treatments-xml/data/4C/F5/F6/4CF5F63B4B065FC035F8653C85BD67DA.xml

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<mods:title id="1CCF9C9740BBA44B3F7A571C49324333">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="3D6B4DC0D184CB8EB74A40663A7EB7BF">Bistroem, Olof</mods:namePart>
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<mods:namePart id="5109AA2F06DD7A621415229B180F8F20">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="B47CD0774CB269E8E0B04A02F3BE49A7">Bergsten, Johannes</mods:namePart>
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<subSubSection id="DB2DE0C660BAEAE02D12EEC5F3C0CEFF" pageId="46" pageNumber="47" type="multiple">
<paragraph id="D76368FDAB0DFD1872564B57A2A85786" pageId="46" pageNumber="47">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
</subSubSection>
<subSubSection id="0981B43B5C5D7D673CB856CFDE358CEC" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="CC510E18688D3A4157875753FC2CDC1A" pageId="46" pageNumber="47">
<taxonomicName id="8C19E4CCDFDD2AD02C27DF66BAB94043" ID-CoL="6NRMN" authority="Gschwendtner, 1935" authorityName="Gschwendtner" authorityYear="1935" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis Gschwendtner, 1935</taxonomicName>
Figs 47-48, 249, 400-401, 535
</paragraph>
</subSubSection>
<subSubSection id="3102B2E04E09635CF63ADD2E8F404D0E" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="64B39092924C33C1CBC2A44AF528B77C" pageId="46" pageNumber="47">
<taxonomicName id="7E8C8FD19CE4E2512E959A52D4271EB4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus posticus" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subspecies" species="posticus" subspecies="continentalis">Laccophilus posticus continentalis</taxonomicName>
<bibRefCitation id="B28CD3256BEAA7129BE9FE259F4A362F" author="Gschwendtner, L" journalOrPublisher="Annals of the Transvaal Museum" pageId="256" pageNumber="257" pagination="15 - 25" title="Wissenschaftliche Ergebnisse der Vernay-Lang Kalahari-Expedition, Maerz bis September, 1930. Haliplidae et Dytiscidae." volume="17" year="1935 a">Gschwendtner 1935a</bibRefCitation>
: 16, 18 (original description, faunistics).
</paragraph>
<paragraph id="EAEEF198D333C9C4F47931468074D36F" pageId="46" pageNumber="47">
<taxonomicName id="32AD787962DEF0CCDAD9ADF118D8986C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschwendter, 
<bibRefCitation id="A1A17DD4A969965036D2B35B69F0EF94" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
: 279, 281, 283, 312, 316 (discussion, description, faunistics); 
<bibRefCitation id="0A5D3F3A560CFF2A3339E89F9772E70D" author="Capra, F" journalOrPublisher="DORIANA, Annali del Museo Civico di Storia Naturale &quot; G. Doria &quot;" pageId="255" pageNumber="256" pagination="6 - 8" title="Appendice." volume="1" year="1952">Capra 1952</bibRefCitation>
: 6: (faunistics); 
<bibRefCitation id="59AC9AB17F6D0A34BB649F87AEE2B311" author="Omer-Cooper, Joyce" journalOrPublisher="Sociedade Estudos de Mocambique" pageId="260" pageNumber="261" pagination="19 - 32" title="Some records of Dytiscidae (Col.) from Mozambique." volume="101" year="1956">Omer-Cooper 1956</bibRefCitation>
: 21 (faunistics, biology); 
<bibRefCitation id="4D58BBD5D1F98E2E31A9ACFE0F6F8D5B" author="Omer-Cooper, Joyce" journalOrPublisher="Memoirs of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="1 - 90" title="Dytiscidae (Coleoptera) collected in the Transvaal (Rhodes University Expedition 1948)." volume="5" year="1957">Omer-Cooper 1957</bibRefCitation>
: 20, 21 (discussion, description); 
<bibRefCitation id="84F43A9EAC852B0CBBE463DEB00090DE" author="Omer-Cooper, Joyce" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="260" pageNumber="261" pagination="57 - 60" title="Some Dytiscidae (Col.) from Swaziland, Southern Africa, with the description of a new species." volume="94" year="1958 a">Omer-Cooper 1958a</bibRefCitation>
: 57, 59 (faunistics, biology); 
<bibRefCitation id="66E0300D7FC4D75B372A51558DEE7082" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper 1958b</bibRefCitation>
: 37, 43, 44, 45 (description, discussion, faunistics, biology); 
<bibRefCitation id="8DDD17922B85D199B284288980139C5B" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 570, 573, 575 (redescription, faunistics); 
<bibRefCitation id="0C41122576D1DA865DCC12642B2E5A51" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
: 77, 81 (description, faunuistics); 
<bibRefCitation id="11B06013FFD8DD611992D69F14999735" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
: 288, 289, 290 (discussion, description, faunistics); 
<bibRefCitation id="C071267B40AD6B20151C92FA10557557" pageId="46" pageNumber="47">Rocchi 1975</bibRefCitation>
: 48 (faunistics); 
<bibRefCitation id="3F59D58785DFAD9A2776F5AC8765DD86" author="Bilardo, A" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="254" pageNumber="255" pagination="188 - 190" title="Mission entomologique du Musee Royal de l'Afrique Centrale aux Monts Uluguru, Tanzanie (L. Berger, N. Leleup et J. Debecker, V-VIII. 1971) 13. ColeopteraDytiscidae." volume="90" year="1976">Bilardo 1976</bibRefCitation>
: 190 (faunistics, biology); 
<bibRefCitation id="5542C44EF3E2B79FD09C300353869789" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali del Museo Civivico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="85 - 106" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Botswana (Coleoptera)." volume="128" year="1987">Bilardo and Rocchi 1987</bibRefCitation>
: 104 (faunistics, biology); 
<bibRefCitation id="1B20D778F84957120FAC163E2D33AF08" author="Rocchi, S" journalOrPublisher="Rendiconti Lincei (Scienze Fisiche e Naturali) (Serie 9)" pageId="262" pageNumber="263" pagination="441 - 445" title="Idroadefagi raccolti nel Kenya nord-orientale e descrizione di Copelatuspardii sp. n." url="10.1007/BF03001779" volume="1" year="1990">Rocchi 1990</bibRefCitation>
:442 (faunistics); 
<bibRefCitation id="91A24787C3D9CDE77B3D2193E8FF29ED" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 242 (catalougue, faunistics); 
<bibRefCitation id="2766DBE1AA6B171E48BB496F01475538" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 210 (catalougue, faunistics).
</paragraph>
<paragraph id="7CCF2F8585AB9CB42FDDE8EA29312D45" pageId="46" pageNumber="47">
<taxonomicName id="F25AD478779D837F5731256BE605A687" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus perplexus" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="perplexus">Laccophilus perplexus</taxonomicName>
<bibRefCitation id="049050552BF3F68A47F11B24381A677C" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
: 287, 288, 289, 290 (original description, faunistics, discussion.); 
<bibRefCitation id="CD73AE6887AFBCE1B18BC025454902E8" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 248 (catalogue, faunistics); 
<bibRefCitation id="9C4EC44AF4D0A60212F80AD6899B5680" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 215 (catalougue, faunistics). New synonym.
