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<mods:title id="971803539D3C134F2630E816B4B6672E">Cybister (s. str.) poblanus sp. n. from Mexico and notes on other species of Cybistrinae (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="1FF789BC291A0A692379BC7F4FF0F02C">Arce-Pérez, Roberto</mods:namePart>
<mods:affiliation id="4C0F8A565D97B7D6BF80D25A8706DFAA">Red de Biodiversidad y Sistemática, Instituto de Ecología, A. C. (INECOL), Carretera Antigua a Coatepec Núm. 351, El Haya, Xalapa, Veracruz 91073, México.</mods:affiliation>
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<mods:namePart id="9D7A2C6CBBE117EA604F0E42F524488F">Novelo-Gutiérrez, Rodolfo</mods:namePart>
<mods:affiliation id="9D74F170C769E5F2F456DBA3FFFE161D">Red de Biodiversidad y Sistemática, Instituto de Ecología, A. C. (INECOL), Carretera Antigua a Coatepec Núm. 351, El Haya, Xalapa, Veracruz 91073, México.</mods:affiliation>
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<mods:namePart id="46C3159FA0077F61A402D6E67C420FCD">Fery, Hans</mods:namePart>
<mods:affiliation id="755637C26B6555E6D0711BDAC6A66FC3">Hans Fery, Räuschstr. 73, D- 13509 Berlin.</mods:affiliation>
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Key to species of 
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occurring in America north of 
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and 
<collectingCountry id="BEC17658FFACFFB7FBE9045CC6C2FCCE" box="[1034,1173,834,860]" name="Guatemala" pageId="8" pageNumber="331">Guatemala</collectingCountry>
(including 
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and 
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)
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<paragraph id="C66936C8FFACFFB7FF7404B1C6B2FB83" blockId="8.[151,1436,943,1041]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FF7404B1C28DFC5B" bold="true" box="[151,218,943,969]" pageId="8" pageNumber="331">Notes</emphasis>
: The distribution area of some of the species studied may be considerably larger and extending far further south into the Neotropical region (see e.g. 
<bibRefCitation id="A2474B39FFACFFB7FD8F04CDC1EDFC7F" author="Arce-Perez, R. &amp; Roughley, R. E." box="[620,954,979,1005]" pageId="8" pageNumber="331" pagination="69 - 104" refId="ref9717" refString="Arce-Perez, R. &amp; Roughley, R. E. (1999) Lista anotada y claves para los Hydradephaga (Coleoptera: Adephaga: Dytiscidae, Noteridae, Haliplidae, Gyrinidae) de Mexico. Dugesiana, 6 (2), 69 - 104." type="journal article" year="1999">Arce-Pérez &amp; Roughley 1999</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FC2704CDC6C4FC7F" author="Blanco Aller, R." box="[964,1171,979,1005]" pageId="8" pageNumber="331" pagination="197 - 205" refId="ref9819" refString="Blanco Aller, R. (2016) Contribucion al conocimiento de los generos Megadytes Sharp, 1882, Thermonectus Dejean, 1833 e Hydaticus Leach, 1817 (Coleoptera: Adephaga: Dytiscidae) en Costa Rica. Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 58, 197 - 205." type="journal article" year="2016">Blanco Aller 2016</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FB7E04CDC77DFC7F" author="Brinck, P." box="[1181,1322,978,1005]" pageId="8" pageNumber="331" pagination="1 - 20" refId="ref9879" refString="Brinck, P. (1945) Nomenklatorische und systematische Studien uber Dytisciden. III. Die Klassifikation der Cybisterinen. Kungliga Fysiografiska Sallskapets Handlingar, 56 (4), 1 - 20." type="journal article" year="1945">Brinck 1945</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FAD704CDC346FB83" author="Hendrich, L. &amp; Manuel, M. &amp; Balke, M." pageId="8" pageNumber="331" pagination="517 - 535" refId="ref10155" refString="Hendrich, L., Manuel, M. &amp; Balke, M. (2019) The return of the Duke - locality data for Megadytes ducalis Sharp, 1882, the world's largest diving beetle, with notes on related species (Coleoptera: Dytiscidae). Zootaxa, 4586 (3), 517 - 535. https: // doi. org / 10.11646 / zootaxa. 4586.3.8" type="journal article" year="2019">
Hendrich 
<emphasis id="F4A2EADAFFACFFB7FF7404E9C298FB83" box="[151,207,1015,1041]" italics="true" pageId="8" pageNumber="331">et al.</emphasis>
2019
</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FEFF04E9C0FCFB83" author="Megna, Y. &amp; Sanchez-Fernandez, D." box="[284,683,1015,1041]" pageId="8" pageNumber="331" pagination="199 - 221" refId="ref10394" refString="Megna, Y. S &amp; Sanchez-Fernandez, D. (2019) Clave para la identificacion de las especies de la familia Dytiscidae (Coleoptera: Adephaga) de Cuba. Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 64, 199 - 221." type="journal article" year="2019">Megna &amp; Sánchez-Fernández 2019</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FD5504E9C169FB83" author="Miller, K. B." box="[694,830,1015,1041]" pageId="8" pageNumber="331" pagination="401 - 410" refId="ref10447" refString="Miller, K. B. (2013) Review of the genus Cybister Curtis, 1827 (Coleoptera: Dytiscidae: Dytiscinae: Cybistrini) in North America. The Coleopterists Bulletin, 67 (4), 401 - 410. https: // doi. org / 10.1649 / 0010 - 065 X- 67.4.401" type="journal article" year="2013">Miller 2013</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FCAA04E9C610FB83" author="Nilsson, A. N. &amp; Hajek, J." box="[841,1095,1015,1041]" pageId="8" pageNumber="331" refId="ref10749" refString="Nilsson, A. N. &amp; Hajek, J. (2021) A world catalogue of the family Dytiscidae (Coleoptera, Adephaga), 315 pp. Internet version 1. I. 2019. Available from: https: // www. waterbeetles. eu / (accessed 22 July 2019)" type="book" year="2021">Nilsson &amp; Hájek 2021</bibRefCitation>
, 
<bibRefCitation id="A2474B39FFACFFB7FBB104E9C68FFB83" author="Wilke, S." box="[1106,1240,1015,1041]" pageId="8" pageNumber="331" pagination="243 - 276" refId="ref11115" refString="Wilke, S. (1920) Beitrage zur Kenntnis der Gattung Cybister Curtis. Archiv fur Naturgeschichte, 85 A (1919), 243 - 276." type="journal article" year="1920">Wilke 1920</bibRefCitation>
).
