293 lines
34 KiB
XML
293 lines
34 KiB
XML
<document id="4222F646FF37A2292470AB0B22DC9E10" ID-CLB-Dataset="4063" ID-DOI="10.12782/specdiv.23.13" ID-GBIF-Dataset="930dbbdf-100c-4d24-8f8d-1e5e7ffe7ddc" ID-ISSN="2189-7301" ID-Zenodo-Dep="4583901" ID-ZooBank="870A4A76-9855-46CE-A0A5-7E7870F4E548" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614893303217" checkinUser="felipe" docAuthor="Kajihara, Hiroshi, Tamura, Kotaro & Tomioka, Shinri" docDate="2018" docId="CF0387C3D501B15FFF2CA025FD06F919" docLanguage="en" docName="SpeciesDiversity.23.1.13-37.pdf" docOrigin="Species Diversity 23 (1)" docSource="http://dx.doi.org/10.12782/specdiv.23.13" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Ototyphlonemertes parmula Diesing 1863, sp. nov." docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="333AFFBBD50EB150FFE0A102FFB1FFE1" masterDocTitle="Histology-free Descriptions for Seven Species of Interstitial Ribbon Worms in the Genus Ototyphlonemertes (Nemertea: Monostilifera) from Vietnam" masterLastPageNumber="37" masterPageNumber="13" pageNumber="28" updateTime="1698932702881" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="8FD0024A4EB03DD08603810C97BFE59B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="06A27DC727660C3AE6EB718C3777576B">
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<mods:title id="18DBE3D09B7FAFC07F2D1F42A6D72F04">Histology-free Descriptions for Seven Species of Interstitial Ribbon Worms in the Genus Ototyphlonemertes (Nemertea: Monostilifera) from Vietnam</mods:title>
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<mods:name id="A23D336DDEEED7B653787AD62D0A490D" type="personal">
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<mods:role id="7FE6B61A133BF1604B2FB48C03782CD7">
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<mods:roleTerm id="A92EC8362E71E09660EDD815970556E6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F75B9E61A8F5C7D6EE047BA90A2B377F">Kajihara, Hiroshi</mods:namePart>
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<mods:affiliation id="F924A1097F6F4FEAF4832DC9FCA90455">Faculty of Science, Hokkaido University, N 10 W 8, Sapporo, Hokkaido 060 - 0810, Japan E-mail: kajihara @ eis. hokudai. ac. jp & Corresponding author</mods:affiliation>
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<mods:nameIdentifier id="065E1DACDF1C0F6856C29751BABE70DC" type="email">kajihara@eis.hokudai.ac.jp</mods:nameIdentifier>
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</mods:name>
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<mods:name id="4752BB8B8419AAA01E9D99AE7B209CEB" type="personal">
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<mods:roleTerm id="9334AB93F4F663935383177B469450AF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="010E63B61236B63080FAADAFA69EFEB1">Tamura, Kotaro</mods:namePart>
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<mods:affiliation id="13062244415749D6850FE597F76E2CE9">Zukosha, W 18 N 1, Obihiro, Hokkaido 080 - 0048, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="4BAD356887701512E079AE8E4A9903AA" type="personal">
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<mods:roleTerm id="6753F3C0357F61D19A6F9BA9D1A56F31">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5FF3B0CAAAA14B90420B5CDE6BEB4A7B">Tomioka, Shinri</mods:namePart>
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<mods:affiliation id="C899C9DE43C7EA0D3155FC89B61A267A">Rishiri Town Museum, Senhoshi, Rishiri Is., Hokkaido 097 - 0311, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="07609848B80E72EAFD722CA57982BAF3">text</mods:typeOfResource>
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<mods:title id="23E895DB6937E73272D646BCAAEC0BA6">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="0B3363E200356E9DCE43E263C5F49B03">
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<mods:date id="99DE339FE45BA57A255B6925AB2689EA">2018</mods:date>
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<mods:number id="61EED11843D002103EAF73C86122A0B7">2018-05-25</mods:number>