</paragraph>
</subSubSection>
<subSubSection id="B693636AF96F74C4B5782381F3A12E03" pageId="46" pageNumber="47" type="type localities">
<paragraph id="F2A30BE0350BC4B20B17D0FFAA9DFB16" pageId="46" pageNumber="47">Type localities.</paragraph>
<paragraph id="6ECAF98135B3398F37D35CFA6E7AD5A1" pageId="46" pageNumber="47">
<taxonomicName id="47D6CC97320CD1C878E056692316899D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
: Botswana: Kalahari, Tsotsorogo Pan.
</paragraph>
<paragraph id="0C40E025AD7AB939EA0F6E47E595D1AD" pageId="46" pageNumber="47">
<taxonomicName id="7D946B2CE6F0F3F34E13B3409C666D3F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus perplexus" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="perplexus">Laccophilus perplexus</taxonomicName>
: Mozambique: Umbeluzi River near Goba.
</paragraph>
</subSubSection>
<subSubSection id="767FBC1E94A9722C7E71E3771255A0CB" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="type material">
<paragraph id="C0845F1D3B59DF07943145BFB03C27A6" pageId="46" pageNumber="47">Type material, studied</paragraph>
<paragraph id="5EC85B86CEC38F990161FD088D0327B0" pageId="46" pageNumber="47">
(21 exs.). 
<taxonomicName id="4B137561B6A664E6F5A3113FFC4873CB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
: Lectotype (by present designation): male: &quot;V.-L. Kal. Exp. Tsotsorogo Pan 17/6-9/7/30 / Type male (symbol) Gschw. / 
<taxonomicName id="CAD31EEC3906E1D655672ADF6437AF82" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus posticus" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subspecies" species="posticus" subspecies="continentalis">Laccophilus posticus continentalis</taxonomicName>
det. Gschwendtner&quot; (TMSA; habitus in Fig. 400). - Paralectotypes: Same data as lectotype, but &quot;Type female (symbol)&quot; (1 ex. TMSA); &quot;V.-L. Kal. Exp. 
<normalizedToken id="3F3E4F2B41B791FD40DD72BBCDD03D51" originalValue="N’Kate">N'Kate</normalizedToken>
Makarikari 6-23/8/1930 / Paratypus 
<taxonomicName id="9B7C08B42C3FBC7527ACCA0055AB254B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus posticus" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subspecies" species="posticus" subspecies="continentalis">Laccophilus posticus continentalis</taxonomicName>
ssp. L. Gschwendtner&quot; (10 exs. TMSA, 1 ex. OLML); same as lectotype but labelled as &quot;Paratype Gschw.&quot; (5 exs. OLML, 2 exs. AMGS).
</paragraph>
<paragraph id="A035DB69D3278D3998194565FBDB5DCB" pageId="47" pageNumber="48">
<taxonomicName id="D7125796E2E797110A27E30CB85DAB6A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus perplexus" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="perplexus">
<pageBreakToken id="F83CF62998E5B5FD6003CFF5AA5DE4A1" pageId="47" pageNumber="48" start="start">Laccophilus</pageBreakToken>
perplexus
</taxonomicName>
: Holotype: male: &quot;Type / 
<taxonomicName id="C5A34630B00F307E48CABDDC0DE3B87C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus perplexus" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="perplexus">Laccophilus perplexus</taxonomicName>
sp. n. / Mozambique Umbeluzi River near Goba 4.12. 1948 J.O.C.&quot; (AMGS; according to original description holotype preserved in BMNH).
</paragraph>
</subSubSection>
<subSubSection id="DBD9BC296083D8A7A056438D0777872A" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="51F183E223DC15583D943D92E7D47E6B" pageId="47" pageNumber="48">Additional material, studied</paragraph>
<paragraph id="B51957A9AC03C6DA8E487C8C8307CC90" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48">
(518 exs.). Senegal: &quot;Sumpf von Peykone, Senegal 9. 08 Riggenbach S.V.&quot; (1 ex. ZMHB). - Gambia-S. Senegal: &quot;Stream of Selety 
<geoCoordinate id="5D00B168F30A3C538999247FA64FF9BD" direction="north" orientation="latitude" precision="925" value="13.166667">13°10'N-</geoCoordinate>
<geoCoordinate id="205A09AD4487EB25AF7CCD349E8C3D9D" direction="west" orientation="longitude" precision="925" value="-16.6">16°36'W</geoCoordinate>
19.2. 1976 Holmen leg.&quot; (2 exs. ZMUC). - Sudan: &quot;Wad Medani a. Bl. Nil 29.10. 1979 Hieke&quot; (1 ex. ZMHB); same but &quot;12.10. 1979 lux&quot; (2 exs. ZMHB); same but &quot;18.10. 1979&quot;(1 ex. ZMHB); same but &quot;8.10. 1979&quot; (2 exs. ZMHB); same but &quot;9.10. 1979&quot; (1 ex. ZMHB); same but &quot;30-31.10. 1979&quot; (1 ex. ZMHB); same but 
<normalizedToken id="3EC382B347E94DD4E75B5B0AE4B490FD" originalValue="“20.10.1979“">&quot;20.10.1979&quot;</normalizedToken>
(1 ex. ZMHB); same but &quot;15.10. 1979&quot; (1 ex. ZMHB); same but 
<normalizedToken id="80A5A4214BC0F2A18190EBC2312CA519" originalValue="“22.10.1979“">&quot;22.10.1979&quot;</normalizedToken>
(2 exs. ZMHB); same but &quot;leg. 
<normalizedToken id="534CED36D6AE0F2FE83528D73D297DCA" originalValue="Königsmann“">Koenigsmann&quot;</normalizedToken>
(1 ex. ZMHB); &quot;Senaar a.Bl. Nil, lux 21.10.1979 Hieke&quot; (5 exs. ZMHB); &quot;Umm Banein, light trap 14.11. 1962 Linnavuori&quot; (1 ex. MZH). - Ghana: &quot;N Region Nyankpala 183 m 
<geoCoordinate id="32E7B67D514C236F073EBDF4D5D6F825" direction="north" orientation="latitude" precision="925" value="9.416667">
N9°25 
<normalizedToken id="7E99B3A00FEA1789C86F373ECDBD6194" originalValue="’-W1°00’">
'- 
<geoCoordinate id="F01BC37D8C8FDACD03F5AFB756B15F24" direction="west" orientation="longitude" precision="925" value="-1.0">W1°00'</geoCoordinate>
</normalizedToken>
</geoCoordinate>
Dr. S. 
