treatments-xml/data/52/71/87/527187C4100BBC690650FC7583B9FD48.xml

<document id="C8BC93AD405A05164E523B5231FB48BA" ID-DOI="10.5281/zenodo.173868" ID-GBIF-Dataset="3057cbcd-031a-4a23-b2df-c3f768da9262" ID-ISSN="1175­5326" ID-Zenodo-Dep="173868" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459785867585" checkinUser="plazi" docAuthor="González, Guillermo &amp; Vandenberg, Natalia J." docDate="2006" docId="527187C4100BBC690650FC7583B9FD48" docLanguage="en" docName="zt01311p050.pdf.imd" docOrigin="Zootaxa 1311" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cycloneda lacrimosa Gonzalez &amp; Vandenberg 2006, new species" docType="treatment" docVersion="14" lastPageNumber="23" masterDocId="AE48FFBC100CBC630758FFE38774FFD6" masterDocTitle="Review of lady beetles in the Cycloneda germainii species complex (Coleoptera; Coccinellidae: Coccinellinae: Coccinellini) with descriptions of new and unusual species from Chile and surrounding countries" masterLastPageNumber="50" masterPageNumber="13" pageNumber="20" updateTime="1698218499776" updateUser="plazi">
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<mods:title id="0DBD5FD2230F38C242487DF75849CD31">Review of lady beetles in the Cycloneda germainii species complex (Coleoptera; Coccinellidae: Coccinellinae: Coccinellini) with descriptions of new and unusual species from Chile and surrounding countries</mods:title>
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<mods:namePart id="4A6E79228088B2FFC6F5708749284CCC">González, Guillermo</mods:namePart>
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<mods:namePart id="412245FCF294569571E5AE0CF1C137CB">Vandenberg, Natalia J.</mods:namePart>
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<mods:date id="A367E4962825DDE949740FB8D13253BB">2006</mods:date>
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<treatment id="527187C4100BBC690650FC7583B9FD48" ID-DOI="http://doi.org/10.5281/zenodo.6257504" ID-GBIF-Taxon="119349074" ID-Zenodo-Dep="6257504" LSID="urn:lsid:plazi:treatment:527187C4100BBC690650FC7583B9FD48" httpUri="http://treatment.plazi.org/id/527187C4100BBC690650FC7583B9FD48" lastPageId="10" lastPageNumber="23" pageId="7" pageNumber="20">
<subSubSection id="92C26559100BBC640650FC758468FC05" pageId="7" pageNumber="20" type="nomenclature">
<paragraph id="DA6736D2100BBC640650FC7584AAFC66" blockId="7.[264,990,918,979]" box="[264,990,918,944]" pageId="7" pageNumber="20">
<heading id="812F81BE100BBC640650FC7584AAFC66" bold="true" box="[264,990,918,944]" fontSize="11" level="1" pageId="7" pageNumber="20" reason="1">
<emphasis id="E8ACEAC0100BBC640650FC7584AAFC66" bold="true" box="[264,990,918,944]" pageId="7" pageNumber="20">
<taxonomicName id="1DD84D51100BBC640650FC758435FC66" authority="Gonzalez &amp; Vandenberg" authorityName="Gonzalez &amp; Vandenberg" authorityYear="2006" box="[264,833,918,944]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="lacrimosa" status="sp. nov.">
<emphasis id="E8ACEAC0100BBC640650FC758571FC66" bold="true" box="[264,517,918,944]" italics="true" pageId="7" pageNumber="20">Cycloneda lacrimosa</emphasis>
González &amp; Vandenberg
</taxonomicName>
, 
<taxonomicNameLabel id="F39F57BB100BBC640416FC7584AAFC66" box="[846,990,918,944]" pageId="7" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DA6736D2100BBC640650FC5A8468FC05" blockId="7.[264,990,918,979]" box="[264,796,953,979]" pageId="7" pageNumber="20">
(
<figureCitation id="42E32A57100BBC640649FC5A860DFC05" box="[273,377,953,979]" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González &amp; Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg &amp; González, new species, female, holotype (3.3 mm); C, C. patagonica González &amp; Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="7" pageNumber="20">Figs. 1A</figureCitation>
; 
<figureCitation id="42E32A57100BBC6406DEFC5A86A4FC05" box="[390,464,953,979]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González &amp; Vandenberg, new species, paratype; B, C. disconsolata Vandenberg &amp; González, new species, holotype; C, C. patagonica González &amp; Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González &amp; Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="7" pageNumber="20">3A, H</figureCitation>
; 
<figureCitation id="42E32A57100BBC640685FC5A8559FC05" box="[477,557,953,979]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González &amp; Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg &amp; González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="7" pageNumber="20">4M–N</figureCitation>
; 
<figureCitation id="42E32A57100BBC640561FC5A8514FC05" box="[569,608,953,979]" captionStart="FIGURE 6" captionStartId="18.[264,368,1741,1765]" captionTargetBox="[266,1322,327,1740]" captionTargetId="figure@18.[264,1323,305,1746]" captionTargetPageId="18" captionText="FIGURE 6. Male genitalia of Cycloneda species (above, ventral and right lateral views of phallobase; below, right lateral view of sipho (=aedeagus)): A, C. lacrimosa González &amp; Vandenberg, new species, holotype; B, C. patagonica González &amp; Vandenberg, new species, holotype; C, C. germainii (Crotch); D, C. eryngii (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924357" httpUri="https://zenodo.org/record/3924357/files/figure.png" pageId="7" pageNumber="20">6A</figureCitation>
; 