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<mods:number id="6B4E157AB2929AE8FC63A10406361143">23</mods:number>
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<mods:start id="86BBE9E2BA71921C0F1EE9FB4D0AF5E7">13</mods:start>
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<treatment id="CF0387C3D501B15FFF2CA025FD06F919" ID-DOI="http://doi.org/10.5281/zenodo.5567322" ID-GBIF-Taxon="188631534" ID-Zenodo-Dep="5567322" LSID="urn:lsid:plazi:treatment:CF0387C3D501B15FFF2CA025FD06F919" httpUri="http://treatment.plazi.org/id/CF0387C3D501B15FFF2CA025FD06F919" lastPageNumber="28" pageId="15" pageNumber="28">
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<subSubSection id="0FB0655ED501B15FFF2CA025FD61FEDE" box="[204,720,295,320]" pageId="15" pageNumber="28" type="nomenclature">
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<paragraph id="471536D5D501B15FFF2CA025FD61FEDE" blockId="15.[204,720,295,353]" box="[204,720,295,320]" pageId="15" pageNumber="28">
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<heading id="1C5D81B9D501B15FFF2CA025FD61FEDE" box="[204,720,295,320]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="28" reason="2">
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<taxonomicName id="80AA4D56D501B15FFF2CA025FDCFFEDE" authority="Kajihara & Tamura & Tomioka, 2018" authorityName="Diesing" authorityYear="1863" box="[204,638,295,320]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="genus" status="sp. nov.">
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<emphasis id="75DEEAC7D501B15FFF2CA025FDCFFEDE" bold="true" box="[204,638,295,320]" pageId="15" pageNumber="28">
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<emphasis id="75DEEAC7D501B15FFF2CA025FE22FEDE" bold="true" box="[204,403,295,319]" italics="true" pageId="15" pageNumber="28">Ototyphlonemertes</emphasis>
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(
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<emphasis id="75DEEAC7D501B15FFE41A025FE4AFEDE" bold="true" box="[417,507,295,319]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
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)
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<emphasis id="75DEEAC7D501B15FFDE9A025FDCFFEDE" bold="true" box="[521,638,295,319]" italics="true" pageId="15" pageNumber="28">tsukagoshii</emphasis>
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EED57BCD501B15FFD64A02AFD61FEDE" box="[644,720,296,319]" pageId="15" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="0FB0655ED501B15FFE43A04AFE49FE80" box="[419,504,328,353]" pageId="15" pageNumber="28" type="description">
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<paragraph id="471536D5D501B15FFE43A04AFE49FE80" blockId="15.[204,720,295,353]" box="[419,504,328,353]" pageId="15" pageNumber="28">
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||
(
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||
<figureCitation id="DF912A50D501B15FFE4AA04AFE41FE80" box="[426,496,328,353]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0FB0655ED501B15FFF55A08BFD5CFE23" pageId="15" pageNumber="28" type="materials_examined">
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<paragraph id="471536D5D501B15FFF55A08BFD5CFE23" blockId="15.[156,768,393,1784]" pageId="15" pageNumber="28">
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||
<emphasis id="75DEEAC7D501B15FFF55A08BFE39FE40" bold="true" box="[181,392,393,417]" pageId="15" pageNumber="28">Material examined.</emphasis>
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<materialsCitation id="F7C23C88D501B15FFE41A088FD58FE23" ID-GBIF-Occurrence="3383262303" collectionCode="ICHUM" pageId="15" pageNumber="28" specimenCode="ICHUM 5371, ICHUM 5372" specimenCount="2" typeStatus="holotype">
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Two specimens, both from Dam Ngoai;
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<specimenCode id="170C9EAED501B15FFF04A0A8FEC1FE23" box="[228,368,426,450]" collectionCode="ICHUM" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="15" pageNumber="28">ICHUM 5371</specimenCode>