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1. Males with only one metatarsal claw; females also with only one claw, but frequently with a short second claw. Posterior metatibial spur simple. Males with stridulatory file at base of metacoxae; females with file absent or weakly developed. Mesotarsomeres I–IV with two separate series of setae near apical margin (one posterodorsally and one posteroventrally; 
<figureCitation id="5EED2A4DFFACFFB7FADA0368C72AFB1E" box="[1337,1405,1142,1164]" captionStart="FIGURES 3" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[211,1362,472,1828]" captionTargetId="figure-149@5.[189,1398,453,1860]" captionTargetPageId="5" captionText="FIGURES 3a–3c. a, b: Part of venter showing ridges on metacoxae (stridulatory organ) and apicoventral angle of metafemora of: (a) Cybister poblanus sp. n.; (b) C. fimbriolatus; c: posterior surface of right mesotarsus of C. fimbriolatus; mesotarsomeres I–IV with two separate series of setae near apical margin: one posterodorsally (on top) and one posteroventrally (on bottom); some further setae removed for better visibility of the setae under consideration." figureDoi="http://doi.org/10.5281/zenodo.5649606" httpUri="https://zenodo.org/record/5649606/files/figure.png" pageId="8" pageNumber="331">Fig. 3c</figureCitation>
) in males and females.* (
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<emphasis id="F4A2EADAFFACFFB7FE50038DC053FB3B" bold="true" box="[435,516,1171,1193]" italics="true" pageId="8" pageNumber="331">Cybister</emphasis>
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).......................................................................... 2
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<keyLead id="7D2795FDFFACFFB7FF7403AFC7CBFA93" pageId="8" pageNumber="331">
- Males and females with two metatarsal claws, though female posterior claw possibly rudimentary. Without stridulatory file at base of metacoxae. Mesotarsomeres I–IV with only one series of setae near apical margin (posteroventrally) in males and females.* (
<taxonomicName id="01D64D4BFFACFFB7FEAF03F5C3E2FA93" ID-CoL="5MNZ" authorityName="Sharp" authorityYear="1882" box="[332,437,1259,1281]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="F4A2EADAFFACFFB7FEAF03F5C3E2FA93" bold="true" box="[332,437,1259,1281]" italics="true" pageId="8" pageNumber="331">Megadytes</emphasis>
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)................................................................................. 5
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<paragraph id="C66936C8FFACFFB7FF740217C7CBFACB" blockId="8.[151,1437,1084,2015]" pageId="8" pageNumber="331">
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2. Apicoventral angle of metafemur sharply pointed (
<figureCitation id="5EED2A4DFFACFFB7FD320217C14EFA8D" box="[721,793,1289,1311]" captionStart="FIGURES 3" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[211,1362,472,1828]" captionTargetId="figure-149@5.[189,1398,453,1860]" captionTargetPageId="5" captionText="FIGURES 3a–3c. a, b: Part of venter showing ridges on metacoxae (stridulatory organ) and apicoventral angle of metafemora of: (a) Cybister poblanus sp. n.; (b) C. fimbriolatus; c: posterior surface of right mesotarsus of C. fimbriolatus; mesotarsomeres I–IV with two separate series of setae near apical margin: one posterodorsally (on top) and one posteroventrally (on bottom); some further setae removed for better visibility of the setae under consideration." figureDoi="http://doi.org/10.5281/zenodo.5649606" httpUri="https://zenodo.org/record/5649606/files/figure.png" pageId="8" pageNumber="331">Fig. 3b</figureCitation>
); male median lobe in lateral view slender and evenly tapering to apex; in ventral view apex truncate and laterally expanded (
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Figs 
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–5h
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). Species from south-western 
<collectingCountry id="BEC17658FFACFFB7FB410238C686FAA9" box="[1186,1233,1318,1339]" name="United States of America" pageId="8" pageNumber="331">USA</collectingCountry>
and western 
<collectingCountry id="BEC17658FFACFFB7FAAE0238C7CEFAAE" box="[1357,1433,1318,1340]" name="Mexico" pageId="8" pageNumber="331">Mexico</collectingCountry>
........................................................................................... 
