus and aedeagal apodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173247/files/figure.png" pageId="14" pageNumber="15">Fig. 6C</figureCitation>
) …… ……………………………………………………………………………………
<taxonomicName id="E58F4D2CFFC1102F5FBF6F83FB4FFC27" ID-CoL="BDCZR" authority="Zhu &amp; Cao &amp; Chen, 2018" authorityName="Zhu &amp; Cao &amp; Chen" authorityYear="2018" box="[1173,1275,926,953]" class="Insecta" family="Drosophilidae" genus="Phortica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="yena" status="sp. nov.">
<emphasis id="10FBEABDFFC1102F5FBF6F83FB4FFC27" bold="true" box="[1173,1275,926,953]" pageId="14" pageNumber="15">
<emphasis id="10FBEABDFFC1102F5FBF6F83FB13FC27" bold="true" box="[1173,1191,927,953]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5FE86F83FB4FFC27" bold="true" box="[1218,1275,927,953]" italics="true" pageId="14" pageNumber="15">yena</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BC857C6FFC1102F5E256F83FAC3FC27" box="[1295,1399,927,953]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<keyStep id="997B2E0AFFC1102F5B976FFCFAC3FBDD" pageId="14" pageNumber="15">
<paragraph id="223036AFFFC1102F5B976FFCFAC2FC64" blockId="14.[189,1399,992,1091]" box="[189,1398,992,1019]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5B976FFCFAC2FC64" box="[189,1398,992,1019]" pageId="14" pageNumber="15">
12. Paramere bifurcated distally (
<figureCitation id="BAB42A2AFFC1102F59656FFCFD1AFC65" box="[591,686,992,1019]" captionStart="Fig.5" captionStartId="10.[189,232,1966,1992]" captionTargetBox="[217,1370,1202,1885]" captionTargetId="figure@10.[189,1398,1169,1912]" captionTargetPageId="10" captionText="Fig.5.Phortica(Phortica)wongdingsp.nov.,♂.A.Epandriumandcercus;B.Surstylus;C.Hypandrium, paramere, aedeagus and aedeagal apodeme. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1173245/files/figure.png" pageId="14" pageNumber="15">Fig. 5C</figureCitation>
); aedeagal median rod without processes subapically …13
</keyLead>
</paragraph>
<paragraph id="223036AFFFC1102F5BEB6819FAC3FBDD" blockId="14.[189,1399,992,1091]" pageId="14" pageNumber="15">
<keyLead id="997E959AFFC1102F5BEB6819FAC3FBDD" pageId="14" pageNumber="15">
Paramere not bifurcated distally (as in
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Chen
<emphasis id="10FBEABDFFC1102F59D26819FC80FB80" box="[760,820,1028,1054]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2005
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, fig. 10C); aedeagal median rod with a pair of triangular processes subapically (as in
<bibRefCitation id="461E4B5EFFC1102F59FB6834FC23FBDD" box="[721,919,1064,1091]" pageId="14" pageNumber="15" refId="ref7424" refString="Chen H. W., Gao J. J. &amp; Wen S. Y. 2005. Species diversity of the subgenus Phortica (s. str.) in Yunnan, China, with descriptions of nine new species (Diptera, Drosophildae). Journal of Natural History 39: 3951 - 3978. https: // doi. org / 10.1080 / 00222930500533534" type="journal article">
Chen
<emphasis id="10FBEABDFFC1102F583C6835FCE5FBDC" box="[790,849,1064,1090]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2005
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, fig.10C) …
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.
<emphasis id="10FBEABDFFC1102F5F636834FB39FBDC" bold="true" box="[1097,1165,1064,1090]" italics="true" pageId="14" pageNumber="15">pangi</emphasis>
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13. Posterior branch of paramere with one triangular process submedially, slender distally (
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) ………………………………………………………………………………
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<emphasis id="10FBEABDFFC1102F5F706892FBD8FB36" bold="true" box="[1114,1132,1166,1192]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5FAD6891FB4AFB36" bold="true" box="[1159,1278,1165,1192]" italics="true" pageId="14" pageNumber="15">wongding</emphasis>
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Posterior branch of paramere with one quadrate process submedially, broadened distally (as in
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, fig. 3B) …………………………………………………
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<emphasis id="10FBEABDFFC1102F5F0D68CAFB8DFB6E" bold="true" box="[1063,1081,1238,1264]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<emphasis id="10FBEABDFFC1102F5F6168CAFB09FB6E" bold="true" box="[1099,1213,1238,1264]" italics="true" pageId="14" pageNumber="15">eparmata</emphasis>
</emphasis>
(
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)
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