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||
(
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||
<typeStatus id="98118877D501B15FFE9DA0A8FE6AFE23" box="[381,475,426,450]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
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||
);
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||
<specimenCode id="170C9EAED501B15FFE0CA0A8FDC9FE23" box="[492,632,426,450]" collectionCode="ICHUM" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="15" pageNumber="28">ICHUM 5372</specimenCode>
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||
(
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||
<typeStatus id="98118877D501B15FFD64A0A8FD53FE23" box="[644,738,426,450]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0FB0655ED501B15FFF55A0C8FE16FAEE" pageId="15" pageNumber="28" type="description">
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||
<paragraph id="471536D5D501B15FFF55A0C8FE16FAEE" blockId="15.[156,768,393,1784]" pageId="15" pageNumber="28">
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<emphasis id="75DEEAC7D501B15FFF55A0C8FE88FE03" bold="true" box="[181,313,458,482]" pageId="15" pageNumber="28">Description.</emphasis>
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||
Body length
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||
<quantity id="80529B30D501B15FFE36A0C8FD93FE03" box="[470,546,458,482]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="28" unit="mm" value="2.5">2.5 mm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFDD0A0C9FD3AFE02" box="[560,651,459,483]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
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) to 4.0 mm (
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||
<typeStatus id="98118877D501B15FFF43A0E9FEB0FDE2" box="[163,257,491,515]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
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), width
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||
<quantity id="80529B30D501B15FFEBAA0E9FE14FDE2" box="[346,421,491,515]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="28" unit="mm" value="0.1">0.1 mm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFE56A0E9FDA3FDE2" box="[438,530,491,515]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
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) to
|
||
<quantity id="80529B30D501B15FFDA0A0E9FD3BFDE2" box="[576,650,491,515]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="28" unit="mm" value="0.2">0.2 mm</quantity>
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||
(
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||
<typeStatus id="98118877D501B15FFD7BA0E9FD49FDE2" box="[667,760,491,515]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
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) (
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||
<emphasis id="75DEEAC7D501B15FFF45A30EFF03FDC2" box="[165,178,524,547]" italics="true" pageId="15" pageNumber="28">n</emphasis>
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||
=2). Epidermis whitish (
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||
<figureCitation id="DF912A50D501B15FFE53A30EFDBEFDC5" box="[435,527,524,548]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11A</figureCitation>
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); cephalic furrow postcerebrally; tissues around brain (and even in precerebral region in one of two specimens examined) reddish (
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<figureCitation id="DF912A50D501B15FFD43A34FFF1EFD64" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11B, C</figureCitation>
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||
); cirri present in both anterior and posterior ends of body; cephalic cirrus formula (
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||
<emphasis id="75DEEAC7D501B15FFE7FA38CFE06FD47" box="[415,439,654,678]" italics="true" pageId="15" pageNumber="28">cf.</emphasis>
|
||