<taxonomicName id="01D64D4BFFACFFB7FAF7025DC7CBFACB" ID-CoL="7SV67" authorityName="LeConte" authorityYear="1852" box="[1300,1436,1347,1369]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="explanatus">
<emphasis id="F4A2EADAFFACFFB7FAF7025DC7CBFACB" bold="true" box="[1300,1436,1347,1369]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FAF7025DC773FACA" bold="true" box="[1300,1316,1347,1368]" italics="true" pageId="8" pageNumber="331">C</emphasis>
. 
<emphasis id="F4A2EADAFFACFFB7FAD3025DC7CBFACB" bold="true" box="[1328,1436,1347,1369]" italics="true" pageId="8" pageNumber="331">explanatus</emphasis>
</emphasis>
</taxonomicName>
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</paragraph>
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- Apicoventral angle of metafemur not pointed (
<figureCitation id="5EED2A4DFFACFFB7FD42027FC0B0FAE5" box="[673,743,1377,1399]" captionStart="FIGURES 3" captionStartId="5.[151,264,1883,1907]" captionTargetBox="[211,1362,472,1828]" captionTargetId="figure-149@5.[189,1398,453,1860]" captionTargetPageId="5" captionText="FIGURES 3a–3c. a, b: Part of venter showing ridges on metacoxae (stridulatory organ) and apicoventral angle of metafemora of: (a) Cybister poblanus sp. n.; (b) C. fimbriolatus; c: posterior surface of right mesotarsus of C. fimbriolatus; mesotarsomeres I–IV with two separate series of setae near apical margin: one posterodorsally (on top) and one posteroventrally (on bottom); some further setae removed for better visibility of the setae under consideration." figureDoi="http://doi.org/10.5281/zenodo.5649606" httpUri="https://zenodo.org/record/5649606/files/figure.png" pageId="8" pageNumber="331">Fig. 3a</figureCitation>
); male median lobe different.................................. 3
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<paragraph id="C66936C8FFACFFB7FF740260C7CBFA23" blockId="8.[151,1437,1084,2015]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF740260C7CBFA23" pageId="8" pageNumber="331">
3. Large species, TL 
<quantity id="012E9B2DFFACFFB7FE740260C389FA06" box="[407,478,1406,1428]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="331" unit="mm" value="38.0">38 mm</quantity>
. Median lobe apically in lateral view moderately broad (
<figureCitation id="5EED2A4DFFACFFB7FBEB0260C605FA06" box="[1032,1106,1406,1428]" captionStart="FIGURES 4" captionStartId="6.[151,264,1936,1960]" captionTargetBox="[151,1436,181,1880]" captionTargetId="figure-32@6.[151,1436,181,1913]" captionTargetPageId="6" captionText="FIGURES 4a–4f: Cybister poblanus sp. n.: a–c: complete aedeagus in (a) ventral view; (b) dorsal view; (c) nearly lateral view; d–e: median lobe in (d) lateral view; (e) ventral view; f: right paramere." figureDoi="http://doi.org/10.5281/zenodo.5649608" httpUri="https://zenodo.org/record/5649608/files/figure.png" pageId="8" pageNumber="331">Fig. 4d</figureCitation>
). Species so far known only from 
<collectingRegion id="0412F82AFFACFFB7FF010282C300FA23" box="[226,343,1436,1458]" country="Mexico" name="Puebla" pageId="8" pageNumber="331">Puebla state</collectingRegion>
, 
<collectingCountry id="BEC17658FFACFFB7FE820282C3FAFA20" box="[353,429,1436,1458]" name="Mexico" pageId="8" pageNumber="331">Mexico</collectingCountry>
..................................................................... 
<emphasis id="F4A2EADAFFACFFB7FB0A0285C7CBFA23" bold="true" box="[1257,1436,1435,1457]" pageId="8" pageNumber="331">
<taxonomicName id="01D64D4BFFACFFB7FB0A0285C737FA23" authority="Arce-Pérez &amp; Novelo-Gutiérrez &amp; Fery, 2021" authorityName="Arce-Pérez &amp; Novelo-Gutiérrez &amp; Fery" authorityYear="2021" box="[1257,1376,1435,1457]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="poblanus" status="sp. nov.">
<emphasis id="F4A2EADAFFACFFB7FB0A0285C6AEFA22" bold="true" box="[1257,1273,1435,1456]" italics="true" pageId="8" pageNumber="331">C</emphasis>
. 