<normalizedToken id="D4656D3F3A7CB17D91D009AADADDE3FB" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
/ shore washing 10.2. 1970&quot; (1 ex. CGW). - Nigeria: &quot;Samaru 17.5. 1959, Sands / light trap&quot; (1 ex. BMNH); &quot;Nt Kano 1.5. 1928 Lindwer Madsen&quot; (1 ex. ZMUC). - Somalia: &quot;Somali Rep. 1961 Roffey&quot;(3 exs. BMNH, 1 ex. MZH). - Kenya: &quot;Kibwezi Scheffler leg.&quot; (1 ex. ZMHB); same as but 
<normalizedToken id="2DE91CE893DCCBFE31044CAA30A6749A" originalValue="“1906“">&quot;1906&quot;</normalizedToken>
(1 ex. ZMHB); &quot;Eastern Mwingi, Nguni env. 28.11. 1999 Snizek&quot; (1 ex. NMW); &quot;SE Kenya ShimbaHills 20 km S Mombasa 5.6. 1985 
<normalizedToken id="7CB690C8FA5EB24620C09CB7147A3425" originalValue="Lödl”">Loedl&quot;</normalizedToken>
(1 ex. NMW, 1 ex. MZH); Mafisini, pond, Kwale District 19.9. 1976 Holmen leg.&quot; (1 ex. ZMUC); &quot;Maji ya Chumvi River, Kwale District 16.9. 1976 Holmen leg.&quot; (1 ex. ZMUC); &quot;Maji-Chumwi (Wa Nyika) Alluaud 7. 1903&quot; (2 exs. NHMB); &quot;Athi River, Machakos District 14.9. 1976 Holmen det&quot; (1 ex. ZMUC); &quot;Kombeni River, Mazeras, Kilifi distr 15.9. 1976 Holmen&quot; (2 exs. ZMUC, 2 exs. MZH); &quot;Mariakani dam, Kilifi District 16.9. 1976 Holmen&quot; (4 exs. MZH); &quot;Mandera R. Dana 23.10. 1970 Brown&quot; (1 ex. BMNH); &quot;Wajir 27.10. 1970 Brown&quot; (1 ex. BMNH); &quot;Malindi, alle luci, 15.11.-5.12. 1989 / 
<taxonomicName id="C07B435D09B052954DCCE2F4C4C98EED" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschwendtner det. 
<bibRefCitation id="7FB8108E1AD16D5DF7EFA95764F2EFEA" author="Rocchi, S" journalOrPublisher="Rendiconti Lincei (Scienze Fisiche e Naturali) (Serie 9)" pageId="262" pageNumber="263" pagination="441 - 445" title="Idroadefagi raccolti nel Kenya nord-orientale e descrizione di Copelatuspardii sp. n." url="10.1007/BF03001779" volume="1" year="1990">Rocchi 1990</bibRefCitation>
&quot; (3 exs. CSR); &quot;Voi 11. 1997 Snizek M. / 
<taxonomicName id="02B23C6EABF96C77BD160786824AD92B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschwendtner det. 
<bibRefCitation id="250895BED340A12A65A308673CF54CA8" author="Rocchi, S" journalOrPublisher="Rendiconti Lincei (Scienze Fisiche e Naturali) (Serie 9)" pageId="262" pageNumber="263" pagination="441 - 445" title="Idroadefagi raccolti nel Kenya nord-orientale e descrizione di Copelatuspardii sp. n." url="10.1007/BF03001779" volume="1" year="1990">Rocchi 1990</bibRefCitation>
&quot; (3 exs. CSR); &quot;Voi Mtito Andei, light trap roof Tsavo Inn 24-25.11. 1990 
<normalizedToken id="5282B4077EE4B9F4349ABF2F1F7D7391" originalValue="Päts">Paets</normalizedToken>
&amp; Viklund&quot; (1 ex. MZH); &quot;Fort Hall / Coll. E. 
<normalizedToken id="5F08A5C4BF52F14A7A6D4C117842BB6E" originalValue="Häuser">Haeuser</normalizedToken>
/ 
<taxonomicName id="CE3FB5371F6EBBCA6CB1CECBD0116F03" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschwendtner det. Brancucci&quot; (1 ex. ZMHB); &quot;Afr. or Jkurha&quot; (1 ex. ZMHB). - Tanzania: &quot;Daressalaam, Pangani und Hinterland Regner&quot; (1 ex. ZMHB); &quot;Daressalaam leg. Methner&quot; (1 ex. NHMB, 1 ex. ZMHB); &quot;Reg. Morogoro Mikumi 17-20.12. 1993 Bednatik&quot; (1 ex. NMW); &quot;Mts Uluguru, Morogoro Campus Fac. Agric., UV, 600 m 5-6. 1971 / 
<taxonomicName id="202174721F5F144D318CE1B734463E8B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. det. Bilardo&quot; (1 ex. NHMB); &quot;Ponds S of Korogwe, Korogwe District 24.9. 1976 Holmen&quot; (2 exs. ZMUC, 1 ex. MZH); &quot;Rice field S of Tanga, Tanga District 26.9. 1976 Holmen&quot; (1 ex. ZMUC); &quot;Tanga / 
<normalizedToken id="34F08A9B4357BACFCBD2BD1B86B865B3" originalValue="Sjöstedt”">Sjoestedt&quot;</normalizedToken>
(1 ex. NHRS); &quot;Tanga Reimer S.&quot; (1 ex. ZMHB); &quot;Tanganyika Pond in stream 103 (?) miles from Dodoma 15.2. 1954 JOC&quot; (1 ex. AMGS); &quot;Tanga Prov. 4-5. 1950 Sweeney / At light&quot; (2 exs. BMNH); &quot;Usa River 3900 feet Dr. J. Szunyoghy / Light trap 15.11.-31.12. 1965&quot; (1 ex. CGW); &quot;2 mi to L. Manyara, seashore 3150 feet Dr. J. Szunyoghy / singled material 1-26. 1965&quot; (1 ex. CGW); 
<normalizedToken id="6662CE38D681FA5D7131963DE928D37C" originalValue="“Usagara”">&quot;Usagara&quot;</normalizedToken>
(1 ex. ZMHB); &quot;Umgb Urumba 15.9. D.O.A. leg. Methner / 
<taxonomicName id="E8AD468EDB13C2A7487B4F922A5AB1E2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. det. M. Brancucci&quot; (3 exs. ZMHB); &quot;J. 