<figureCitation id="42E32A57100BBC640535FC5A85E0FC05" box="[621,660,953,979]" captionStart="FIGURE 8" captionStartId="25.[264,368,1723,1747]" captionTargetBox="[294,1301,321,1698]" captionTargetId="figure@25.[284,1309,311,1730]" captionTargetPageId="25" captionText="FIGURE 8. Female genitalia of Cycloneda species (oriented with anterior end pointing left): A, C. lacrimosa González &amp; Vandenberg, new species, allotype; B, C. disconsolata Vandenberg &amp; González, new species, holotype; C, C. patagonica González &amp; Vandenberg, new species; D, Cycloneda germainii (Crotch); E, C. eryngii (Mulsant); F, C. sicardi (Brèthes), specimen from Tacuil, Salta; G, C. boliviana (Mulsant), specimen from Molinos, Salta." figureDoi="http://doi.org/10.5281/zenodo.3924361" httpUri="https://zenodo.org/record/3924361/files/figure.png" pageId="7" pageNumber="20">8A</figureCitation>
; 
<figureCitation id="42E32A57100BBC6405F8FC5A859DFC05" box="[672,745,953,979]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: A­C, C. lacrimosa Gonzlez &amp; Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez &amp; Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="7" pageNumber="20">9A–C</figureCitation>
; 
<figureCitation id="42E32A57100BBC6405ADFC5A8467FC05" box="[757,787,953,979]" captionStart="FIGURE 10" captionStartId="27.[264,368,1868,1892]" captionTargetBox="[287,1302,680,1870]" captionTargetId="figure@27.[287,1312,671,1870]" captionTargetPageId="27" captionText="FIGURE 10. Distributions of Cycloneda species." figureDoi="http://doi.org/10.5281/zenodo.3924342" httpUri="https://zenodo.org/record/3924342/files/figure.png" pageId="7" pageNumber="20">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="92C26559100BBC640650FBEA846FFA0D" pageId="7" pageNumber="20" type="diagnosis">
<paragraph id="DA6736D2100BBC640650FBEA846FFA0D" blockId="7.[264,1323,1033,1899]" pageId="7" pageNumber="20">
<emphasis id="E8ACEAC0100BBC640650FBEA86FEFBF5" bold="true" box="[264,394,1033,1059]" pageId="7" pageNumber="20">Diagnosis:</emphasis>
Distinguished from other 
<taxonomicName id="1DD84D51100BBC64059FFBEA8430FBF4" authorityName="Crotch" authorityYear="1871" box="[711,836,1033,1058]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="E8ACEAC0100BBC64059FFBEA8430FBF4" box="[711,836,1033,1058]" italics="true" pageId="7" pageNumber="20">Cycloneda</emphasis>
</taxonomicName>
species by the bicolored or very weakly tricolored elytra with a large ocher­colored teardrop­shaped sutural macula, and by the robust convex oblong body form (
<figureCitation id="42E32A57100BBC6405CCFBBA858FFBA5" box="[660,763,1113,1139]" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González &amp; Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg &amp; González, new species, female, holotype (3.3 mm); C, C. patagonica González &amp; Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="7" pageNumber="20">Figs. 1A</figureCitation>
, 
<figureCitation id="42E32A57100BBC64045FFBBA845BFBA5" box="[775,815,1113,1139]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González &amp; Vandenberg, new species, paratype; B, C. disconsolata Vandenberg &amp; González, new species, holotype; C, C. patagonica González &amp; Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González &amp; Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="7" pageNumber="20">3A</figureCitation>
). This species appears to be closely related to 
<taxonomicName id="1DD84D51100BBC64067FFB6286ABFB4C" authority="González &amp; Vandenberg, 2006" authorityName="González &amp; Vandenberg" authorityYear="2006" box="[295,479,1153,1178]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="disconsolata" status="sp. nov.">
<emphasis id="E8ACEAC0100BBC64067FFB6286ABFB4C" box="[295,479,1153,1178]" italics="true" pageId="7" pageNumber="20">C. disconsolata</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="F39F57BB100BBC6406B7FB6285F4FB4D" box="[495,640,1153,1179]" pageId="7" pageNumber="20" rank="species">new species</taxonomicNameLabel>
, but the latter has the elytron distinctly tricolored and the dark discal figure extended obliquely toward the humeral angle. If females of the two species are compared, 
<taxonomicName id="1DD84D51100BBC64054DFB3285DAFB3C" authorityName="Gonzalez &amp; Vandenberg" authorityYear="2006" box="[533,686,1233,1258]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="lacrimosa">
<emphasis id="E8ACEAC0100BBC64054DFB3285DAFB3C" box="[533,686,1233,1258]" italics="true" pageId="7" pageNumber="20">C. lacrimosa</emphasis>
</taxonomicName>
is distinctly broader. 
<taxonomicName id="1DD84D51100BBC6404E1FB3283C0FB3C" authorityName="Gonzalez &amp; Vandenberg" authorityYear="2006" box="[953,1204,1233,1258]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="lacrimosa">
<emphasis id="E8ACEAC0100BBC6404E1FB3283C0FB3C" box="[953,1204,1233,1258]" italics="true" pageId="7" pageNumber="20">Cycloneda lacrimosa</emphasis>
</taxonomicName>
is similar in color and convexity to 
<taxonomicName id="1DD84D51100BBC64056DFB1A845CFAC5" authority="(Mulsant)" baseAuthorityName="Mulsant" box="[565,808,1273,1299]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="eryngii">
<emphasis id="E8ACEAC0100BBC64056DFB1A85DFFAC4" box="[565,683,1273,1298]" italics="true" pageId="7" pageNumber="20">C. eryngii</emphasis>
(Mulsant)
</taxonomicName>
, but the latter has an ovoid body form, with the lateral elytral margin more evenly arcuate (
<figureCitation id="42E32A57100BBC640461FAC284E6FAED" box="[825,914,1313,1339]" captionStart="FIGURE 2" captionStartId="5.[264,368,1403,1427]" captionTargetBox="[302,1293,302,1387]" captionTargetId="figure@5.[264,1323,280,1401]" captionTargetPageId="5" captionText="FIGURE 2. Habitus views of Cycloneda species: A, C. eryngii (Mulsant), female, common form (3.8 mm); B, C. sicardi (Brèthes), female from Salta, Argentina (3.8 mm); C, C. boliviana (Mulsant), female, holotype (4.6 mm)." httpUri="https://zenodo.org/record/173870/files/figure.png" pageId="7" pageNumber="20">Fig 2A</figureCitation>