<emphasis id="F4A2EADAFFACFFB7FAE60285C737FA23" bold="true" box="[1285,1376,1435,1457]" italics="true" pageId="8" pageNumber="331">poblanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF9157A1FFACFFB7FA850282C7CBFA23" box="[1382,1436,1436,1457]" pageId="8" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="C66936C8FFACFFB7FF7402A7C7CBFA5C" blockId="8.[151,1437,1084,2015]" box="[151,1436,1465,1487]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF7402A7C7CBFA5C" box="[151,1436,1465,1487]" pageId="8" pageNumber="331">
- Smaller species, TL &lt;
<quantity id="012E9B2DFFACFFB7FE5A02A7C3A8FA5C" box="[441,511,1465,1487]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="331" unit="mm" value="35.0">35 mm</quantity>
. Median lobe apically in lateral view broader (
<figureCitation id="5EED2A4DFFACFFB7FC5C02A7C667FA5D" box="[959,1072,1465,1487]" captionStart="FIGURES 5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[165,1423,181,1920]" captionTargetId="figure-18@7.[165,1423,181,1920]" captionTargetPageId="7" captionText="FIGURES 5a–5i: Median lobe in lateral view (a, d, g), in ventral view (b, e, h) and paramere (c, f, i) of: (a–c) Cybister fimbriolatus; (d–f) C. occidentalis; (g–i) C. explanatus." figureDoi="http://doi.org/10.5281/zenodo.5649612" httpUri="https://zenodo.org/record/5649612/files/figure.png" pageId="8" pageNumber="331">Figs. 5a, 5d</figureCitation>
)............................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D222E6DFFACFFB7FF7402C8C7CBF9EC" pageId="8" pageNumber="331">
<paragraph id="C66936C8FFACFFB7FF7402C8C7CBF9B4" blockId="8.[151,1437,1084,2015]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF7402C8C7CBF9B4" pageId="8" pageNumber="331">
4. Male median lobe as in 
<figureCitation id="5EED2A4DFFACFFB7FE3102C8C016FA7E" box="[466,577,1494,1516]" captionStart="FIGURES 5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[165,1423,181,1920]" captionTargetId="figure-18@7.[165,1423,181,1920]" captionTargetPageId="7" captionText="FIGURES 5a–5i: Median lobe in lateral view (a, d, g), in ventral view (b, e, h) and paramere (c, f, i) of: (a–c) Cybister fimbriolatus; (d–f) C. occidentalis; (g–i) C. explanatus." figureDoi="http://doi.org/10.5281/zenodo.5649612" httpUri="https://zenodo.org/record/5649612/files/figure.png" pageId="8" pageNumber="331">Figs 5d–5e</figureCitation>
; apex in lateral view truncate and more or less rectangular apicodorsally. Last abdominal ventrite of males and females more or less evenly rounded and flat. Species from 
<collectingCountry id="BEC17658FFACFFB7FBD202EAC631F998" box="[1073,1126,1524,1546]" name="Cuba" pageId="8" pageNumber="331">Cuba</collectingCountry>
, 
<collectingCountry id="BEC17658FFACFFB7FB9102EAC69DF998" box="[1138,1226,1524,1546]" name="Bahamas" pageId="8" pageNumber="331">Bahamas</collectingCountry>
and southern 
<collectingRegion id="0412F82AFFACFFB7FAB402EAC7CBF998" box="[1367,1436,1524,1546]" country="United States of America" name="Florida" pageId="8" pageNumber="331">Florida</collectingRegion>
(
<collectingCountry id="BEC17658FFACFFB7FF0A010FC349F9B5" box="[233,286,1553,1575]" name="United States of America" pageId="8" pageNumber="331">USA</collectingCountry>
).**............................................................................... 
<taxonomicName id="01D64D4BFFACFFB7FAE8010FC7CBF9B4" ID-CoL="7SV8D" authorityName="Aube" authorityYear="1838" box="[1291,1436,1552,1574]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="F4A2EADAFFACFFB7FAE8010FC7CBF9B4" bold="true" box="[1291,1436,1552,1574]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FAE8010FC74CF9B4" bold="true" box="[1291,1307,1553,1574]" italics="true" pageId="8" pageNumber="331">C</emphasis>
. 