<bibRefCitation id="233B4B24D501B15FFE5EA38CFDC7FD47" author="Norenburg, J. L." box="[446,630,654,678]" pageId="15" pageNumber="28" pagination="87 - 92" refId="ref10887" refString="Norenburg, J. L. 1988 b. Remarks on marine interstitial nemertines and key to the species. Hydrobiologia 156: 87 - 92." type="journal article" year="1988">Norenburg 1988b</bibRefCitation>
|
||
) observed in two specimens: A=2, B=0, C=8, D=?, E=?, tail=
|
||
<quantity id="80529B30D501B15FFD27A3ACFD4EFD27" box="[711,767,686,710]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="15" pageNumber="28" unit="in" value="12.0">12 in</quantity>
|
||
<typeStatus id="98118877D501B15FFF7CA3CDFF46FD06" box="[156,247,719,743]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
; A=2, B=0, C=5+6, D=6+10, E=3, tail=
|
||
<quantity id="80529B30D501B15FFD29A3CDFD4EFD06" box="[713,767,719,743]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="28" unit="in" value="10.0">10 in</quantity>
|
||
<typeStatus id="98118877D501B15FFF7CA3F2FF46FCE9" box="[156,247,752,776]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
. Foregut–intestine junction indistinct. Statolith polygranular (
|
||
<figureCitation id="DF912A50D501B15FFECDA212FE14FCC9" box="[301,421,784,808]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11D, E</figureCitation>
|
||
),
|
||
<quantity id="80529B30D501B15FFE59A212FE4DFCC9" box="[441,508,784,808]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="28" unit="mm" value="9.5">9.5 µm</quantity>
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||
in diameter (
|
||
<emphasis id="75DEEAC7D501B15FFD6CA213FD28FCC9" box="[652,665,785,808]" italics="true" pageId="15" pageNumber="28">n</emphasis>
|
||
=2), consisting of 6 granules; statocyst
|
||
<quantity id="80529B30D501B15FFE2FA232FDAEFCA8" box="[463,543,816,841]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="15" pageNumber="28" unit="mm" value="23.5">23.5 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFDCCA232FD38FCA9" box="[556,649,816,840]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
) to
|
||
<quantity id="80529B30D501B15FFD50A233FCB1FCA8" box="[688,768,817,841]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="15" pageNumber="28" unit="mm" value="28.4">28.4 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFF43A253FF4EFC88" box="[163,255,849,873]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
) in diameter (
|
||
<emphasis id="75DEEAC7D501B15FFE7AA250FE16FC88" box="[410,423,850,873]" italics="true" pageId="15" pageNumber="28">n</emphasis>
|
||
=2). Proboscis anterior chamber with wart-like papillae (
|
||
<figureCitation id="DF912A50D501B15FFE40A273FE4AFC6B" box="[416,507,881,906]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11F</figureCitation>
|
||
); diaphragm short (
|
||
<figureCitation id="DF912A50D501B15FFD38A273FF75FC4B" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11F</figureCitation>
|
||
); gland secretion ring situated posterior to hind end of basis; accessory-stylet pouches two, each containing 2–3 accessory stylets directing both forward and backward; central stylet (as well as accessory stylets) spiral (
|
||
<figureCitation id="DF912A50D501B15FFD96A2F6FD43FBED" box="[630,754,1012,1036]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11G, H</figureCitation>
|
||
), 35.0 µm (
|
||
<typeStatus id="98118877D501B15FFF18A516FEE5FBCD" box="[248,340,1044,1068]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
) to
|
||
<quantity id="80529B30D501B15FFE9BA516FE7BFBCC" box="[379,458,1044,1069]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="15" pageNumber="28" unit="mm" value="40.2">40.2 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFE37A516FD82FBCD" box="[471,563,1044,1068]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
) long; basis
|
||
<quantity id="80529B30D501B15FFD50A516FD4EFBCC" box="[688,767,1044,1069]" metricMagnitude="-2" metricUnit="m" metricValue="3.71" pageId="15" pageNumber="28" unit="mm" value="37.1">37.1 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFF43A537FF4EFBAC" box="[163,255,1077,1101]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