), and the elytron bearing a pair of transverse discal maculae that are separated at the midline, or joined only at the inner edges (
<figureCitation id="42E32A57100BBC640606FA928684FA5D" box="[350,496,1393,1419]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González &amp; Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg &amp; González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="7" pageNumber="20">Figs. 4A–H</figureCitation>
). The male genitalia of 
<taxonomicName id="1DD84D51100BBC640478FA9284CFFA5C" authorityName="Gonzalez &amp; Vandenberg" authorityYear="2006" box="[800,955,1393,1418]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="lacrimosa">
<emphasis id="E8ACEAC0100BBC640478FA9284CFFA5C" box="[800,955,1393,1418]" italics="true" pageId="7" pageNumber="20">C. lacrimosa</emphasis>
</taxonomicName>
(
<figureCitation id="42E32A57100BBC640496FA92835AFA5D" box="[974,1070,1393,1419]" captionStart="FIGURE 6" captionStartId="18.[264,368,1741,1765]" captionTargetBox="[266,1322,327,1740]" captionTargetId="figure@18.[264,1323,305,1746]" captionTargetPageId="18" captionText="FIGURE 6. Male genitalia of Cycloneda species (above, ventral and right lateral views of phallobase; below, right lateral view of sipho (=aedeagus)): A, C. lacrimosa González &amp; Vandenberg, new species, holotype; B, C. patagonica González &amp; Vandenberg, new species, holotype; C, C. germainii (Crotch); D, C. eryngii (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924357" httpUri="https://zenodo.org/record/3924357/files/figure.png" pageId="7" pageNumber="20">Fig. 6A</figureCitation>
) have the basal lobe more strongly dilated in apical 1/4 compared with the other species in this complex (except males not known in 
<taxonomicName id="1DD84D51100BBC64050BFA22847DFA0C" authorityName="González &amp; Vandenberg" authorityYear="2006" box="[595,777,1473,1498]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="disconsolata">
<emphasis id="E8ACEAC0100BBC64050BFA22847DFA0C" box="[595,777,1473,1498]" italics="true" pageId="7" pageNumber="20">C. disconsolata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="92C26559100BBC6A0660FA0A86E8FCB0" lastPageId="9" lastPageNumber="22" pageId="7" pageNumber="20" type="description">
<paragraph id="DA6736D2100BBC640660FA0A8488F975" blockId="7.[264,1323,1033,1899]" pageId="7" pageNumber="20">
<emphasis id="E8ACEAC0100BBC640660FA0A85C4F9D5" bold="true" box="[312,688,1513,1539]" pageId="7" pageNumber="20">
Description (
<typeStatus id="05638870100BBC640687FA0A8527F9D5" box="[479,595,1513,1539]" pageId="7" pageNumber="20" type="holotype">Holotype</typeStatus>
male):
</emphasis>
Length 3.0 mm, width 
<quantity id="1D209B37100BBC6404BFFA0A833FF9D5" box="[999,1099,1513,1539]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="20" unit="mm" value="2.3">2.3 mm</quantity>
. Form shortened oblong, nearly parallel­sided, convex, apically rounded; elytral, pronotal margins very narrowly reflexed. Punctation on dorsal surfaces fine, regular, with each puncture separated by 2.0–3.0X its diameter; surface between punctures shiny, reticulate on head, pronotum, with only faint trace of reticulation visible on elytron.
</paragraph>
<paragraph id="DA6736D2100BBC6B0660F95286A3FD38" blockId="7.[264,1323,1033,1899]" lastBlockId="8.[264,1325,284,1870]" lastPageId="8" lastPageNumber="21" pageId="7" pageNumber="20">
Dorsal color pattern as follows: Head black with two irregular cream­colored spots, one at inner margin of each eye extending from eye canthus to just below level of upper 1/3 of eye; eye silvery; labrum brown. Pronotum predominantly dark reddish brown, nearly black; lateral margins yellow cream­colored, beginning at posterior angle as narrow band, widened apically to enclose entire anterior angle, continued as broken trace on anterior margin (
<figureCitation id="42E32A571004BC6B0688FEFF855BFEE0" box="[464,559,284,310]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González &amp; Vandenberg, new species, paratype; B, C. disconsolata Vandenberg &amp; González, new species, holotype; C, C. patagonica González &amp; Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González &amp; Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="8" pageNumber="21">Fig. 3H</figureCitation>
). Scutellum black. Elytron with ground color pale ocherous or straw with dark brown irregular figure filling most of disc (
<figureCitation id="42E32A571004BC6B0484FEA7834AFE88" box="[988,1086,324,350]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González &amp; Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg &amp; González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="8" pageNumber="21">Fig. 4N</figureCitation>
); discal figure with incised borders as follows: outer border with anteromedial triangular emargination pointing toward sutural apex; inner border with large teardrop­shaped emargination followed by smaller semicircular emargination, the latter deeply penetrating the dark zone leaving only a slender dark hook; union of dark, light areas irregular, suffused, somewhat reddish. Each puncture of dorsal surfaces with pinpoint of brown at center, visible only in areas with light to medium background coloration; staggered double row of punctures nearest suture, single to double staggered row along lateral margin beginning just outside of humeral bulge with more pronounced pigmentation. Anterior, lateral margins of pronotum, all margins of elytron narrowly transparent to light amber, sutural margin somewhat darker.