<emphasis id="F4A2EADAFFACFFB7FAC4010EC7CBF9B4" bold="true" box="[1319,1436,1552,1574]" italics="true" pageId="8" pageNumber="331">occidentalis</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C66936C8FFACFFB7FF740131C7CBF9EC" blockId="8.[151,1437,1084,2015]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF740131C7CBF9EC" pageId="8" pageNumber="331">
- Male median lobe as in 
<figureCitation id="5EED2A4DFFACFFB7FE2D0130C06BF9D6" box="[462,572,1582,1604]" captionStart="FIGURES 5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[165,1423,181,1920]" captionTargetId="figure-18@7.[165,1423,181,1920]" captionTargetPageId="7" captionText="FIGURES 5a–5i: Median lobe in lateral view (a, d, g), in ventral view (b, e, h) and paramere (c, f, i) of: (a–c) Cybister fimbriolatus; (d–f) C. occidentalis; (g–i) C. explanatus." figureDoi="http://doi.org/10.5281/zenodo.5649612" httpUri="https://zenodo.org/record/5649612/files/figure.png" pageId="8" pageNumber="331">Figs 5a–5b</figureCitation>
; apex in lateral view less truncate and rounded apicodorsally. Last abdominal ventrite of males truncate and with some transverse ridges (
<figureCitation id="5EED2A4DFFACFFB7FD260152C158F9F0" box="[709,783,1612,1634]" captionStart="FIGURES 2" captionStartId="3.[151,264,1937,1961]" captionTargetBox="[163,1421,453,1885]" captionTargetId="figure-144@3.[151,1436,438,1914]" captionTargetPageId="3" captionText="FIGURES 2a–1c: a, b: Last abdominal ventrite of: (a) Cybister poblanus sp. n.; (b) C. fimbriolatus; c: oblique view on head of Cybister poblanus sp. n.; d–e: last left metatarsomere of male of (d) Megadytes lherminieri (two claws); (e) Cybister poblanus sp. n. (one claw)." figureDoi="http://doi.org/10.5281/zenodo.5649602" httpUri="https://zenodo.org/record/5649602/files/figure.png" pageId="8" pageNumber="331">Fig. 2b</figureCitation>
); in females ventrite rounded and flat. Species from south-eastern 
<collectingCountry id="BEC17658FFACFFB7FF010177C37BF9ED" box="[226,300,1641,1663]" name="Canada" pageId="8" pageNumber="331">Canada</collectingCountry>
, large parts of 
<collectingCountry id="BEC17658FFACFFB7FE590177C3BBF9EC" box="[442,492,1641,1662]" name="United States of America" pageId="8" pageNumber="331">USA</collectingCountry>
, 
<collectingCountry id="BEC17658FFACFFB7FE150177C019F9ED" box="[502,590,1641,1663]" name="Bahamas" pageId="8" pageNumber="331">Bahamas</collectingCountry>
and 
<collectingCountry id="BEC17658FFACFFB7FD9E0177C09EF9ED" box="[637,713,1641,1663]" name="Mexico" pageId="8" pageNumber="331">Mexico</collectingCountry>
................................................ 
<taxonomicName id="01D64D4BFFACFFB7FAE40177C7CBF9EC" ID-CoL="7SV6B" baseAuthorityName="Say" baseAuthorityYear="1823" box="[1287,1436,1640,1662]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="fimbriolatus">
<emphasis id="F4A2EADAFFACFFB7FAE40177C7CBF9EC" bold="true" box="[1287,1436,1640,1662]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FAE40177C740F9EC" bold="true" box="[1287,1303,1641,1662]" italics="true" pageId="8" pageNumber="331">C</emphasis>
. 
<emphasis id="F4A2EADAFFACFFB7FAC00176C7CBF9EC" bold="true" box="[1315,1436,1640,1662]" italics="true" pageId="8" pageNumber="331">fimbriolatus</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D222E6DFFACFFB7FF740198C7CBF92B" pageId="8" pageNumber="331">
<paragraph id="C66936C8FFACFFB7FF740198C7CBF90E" blockId="8.[151,1437,1084,2015]" box="[151,1436,1670,1692]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF740198C7CBF90E" box="[151,1436,1670,1692]" pageId="8" pageNumber="331">
5. Posterior metatibial spur simple; elytra without lateral yellow margin. Smaller species, TL: 
<quantity id="012E9B2DFFACFFB7FBB20198C6EFF90E" box="[1105,1208,1670,1692]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="8" pageNumber="331" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
(
<emphasis id="F4A2EADAFFACFFB7FB250198C73DF909" bold="true" box="[1222,1386,1670,1692]" pageId="8" pageNumber="331">
<taxonomicName id="01D64D4BFFACFFB7FB250198C778F90E" ID-CoL="5MNZ" authorityName="Sharp" authorityYear="1882" box="[1222,1327,1670,1692]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="F4A2EADAFFACFFB7FB250198C778F90E" bold="true" box="[1222,1327,1670,1692]" italics="true" pageId="8" pageNumber="331">Megadytes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF9157A1FFACFFB7FAD60198C73DF909" box="[1333,1386,1670,1691]" pageId="8" pageNumber="331" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
).. 6
</keyLead>
</paragraph>
<paragraph id="C66936C8FFACFFB7FF7401BAC7CBF92B" blockId="8.[151,1437,1084,2015]" box="[151,1436,1700,1722]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF7401BAC7CBF92B" box="[151,1436,1700,1722]" pageId="8" pageNumber="331">
- Posterior metatibial spur bi- or trifurcate; elytra with lateral yellow margin. Larger species, TL: 
<quantity id="012E9B2DFFACFFB7FB9901BAC6B6F92B" box="[1146,1249,1700,1721]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.3" metricValueMin="2.7" pageId="8" pageNumber="331" unit="mm" value="35.0" valueMax="43.0" valueMin="27.0">27–43 mm</quantity>
.............. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D222E6DFFACFFB7FF7401DFC7CBF836" pageId="8" pageNumber="331">