) to
|
||
<quantity id="80529B30D501B15FFECAA537FEC8FBAC" box="[298,377,1077,1101]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="15" pageNumber="28" unit="mm" value="40.2">40.2 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFE68A537FE54FBAC" box="[392,485,1077,1101]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
) long,
|
||
<quantity id="80529B30D501B15FFDCEA537FDC0FBAC" box="[558,625,1077,1101]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="15" pageNumber="28" unit="mm" value="8.3">8.3 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFD60A537FD6DFBAC" box="[640,732,1077,1101]" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
) to
|
||
<quantity id="80529B30D501B15FFF7CA557FF5AFB8F" box="[156,235,1109,1134]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="15" pageNumber="28" unit="mm" value="11.2">11.2 µm</quantity>
|
||
(
|
||
<typeStatus id="98118877D501B15FFF1BA557FEE9FB8C" box="[251,344,1109,1133]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
) wide (
|
||
<emphasis id="75DEEAC7D501B15FFE49A554FE07FB8C" box="[425,438,1110,1133]" italics="true" pageId="15" pageNumber="28">n</emphasis>
|
||
=2), with each ridge twisting 3 times; basis length/width ratio 3.59 (
|
||
<typeStatus id="98118877D501B15FFDECA574FDD8FB6F" box="[524,617,1142,1166]" pageId="15" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
) to 4.47 (
|
||
<typeStatus id="98118877D501B15FFD27A574FF7AFB4F" pageId="15" pageNumber="28" type="paratype">paratype</typeStatus>
|
||
); middle chamber bulbous (
|
||
<figureCitation id="DF912A50D501B15FFE0EA594FDF7FB4F" box="[494,582,1174,1198]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11F</figureCitation>
|
||
); posterior chamber opaque, with specialized anterior part (
|
||
<figureCitation id="DF912A50D501B15FFDB6A5B5FD7CFB2E" box="[598,717,1207,1231]" captionStart="Fig" captionStartId="14.[156,187,1607,1629]" captionTargetBox="[188,1398,206,1567]" captionTargetId="figure-214@14.[187,1399,205,1568]" captionTargetPageId="14" captionText="Fig. 11. Ototyphlonemertes (parmula) tsukagoshii sp. nov. A, Entire body; B, C, composite photomicrographs of squeezed specimens; D, E, statoliths; F, partially everted proboscis; G, H, central stylets and bases. A, B, D, F, G, ICHUM 5371 (holotype); C, E, H, ICHUM 5372 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.4583939" httpUri="https://zenodo.org/record/4583939/files/figure.png" pageId="15" pageNumber="28">Fig. 11B, C</figureCitation>
|
||
). Intestinal diverticula deep. Posterior adhesive plate containing cells with orange contents.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FB0655ED501B15FFF55A41AFE2FFAB0" pageId="15" pageNumber="28" type="distribution">
|
||
<paragraph id="471536D5D501B15FFF55A41AFE2FFAB0" blockId="15.[156,768,393,1784]" pageId="15" pageNumber="28">
|
||
<emphasis id="75DEEAC7D501B15FFF55A41AFEF0FAD1" bold="true" box="[181,321,1304,1328]" pageId="15" pageNumber="28">Distribution.</emphasis>
|
||
So far only recorded from the
|
||
<typeStatus id="98118877D501B15FFD73A41BFD0EFAD0" box="[659,703,1305,1329]" pageId="15" pageNumber="28">type</typeStatus>
|
||
locality, Dam Ngoai,
|
||
<collectingCountry id="3FBD7645D501B15FFEA0A43BFE2BFAB0" box="[320,410,1337,1361]" name="Vietnam" pageId="15" pageNumber="28">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FB0655ED501B15FFF55A458FDA6FA32" pageId="15" pageNumber="28" type="etymology">
|
||
<paragraph id="471536D5D501B15FFF55A458FDA6FA32" blockId="15.[156,768,393,1784]" pageId="15" pageNumber="28">
|
||
<emphasis id="75DEEAC7D501B15FFF55A458FE9CFA93" bold="true" box="[181,301,1370,1394]" pageId="15" pageNumber="28">Etymology.</emphasis>
|
||
The specific name is a noun in the genitive case, after the Japanese ostracode systematist Professor Akira Tsukagoshi at
|
||
<collectingRegion id="856EF837D501B15FFE93A498FE61FA53" box="[371,464,1434,1458]" country="Japan" name="Shizuoka" pageId="15" pageNumber="28">Shizuoka</collectingRegion>
|
||
University,
|
||