</paragraph>
<paragraph id="DA6736D21004BC6B0660FD1F832BFC08" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="21">Ground color of ventral surfaces dark brown; elytral epipleuron, pronotal hypomeron except basally at inner margin straw­colored; mesepimeron cream­colored; antenna, mouthparts yellow brown with antennal club slightly darker; legs with coxa brown, femur dark brown to blackish; tibia yellow brown with darker brown narrowly along outer margin; tarsus translucent yellow brown with last tarsomere, base of claw darker. Ventral surfaces including appendages clothed in decumbent silvery pubescence.</paragraph>
<paragraph id="DA6736D21004BC6B0660FC0F8303F928" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="21">
Eyes finely facetted, separated by 2½X eye diameter; inner orbits nearly parallel in lower half, diverging at upper level. Antenna of 10 antennomeres (
<figureCitation id="42E32A571004BC6B0366FBF783E9FBF8" box="[1086,1181,1044,1070]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: A­C, C. lacrimosa Gonzlez &amp; Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez &amp; Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="8" pageNumber="21">Fig. 9B</figureCitation>
), combined length slightly greater than distance between eyes; third antennomere elongate, 1½X length of second, subequal to four plus five combined. Pronotum (
<figureCitation id="42E32A571004BC6B034FFB878306FBA8" box="[1047,1138,1124,1150]" captionStart="FIGURE 3" captionStartId="15.[264,367,1565,1589]" captionTargetBox="[272,1316,323,1530]" captionTargetId="figure@15.[256,1331,310,1551]" captionTargetPageId="15" captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González &amp; Vandenberg, new species, paratype; B, C. disconsolata Vandenberg &amp; González, new species, holotype; C, C. patagonica González &amp; Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González &amp; Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range." httpUri="https://zenodo.org/record/173871/files/figure.png" pageId="8" pageNumber="21">Fig. 3H</figureCitation>
) evenly convex except for very narrowly reflexed transparent lateral margin; in outline with basal margin subsinuate, lateral margin strongly evenly arcuate, anterior margin subtrapezoidally emarginate, medially slightly arcuately produced; anterior angles subtriangulate, conspicuous in dorsal view, projecting anteroventrally. Elytron broad (
<figureCitation id="42E32A571004BC6B033CFAE7838FFAC8" box="[1124,1275,1284,1310]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González &amp; Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg &amp; González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="8" pageNumber="21">Figs. 4M–N</figureCitation>
), in dorsal view with humeral angle rounded, weakly arcuate from beyond humeral angle to apical 1/3, broadest near apical 1/3, roundly tapered distally; epipleuron flat, horizontal in anterior half, progressively inwardly sloping in posterior half. Prosternum T­shaped, with lateral arms gently folded back from stem, in cross section forming shallow arc; intercoxal process weakly convex, bearing fine superficial median sulcus on distal 1/4, apex truncate. Mesosternum trapezoidal; anterior border approximately linear with raised margin. Metasternum broad, with postmesocoxal line reaching lateral margin; surface polished, obsoletely rugulose; discrimen present, indistinct. Abdomen shortened semi­oval, broadest between first, second ventrites; posterior margin of ventrites 1–4 linear, of 5 broadly shallowly emarginate, of 6 apically rounded; postmetacoxal line of first abdominal ventrite curved posterolaterad, closely paralleling posterior margin for much of length, not attaining lateral margin. Tarsal claw with shallow subquadrate basal tooth.
</paragraph>
<paragraph id="DA6736D21004BC6A0660F8EF83ADFEE0" blockId="8.[264,1325,284,1870]" lastBlockId="9.[264,1323,284,1870]" lastPageId="9" lastPageNumber="22" pageId="8" pageNumber="21">
Male genitalia as shown (
<figureCitation id="42E32A571004BC6B053CF8EF85B0F8F0" box="[612,708,1804,1830]" captionStart="FIGURE 6" captionStartId="18.[264,368,1741,1765]" captionTargetBox="[266,1322,327,1740]" captionTargetId="figure@18.[264,1323,305,1746]" captionTargetPageId="18" captionText="FIGURE 6. Male genitalia of Cycloneda species (above, ventral and right lateral views of phallobase; below, right lateral view of sipho (=aedeagus)): A, C. lacrimosa González &amp; Vandenberg, new species, holotype; B, C. patagonica González &amp; Vandenberg, new species, holotype; C, C. germainii (Crotch); D, C. eryngii (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924357" httpUri="https://zenodo.org/record/3924357/files/figure.png" pageId="8" pageNumber="21">Fig. 6A</figureCitation>
): basal lobe elongate, roughly parallel­sided in basal half, subapically distinctly swollen with greatest width at apical 1/4; apex tapered, slightly attenuate; parameres slender, reaching three quarters distance to apex of basal lobe.
</paragraph>
<paragraph id="DA6736D21005BC6A0660FEA08419FE78" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="22">
<emphasis id="E8ACEAC01005BC6A0660FEA086E8FE8B" bold="true" box="[312,412,323,349]" pageId="9" pageNumber="22">Female:</emphasis>
Similar to male except slightly larger on average and proportionally broader, apex slightly pointed. Abdomen with posterior margin of ventrite 5 nearly linear, apex of ventrite 6 pointed. Female genitalia as in figure 8A.
</paragraph>
<paragraph id="DA6736D21005BC6A0660FE5886E8FCB0" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="22">
<emphasis id="E8ACEAC01005BC6A0660FE5886C0FE03" bold="true" box="[312,436,443,469]" pageId="9" pageNumber="22">Variation:</emphasis>
Length 3.0 to 
<quantity id="1D209B371005BC6A0532FE5F85BCFE00" box="[618,712,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="22" unit="mm" value="3.6">3.6 mm</quantity>
. Pale markings on head variable in size but always well separated by dark frons; not gender­specific. Pale anterior margin of pronotum obsolete to narrow but entire. Elytron may have additional dark spot at apex (
<figureCitation id="42E32A571005BC6A03C7FDEF8388FDF0" box="[1183,1276,524,550]" captionStart="FIGURE 1" captionStartId="4.[264,367,1784,1808]" captionTargetBox="[344,1263,723,1752]" captionTargetId="figure@4.[279,1332,713,1777]" captionTargetPageId="4" captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González &amp; Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg &amp; González, new species, female, holotype (3.3 mm); C, C. patagonica González &amp; Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)." httpUri="https://zenodo.org/record/173869/files/figure.png" pageId="9" pageNumber="22">Fig. 1A</figureCitation>
), or faint dark spot at humeral bulge; sometimes with hook shaped mark at apex of dark discal figure disconnected (
<figureCitation id="42E32A571005BC6A055BFDBF8510FDA0" box="[515,612,604,630]" captionStart="FIGURE 4" captionStartId="16.[264,367,1625,1649]" captionTargetBox="[395,1202,315,1596]" captionTargetId="figure@16.[316,1279,307,1620]" captionTargetPageId="16" captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González &amp; Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg &amp; González, new species (O, paratype; P, holotype)." httpUri="https://zenodo.org/record/173872/files/figure.png" pageId="9" pageNumber="22">Fig. 4M</figureCitation>
) or entirely absent; some specimens with suffused creamcolored maculae nested within 3 incised areas of dark discal figure; more commonly with ground color only slightly lighter on disc or apparently unicolorous. Dark areas on dorsal surfaces nearly black in many specimens. Pronotal hypomeron unicolorous or basally darkened as in 
<typeStatus id="056388701005BC6A06E4FD1F8552FCC0" box="[444,550,764,790]" pageId="9" pageNumber="22" type="holotype">holotype</typeStatus>
. Tibia totally yellowish in some. Antenna may have 9, 10 or 11 antennomeres (
<figureCitation id="42E32A571005BC6A06E5FCC7853FFCE8" box="[445,587,804,830]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: A­C, C. lacrimosa Gonzlez &amp; Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez &amp; Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="9" pageNumber="22">Figs. 9A–C</figureCitation>
). Sulcus on prosternal intercoxal process may be less distinct or wavering.