<paragraph id="C66936C8FFACFFB7FF7401DFC7CBF8BD" blockId="8.[151,1437,1084,2015]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF7401DFC7CBF8BD" pageId="8" pageNumber="331">
6. Male protarsal palette more elongate, ratio long to short axis ca. 1.65; aedeagus as in 
<figureCitation id="5EED2A4DFFACFFB7FBF701DFC6D7F945" box="[1044,1152,1729,1751]" captionStart="FIGURES 6" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[192,1397,196,1877]" captionTargetId="figure-32@10.[177,1410,181,1885]" captionTargetPageId="10" captionText="FIGURES 6a–6j: a–f: Median lobe in lateral view (a, d), ventral view (b, e) and paramere (c, f) of (a–c) Megadytes fraternus; (d–f) M. flohri; g–j: median lobe in lateral view (g, i) and ventral view (h, j) of (g, h) M. lherminieri and (i, j) M. magnus. Figures 6g–6j adapted from figs 5 and 6 in Hendrich et al. (2019) (licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0)." figureDoi="http://doi.org/10.5281/zenodo.5649614" httpUri="https://zenodo.org/record/5649614/files/figure.png" pageId="8" pageNumber="331">Figs 6a–6c</figureCitation>
; males with anterior protarsal claw longer and thicker than posterior, same with mesotarsal claws; females with elytra covered with striae, not smooth as in males; TL: 18,5– 
<quantity id="012E9B2DFFACFFB7FE6501E5C39BF883" box="[390,460,1787,1809]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="331" unit="mm" value="22.0">22 mm</quantity>
. Species from 
<collectingCountry id="BEC17658FFACFFB7FDBA01E2C0DFF883" box="[601,648,1788,1809]" name="United States of America" pageId="8" pageNumber="331">USA</collectingCountry>
(
<collectingRegion id="0412F82AFFACFFB7FD7601E2C088F880" box="[661,735,1788,1810]" country="United States of America" name="Florida" pageId="8" pageNumber="331">Florida</collectingRegion>
), 
<collectingCountry id="BEC17658FFACFFB7FD0C01E2C16CF880" box="[751,827,1788,1810]" name="Mexico" pageId="8" pageNumber="331">Mexico</collectingCountry>
, 
<collectingCountry id="BEC17658FFACFFB7FCA601E2C12DF880" box="[837,890,1788,1810]" name="Cuba" pageId="8" pageNumber="331">Cuba</collectingCountry>
, parts of Central and South America................................................................................................... 
<taxonomicName id="01D64D4BFFACFFB7FB370007C7CBF8BD" ID-CoL="7VSTS" authorityName="Sharp" authorityYear="1882" box="[1236,1436,1817,1839]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="F4A2EADAFFACFFB7FB370007C7CBF8BD" bold="true" box="[1236,1436,1817,1839]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FB370007C6BEF8BC" bold="true" box="[1236,1257,1817,1838]" italics="true" pageId="8" pageNumber="331">M</emphasis>
. (s. str.) 
<emphasis id="F4A2EADAFFACFFB7FAA30007C7CBF8BD" bold="true" box="[1344,1436,1817,1839]" italics="true" pageId="8" pageNumber="331">fraternus</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C66936C8FFACFFB7FF740029C7CBF836" blockId="8.[151,1437,1084,2015]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF740029C7CBF836" pageId="8" pageNumber="331">
- Male protarsal palette less elongate, ratio long to short axis ca. 1.4; aedeagus as in 
<figureCitation id="5EED2A4DFFACFFB7FBED0028C62DF8DE" box="[1038,1146,1846,1868]" captionStart="FIGURES 6" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[192,1397,196,1877]" captionTargetId="figure-32@10.[177,1410,181,1885]" captionTargetPageId="10" captionText="FIGURES 6a–6j: a–f: Median lobe in lateral view (a, d), ventral view (b, e) and paramere (c, f) of (a–c) Megadytes fraternus; (d–f) M. flohri; g–j: median lobe in lateral view (g, i) and ventral view (h, j) of (g, h) M. lherminieri and (i, j) M. magnus. Figures 6g–6j adapted from figs 5 and 6 in Hendrich et al. (2019) (licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0)." figureDoi="http://doi.org/10.5281/zenodo.5649614" httpUri="https://zenodo.org/record/5649614/files/figure.png" pageId="8" pageNumber="331">Figs 6d–6f</figureCitation>
; males with anterior protarsal claw longer, but only slightly thicker than posterior, differences in mesotarsal claws less distinct; females with elytra smooth as in males or with a few very fine strioles (striae) near base; TL: 
<quantity id="012E9B2DFFACFFB7FCDC006FC19AF814" box="[831,973,1905,1927]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.25" metricValueMin="1.95" pageId="8" pageNumber="331" unit="mm" value="21.0" valueMax="22.5" valueMin="19.5">19.5–22.5 mm</quantity>
. Species from 
<collectingCountry id="BEC17658FFACFFB7FBB8006FC6F3F815" box="[1115,1188,1905,1927]" name="Mexico" pageId="8" pageNumber="331">Mexico</collectingCountry>
[
<collectingCountry id="BEC17658FFACFFB7FB53006EC6BCF814" box="[1200,1259,1904,1926]" name="Cuba" pageId="8" pageNumber="331">Cuba</collectingCountry>
?].................................................................................................... 
<taxonomicName id="01D64D4BFFACFFB7FB1B0091C7CBF836" ID-CoL="7VSTR" authorityName="Sharp" authorityYear="1882" box="[1272,1436,1934,1956]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="flohri">
<emphasis id="F4A2EADAFFACFFB7FB1B0091C7CBF836" bold="true" box="[1272,1436,1934,1956]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FB1B0091C75AF836" bold="true" box="[1272,1293,1935,1956]" italics="true" pageId="8" pageNumber="331">M</emphasis>
. (s. str.) 