<pageBreakToken id="8703CA9C285EA512340C52E5E7657AFD" pageId="48" pageNumber="49" start="start">Kurha</pageBreakToken>
&quot; (1 ex. ZMHB); &quot;Zanzibar Pemba 23. Sept. 1955 Fowler&quot; (5 exs. AMGS); &quot;Zanzibar 13th Sept. 1955 JOC.&quot; (2 exs. AMGS); &quot;Zanzibar 5. 1954 Brown&quot; (2 exs. BMNH, 1 ex. MZH); &quot;Zanzibar / Reimer S. &amp; Schultz. - Zambia: &quot;S Luangwa NP, Mfuwe Crocodile Farm, 13.06.03S, 31.47.32E, 450 m, lux 23.3. 1993 Uhlig&quot; (45 exs. ZMHB, 2 exs. MZH); same but &quot;21.3. 1993&quot; (4 exs. ZMHB); same but &quot;24.3. 1993&quot; (6 exs. ZMHB); same but &quot; 
<geoCoordinate id="512DC4AED5C1DC5CA406135034581E7B" direction="south" orientation="latitude" precision="555" value="-13.06">13.06S</geoCoordinate>
, 
<geoCoordinate id="274B096A2D83C1CAD1C4B5F7EEA1A89E" direction="south" orientation="latitude" precision="555" value="-31.47">31.47S</geoCoordinate>
, 21-24.3. 1993 
<normalizedToken id="357E2CF5E119380E543CA480237B1027" originalValue="Göllner”">Goellner&quot;</normalizedToken>
(2 exs. ZMHB); same but 
<normalizedToken id="3A9D4C137D1064DE1FD8156B6A8BEEC3" originalValue="“Deckert”">&quot;Deckert&quot;</normalizedToken>
(1 ex. ZMHB). - Namibia: &quot;E Caprivi: Katima Mulilo, lux, 
<geoCoordinate id="E01790BD50C1DF494FA6699BAFEF4CFA" direction="south" orientation="latitude" precision="555" value="-17.29">17.29S</geoCoordinate>
, 
<geoCoordinate id="6C8CA2F75805C3D409D4022917CB5D91" direction="east" orientation="longitude" precision="555" value="24.17">24.17E</geoCoordinate>
, 3-8.3. 1992 Uhlig&quot; (65 exs. ZMHB, 5 exs. MZH; habitus in Fig. 401); same but &quot;Deckert leg.&quot; (2 exs. ZMHb); same but 
<normalizedToken id="F731A9515C2B375BCB20C74C4556C43B" originalValue="“Göllner">&quot;Goellner</normalizedToken>
leg.&quot; (2 exs. ZMHB); &quot;E Caprivi: 3 km E Katima Mulilo, 
<geoCoordinate id="E8762FEB0C9D956726CB7A0821D13B11" direction="south" orientation="latitude" precision="555" value="-17.29">17.29S</geoCoordinate>
, 
<geoCoordinate id="4F7B86EE10932E2DC41072D0E5F8CC57" direction="east" orientation="longitude" precision="555" value="24.18">24.18E</geoCoordinate>
, Hippo Camp, in Swimming Pool 6.3.1992 Uhlig&quot; (1 ex. ZMHB); &quot;E Caprivi: 30 km SE Katima Mulilo 
<geoCoordinate id="CCBD3D7F36791FA48BFB872B2F222670" direction="south" orientation="latitude" precision="555" value="-17.31">17.31S</geoCoordinate>
, 
<geoCoordinate id="720E59A9A60C00AC919622F7250D6A03" direction="east" orientation="longitude" precision="555" value="24.25">24.25E</geoCoordinate>
, Zambesi - Altwasserarm, lux 6.3. 1992 Uhlig&quot; (7 exs. ZMHB, 1 ex. NMNW); &quot;E Caprivi: Mudumu NP, Nakatwa, 
<geoCoordinate id="63B71518AAA6AB971F8BD68A868F1B34" direction="south" orientation="latitude" precision="555" value="-18.1">18.10S</geoCoordinate>
, 
<geoCoordinate id="C5D5CDDD4887512267315A3D31297902" direction="east" orientation="longitude" precision="555" value="23.26">23.26E</geoCoordinate>
, 8-13.3. 1992 Uhlig&quot; (3 exs. ZMHB); same but &quot;Buffalo Trails Camp lux, ca. 
<geoCoordinate id="F846F9F1F5F785673C9DB720F2EC990A" direction="south" orientation="latitude" precision="555" value="-18.1">18.10S</geoCoordinate>
, 
<geoCoordinate id="EE303261152EABC74CD749E2E3B94CCE" direction="east" orientation="longitude" precision="555" value="23.26">23.26E</geoCoordinate>
, 12.3. 1992&quot; (3 exs. ZMHB); same but &quot;Kwando-Ufer, 
<taxonomicName id="91FF0A114CF9B3330826322D0C9FD83B" class="Liliopsida" family="Poaceae" genus="Phragmites" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phragmites" order="Poales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="genus">Phragmites</taxonomicName>
, schlammig&quot; (1 ex. ZMHB); &quot;Kavango Popa Falls 
<geoCoordinate id="AE18AB310C2166B97B1694EFC2C91561" direction="south" orientation="latitude" precision="555" value="-18.07">18.07S</geoCoordinate>
, 
<geoCoordinate id="478CACF91716F0A4521C16C91596AA93" direction="east" orientation="longitude" precision="555" value="21.35">21.35E</geoCoordinate>
, lux 26.2.-3.3. 1992 lux Deckert&quot; (1 ex. ZMHB); same but 
<normalizedToken id="00801D810E02D9F54BA4854875D9EBD9" originalValue="“Uhlig”">&quot;Uhlig&quot;</normalizedToken>
(1 ex. ZMHB); &quot;Kavango Mahango Game Reserve 
<geoCoordinate id="CB65E52D33138296B69A9984050EDA8C" direction="south" orientation="latitude" precision="555" value="-18.17">18.17S</geoCoordinate>
, 
<geoCoordinate id="1399A221BD1D2DFF47B2B08B361FC6DA" direction="east" orientation="longitude" precision="555" value="21.43">21.43E</geoCoordinate>
, lux 2.3. 1992 
<normalizedToken id="232DDA711B8A9A83D15A2836E3680B75" originalValue="Göllner”">Goellner&quot;</normalizedToken>
(1 ex. ZMHB); Okavango Distr., Mutompo, 60 km S Rundu, 18.18.38,7S, 19.15.29,4E, 1180 m NN, 13.3. 2003, hand light trap, Frisch &amp; Voland&quot; (1 ex. ZMHB). - Botswana: &quot;Serowe, sevage ponds, 
<normalizedToken id="1439762A98DD9DA4214C14CE4B8BC499" originalValue="Farmer’s">Farmer's</normalizedToken>
Brigade 1.6. 1987, SE22 26 BD, Forchhammer leg.&quot; (3 exs. MZH); same but &quot;7.6. 1987&quot; (3 exs. MZH); same data but &quot;4. 1988 / 
<taxonomicName id="71C0C85944C1C9916553E0AD9B302EB3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschwendtner det. Rocchi 1993&quot; (2 exs. CSR); &quot;Kasane, Chobe Safari lodge, 17.48.32S, 25.08.39E, 26.11.1993 lux, Uhlig&quot; (5 exs. ZMHB, 3 exs. MZH); &quot;5 km NW San-ta-wani Safari Lodge, 19.27.01S, 23.38.46E, lux 8-9.3.1993 Uhlig&quot; (27 exs. ZMHB); &quot;5 km NW San-ta-wani Safari Lodge, 
<geoCoordinate id="4A88C9D744BFF7BEBDE1F57A77526034" direction="south" orientation="latitude" precision="555" value="-19.27">19.27S</geoCoordinate>
, 
<geoCoordinate id="FF5059705D42078EB2EC0F321CC8573D" direction="east" orientation="longitude" precision="555" value="23.38">23.38E</geoCoordinate>
, lux 8-9.3. 1993 
<normalizedToken id="B77F731B1F07EC7BC5B5AA1F3934CC2E" originalValue="Göllner”">Goellner&quot;</normalizedToken>
(1 ex. ZMHB); &quot;6 km E Kalkfontein, 