</paragraph>
</subSubSection>
<subSubSection id="92C265591005BC6A0660FC9785CFFB18" pageId="9" pageNumber="22" type="materials_examined">
<paragraph id="DA6736D21005BC6A0660FC9785CFFB18" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="22">
<emphasis id="E8ACEAC01005BC6A0660FC978686FC58" bold="true" box="[312,498,884,910]" pageId="9" pageNumber="22">Type material:</emphasis>
<materialsCitation id="6AB03C8F1005BC6A06A7FC978568FC60" ID-GBIF-Occurrence="2651531319" collectingDate="1986-12-25" collectionCode="MNHN" collectorName="G. Gonzalez" country="Chile" location="Agua Verde" pageId="9" pageNumber="22" specimenCount="1" stateProvince="Antofagasta" typeStatus="holotype">
<typeStatus id="056388701005BC6A06A7FC978518FC58" box="[511,620,884,910]" pageId="9" pageNumber="22" type="holotype">Holotype</typeStatus>
(male) “ 
<location id="DF0760091005BC6A0581FC97841AFC58" LSID="urn:lsid:plazi:treatment:527187C4100BBC690650FC7583B9FD48:DF0760091005BC6A0581FC97841AFC58" box="[729,878,884,910]" country="Chile" name="Agua Verde" pageId="9" pageNumber="22" stateProvince="Antofagasta">Agua Verde</location>
, 
<collectingRegion id="181CF8301005BC6A0425FC97837AFC58" box="[893,1038,884,910]" country="Chile" name="Antofagasta" pageId="9" pageNumber="22">Antofagasta</collectingRegion>
, 
<collectingCountry id="A2CF76421005BC6A0346FC978314FC58" box="[1054,1120,884,910]" name="Chile" pageId="9" pageNumber="22">Chile</collectingCountry>
, 
<date id="AE6610121005BC6A0328FC978277FC58" box="[1136,1283,884,910]" pageId="9" pageNumber="22" value="1986-12-25">
<collectingDate id="BE22E9FA1005BC6A0328FC978277FC58" box="[1136,1283,884,910]" pageId="9" pageNumber="22" value="1986-12-25">25­dic­1986</collectingDate>
</date>
. 
<collectorName id="772D53041005BC6A024AFC978602FC60" pageId="9" pageNumber="22">G. González</collectorName>
F.”(
<collectionCode id="BCC9AE171005BC6A06EAFC7F8560FC60" LSID="urn:lsid:biocol.org:col:34988" box="[434,532,924,950]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="22">MNHN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6AB03C8F1005BC6A056CFC7F845DFC08" ID-GBIF-Occurrence="2651531320" collectingDate="1991-12-21" collectionCode="MNHN" collectorName="G. Gonzalez." country="Chile" location="Agua Verde" pageId="9" pageNumber="22" specimenCount="1" stateProvince="Antofagasta" typeStatus="allotype">
<typeStatus id="056388701005BC6A056CFC7F85EFFC60" box="[564,667,924,950]" pageId="9" pageNumber="22" type="allotype">Allotype</typeStatus>
(female), “ 
<collectingCountry id="A2CF76421005BC6A046EFC7F84E6FC60" box="[822,914,924,950]" name="Chile" pageId="9" pageNumber="22">CHILE</collectingCountry>
, 
<collectingRegion id="181CF8301005BC6A04F2FC7F834FFC60" box="[938,1083,924,950]" country="Chile" name="Antofagasta" pageId="9" pageNumber="22">Antofagasta</collectingRegion>
, 
<location id="DF0760091005BC6A030AFC7F8399FC60" LSID="urn:lsid:plazi:treatment:527187C4100BBC690650FC7583B9FD48:DF0760091005BC6A030AFC7F8399FC60" box="[1106,1261,924,950]" country="Chile" name="Agua Verde" pageId="9" pageNumber="22" stateProvince="Antofagasta">Agua Verde</location>
. 
<date id="AE6610121005BC6A025BFC7F86B0FC08" pageId="9" pageNumber="22" value="1991-12-21">
<collectingDate id="BE22E9FA1005BC6A025BFC7F86B0FC08" pageId="9" pageNumber="22" value="1991-12-21">21­ diciembre­1991</collectingDate>
</date>
, leg. 