<emphasis id="F4A2EADAFFACFFB7FA870090C7CBF836" bold="true" box="[1380,1436,1934,1956]" italics="true" pageId="8" pageNumber="331">flohri</emphasis>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D222E6DFFACFFB7FF7400B2C7CBF84C" pageId="8" pageNumber="331">
<paragraph id="C66936C8FFACFFB7FF7400B2C7CBF853" blockId="8.[151,1437,1084,2015]" box="[151,1436,1963,1986]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF7400B2C7CBF853" box="[151,1436,1963,1986]" pageId="8" pageNumber="331">
7. Posterior metatibial spur trifurcate. TL: 
<quantity id="012E9B2DFFACFFB7FD8200B2C09FF853" box="[609,712,1964,1985]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="8" pageNumber="331" unit="mm" value="28.0" valueMax="29.0" valueMin="27.0">27–29 mm</quantity>
(
<emphasis id="F4A2EADAFFACFFB7FD3400B5C1E6F853" bold="true" box="[727,945,1963,1985]" italics="true" pageId="8" pageNumber="331">
<taxonomicName id="01D64D4BFFACFFB7FD3400B5C117F853" ID-CoL="5MNZ" authorityName="Sharp" authorityYear="1882" box="[727,832,1963,1985]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="genus">Megadytes</taxonomicName>
Trifurcitus
</emphasis>
). Species from 
<collectingCountry id="BEC17658FFACFFB7FBAA00B2C6C2F850" box="[1097,1173,1964,1986]" name="Mexico" pageId="8" pageNumber="331">Mexico</collectingCountry>
.......... 
<emphasis id="F4A2EADAFFACFFB7FAF400B2C7CBF853" bold="true" box="[1303,1436,1963,1985]" pageId="8" pageNumber="331">
<emphasis id="F4A2EADAFFACFFB7FAF400B2C77BF853" bold="true" box="[1303,1324,1964,1985]" italics="true" pageId="8" pageNumber="331">M</emphasis>
. 
<taxonomicName id="01D64D4BFFACFFB7FADB00B5C7CBF853" authorityName="Aube" authorityYear="1883" box="[1336,1436,1963,1985]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="species" species="fallax" subGenus="Trifurcitus">
(
<emphasis id="F4A2EADAFFACFFB7FAA300B2C718F853" bold="true" box="[1344,1359,1964,1985]" italics="true" pageId="8" pageNumber="331">T</emphasis>
.) 
<emphasis id="F4A2EADAFFACFFB7FA8000B5C7CBF853" bold="true" box="[1379,1436,1963,1985]" italics="true" pageId="8" pageNumber="331">fallax</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="C66936C8FFACFFB7FF7400D7C7CBF84C" blockId="8.[151,1437,1084,2015]" box="[151,1436,1992,2015]" pageId="8" pageNumber="331">
<keyLead id="7D2795FDFFACFFB7FF7400D7C7CBF84C" box="[151,1436,1992,2015]" pageId="8" pageNumber="331">
- Posterior metatibial spur bifurcate. TL: 
<quantity id="012E9B2DFFACFFB7FDBC00D7C091F84C" box="[607,710,1993,2015]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="8" pageNumber="331" unit="mm" value="39.5" valueMax="43.0" valueMin="36.0">36–43 mm</quantity>
(
<emphasis id="F4A2EADAFFACFFB7FD3700D6C1F0F84D" bold="true" box="[724,935,1992,2015]" italics="true" pageId="8" pageNumber="331">
<taxonomicName id="01D64D4BFFACFFB7FD3700D6C16AF84C" ID-CoL="5MNZ" authorityName="Sharp" authorityYear="1882" box="[724,829,1992,2014]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="331" phylum="Arthropoda" rank="genus">Megadytes</taxonomicName>
Bifurcitus
</emphasis>
)........................................ 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7D222E6DFFADFFB6FF740786C7CAFE97" pageId="9" pageNumber="332">
<paragraph id="C66936C8FFADFFB6FF740786C7CBFF58" blockId="9.[151,1437,151,261]" pageId="9" pageNumber="332">
<keyLead id="7D2795FDFFADFFB6FF740786C7CBFF58" pageId="9" pageNumber="332">
8. Median lobe of aedeagus in lateral and ventral view as in 
<figureCitation id="5EED2A4DFFADFFB6FCED0789C12BFF3F" box="[782,892,151,173]" captionStart="FIGURES 6" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[192,1397,196,1877]" captionTargetId="figure-32@10.[177,1410,181,1885]" captionTargetPageId="10" captionText="FIGURES 6a–6j: a–f: Median lobe in lateral view (a, d), ventral view (b, e) and paramere (c, f) of (a–c) Megadytes fraternus; (d–f) M. flohri; g–j: median lobe in lateral view (g, i) and ventral view (h, j) of (g, h) M. lherminieri and (i, j) M. magnus. Figures 6g–6j adapted from figs 5 and 6 in Hendrich et al. (2019) (licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0)." figureDoi="http://doi.org/10.5281/zenodo.5649614" httpUri="https://zenodo.org/record/5649614/files/figure.png" pageId="9" pageNumber="332">
Figs 
<quantity id="012E9B2DFFADFFB6FCDD0789C101FF3F" box="[830,854,151,173]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="9" pageNumber="332" unit="g" value="6.0">6g</quantity>
–6h
</figureCitation>
; TL: 
<quantity id="012E9B2DFFADFFB6FC530789C67AFF3F" box="[944,1069,151,173]" metricMagnitude="-2" metricUnit="m" metricValue="3.925" metricValueMax="4.25" metricValueMin="3.6" pageId="9" pageNumber="332" unit="mm" value="39.25" valueMax="42.5" valueMin="36.0">36–42.5 mm</quantity>
. Species from 
<collectingCountry id="BEC17658FFADFFB6FB590789C751FF3F" box="[1210,1286,151,173]" name="Mexico" pageId="9" pageNumber="332">Mexico</collectingCountry>
, the Caribbean, large parts of Central and South America to southern 
<collectingCountry id="BEC17658FFADFFB6FD0107ABC14AFF59" box="[738,797,181,203]" name="Brazil" pageId="9" pageNumber="332">Brazil</collectingCountry>
..................................... 
<emphasis id="F4A2EADAFFADFFB6FB3C07ABC7CBFF58" bold="true" box="[1247,1436,180,202]" pageId="9" pageNumber="332">
<emphasis id="F4A2EADAFFADFFB6FB3C07ABC6A3FF58" bold="true" box="[1247,1268,181,202]" italics="true" pageId="9" pageNumber="332">M</emphasis>
. 
<taxonomicName id="01D64D4BFFADFFB6FAE307AAC7CBFF58" ID-CoL="7VSTM" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[1280,1436,180,202]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="lherminieri" subGenus="Bifurcitus">
(
<emphasis id="F4A2EADAFFADFFB6FAEB07ABC74FFF58" bold="true" box="[1288,1304,181,202]" italics="true" pageId="9" pageNumber="332">B</emphasis>
.) 
<emphasis id="F4A2EADAFFADFFB6FACF07AAC7CBFF58" bold="true" box="[1324,1436,180,202]" italics="true" pageId="9" pageNumber="332">lherminieri</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="C66936C8FFADFFB6FF7407CDC7CAFE97" blockId="9.[151,1437,151,261]" pageId="9" pageNumber="332">
<keyLead id="7D2795FDFFADFFB6FF7407CDC7CAFE97" pageId="9" pageNumber="332">
- Median lobe of aedeagus in lateral and ventral view as in 
<figureCitation id="5EED2A4DFFADFFB6FCC307CCC1D2FF7A" box="[800,901,210,232]" captionStart="FIGURES 6" captionStartId="10.[151,264,1909,1933]" captionTargetBox="[192,1397,196,1877]" captionTargetId="figure-32@10.[177,1410,181,1885]" captionTargetPageId="10" captionText="FIGURES 6a–6j: a–f: Median lobe in lateral view (a, d), ventral view (b, e) and paramere (c, f) of (a–c) Megadytes fraternus; (d–f) M. flohri; g–j: median lobe in lateral view (g, i) and ventral view (h, j) of (g, h) M. lherminieri and (i, j) M. magnus. Figures 6g–6j adapted from figs 5 and 6 in Hendrich et al. (2019) (licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0)." figureDoi="http://doi.org/10.5281/zenodo.5649614" httpUri="https://zenodo.org/record/5649614/files/figure.png" pageId="9" pageNumber="332">Figs 6i–6j</figureCitation>
; TL: 
<quantity id="012E9B2DFFADFFB6FC5F07CCC67FFF7A" box="[956,1064,210,232]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="9" pageNumber="332" unit="mm" value="41.0" valueMax="43.0" valueMin="39.0">39–43 mm</quantity>
. Species from 
<collectingCountry id="BEC17658FFADFFB6FB5807CCC750FF7A" box="[1211,1287,210,232]" name="Mexico" pageId="9" pageNumber="332">Mexico</collectingCountry>
, parts of South America until south-eastern 
<collectingCountry id="BEC17658FFADFFB6FE1107F1C067FE97" box="[498,560,239,261]" name="Brazil" pageId="9" pageNumber="332">Brazil</collectingCountry>
; most probably also from Central America......................... 
<emphasis id="F4A2EADAFFADFFB6FAE307EEC7CAFE97" bold="true" box="[1280,1437,239,261]" pageId="9" pageNumber="332">
<emphasis id="F4A2EADAFFADFFB6FAE307EEC742FE97" bold="true" box="[1280,1301,240,261]" italics="true" pageId="9" pageNumber="332">M</emphasis>
. 
<taxonomicName id="01D64D4BFFADFFB6FAC107F1C7CAFE97" ID-CoL="7VSTN" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[1314,1437,239,261]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="magnus" subGenus="Bifurcitus">
(
<emphasis id="F4A2EADAFFADFFB6FAC907EEC76DFE97" bold="true" box="[1322,1338,240,261]" italics="true" pageId="9" pageNumber="332">B</emphasis>
.) 
<emphasis id="F4A2EADAFFADFFB6FAAD07EEC7CAFE97" bold="true" box="[1358,1437,240,261]" italics="true" pageId="9" pageNumber="332">magnus</emphasis>
</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>