<collectingCountry id="3FBD7645D501B15FFDAEA499FD38FA53" box="[590,649,1435,1458]" name="Japan" pageId="15" pageNumber="28">Japan</collectingCountry>
|
||
, who organized our sampling trip to Phu Quoc.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FB0655ED501B15FFF55A4DEFD06F919" pageId="15" pageNumber="28" type="discussion">
|
||
<paragraph id="471536D5D501B15FFF55A4DEFD06F919" blockId="15.[156,768,393,1784]" pageId="15" pageNumber="28">
|
||
<emphasis id="75DEEAC7D501B15FFF55A4DEFEA8FA15" bold="true" box="[181,281,1500,1524]" pageId="15" pageNumber="28">Remarks.</emphasis>
|
||
In the phylogenetic tree,
|
||
<taxonomicName id="80AA4D56D501B15FFDA3A4D9FD4EFA12" authorityName="Diesing" authorityYear="1863" box="[579,767,1499,1523]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="genus">
|
||
<emphasis id="75DEEAC7D501B15FFDA3A4D9FD4EFA12" box="[579,767,1499,1523]" italics="true" pageId="15" pageNumber="28">Ototyphlonemertes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="80AA4D56D501B15FFF45A4FEFF4BF9F5" authorityName="Correa" authorityYear="1950" box="[165,250,1532,1556]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="parmula">
|
||
<emphasis id="75DEEAC7D501B15FFF45A4FEFF4BF9F5" box="[165,250,1532,1556]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName id="80AA4D56D501B15FFEEEA4FEFECDF9F5" authority="Kajihara & Tamura & Tomioka, 2018" authorityName="Kajihara & Tamura & Tomioka" authorityYear="2018" box="[270,380,1532,1556]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="tsukagoshii" status="sp. nov.">
|
||
<emphasis id="75DEEAC7D501B15FFEEEA4FEFECDF9F5" box="[270,380,1532,1556]" italics="true" pageId="15" pageNumber="28">tsukagoshii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EED57BCD501B15FFE67A4FFFE68F9F5" box="[391,473,1533,1556]" pageId="15" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was sister to
|
||
<emphasis id="75DEEAC7D501B15FFD95A4FEFD3CF9F2" box="[629,653,1532,1555]" italics="true" pageId="15" pageNumber="28">O.</emphasis>
|
||
(
|
||
<taxonomicName id="80AA4D56D501B15FFD41A4FEFD47F9F5" authorityName="Correa" authorityYear="1950" box="[673,758,1532,1556]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="parmula">
|
||
<emphasis id="75DEEAC7D501B15FFD41A4FEFD47F9F5" box="[673,758,1532,1556]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName id="80AA4D56D501B15FFF7CA71FFF41F9D4" authority="Kajihara & Tamura & Tomioka, 2018" authorityName="Kajihara & Tamura & Tomioka" authorityYear="2018" box="[156,240,1565,1589]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="nakaoae" status="sp. nov.">
|
||
<emphasis id="75DEEAC7D501B15FFF7CA71FFF41F9D4" box="[156,240,1565,1589]" italics="true" pageId="15" pageNumber="28">nakaoae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EED57BCD501B15FFF18A71FFEF7F9D5" box="[248,326,1565,1588]" pageId="15" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="DF912A50D501B15FFEB4A71FFE23F9D4" box="[340,402,1565,1589]" captionStart="Fig" captionStartId="3.[156,187,2023,2045]" captionTargetBox="[228,1359,206,1985]" captionTargetId="figure-7@3.[227,1360,205,1985]" captionTargetPageId="3" captionText="Fig. 2. Part of the phylogenetic tree resulting from the maximum-likelihood analysis of COI (ln L=−15264.650301), along with results of three species-delimitation analyses, ABGD, PTP, and TCS, as indicated by vertical bars. Numbers near nodes indicate maximum-likelihood bootstrap support value." figureDoi="http://doi.org/10.5281/zenodo.4583907" httpUri="https://zenodo.org/record/4583907/files/figure.png" pageId="15" pageNumber="28">Fig. 2</figureCitation>
|
||
).
|
||
<taxonomicName id="80AA4D56D501B15FFE46A71FFDD3F9D4" authorityName="Diesing" authorityYear="1863" box="[422,610,1565,1589]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="genus">
|
||
<emphasis id="75DEEAC7D501B15FFE46A71FFDD3F9D4" box="[422,610,1565,1589]" italics="true" pageId="15" pageNumber="28">Ototyphlonemertes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="80AA4D56D501B15FFD93A71FFD79F9D4" authorityName="Correa" authorityYear="1950" box="[627,712,1565,1589]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="parmula">