<collectorName id="772D53041005BC6A0550FC2785D0FC08" box="[520,676,964,990]" pageId="9" pageNumber="22">G. Gonzalez.</collectorName>
” (
<collectionCode id="BCC9AE171005BC6A0598FC278454FC08" LSID="urn:lsid:biocol.org:col:34988" box="[704,800,964,990]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="22">MNHN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6AB03C8F1005BC6A046FFC27869FFBD0" ID-GBIF-Occurrence="2651531387" collectingDate="1986-12-25" collectionCode="MNHN" collectorName="G. Gonzalez" country="Chile" location="Agua Verde" pageId="9" pageNumber="22" specimenCount="1" stateProvince="Antofagasta" typeStatus="paratype">
<typeStatus id="056388701005BC6A046FFC2784DDFC08" box="[823,937,964,990]" pageId="9" pageNumber="22" type="paratype">Paratypes</typeStatus>
(total=24), 
<specimenCount id="CCDEFD5B1005BC6A0362FC27833DFC08" box="[1082,1097,964,990]" pageId="9" pageNumber="22">1</specimenCount>
with same data as holotype (
<collectionCode id="BCC9AE171005BC6A06DBFC0F8697FBD0" LSID="urn:lsid:biocol.org:col:34988" box="[387,483,1004,1030]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="22">MNHN</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="6AB03C8F1005BC6A06A3FC0F861DFBF8" ID-GBIF-Occurrence="2651531355" collectingDate="1973-01-06" collectionCode="AMNH, CAS, MNHN, USNM" collectorName="W. Wittmer" country="Argentina" county="Nevado de Cachi" elevation="3700" location="Stephan Halloy" municipality="Col" pageId="9" pageNumber="22" specimenCount="36" stateProvince="Salta" typeStatus="paratype">
<specimenCount id="CCDEFD5B1005BC6A06A3FC0F856CFBD0" box="[507,536,1004,1030]" pageId="9" pageNumber="22">18</specimenCount>
with same data as allotype (
<specimenCount id="CCDEFD5B1005BC6A04DCFC0F84E1FBD0" box="[900,917,1004,1030]" pageId="9" pageNumber="22">4</specimenCount>
, 
<collectionCode id="BCC9AE171005BC6A04FEFC0F8377FBD0" LSID="urn:lsid:biocol.org:col:34925" box="[934,1027,1004,1030]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="22">AMNH</collectionCode>
; 
<specimenCount id="CCDEFD5B1005BC6A034CFC0F8350FBD0" box="[1044,1060,1004,1030]" pageId="9" pageNumber="22">3</specimenCount>
, 
<collectionCode id="BCC9AE171005BC6A036FFC0F8300FBD0" box="[1079,1140,1004,1030]" pageId="9" pageNumber="22">CAS</collectionCode>
; 
<specimenCount id="CCDEFD5B1005BC6A03DCFC0F83E1FBD0" box="[1156,1173,1004,1030]" pageId="9" pageNumber="22">7</specimenCount>
, 
<collectionCode id="BCC9AE171005BC6A03FFFC0F8270FBD0" LSID="urn:lsid:biocol.org:col:34988" box="[1191,1284,1004,1030]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="22">MNHN</collectionCode>
; 
<specimenCount id="CCDEFD5B1005BC6A024DFC0F8252FBD0" box="[1301,1318,1004,1030]" pageId="9" pageNumber="22">4</specimenCount>
, 
<collectionCode id="BCC9AE171005BC6A0650FBF78615FBF8" LSID="urn:lsid:biocol.org:col:34871" box="[264,353,1044,1070]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="22" type="Museum">USNM</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="6AB03C8F1005BC6A062CFBF78602FBA8" ID-GBIF-Occurrence="2651531364" collectingDate="1973-01-06" collectionCode="BMNH" collectorName="W. Wittmer" country="Bolivia" county="Nevado de Cachi" elevation="3700" location="Oruro" municipality="Col" pageId="9" pageNumber="22" specimenCount="4" stateProvince="Salta" typeStatus="paratype">
<specimenCount id="CCDEFD5B1005BC6A062CFBF786F7FBF8" box="[372,387,1044,1070]" pageId="9" pageNumber="22">4</specimenCount>
“ 
<location id="DF0760091005BC6A06CCFBF786ABFBF8" LSID="urn:lsid:plazi:treatment:527187C4100BBC690650FC7583B9FD48:DF0760091005BC6A06CCFBF786ABFBF8" box="[404,479,1044,1070]" country="Bolivia" county="Nevado de Cachi" municipality="Col" name="Oruro" pageId="9" pageNumber="22" stateProvince="Salta">Oruro</location>
, 
<collectingCountry id="A2CF76421005BC6A06B3FBF78530FBF8" box="[491,580,1044,1070]" pageId="9" pageNumber="22">Bolivia</collectingCountry>
, 
<collectingDate id="BE22E9FA1005BC6A0517FBF785D3FBF8" box="[591,679,1044,1070]" pageId="9" pageNumber="22">14.1.40</collectingDate>
<quantity id="1D209B371005BC6A05F6FBF7847DFBF8" box="[686,777,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="22" unit="m" value="3700.0">
<elevation id="51F5D1E11005BC6A05F6FBF7847DFBF8" box="[686,777,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="22" unit="m" value="3700.0">3700 m</elevation>
</quantity>
, 
<collectorName id="772D53041005BC6A044CFBF784D7FBF8" box="[788,931,1044,1070]" pageId="9" pageNumber="22">W. Wittmer</collectorName>
/Brit.Mus., 1945­33.[one paratype mounted along with an incomplete specimen, the latter not designated as a paratype] (
<collectionCode id="BCC9AE171005BC6A0648FB87861AFBA8" box="[272,366,1124,1150]" pageId="9" pageNumber="22">BMNH</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="6AB03C8F1005BC6A06D1FB8785CFFB18" ID-GBIF-Occurrence="2651531388" collectingDate="1973-01-06" collectionCode="IML" collectorName="Stephan Halloy" country="Argentina" county="Nevado de Cachi" elevation="5200" location="Nevado de Cachi" municipality="Col" pageId="9" pageNumber="22" specimenCount="1" stateProvince="SALTA" typeStatus="paratype">
<specimenCount id="CCDEFD5B1005BC6A06D1FB8786ECFBA8" box="[393,408,1124,1150]" pageId="9" pageNumber="22">1</specimenCount>
“ 
<collectingCountry id="A2CF76421005BC6A06E9FB878516FBA8" box="[433,610,1124,1150]" name="Argentina" pageId="9" pageNumber="22">ARGENTINA</collectingCountry>
, 
<collectingRegion id="181CF8301005BC6A052DFB8785A6FBA8" box="[629,722,1124,1150]" country="Argentina" name="Salta" pageId="9" pageNumber="22">SALTA</collectingRegion>
, 
<collectingCounty id="33064E5E1005BC6A05BCFB8784CBFBA8" box="[740,959,1124,1150]" pageId="9" pageNumber="22">Nevado de Cachi</collectingCounty>
, 
<quantity id="1D209B371005BC6A048BFB878340FBA8" box="[979,1076,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="22" unit="m" value="5200.0">
<elevation id="51F5D1E11005BC6A048BFB878340FBA8" box="[979,1076,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="22" unit="m" value="5200.0">5200 m</elevation>
</quantity>
_ 
<quantity id="1D209B371005BC6A031CFB87831EFBA8" box="[1092,1130,1124,1150]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="22" unit="m" value="6.0">
<elevation id="51F5D1E11005BC6A031CFB87831EFBA8" box="[1092,1130,1124,1150]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="22" unit="m" value="6.0">6m</elevation>
</quantity>
, 
<date id="AE6610121005BC6A0325FB878397FBA8" box="[1149,1251,1124,1150]" pageId="9" pageNumber="22" value="1973-01-06">
<collectingDate id="BE22E9FA1005BC6A0325FB878397FBA8" box="[1149,1251,1124,1150]" pageId="9" pageNumber="22" value="1973-01-06">6.I­1973</collectingDate>
</date>
, Col: 
<collectorName id="772D53041005BC6A0650FB6F86B7FB70" box="[264,451,1164,1190]" pageId="9" pageNumber="22">Stephan Halloy</collectorName>
/COLECCION, INST.– FUND M. LILLO (4000) – S.M.TUCUMAN, TUCUMAN – ARGENTINA”(
<collectionCode id="BCC9AE171005BC6A0524FB5785C7FB18" box="[636,691,1204,1230]" country="Argentina" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="9" pageNumber="22">IML</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="92C265591005BC6A0660FB3F831AFAC8" pageId="9" pageNumber="22" type="etymology">
<paragraph id="DA6736D21005BC6A0660FB3F831AFAC8" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="22">
<emphasis id="E8ACEAC01005BC6A0660FB3F86BCFB20" bold="true" box="[312,456,1244,1270]" pageId="9" pageNumber="22">Etymology:</emphasis>
From the Latin 
<emphasis id="E8ACEAC01005BC6A05D3FB3F847FFB23" box="[651,779,1244,1269]" italics="true" pageId="9" pageNumber="22">lacrimosus</emphasis>
(adj.) meaning “prone to tears or crying,” the name is a reference to the large teardrop­shaped mark at the elytral suture.
</paragraph>
</subSubSection>
<subSubSection id="92C265591005BC690660FACF84FCFDA0" lastPageId="10" lastPageNumber="23" pageId="9" pageNumber="22" type="discussion">
<paragraph id="DA6736D21005BC690660FACF86FAFE88" blockId="9.[264,1323,284,1870]" lastBlockId="10.[264,1323,284,670]" lastPageId="10" lastPageNumber="23" pageId="9" pageNumber="22">
<emphasis id="E8ACEAC01005BC6A0660FACF86C7FA90" bold="true" box="[312,435,1324,1350]" pageId="9" pageNumber="22">Remarks:</emphasis>
The specimens from Agua Verde were collected at an isolated gas station in the 
<collectingRegion id="181CF8301005BC6A066CFAB78682FAB8" box="[308,502,1364,1390]" country="Chile" name="Atacama" pageId="9" pageNumber="22">Atacama Desert</collectingRegion>
, on plants in flower­pots, about 800 kilometers north of the northern most locality for 
<emphasis id="E8ACEAC01005BC6A06BEFA9F85FDFA43" box="[486,649,1404,1429]" italics="true" pageId="9" pageNumber="22">
<taxonomicName id="1DD84D511005BC6A06BEFA9F85F2FA43" authorityName="González &amp; Vandenberg" authorityYear="2006" baseAuthorityName="González &amp; Vandenberg" baseAuthorityYear="2006" box="[486,646,1404,1429]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="22" phylum="Arthropoda" rank="species" species="germainii">C. germainii</taxonomicName>
.
</emphasis>
From this single collection locality specimens were obtained with either nine, ten, or eleven antennomeres. These variants are assumed to be conspecific as they show no other apparent differences. 
<figureCitation id="42E32A571005BC6A0487FA2F83F4FA30" box="[991,1152,1484,1510]" captionStart="FIGURE 9" captionStartId="26.[264,368,885,909]" captionTargetBox="[276,1316,321,884]" captionTargetId="figure@26.[264,1323,305,886]" captionTargetPageId="26" captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: A­C, C. lacrimosa Gonzlez &amp; Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez &amp; Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)." figureDoi="http://doi.org/10.5281/zenodo.3924340" httpUri="https://zenodo.org/record/3924340/files/figure.png" pageId="9" pageNumber="22">Figure 9A–C</figureCitation>
suggests one possible interpretation for the observed variation based on fusion in the region between the basal two and distal six antennomeres. This hypothesis is supported by a corresponding increase in the length of antennomere four in specimens with only ten antennomeres, or antennomere three in specimens with only nine antennomeres. Antennal polymorphism was previously reported in the coccinellid species 
<taxonomicName id="1DD84D511005BC6A0434F9778252F978" authority="Chapin" authorityName="Chapin" box="[876,1318,1684,1710]" class="Insecta" family="Coccinellidae" genus="Catana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="22" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="E8ACEAC01005BC6A0434F9778359F97B" box="[876,1069,1684,1709]" italics="true" pageId="9" pageNumber="22">Catana clauseni</emphasis>
Chapin (Serangiini)
</taxonomicName>
. 
<bibRefCitation id="BE494B231005BC6A0650F95F86CCF900" author="Chapin" box="[264,440,1724,1750]" firstAuthor="Chapin" pageId="9" pageNumber="49" pagination="263" refId="ref18087" refString="Chapin, E. A. (1940) New genera and species of lady-beetles related to Serangium Blackburn (Coleoptera: Coccinellidae). Journal of the Washington Academy of Sciences, 30, 263 - 272." type="journal article" year="1940">Chapin (1940)</bibRefCitation>
distinguished 
<taxonomicName id="1DD84D511005BC6A0528F95F845DF900" authority="Chapin" authorityName="Chapin" authorityYear="1940" box="[624,809,1724,1750]" class="Insecta" family="Coccinellidae" genus="Catana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="E8ACEAC01005BC6A0528F95F85B2F903" box="[624,710,1724,1749]" italics="true" pageId="9" pageNumber="22">Catana</emphasis>
Chapin
</taxonomicName>
from other members of the tribe by the possession of only eight antennomeres, but he noted that the fifth antennomere sometimes shows the beginning of a division and is therefore morphologically equivalent to the fourth and fifth combined. 
<bibRefCitation id="BE494B231005BC6A055AF8D785B3F898" author="Miyatake, M." box="[514,711,1844,1870]" firstAuthor="Miyatake" journalOrPublisher="Memoirs of the Ehime University, Sect. VI (Agriculture)" pageId="9" pageNumber="49" pagination="135 - 146" part="6" refId="ref18280" refString="Miyatake, M. (1961) The East-Asian coccinellid-beetles preserved in the California Academy of Sciences, tribe Serangiini. Memoirs of the Ehime University, Sect. VI (Agriculture), 6 (2), 135 - 146." title="The East-Asian coccinellid-beetles preserved in the California Academy of Sciences, tribe Serangiini" type="journal article" year="1961">Miyatake (1961)</bibRefCitation>
subsequently discovered a specimen of 
<taxonomicName id="1DD84D511005BC6A03FDF8D7825FF89B" authorityName="Chapin" box="[1189,1323,1844,1869]" class="Insecta" family="Coccinellidae" genus="Catana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="22" phylum="Arthropoda" rank="species" species="clauseni">
<emphasis id="E8ACEAC01005BC6A03FDF8D7825FF89B" box="[1189,1323,1844,1869]" italics="true" pageId="9" pageNumber="22">C. clauseni</emphasis>
</taxonomicName>
with a complete division resulting in nine distinct antennomeres as in the related genus 
<taxonomicName id="1DD84D511006BC690650FEA786F3FE8B" authorityName="W.Wood ex R.A.Salisbury" authorityYear="1866" box="[264,391,324,349]" class="Insecta" family="Coccinellidae" genus="Serangium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="E8ACEAC01006BC690650FEA786F3FE8B" box="[264,391,324,349]" italics="true" pageId="10" pageNumber="23">Serangium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DA6736D21006BC690660FE8F84FCFDA0" blockId="10.[264,1323,284,670]" pageId="10" pageNumber="23">
It is probable that Weise was familiar with the new species described here, but regarded it as a variation of 
<emphasis id="E8ACEAC01006BC69050EFE7785A7FE7B" box="[598,723,404,429]" italics="true" pageId="10" pageNumber="23">
<taxonomicName id="1DD84D511006BC69050EFE7785A4FE7B" authorityName="González &amp; Vandenberg" authorityYear="2006" baseAuthorityName="González &amp; Vandenberg" baseAuthorityYear="2006" box="[598,720,404,429]" class="Insecta" family="Coccinellidae" genus="Cycloneda" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="eryngii">C. eryngii</taxonomicName>
.
</emphasis>
The Smithsonian Entomology Library has a copy of 
<bibRefCitation id="BE494B231006BC690671FE5F86BBFE00" author="Crotch" box="[297,463,444,470]" firstAuthor="Crotch" pageId="10" pageNumber="49" refId="ref18126" refString="Crotch, G. R. (1874) A revision of the coleopterous family Coccinellidae. University Press, London, 311 pp." type="book" year="1874">Crotch (1874)</bibRefCitation>
that was signed by Weise and apparently extensively annotated by him. In the left hand margin of page 107, below the caption for 
<emphasis id="E8ACEAC01006BC690492FE0783D9FE2B" box="[970,1197,484,509]" italics="true" pageId="10" pageNumber="23">
<taxonomicName id="1DD84D511006BC690492FE0783DCFE2B" authorityName="Mulsant" authorityYear="1850" box="[970,1192,484,509]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="eryngii">Coccinella eryngii</taxonomicName>
,
</emphasis>
is a handpenned figure (left half of pronotum, left elytron) of a specimen with a pattern identical to the example illustrated in figure 
<specimenCount id="CCDEFD5B1006BC6905DCFDD785C4FD98" box="[644,688,564,590]" pageId="10" pageNumber="23" type="generic">4N.</specimenCount>
The sketch is labeled with the female symbol and the words “var. Moreno, 
<collectingCountry id="A2CF76421006BC69055AFDBF851AFDA0" box="[514,622,604,630]" name="Argentina" pageId="10" pageNumber="23">Argentin</collectingCountry>
. Mus. [illegible word]”.
</paragraph>
</subSubSection>
<subSubSection id="92C265591006BC690660FD6783B9FD48" box="[312,1229,644,670]" pageId="10" pageNumber="23" type="materials_examined">
<paragraph id="DA6736D21006BC690660FD6783B9FD48" blockId="10.[264,1323,284,670]" box="[312,1229,644,670]" pageId="10" pageNumber="23">
<emphasis id="E8ACEAC01006BC690660FD6784F6FD48" bold="true" box="[312,898,644,670]" pageId="10" pageNumber="23">
Data from specimens examined (Map, 
<figureCitation id="42E32A571006BC690442FD678406FD48" box="[794,882,644,670]" captionStart="FIGURE 10" captionStartId="27.[264,368,1868,1892]" captionTargetBox="[287,1302,680,1870]" captionTargetId="figure@27.[287,1312,671,1870]" captionTargetPageId="27" captionText="FIGURE 10. Distributions of Cycloneda species." figureDoi="http://doi.org/10.5281/zenodo.3924342" httpUri="https://zenodo.org/record/3924342/files/figure.png" pageId="10" pageNumber="23">Fig. 10</figureCitation>
):
</emphasis>
see “ 
<typeStatus id="056388701006BC6904E7FD678488FD48" box="[959,1020,644,670]" pageId="10" pageNumber="23">Type</typeStatus>
material,” above.
</paragraph>
</subSubSection>
</treatment>
</document>