<geoCoordinate id="9143716499F5BBDBC800A3B20EE476E2" direction="south" orientation="latitude" precision="555" value="-22.04">22.04S</geoCoordinate>
, 
<geoCoordinate id="6C917FFCB3A5F6F20287E2019931BA71" direction="east" orientation="longitude" precision="555" value="20.56">20.56E</geoCoordinate>
, 7.3. 1993 lux 
<normalizedToken id="107496B5F53832721AAB9B1B8E63ED75" originalValue="Göllner”">Goellner&quot;</normalizedToken>
(1 ex. ZMHB); &quot;Okavango Delta, Moremi Wildlife Res. Third Bridge Campsite, lux 10.3. 1993, 19.14.22S, 23.21.24E, 10.3. 1993 Uhlig&quot; (3 exs. ZMHB); &quot;Chobe NP Savuti Camp 18.33.55S, 24.03.53E, lux 11.3. 1993 Uhlig&quot; (5 exs. ZMHB); &quot;R. Thamalakane 7 mls NE Maun 20.4. 1972/at light&quot; (1 ex. BMNH). - Zimbabwe: &quot;Victoria Falls Zambezi NP -Camp 11-12.12. 1993, 
<geoCoordinate id="D4A76A262C0A93C404F0971A0C602181" direction="south" orientation="latitude" precision="555" value="-17.53">17.53S</geoCoordinate>
, 
<geoCoordinate id="C50685610AD070471DBBBB14221D0729" direction="east" orientation="longitude" precision="555" value="25.49">25.49E</geoCoordinate>
, lux, Uhlig&quot; (1 ex. ZMHB); &quot;S. Rhodesia Pool Lundi 22. N. 1948 JOC.&quot; (6 exs. AMGS); &quot;S. Rhodesia Wankie Reserve water holes 3.9. 1948 / 
<taxonomicName id="7BE36C016B40D3AC662A511F690917A6" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (1 ex. AMGS); &quot;Wankie Game Res. 5 Sept. 1948 JOC. Pools at Robins rest camp / 
<taxonomicName id="D6AFFFAF3F602EAF562BAB6B2C6122F8" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (1 ex. AMGS); &quot;Wankie Game Res. 4 Sept. 1948 JOC.&quot; (4 exs. AMGS, 1 ex. BMNH); &quot;Wankie Reserve water hole Sept. 1948&quot; (13 exs. AMGS); &quot;Wankie Game Reserve Shapi pond 5.IX. 1948 JOC. / 
<taxonomicName id="EC4C2C20E3685E6862365625EDBCCE74" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (9 exs. AMGS); &quot;Wankie Game Reserve Shapi pan 5.IX. 1948&quot; (3 exs. AMGS); &quot;Wankie Reserve Musumu dam 14.IX. 1948 JOC. / 
<taxonomicName id="E2088319D04C56A4B8A4E2BEC5DFF249" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (1 ex. AMGS); &quot;Wankie Game Reserve Sept. waterhole 1948 JOC. / 
<taxonomicName id="D69342B7D6E41244F388CF507E43D59E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (7 exs. AMGS);&quot;5 mi SE Wankie 7.4.1968 Spangler&quot; (34 exs. USNM, 6 exs. MZH); &quot;Marandellas 2. N. 1948 JOC. / 
<taxonomicName id="DC02CF2B6566A6CA8A0BA1DFA5C9AE92" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (1 ex. AMGS); &quot;Gwai 
<pageBreakToken id="AC3E2E0A3A8C5EB756BD23BC959CAF83" pageId="49" pageNumber="50" start="start">River</pageBreakToken>
3.4. 1968 Spangler&quot; (8 exs. USNM); &quot;Shamgani 60 km SW of Gweru 2.12. 1998 F. Kantner leg.&quot; (1 ex. NMPC);&quot;Tongwe 30 km N Beitbridge 7.12. 1998 Kantner leg.&quot; (1 ex. NMPC). - Mozambique: &quot;Mozambique Beira 7. Sept. 1955 JOC.&quot; (12 exs. AMGS); &quot;Moz., Dambo Pan 30.6. 1960&quot; (2 exs. AMGS); &quot;Port. E Afr. Lorenco Marques 3.12. 1948 JOC.&quot; (2 exs. AMGS); &quot;Lourenco Marques Dec. 1948 JOC.&quot; (1 ex. AMGS); &quot;Umbuluzi R. nr. Goba 4.12. 1948 J.O-C.&quot; (2 exs. AMGS). - Swaziland: &quot;Muddy pond nr Stegi 4.12. 1948 / 
<taxonomicName id="702D29851DB76BD9E261EB8C4BB41365" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. Det. JOC.&quot; (1 ex. AMGS). - South Africa: &quot;Kruger Nat. Pk. Mathlakusapan 
<geoCoordinate id="09BE255C377A12CD6C96D476443CFEB3" direction="south" orientation="latitude" precision="555" value="-22.37">22.37S-</geoCoordinate>
<geoCoordinate id="9C7412494D178F4ACA8D26CF34347ABB" direction="east" orientation="longitude" precision="555" value="31.22">31.22E</geoCoordinate>
/ shorewashing 9.2. 1994 
<normalizedToken id="56743DF2A00C2F35FF7E9C30BB938462" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (9 exs. TMSA, 2 exs. MZH); &quot;Kruger Nat. Pk. Skukuza 12 km S, 
<geoCoordinate id="92FA7CEC04C891956254E16ECFF6C3C9" direction="south" orientation="latitude" precision="555" value="-25.04">25.04S-</geoCoordinate>
<geoCoordinate id="2C87454D9423B197259CC360F111035E" direction="east" orientation="longitude" precision="555" value="31.37">31.37E</geoCoordinate>
/ UV light, 6.3. 1996 
<normalizedToken id="F6C169A8766EF39FD03B7D07C200B0F3" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (10 exs. TMSA, 2 exs. MZH); &quot;Kruger Nat. Pk, Skukuza Res. camp 
<geoCoordinate id="CA6FA9319C5B76728C85E997A32D686E" direction="south" orientation="latitude" precision="555" value="-24.59">24.59S-</geoCoordinate>
<geoCoordinate id="E188AF3B874EAD994C1DBC472EDCAA59" direction="east" orientation="longitude" precision="555" value="31.36">31.36E</geoCoordinate>
/ UV light &amp; trap 25.2. 1995 
<normalizedToken id="0B622560415ED4F3FA95021DB4876770" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA); &quot;Kruger Nat. Pk, Skukuza Res. camp 
<geoCoordinate id="310BE3C56F55FADCF0F6307EB5BB6F50" direction="south" orientation="latitude" precision="555" value="-24.59">24.59S-</geoCoordinate>
<geoCoordinate id="CC1A3C68EEFAC66D8AD2D2E15CE85B2D" direction="east" orientation="longitude" precision="555" value="31.35">31.35E</geoCoordinate>
/ UV light 3.3. 1996 
<normalizedToken id="6D76B3DECF6C52426A3524D79EF32F3E" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA); &quot;Kruger Nat. Pk, Skukuza Res. camp 
<geoCoordinate id="C2D237457570EDD3D65E50B142CECBC8" direction="south" orientation="latitude" precision="555" value="-25.0">25.00S-</geoCoordinate>
<geoCoordinate id="77F18AF2FCEEDCD3920773F1BD4787B8" direction="east" orientation="longitude" precision="555" value="31.35">31.35E</geoCoordinate>
/ UV light &amp; trap 19.2. 1995 
<normalizedToken id="1D1476D6E4C956419BE1EA3DCDC6A791" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA); &quot;Kruger Nat. Pk, Skukuza, 40 km S dam 
<geoCoordinate id="F016FC253FD5F34FEADDC7CD9D06E48A" direction="south" orientation="latitude" precision="555" value="-20.04">20.04S-</geoCoordinate>
<geoCoordinate id="A78D2E0FC9AE5D142E8C5A2D7368296A" direction="east" orientation="longitude" precision="555" value="31.36">31.36E</geoCoordinate>
/Shorewashing 23.2. 1995 
<normalizedToken id="10D7A3C0F9FB0FF2C18A235A4547B351" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA); &quot;Kruger Nat. Pk, Skukuza Res. camp 
<geoCoordinate id="BBB5826F80F9C0A42F6E4F8217883348" direction="south" orientation="latitude" precision="555" value="-24.59">24.59S-</geoCoordinate>
<geoCoordinate id="74F65E5486D469EB07AB3DC3E7B19F66" direction="east" orientation="longitude" precision="555" value="31.35">31.35E</geoCoordinate>
/ UV light 7.3. 1996 
<normalizedToken id="69124A5E83051D67AD336603FA76AC1B" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA); &quot;Kruger Nat. Pk, Pafuri research camp 4 km W, 
<geoCoordinate id="7E4E68978974D5EF286B83A184200162" direction="south" orientation="latitude" precision="555" value="-22.25">22.25S-</geoCoordinate>
<geoCoordinate id="57719704E66FC472E22710EA0019E9AF" direction="east" orientation="longitude" precision="555" value="31.09">31.09E</geoCoordinate>
/ 1.2.1994 UV light &amp; trap 
<normalizedToken id="8B0CC38A443CDD2A5966CADC2927B6E6" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (4 exs. TMSA); &quot;Kruger Nat. Pk, Pafuri res. camp, 
<geoCoordinate id="CE7E1F3D39E03482D2FF5525C654C18A" direction="south" orientation="latitude" precision="555" value="-22.25">22.25S-</geoCoordinate>
<geoCoordinate id="8E42A0513757F10AE8324964ADBDC154" direction="east" orientation="longitude" precision="555" value="31.12">31.12E</geoCoordinate>
/30.1. 1994 UV light &amp; trap 
<normalizedToken id="4DCC5C794632A0FFB305F8FB6B285012" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (3 exs. TMSA, 1 ex. MZH); &quot;Kruger Nat. Pk, Pafuri research camp, 
<geoCoordinate id="0E1919FEE9AA9249DFA3CA3EFB838673" direction="south" orientation="latitude" precision="555" value="-22.25">22.25S-</geoCoordinate>
<geoCoordinate id="705D86B1B68A1948766CB70E4F830D96" direction="east" orientation="longitude" precision="555" value="31.1">31.10E</geoCoordinate>
/ 2.2. 1994 UV light &amp; trap 
<normalizedToken id="F246E1A4FB2B9C70C8C069E4670ABD39" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (1 ex. TMSA); &quot;Kruger Nat. Pk Levuvu River 
<geoCoordinate id="1C662D37C2F88A39B1988B57B7FD167A" direction="south" orientation="latitude" precision="555" value="-22.27">22.27S-</geoCoordinate>
<geoCoordinate id="2127FF1F95862AD9EDD7676769AB3617" direction="east" orientation="longitude" precision="555" value="31.1">31.10E</geoCoordinate>
/ 12.2. 1994 shorewashing 
<normalizedToken id="AD68761A30D49EAB79F76B41786A9924" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (4 exs. TMSA); &quot;Kruger Nat. Pk, Malonga springs 
<geoCoordinate id="AD5698D4137749B5A361BD26D91C1C4E" direction="south" orientation="latitude" precision="555" value="-22.36">22.36S-</geoCoordinate>
<geoCoordinate id="2A2DB86835A2B3720AD5161E3270D1FE" direction="east" orientation="longitude" precision="555" value="31.2">31.20E</geoCoordinate>
/ 8.2. 1994 shorewashing 
<normalizedToken id="8E128CE490682015CE47C3BED2BC4BEE" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (1 ex. TMSA); &quot;Kruger Nat. Pk Punda Maria Ngots Dam 
<geoCoordinate id="C5BF4018123D3AFE103DA7B26C5FF49B" direction="south" orientation="latitude" precision="555" value="-21.26">21.26S-</geoCoordinate>
<geoCoordinate id="79DD1979186AF9B9411FAA93739D3427" direction="east" orientation="longitude" precision="555" value="31.14">31.14E</geoCoordinate>
/ 7.2. 1994 shorewashing 
<normalizedToken id="07E4E816B1F9E02241BEAD9F4DCD0F50" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (1 ex. TMSA); &quot;Kruger Nat. Pk Letaba Riv. bel. dam 
<geoCoordinate id="DC968E699AA22C96A2A1AA5D2B8698B5" direction="south" orientation="latitude" precision="555" value="-23.46">23.46S-</geoCoordinate>
<geoCoordinate id="4743ECA304F5D32A9CC4426E412A5449" direction="east" orientation="longitude" precision="555" value="31.3">31.30E</geoCoordinate>
/1.3. 1995 shorewashing 
<normalizedToken id="B4827567D969260121F24D71DE779C39" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
leg.&quot; (1 ex. TMSA); &quot;Transvaal Kruger National Park, Leeu Pan NE Skukuza 1.5. 1951 / 
<taxonomicName id="BC3EF5865F0B26C780A27624B0E136D2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
Gschw. det. J. Omer-Cooper&quot; (1 ex. MZLU); &quot;Trsvl, Waterhole nr Police Picket, KNP. 30.6. 1960&quot; (25 exs. AMGS); &quot;Trsvl, Kumana Pan 24 km S Satara Camp, KNP, 
<geoCoordinate id="212B89165C1775B167B1FCF00DA82640" direction="north" orientation="latitude" precision="55" value="24.61">N-24.610</geoCoordinate>
, 
<geoCoordinate id="5ABF77E6E244B708A81D491B04D79520" direction="east" orientation="longitude" precision="55" value="31.8">E31.800</geoCoordinate>
, 18.6. 1960&quot; (1 ex. AMGS); &quot;Gauteng Tswaing 
<geoCoordinate id="66608E25FF00979927BCBDA46D5083DE" direction="south" orientation="latitude" precision="555" value="-25.24">25.24S-</geoCoordinate>
<geoCoordinate id="3BC48240C6998E41D6CC1C615EC0E79B" direction="east" orientation="longitude" precision="555" value="28.06">28.06E</geoCoordinate>
/ 16.2. 2003 light trap TMSA staff leg.&quot; (2 exs. TMSA); &quot;Trsvl Mmabolela Estate 
<geoCoordinate id="32E6401CBE92DC1E83B4CC2A4395A2D3" direction="south" orientation="latitude" precision="555" value="-22.4">22.40S-</geoCoordinate>
<geoCoordinate id="C38FC5F1BBD1C05EF5B2BDE25AFECB00" direction="east" orientation="longitude" precision="555" value="28.15">28.15E</geoCoordinate>
/ 21.11. 1991 shorew. Limpopo Riv. Klimazewski leg.&quot; (1 ex. TMSA); &quot;Trsvl 5 mi W Warmbad 24-25.2. 1968 Spangler&quot; (1 ex. USNM); &quot;Trsvl, Hartebeespoortdam, 
<geoCoordinate id="76721EC9F7AB70FA57A8F1BBF838DDB3" direction="north" orientation="latitude" precision="55" value="25.73">N-25.730</geoCoordinate>
, 
<geoCoordinate id="94DE4FC7B8BFFD671FED11AB7C3DAA1D" direction="east" orientation="longitude" precision="55" value="27.82">E27.820</geoCoordinate>
, 30.5. 1971 Reavell&quot; (1 ex. AMGS); &quot;N. Prov. Messina Nat. Res. 
<geoCoordinate id="63FE4D0E2CA9803DE240D61946B8F40D" direction="south" orientation="latitude" precision="555" value="-22.21">22.21S-</geoCoordinate>
<geoCoordinate id="F8B24B6B4F8444D0B8CC29BB1B58156A" direction="east" orientation="longitude" precision="555" value="30.03">30.03E</geoCoordinate>
/ 13.12. 2000 light trap 
<normalizedToken id="754B0DFDE8C8A2FD23620EECB20FDFF7" originalValue="Müller">Mueller</normalizedToken>
, Burger leg.&quot; (1 ex. TMSA); &quot;Kw. Natal, Sivai Lagoon 10.3. 1981 Reavell&quot; (1 ex. AMGS); &quot;Zululd. Ndumu Banzi fresh. wat. pan 
<geoCoordinate id="F0FD03ABFE48C5BB313741A3B8702311" direction="south" orientation="latitude" precision="555" value="-26.53">26.53S-</geoCoordinate>
<geoCoordinate id="A002BCA790EEA964EAD11E8D9BF15A3A" direction="east" orientation="longitude" precision="555" value="32.16">32.16E</geoCoordinate>
/ 16.2. 1989 shorewashing 
<normalizedToken id="E075AC3D663C456B7E827CB47893BD15" originalValue="Endrödy">Endroedy</normalizedToken>
&amp; Klimazew&quot; (1 ex. TMSA); &quot;Natal Zululand Mtuba-tuba 23.9. 1947 JOC&quot;. (1 ex. AMGS); &quot;Natal Durban Umgeni 
<normalizedToken id="1E0C12A9538FDD868CB30EAFA8151023" originalValue="Trägårdh”">Traegardh&quot;</normalizedToken>
(1 ex. MZLU); &quot;Zululand Hhu-Hluve 18.IX. 1947&quot; (1 ex. AMGS); &quot;Natal, roadside puddles, ca. 2 km S Mbazwana to Hluhluwe nr Sodwana 9.1. 1997 Turner&quot; (9 exs. NHMB); same data but &quot;5.3. 1997&quot; (6 exs. NHMB); &quot;Natal, Mkuze NP 
<geoCoordinate id="79983106997B217052D12C5533B1DB3E" direction="south" orientation="latitude" precision="555" value="-17.36">17.36S</geoCoordinate>
, 
<geoCoordinate id="CA76A5B1A4CC522D2BA5F4A6F60DF6E1" direction="east" orientation="longitude" precision="555" value="32.13">32.13E</geoCoordinate>
, 2-3.2. 1994 lux 
<pageBreakToken id="BB17CE7116DA3073CF7262F32F27B6DD" pageId="50" pageNumber="51" start="start">Uhlig</pageBreakToken>
(1 ex. ZMHB);&quot;Natal, Waterton Timber Co. N-28.20.5, 
<geoCoordinate id="09E8C5F91A45A6EECC5A19DB1FD99308" direction="east" orientation="longitude" precision="555" value="32.14">E32.14</geoCoordinate>
, at light Atkinson&quot; (5 exs. NHMB).
</paragraph>
</subSubSection>
<subSubSection id="29B027283A903A1F462C9324F993899E" pageId="50" pageNumber="51" type="specimens with uncertain location">
<paragraph id="00C2D3AFFA6F9B950DECABACD27E5D2E" pageId="50" pageNumber="51">Specimens with uncertain location.</paragraph>
<paragraph id="B9D21ED89EDECB723D230BC502E83FB2" pageId="50" pageNumber="51">&quot;Lowrie 17.5. 1955&quot; (1 ex. AMGS); &quot;Kotsch Lu.(?) Kohu&quot; (1 ex. NMW).</paragraph>
</subSubSection>
<subSubSection id="1909F5385D758E8DFFFB18351C58617E" pageId="50" pageNumber="51" type="comments on the synonymy">
<paragraph id="0E648B39E69C861DBB536A790F6C6977" pageId="50" pageNumber="51">Comments on the synonymy.</paragraph>
<paragraph id="F6B9042564B299355FE58F05F5D3EF69" pageId="50" pageNumber="51">
The lectotype of 
<taxonomicName id="5C4234E8E015013C1B8521C4879883DB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
and the holotype of 
<taxonomicName id="5FF20553269894209FD9C5EFE927E103" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus perplexus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="perplexus">Laccophilus perplexus</taxonomicName>
have been examined and compared. Minor difference is present in appearance of the elytral colour pattern and shape of the penis. Differences observed are, however, superficial and clearly falls within the variation exhibited by one species. 
<taxonomicName id="6058DB0F80C20C52B7E8157653206732" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
being the older name is the valid name of the species.
</paragraph>
</subSubSection>
<subSubSection id="129D5B22D2AD3AD97B1E8048DBDD1FC5" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="8EAC451D0BBCCDB341B51603C77CD7B7" pageId="50" pageNumber="51">Diagnosis.</paragraph>
<paragraph id="EB6F154381A24484568B2A189E636EF2" pageId="50" pageNumber="51">
Externally 
<taxonomicName id="F889A7969B2DBC49BAF9CA1E1E1B49C9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
resembles much of 
<taxonomicName id="ADA3849C96F106370487D62EB85928DB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus posticus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="posticus">Laccophilus posticus</taxonomicName>
. Useful diagnostic character is the shape of the penis. In 
<taxonomicName id="BCD1FD447F54D97E44E0D0F02A667F61" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
apical part of penis is less prominent in comparison with rest of the penis when 
<taxonomicName id="CD646EC8A3E0705BBF977EFCB021EB0C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus posticus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="posticus">Laccophilus posticus</taxonomicName>
is characterized by distinctly more prominent apex of penis. The shape of penis separates the two species from all other African 
<taxonomicName id="369B216C5D8AFF5F4612B7F1F8789ABB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="CABB2D7E888E0D547009A8E05F10C4D4" pageId="50" pageNumber="51" type="description">
<paragraph id="4898D5F59AA0602BDEE8547CD78C14A9" pageId="50" pageNumber="51">Description.</paragraph>
<paragraph id="54A9F64A8A072BE0886B483986ACBA45" pageId="50" pageNumber="51">Body length 3.6-4.1 mm, width 1.9-2.2 mm. Pale ferrugineous, with dark ferrugineous, extensive but variable elytral irrorations (Figs 400-401). Sometimes single irroration is formed only by distinct outlines leaving the middle pale coloured. Other extreme is that single irroration is solid and totally dark.</paragraph>
<paragraph id="08E4B4FC705EBAEBAB972F1D42D2C842" pageId="50" pageNumber="51">Head: Frontally pale ferrugineous; posteriorly head becomes gradually slightly darker; at pronotum ferrugineous to pale brown. Head often uniformly pale coloured. Submat, finely microsculptured; reticulation almost uniform. Indistinct fragments of double reticulation discernible (large meshes incomplete). Almost impunctate; at eyes with fine, hardly discernible punctures.</paragraph>
<paragraph id="D9CFEEAB0266FD0E4C20166D531503F3" pageId="50" pageNumber="51">Pronotum: Pale ferrugineous, frontally with a vague dark ferruginous to dark brownish marking. Darker marking on disc sometimes reduced and visible in frontal part of pronotum. Submat, finely microsculptured. Reticulation almost simple; indistinct fragments of coarse reticulation discernible: large meshes incomplete. Almost impunctate; frontally and laterally with scattered, very fine punctures discernible.</paragraph>
<paragraph id="F9FF039D59B3FF4F780EA272D3B4BE3D" pageId="50" pageNumber="51">Elytra: Pale ferrugineous, with variable, dark ferrugineous irrorations. Irrorations sometimes in part reduced; outlines only distinct (Figs 400-401). Submat, finely microsculpture; reticulation of one kind. Discally with sparse, irregular row of very fine punctures. Laterally with sparsely scattered, very fine punctures. Fine lateral furrow formed of fine punctures located somewhat posterior to middle towards elytral apex.</paragraph>
<paragraph id="DD2E6107C330A6F58B7F417165E6D9EC" pageId="50" pageNumber="51">Ventral aspect: Pale ferrugineous. Abdomen in part ferrugineous to dark ferrugineous, with lateral and apical areas paler. Almost impunctate. Extensively very finely microsculptured. Prosternal process slender; apex extended and pointed. Metacoxal plates in anterior half with some transversely located, shallow furrows. Abdomen with sparse but distinct striae. Apical ventrite (Fig. 47).</paragraph>
<paragraph id="E7DD1AFB6F70546A1F7651E2A7F24470" pageId="50" pageNumber="51">Legs: Pro- and mesotarsus rather long and slender. Pro- and mesotarsus provided with distinct suckers.</paragraph>
<paragraph id="98962C1421DC532090C270C632A96618" pageId="50" pageNumber="51">Male genitalia: Extreme apex of penis forms a short, sharp and small extension (Fig. 249).</paragraph>
<paragraph id="B7B92FD13BA83B5A4BFA654719284EBC" pageId="50" pageNumber="51">Female: Apical ventrite (Fig. 48).</paragraph>
</subSubSection>
<subSubSection id="5AE1ACA4BFACAAF94B855E8027F4A3DE" pageId="51" pageNumber="52" type="distribution">
<paragraph id="C8D942BA2C03FA3074C27877606A953B" pageId="51" pageNumber="52">
<pageBreakToken id="8B2B546E4686D133C13BBD566D7E5CD3" pageId="51" pageNumber="52" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph id="3A40FD7B391AC3DB60EF17CB0ED80F65" pageId="51" pageNumber="52">
Gambia, Senegal, Sudan, Ghana, Nigeria, Somalia, Kenya, Tanzania, Namibia, Botswana, Zimbabwe, Mozambique, South Africa (Fig. 535). Additionally, 
<taxonomicName id="D10F6DD89ACF583442AEEBA8B73BC25F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
is recorded from Swaziland (
<bibRefCitation id="5E120A2EA1BCFE3AE4A0A51B44B5277C" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper 1958b</bibRefCitation>
). Record outside Africa but in close neigborhood is Yemen (Socotra) by 
<bibRefCitation id="72DB2C51BFD3B218C86A57CD69EB4504" author="Hajek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="258" pageNumber="259" pagination="63 - 99" title="Adephagous water beetles (Coleoptera: Gyrinidae, Haliplidae, Noteridae, Dytiscidae) of Yemen and Dhofar region (Oman) with description of a new Hyphydrus from Socotra Island." volume="54" year="2014">
<normalizedToken id="9CD208EED66363AFB5224BE05263F829" originalValue="Hájek">Hajek</normalizedToken>
and Reiter (2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="06D1BB186D5B325A7AFF753D8A225BB5" pageId="51" pageNumber="52" type="collecting circumstances">
<paragraph id="D074F9B3A47B57FD8E322E0C89E3C913" pageId="51" pageNumber="52">Collecting circumstances.</paragraph>
<paragraph id="CAF30762F79872FC7B6C69E476EF7187" pageId="51" pageNumber="52">
The biology and habitats of 
<taxonomicName id="9A7C4BF3151DDC82B0A605B2554A45F7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
are not well documented. Scattered observations can be gathered from literature and from collection data written on the labels. Label data give that the species is capable of flying and attracted by light. 
<taxonomicName id="DB2E13B90A567B01CF9F75A1D13B9090" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus continentalis" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="continentalis">Laccophilus continentalis</taxonomicName>
has also been sampled from various water-bodies as ponds and streams. 
<bibRefCitation id="2126796591272817F029B2650C24C9BC" author="Omer-Cooper, Joyce" journalOrPublisher="Sociedade Estudos de Mocambique" pageId="260" pageNumber="261" pagination="19 - 32" title="Some records of Dytiscidae (Col.) from Mozambique." volume="101" year="1956">Omer-Cooper (1956)</bibRefCitation>
reports the species in Mozambique from pools, a ditch and a slow flowing stream with vegetation as 
<taxonomicName id="C2D1543B52732CC560B40BDF720C4564" class="Marchantiopsida" family="Lunulariaceae" genus="Marsilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Marsilia" order="Lunulariales" pageId="51" pageNumber="52" phylum="Marchantiophyta" rank="genus">Marsilia</taxonomicName>
sp., 
<taxonomicName id="3713B2C52CFDCF401A6573C58B491B08" class="Liliopsida" family="Hydrocharitaceae" genus="Lagarosiphon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lagarosiphon" order="Alismatales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="genus">Lagarosiphon</taxonomicName>
sp., 
<taxonomicName id="F6F0CD978C87899D929265792DD803D3" class="Liliopsida" family="Alismataceae" genus="Limnophyton" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Limnophyton" order="Alismatales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="genus">Limnophyton</taxonomicName>
sp., water lilies and duckweed. Also reported from Zimbabwe in water holes, springs and dams used by the game. Additionally, taken from streams e.g. with a pool in the river bed, in ponds with rock and gravel bottom and some mud deposition; blue water lilies and weed growing in the pools. 
<bibRefCitation id="AEA4E65C3D41ACA7DA6BE49E4143D734" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot (1946c)</bibRefCitation>
assumed that the species is a character-species of steppes and savannes.
</paragraph>
</subSubSection>
</treatment>
</document>