|
||
<emphasis id="75DEEAC7D501B15FFD93A71FFD79F9D4" box="[627,712,1565,1589]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName id="80AA4D56D501B15FFD39A71FFF5DF9B4" authorityName="Kajihara & Tamura & Tomioka" authorityYear="2018" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="tsukagoshii">
|
||
<emphasis id="75DEEAC7D501B15FFD39A71FFF5DF9B4" italics="true" pageId="15" pageNumber="28">tsukagoshii</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="75DEEAC7D501B15FFE9BA73FFE22F9B5" box="[379,403,1597,1620]" italics="true" pageId="15" pageNumber="28">O.</emphasis>
|
||
(
|
||
<taxonomicName id="80AA4D56D501B15FFE44A73FFE48F9B4" authorityName="Correa" authorityYear="1950" box="[420,505,1597,1621]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="parmula">
|
||
<emphasis id="75DEEAC7D501B15FFE44A73FFE48F9B4" box="[420,505,1597,1621]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName id="80AA4D56D501B15FFDECA73FFDD1F9B4" authorityName="Kajihara & Tamura & Tomioka" authorityYear="2018" box="[524,608,1597,1621]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="nakaoae">
|
||
<emphasis id="75DEEAC7D501B15FFDECA73FFDD1F9B4" box="[524,608,1597,1621]" italics="true" pageId="15" pageNumber="28">nakaoae</emphasis>
|
||
</taxonomicName>
|
||
by the smaller body length (2.5–4.0 mm
|
||
<emphasis id="75DEEAC7D501B15FFE45A75CFE71F994" box="[421,448,1630,1653]" italics="true" pageId="15" pageNumber="28">vs.</emphasis>
|
||
<quantity id="80529B30D501B15FFE27A75CFDF3F997" box="[455,578,1630,1654]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="6.2" metricValueMin="4.1" pageId="15" pageNumber="28" unit="mm" value="5.15" valueMax="6.2" valueMin="4.1">4.1–6.2 mm</quantity>
|
||
), fewer number of statolith granules (6
|
||
<emphasis id="75DEEAC7D501B15FFE91A77DFE3AF977" box="[369,395,1663,1686]" italics="true" pageId="15" pageNumber="28">vs.</emphasis>
|
||
7–8), and greater number of styletridge twists (3
|
||
<emphasis id="75DEEAC7D501B15FFEA3A79DFEEFF957" box="[323,350,1695,1718]" italics="true" pageId="15" pageNumber="28">vs.</emphasis>
|
||
1.5–2). Also,
|
||
<emphasis id="75DEEAC7D501B15FFE1BA79DFDA3F957" box="[507,530,1695,1718]" italics="true" pageId="15" pageNumber="28">O.</emphasis>
|
||
(
|
||
<taxonomicName id="80AA4D56D501B15FFDC7A79DFDCDF956" authorityName="Correa" authorityYear="1950" box="[551,636,1695,1719]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="parmula">
|
||
<emphasis id="75DEEAC7D501B15FFDC7A79DFDCDF956" box="[551,636,1695,1719]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName id="80AA4D56D501B15FFD71A79DFD4EF956" authorityName="Kajihara & Tamura & Tomioka" authorityYear="2018" box="[657,767,1695,1719]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="tsukagoshii">
|
||
<emphasis id="75DEEAC7D501B15FFD71A79DFD4EF956" box="[657,767,1695,1719]" italics="true" pageId="15" pageNumber="28">tsukagoshii</emphasis>
|
||
</taxonomicName>
|
||
tends to be more reddish in the external appearance, especially in the brain region, than
|
||
<emphasis id="75DEEAC7D501B15FFE33A7E2FE5AF916" box="[467,491,1760,1783]" italics="true" pageId="15" pageNumber="28">O.</emphasis>
|
||
(
|
||
<taxonomicName id="80AA4D56D501B15FFE19A7E2FDFFF919" authorityName="Correa" authorityYear="1950" box="[505,590,1760,1784]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="parmula">
|
||
<emphasis id="75DEEAC7D501B15FFE19A7E2FDFFF919" box="[505,590,1760,1784]" italics="true" pageId="15" pageNumber="28">parmula</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<emphasis id="75DEEAC7D501B15FFDBCA7E2FD06F919" box="[604,695,1760,1784]" italics="true" pageId="15" pageNumber="28">
|
||
<taxonomicName id="80AA4D56D501B15FFDBCA7E2FD02F919" authorityName="Kajihara & Tamura & Tomioka" authorityYear="2018" box="[604,691,1760,1784]" class="Enopla" family="Ototyphlonemertidae" genus="Ototyphlonemertes" kingdom="Animalia" order="Monostilifera" pageId="15" pageNumber="28" phylum="Nemertea" rank="species" species="nakaoae">nakaoae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |