301 lines
48 KiB
XML
301 lines
48 KiB
XML
<document ID-CLB-Dataset="5072" ID-DOI="http://dx.doi.org/10.3897/zookeys.1055.68911" ID-GBIF-Dataset="fc0cd46a-d486-442b-ad09-fe9b906da4ac" ID-PMC="PMC8360822" ID-Pensoft-Pub="1313-2970-1055-69" ID-Pensoft-UUID="990477668F06527BB3C5B25F397886CF" ID-PubMed="34393573" ID-ZooBank="EDAA550DC7AB4C4D8CA91D988F4F3940" ModsDocID="1313-2970-1055-69" checkinTime="1628190754421" checkinUser="pensoft" docAuthor="Strong, Ellen E., Sirenko, Boris I. & McLean, James H." docDate="2021" docId="4E6C39F1EBAE5C71B7F4442F99FD7EFF" docLanguage="en" docName="ZooKeys 1055: 69-87" docOrigin="ZooKeys 1055" docPubDate="2021-08-05" docSource="http://dx.doi.org/10.3897/zookeys.1055.68911" docTitle="Pseudomohnia sp." docType="treatment" docVersion="6" id="990477668F06527BB3C5B25F397886CF" lastPageNumber="69" masterDocId="990477668F06527BB3C5B25F397886CF" masterDocTitle="The only species of Mohnia Friele, 1879 (Caenogastropoda, Buccinoidea, Buccinidae) in the North Pacific represents an unrecognized new genus of Newtoniellidae (Triphoroidea)" masterLastPageNumber="87" masterPageNumber="69" pageNumber="69" updateTime="1720490775729" updateUser="ExternalLinkService">
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<mods:mods id="C331ADCC73DD22597284ECE03A789EAF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1DC7463F1B08FBBB71160E7E17C6D584">
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<mods:title id="201EF37F0C9117CEB2DD146F28EE1E1C">The only species of Mohnia Friele, 1879 (Caenogastropoda, Buccinoidea, Buccinidae) in the North Pacific represents an unrecognized new genus of Newtoniellidae (Triphoroidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="40CB40188B95855C81998B0EF41188F6" type="personal">
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<mods:role id="A75448B953215CAB3211A0BD3C852D44">
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<mods:roleTerm id="02D90E7CC810F5CA5F2A6634ABA9342D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA0BDB46572070E937909141A53517F0">Strong, Ellen E.</mods:namePart>
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<mods:nameIdentifier id="06469E0A85A407F12363F2F88A149E22" type="ORCID">https://orcid.org/0000-0001-7181-4114</mods:nameIdentifier>
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<mods:affiliation id="ECE24D10DE77A2B27CAE3E80440FFA93">National Museum of Natural History, Smithsonian Institution, PO Box 37012, MRC 163, Washington, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier id="13244C7FB023314BF29071E933D5F699" type="email">stronge@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="BA5A8FD2845F62B574652EC5822FEEB9" type="personal">
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<mods:role id="0B08ACF3545267872F589474645DF707">
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<mods:roleTerm id="D731601CC045935E3E24BA531AF17CA1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A52B0FD5FCABD98D27F9BC3D51B34921">Sirenko, Boris I.</mods:namePart>
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<mods:affiliation id="B584B88CB9AC53ED30B8681F0E4EE2F9">Zoological Institute, Russian Academy of Sciences, St. Petersburg, 199034, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="AFDD47B105983CF3CA17FF778AD7E403" type="personal">
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<mods:role id="83D3D6570DFA08792A432C0F307705BE">
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<mods:roleTerm id="D44C63B5AE1C90C82A203B4665C75108">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F4CCA813ED419FD31248EFB80B9B1542">McLean, James H.</mods:namePart>
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<mods:affiliation id="199D58FCE9F176C8EB46BD1F524F093C">Deceased; formerly of Natural History Museum of Los Angeles County, 900 Exposition Blvd, Los Angeles, CA 90007, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="DCF3BFBF32F858493AD30EA908E73E50">text</mods:typeOfResource>
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<mods:titleInfo id="C31A21155FA584578DAAAF3015BCF47A">
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<mods:title id="6F40A871110129B3234B9BE5B9BE9326">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="C0AE852398811030B29720A93A0EA78C">
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<mods:date id="FA40A0C986C017E05E24701B63E33B92">2021</mods:date>
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<mods:number id="2CFE98A8634439E2C608FCD1348FD401">2021-08-05</mods:number>
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<mods:detail id="DD6A96798A9EA8522542659774C6A743" type="volume">
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<mods:number id="E98628EFC294AEB1A1B14984F8954256">1055</mods:number>
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<mods:start id="0C625643D8B35D64FA90A1AB61EDCA5C">69</mods:start>
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<mods:end id="C336AA31DA08B031046FC8711B839C98">87</mods:end>
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<mods:location id="D700FDAB99E4072694CF5B3BE76673BE">
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<mods:url id="27C2F194B0C83DDE8D68342A20E192FB">http://dx.doi.org/10.3897/zookeys.1055.68911</mods:url>
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<mods:classification id="016CF8034C71F78A2FE77C2A310B1098">journal article</mods:classification>
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<mods:identifier id="AA1D0D83BB15F45731A0DF18E5DA7DCB" type="DOI">http://dx.doi.org/10.3897/zookeys.1055.68911</mods:identifier>
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<mods:identifier id="F05F2D2348D8B4465CCF07F5A63855B4" type="Pensoft-Pub">1313-2970-1055-69</mods:identifier>
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<mods:identifier id="E9E48CCD62BFAC5D1FAA79C73EC5A468" type="ZooBank">EDAA550DC7AB4C4D8CA91D988F4F3940</mods:identifier>
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<mods:identifier id="CCA50FD76D754D4C76B9FE2CF26DB569" type="Pensoft-UUID">990477668F06527BB3C5B25F397886CF</mods:identifier>
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</mods:mods>
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<subSection id="300C96AA5A2947280BA2B4773404CEEA" pageId="0" pageNumber="69" type="results">
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<treatment id="4E6C39F1EBAE5C71B7F4442F99FD7EFF" ID-GBIF-Taxon="183925355" LSID="urn:lsid:plazi:treatment:4E6C39F1EBAE5C71B7F4442F99FD7EFF" httpUri="http://treatment.plazi.org/id/4E6C39F1EBAE5C71B7F4442F99FD7EFF" lastPageNumber="69" pageId="0" pageNumber="69">
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<subSubSection id="45B6D6D57FF35731FF1796697FFF017F" pageId="0" pageNumber="69" type="nomenclature">
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<paragraph id="D11A162FB53EC320DA287C43347A925F" pageId="0" pageNumber="69">
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<taxonomicName id="5F4C02FE31741AB1B17F2BB61E5BA0E6" ID-CoL="8NCRZ" LSID="4E6C39F1-EBAE-5C71-B7F4-442F99FD7EFF" authority="sp." authorityName="sp." class="Gastropoda" family="Newtoniellidae" genus="Pseudomohnia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pseudomohnia" pageId="0" pageNumber="69" phylum="Mollusca" rank="genus">Pseudomohnia sp.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="8D621CE2D047EFC3118EBE915C73D8BA" pageId="0" pageNumber="69" type="description">
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<paragraph id="894119DC37460610103AE49AF80FC84F" pageId="0" pageNumber="69">
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<figureCitation id="D9E7AA2CE0F50F1355DA5B226D572BC4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shell morphology of Pseudomohnia gen. nov. A, B Pseudomohnia kurilana (Dall, 1913) comb. nov A lectotype, USNM 205224. Kuril Islands B ZIN 57494 / 1. Kuril Islands C, D Pseudomohnia sp. LACM 1997 - 156.7 Aleutian Islands E-I Pseudomohnia rogerclarki sp. nov. Aleutian Islands E holotype, LACM 3776 F LACM 1997 - 174 G LACM 1997 - 174 H LACM 1997 - 163.21 I LACM 1997 - 165.20. Scale bar: 1 cm." figureDoi="10.3897/zookeys.1055.68911.figure1" httpUri="https://binary.pensoft.net/fig/574832" pageId="0" pageNumber="69">Figs 1C, D</figureCitation>
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<figureCitation id="806DFDBAC3AB2D996542EEB86A6F1DDC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">, 5B-E</figureCitation>
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<figureCitation id="4B3527E80429BF97A1D8B5BF0D0949EA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula morphology of Pseudomohnia sp., LACM 1997 - 168.10 A radular ribbon B detail of rachidian and lateral teeth C detail of rachidian D, E detail of lateral and marginal teeth F detail of inner and outer marginal teeth. Scale bars: 20 µm (C, F); 100 µm (A, B, D, E)." figureDoi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DE690C175779F758E0CBF6D6657E67B3" pageId="0" pageNumber="69" type="materials_examined">
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<paragraph id="6DDBBA7F1D316E478C15890B15A78CA1" pageId="0" pageNumber="69">Material examined.</paragraph>
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<paragraph id="AEE8C5148E8F91E7459E3864FB314970" pageId="0" pageNumber="69">
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<materialsCitation id="D701BC5BF1AE9D63AEBFDA7776398BEB" collectingDate="1997-07-27" collectingDateMax="1997-08-09" collectingDateMin="1997-07-27" collectorName="V Dominator, RN Clark" county="Buldir Island" elevation="384" latitude="52.308334" location="Aleutian Islands" longLatPrecision="57" longitude="175.81667" municipality="Amchitka Island" specimenCount="1">
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<location id="3A22CA00507FAB0F8F275EA0DE72E242" LSID="urn:lsid:plazi:treatment:4E6C39F1EBAE5C71B7F4442F99FD7EFF:3A22CA00507FAB0F8F275EA0DE72E242" county="Buldir Island" latitude="52.308334" longLatPrecision="57" longitude="175.81667" municipality="Amchitka Island" name="Aleutian Islands">Aleutian Islands</location>
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• 2 spms;
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<location id="A317CA8A84D11620A1EAA0997B371A85" LSID="urn:lsid:plazi:treatment:4E6C39F1EBAE5C71B7F4442F99FD7EFF:A317CA8A84D11620A1EAA0997B371A85" county="Buldir Island" latitude="52.308334" longLatPrecision="57" longitude="175.81667" municipality="Amchitka Island" name="Rat Islands">Rat Islands</location>
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, southwest of
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<collectingMunicipality id="DAC06569FD34FEA61CE9BF7761B7A109">Amchitka Island</collectingMunicipality>
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;
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<geoCoordinate id="69906C45A7CA02D7A9548F0179074F6E" degrees="51" direction="north" minutes="27.70" orientation="latitude" precision="9" value="51.461666">51°27.70'N</geoCoordinate>
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,
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<geoCoordinate id="4A27D38B91D80F288CAAFEB384F95C8C" degrees="178" direction="east" minutes="35.0" orientation="longitude" precision="92" value="178.58333">178°35.0'E</geoCoordinate>
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;
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<quantity id="76F0C3D661CC3DA5ABABD3AB4BA38E24" metricMagnitude="2" metricUnit="m" metricValue="3.84" unit="m" value="384.0">
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<elevation id="98CC80BA28CC546897BF7E023E3F45F0" metricMagnitude="2" metricUnit="m" metricValue="3.84" unit="m" value="384.0">384 m</elevation>
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</quantity>
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;
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<collectingDate id="A7CE4899416CDA1660D1BD0C6194B0AD" value="1997-07-27">27 July 1997</collectingDate>
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; R/
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<collectorName id="E96A26F406A07518408F4DAC466A7A57">V Dominator</collectorName>
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stn 23-971-181;
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<collectorName id="04A23A2BBC4328854F89ACF9B9DDC169">RN Clark</collectorName>
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leg.; LACM 1997-156.7 (Fig.
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<figureCitation id="EFC1B9FAE151CEA49DA7A339A8CE949F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shell morphology of Pseudomohnia gen. nov. A, B Pseudomohnia kurilana (Dall, 1913) comb. nov A lectotype, USNM 205224. Kuril Islands B ZIN 57494 / 1. Kuril Islands C, D Pseudomohnia sp. LACM 1997 - 156.7 Aleutian Islands E-I Pseudomohnia rogerclarki sp. nov. Aleutian Islands E holotype, LACM 3776 F LACM 1997 - 174 G LACM 1997 - 174 H LACM 1997 - 163.21 I LACM 1997 - 165.20. Scale bar: 1 cm." figureDoi="10.3897/zookeys.1055.68911.figure1" httpUri="https://binary.pensoft.net/fig/574832" pageId="0" pageNumber="69">1C, D</figureCitation>
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); • 1 spm; southwest of
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<collectingCounty id="C2B6CEB938F5A46EB74C51D806CE4F93">Buldir Island</collectingCounty>
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;
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<geoCoordinate id="A9482EEECDEEB5D72C5DE437FCA2BD59" degrees="52" direction="north" minutes="18.50" orientation="latitude" precision="9" value="52.308334">52°18.50'N</geoCoordinate>
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,
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<geoCoordinate id="884D34B6C36BB540C3EEED3847E605C3" degrees="175" direction="east" minutes="49.0" orientation="longitude" precision="92" value="175.81667">175°49.0'E</geoCoordinate>
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;
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<quantity id="FE4EA1F41F7AD99B1F6ED4344096C964" metricMagnitude="2" metricUnit="m" metricValue="3.25" unit="m" value="325.0">
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<elevation id="1A7BB7AAB6E35009034AA0B78FD2D649" metricMagnitude="2" metricUnit="m" metricValue="3.25" unit="m" value="325.0">325 m</elevation>
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</quantity>
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;
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<collectingDate id="4910197C53C637A077CAEDCA442F4355" value="1997-08-09">9 August 1997</collectingDate>
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; R/
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<collectorName id="BBFEC9C040E61ACDB6ABADE63EC33DC5">V Dominator</collectorName>
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stn 23-971-243;
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<collectorName id="05369C419CA9519B7B70CECEAC66403F">RN Clark</collectorName>
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leg.; LACM 1997-168.10 (Figs
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<figureCitation id="65711E29DB368971167D0AC4C8E96E28" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5B-E</figureCitation>
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,
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<figureCitation id="A9F81D6E2809CF4965EEE7FC1E0C7166" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula morphology of Pseudomohnia sp., LACM 1997 - 168.10 A radular ribbon B detail of rachidian and lateral teeth C detail of rachidian D, E detail of lateral and marginal teeth F detail of inner and outer marginal teeth. Scale bars: 20 µm (C, F); 100 µm (A, B, D, E)." figureDoi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69">7</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="999DEB8A667C0AEB57F076047E74E5B9" pageId="0" pageNumber="69" type="description">
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<paragraph id="5D085EA6E888EEEC4F7DF19CB356C36E" pageId="0" pageNumber="69">Description.</paragraph>
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<paragraph id="B4A34689BB1C9FC1D2E7A348C2447088" pageId="0" pageNumber="69">
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<emphasis id="9B50FF4114C6E57B5CECD4F049D44CBF" bold="true" italics="true" pageId="0" pageNumber="69">Shell</emphasis>
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[LACM 1997-156.7]. Shell broadly turreted, spire angle ca. 41°, ~ 24 mm in adult shell length, consisting of approximately eight, thin, convex whorls, separated by deeply impressed suture (Fig.
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<figureCitation id="986E1AEE3F7E2A5B27BBA907F5EFCA7D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shell morphology of Pseudomohnia gen. nov. A, B Pseudomohnia kurilana (Dall, 1913) comb. nov A lectotype, USNM 205224. Kuril Islands B ZIN 57494 / 1. Kuril Islands C, D Pseudomohnia sp. LACM 1997 - 156.7 Aleutian Islands E-I Pseudomohnia rogerclarki sp. nov. Aleutian Islands E holotype, LACM 3776 F LACM 1997 - 174 G LACM 1997 - 174 H LACM 1997 - 163.21 I LACM 1997 - 165.20. Scale bar: 1 cm." figureDoi="10.3897/zookeys.1055.68911.figure1" httpUri="https://binary.pensoft.net/fig/574832" pageId="0" pageNumber="69">1C, D</figureCitation>
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); growth indeterminate. Shell whitish, with thick, velvety periostracum. Larval shell non-planktotrophic, ~ 3 elevated, constricted whorls, with smooth, blunt nucleus; well-defined opisthocyrt riblets and spiral threads producing cancellate sculpture on subsequent whorls. Axial elements becoming more closely spaced toward teleoconch transition. Inferred transition to teleoconch marked by change in orientation of axial sculpture and slight expansion in whorl diameter. Teleoconch initially with six or seven thin spiral cords, somewhat irregular in width and spacing; cords becoming flatter, broader and less distinct on later whorls and on base and intercalated by additional cords. Spiral ornament crossed by variably developed, well separated, weakly prosocline axial threads and growth increments; axial threads becoming obsolete on body whorl. Aperture broad, outer lip thin, sharp. Axis weakly gyrate, pervious; columellar plait lacking. Anterior canal short, slightly recurved.
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</paragraph>
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<paragraph id="194FA7E31CDE12A4F1F15A7DAFAEF0ED" pageId="0" pageNumber="69">
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<emphasis id="13F016309BC814EA81EC8FA792C58F33" bold="true" italics="true" pageId="0" pageNumber="69">Operculum</emphasis>
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[LACM 1997-156.7]. Operculum thin, corneous, honey in color, thinning toward edges; paucispiral, nucleus eccentric, occupying ca. 42% of operculum length.
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</paragraph>
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<paragraph id="50068F865CA127BACE6EAB15F3A4515A" pageId="0" pageNumber="69">
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<emphasis id="D26F0F73D3C203E5E259A09FE9779773" bold="true" italics="true" pageId="0" pageNumber="69">Radula</emphasis>
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[LACM 1997-168.10]. Radular ribbon comprising 37 rows, ~ 5.7 mm in length. Radula taenioglossate (Fig.
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<figureCitation id="3C4BDA52218882086CC9911B87F0D308" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula morphology of Pseudomohnia sp., LACM 1997 - 168.10 A radular ribbon B detail of rachidian and lateral teeth C detail of rachidian D, E detail of lateral and marginal teeth F detail of inner and outer marginal teeth. Scale bars: 20 µm (C, F); 100 µm (A, B, D, E)." figureDoi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69">7A</figureCitation>
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). Rachidian small, concave, with slight constriction below broad cutting edge, tapering to flat, narrow base. Cutting edge bearing single prominent, broadly triangular, bluntly pointed, finely serrated cusp, and single smaller, irregular denticle on each side (Fig.
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<figureCitation id="844CF8410D6F701801BE7DAD6C21C2F1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula morphology of Pseudomohnia sp., LACM 1997 - 168.10 A radular ribbon B detail of rachidian and lateral teeth C detail of rachidian D, E detail of lateral and marginal teeth F detail of inner and outer marginal teeth. Scale bars: 20 µm (C, F); 100 µm (A, B, D, E)." figureDoi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69">7B, C</figureCitation>
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). Radular membrane diagonally creased between rachidian and lateral teeth of each row. Lateral teeth robust, broad, with smooth inner edge of shaft curving posteriorly; cutting edge with prominent, dagger-like, pointed cusp and small, blunt inner cusp (Fig.
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<figureCitation id="BC2786762C45B4165B19AB2B6B01F8D2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula morphology of Pseudomohnia sp., LACM 1997 - 168.10 A radular ribbon B detail of rachidian and lateral teeth C detail of rachidian D, E detail of lateral and marginal teeth F detail of inner and outer marginal teeth. Scale bars: 20 µm (C, F); 100 µm (A, B, D, E)." figureDoi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69">7B, D, E</figureCitation>
|
|
). Marginal teeth long, slender, with cylindrical shafts and constriction below claw-like tips; cutting edges of inner and outer marginal teeth bearing three to five elongate, curving, pointed cusps, somewhat angular in cross section (Fig.
|
|
<figureCitation id="E6A59701444E34A0F41021D9CBDFFA37" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula morphology of Pseudomohnia sp., LACM 1997 - 168.10 A radular ribbon B detail of rachidian and lateral teeth C detail of rachidian D, E detail of lateral and marginal teeth F detail of inner and outer marginal teeth. Scale bars: 20 µm (C, F); 100 µm (A, B, D, E)." figureDoi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69">7E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="8F6435F00D87A243194E774E668773E1" doi="10.3897/zookeys.1055.68911.figure7" httpUri="https://binary.pensoft.net/fig/574838" pageId="0" pageNumber="69" start="Figure 7" startId="F7">
|
|
<paragraph id="E9F349F2751AF2ACC3BE5773471561A6" pageId="0" pageNumber="69">
|
|
<emphasis id="2FBA4B998DA2464C0109536EC2E69442" bold="true" pageId="0" pageNumber="69">Figure 7.</emphasis>
|
|
Radula morphology of
|
|
<taxonomicName id="FFE1378F14B459F6F3FC4C0C1C130B90" class="Gastropoda" family="Newtoniellidae" genus="Pseudomohnia" kingdom="Animalia" lsidName="Pseudomohnia" pageId="0" pageNumber="69" phylum="Mollusca" rank="genus">
|
|
<emphasis id="14F5A21446E6902290D0A226EEC8632A" italics="true" pageId="0" pageNumber="69">Pseudomohnia</emphasis>
|
|
</taxonomicName>
|
|
sp., LACM 1997-168.10
|
|
<emphasis id="68866D70A85C8CE7E171A61C60286A5F" bold="true" pageId="0" pageNumber="69">A</emphasis>
|
|
radular ribbon
|
|
<emphasis id="E275D104E47D78596D80D07CA82E9D44" bold="true" pageId="0" pageNumber="69">B</emphasis>
|
|
detail of rachidian and lateral teeth
|
|
<emphasis id="70ED30E98202348126916D733F1E3D74" bold="true" pageId="0" pageNumber="69">C</emphasis>
|
|
detail of rachidian
|
|
<emphasis id="26579294D98A8BF18042A97DFB71CB7A" bold="true" pageId="0" pageNumber="69">D, E</emphasis>
|
|
detail of lateral and marginal teeth
|
|
<emphasis id="EE0085E4B1CB5F4F41AAA3DFBC52AD07" bold="true" pageId="0" pageNumber="69">F</emphasis>
|
|
detail of inner and outer marginal teeth. Scale bars: 20
|
|
<normalizedToken id="02B83475D74416AD6024E2AED9A18E57" originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis id="A333116924633C9581503F07C3B0014D" bold="true" pageId="0" pageNumber="69">C, F</emphasis>
|
|
); 100
|
|
<normalizedToken id="A8038B32C98ADD15A7508EFE9DB5ACA8" originalValue="µm">µm</normalizedToken>
|
|
(
|
|
<emphasis id="BAE4A12E2E29AA33EC4768D442EABE9A" bold="true" pageId="0" pageNumber="69">A, B, D, E</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="C55B28E22E06AD533B7FE8D943CB927F" pageId="0" pageNumber="69">
|
|
<emphasis id="82A816A72CA3F8B843B4B2E9E2AD0840" bold="true" italics="true" pageId="0" pageNumber="69">Anatomy</emphasis>
|
|
[LACM 1997-168.10]. Foot elongate oval. Propodium large, crescent shaped, with deep, triangular propodial groove (Fig.
|
|
<figureCitation id="87FC2DEF78CE21803CBC940CBA58F63E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5B</figureCitation>
|
|
). Shallow furrow (= epipodial skirt) continuous with opercular lobe, evident along sides of foot, becoming obsolete in deep groove where propodium joins neck. Foot sole, particular that of metapodium, deeply wrinkled with many deep transverse grooves. Sole divided longitudinally by deep medial cleft. Rather large bilobed mesopodial pedal gland within foot below and in front of pedal ganglia on both sides of cleft, opening via large pore near center of sole.
|
|
</paragraph>
|
|
<paragraph id="29F8143CA28316440E65207B0E907FD7" pageId="0" pageNumber="69">
|
|
Head with short, broad, muscular snout and long, tapering cephalic tentacles (Fig.
|
|
<figureCitation id="180906A3BE7EDFE4DFD14186D45E0417" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5B</figureCitation>
|
|
). Eyes conspicuous, on prominent ocular peduncles at outer bases of tentacles. Mantle edge smooth, with short, clearly defined siphon at left. Columellar muscle short, broad, extending roughly one-half whorl to base of mantle cavity. Ctenidium long, extending from siphon to base of mantle cavity, with long, narrowly triangular leaflets. Osphradium forming tall, narrow, undulating ridge, extending almost entire length of gill, from near anterior end almost to base. Hypobranchial gland well developed. Rectum broad, filled with sponge spicules, terminating in papillate anus near mantle edge at right. Rectum bordering pallial glandular gonoduct. Pallial gonoduct open for much of its length, lacking accessory pouches; thick, highly glandular tissue subdivided by deep transverse grooves (Fig.
|
|
<figureCitation id="58C4E9DDBFC637CFA0668F28D819D6D8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5E</figureCitation>
|
|
). Penis lacking.
|
|
</paragraph>
|
|
<paragraph id="95537510FC249BA90A7C58BCFB2289F6" pageId="0" pageNumber="69">
|
|
Proboscis acrembolic. Introvert rather short, muscular, oral tube not cuticularized. Jaws large, robust, surrounding anterior end of odontophore (Fig.
|
|
<figureCitation id="66909F5642AC2FE020454B9EF0EF520B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5C</figureCitation>
|
|
). Buccal mass large with long radular sac (Fig.
|
|
<figureCitation id="47B2C0729268D7E6B32119B1499CFF83" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5C, D</figureCitation>
|
|
) emerging mid-ventrally near posterior end, continuing to right before arcing dorsally across anterior esophagus just behind buccal mass with posterior, weakly-bifid tip lying on left side of esophagus near supra-esophageal ganglion. Posterior buccal cavity with broad, deep, subtriangular, acinous salivary glands on either side of dorsal food groove. Anterior esophagus not cuticularized. Large mid-esophageal gland (Fig.
|
|
<figureCitation id="5084782124AFF1E9FCE8B848EC1BD3D6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5C</figureCitation>
|
|
) with shallow, glandular septae and voluminous lumen, narrowing to posterior esophagus near end of mantle cavity, with ca. seven low, longitudinal folds.
|
|
</paragraph>
|
|
<paragraph id="5E65ABB9A5F35A01EC1E90597D81D514" pageId="0" pageNumber="69">
|
|
Nervous system epiathroid. Circum-esophageal nerve ring surrounding anterior esophagus (Fig.
|
|
<figureCitation id="40876E0E47101710450C80FEB60CA421" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5C</figureCitation>
|
|
) just behind buccal apparatus. Nerve ring highly asymmetrical, with both cerebral ganglia lying on left side of esophagus; left cerebral ganglion below and slightly in front of right ganglion, joined by very short but distinct commissure. Buccal ganglia (Fig.
|
|
<figureCitation id="D12D71C9C86959E02337E5A04FD04448" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5D</figureCitation>
|
|
) joined by short commissure, lying on either side of posterior buccal mass at emergence of anterior esophagus, just below salivary glands. Small pleural ganglia lying immediately behind cerebral ganglia, separated by narrow constrictions. Long connective joining right pleural with supra-esophageal ganglion (Fig.
|
|
<figureCitation id="A2D1BAA7BF524545B7B8E9540E3FB4FF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anatomy of Pseudomohnia gen. nov. A Pseudomohnia kurilana (Dall, 1913) comb. nov., lectotype, USNM 205224 B-E Pseudomohnia sp., LACM 1997 - 168.10 A, B external anatomy, dorsal view, mantle cavity opened via mid-dorsal incision C foregut, dorsal view D foregut, left-lateral view E pallial gonoduct, left-lateral view. Scale bars 1 mm. Abbreviations: ae, anterior esophagus; bg, buccal ganglion; cs, longitudinal cleft in foot sole; ct, ctendium; dg, digestive gland; go, gonad; hg, hypobranchial gland; meg, mid-esophageal gland; od, odontophore; op, opercular lobe; os, osphradium; pg, pallial gonoduct; ppg, propodeal pedal gland; r, rectum; rm, retractor muscles; rs, radular sac; sg, salivary gland; si, siphon; sp, supra-esophageal ganglion; sto, stomach; t, cephalic tentacle." figureDoi="10.3897/zookeys.1055.68911.figure5" httpUri="https://binary.pensoft.net/fig/574836" pageId="0" pageNumber="69">5C</figureCitation>
|
|
) at left side of cephalic hemocoel near tip of radular ribbon. Sub-esophageal ganglion lying below right side of anterior esophagus, separated from left pleural ganglion by slight constriction. Long, highly asymmetric connectives joining cerebral and pleural ganglia with pedal ganglia lying within foot at short distance anterior to cerebral ganglia. Small statocysts with numerous, tiny statoconia on postero-dorsal surface of pedal ganglia. Pedal ganglia joined by short, thick commissure.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0D899E4A1DA24C2E5A5A514258DCB905" pageId="0" pageNumber="69" type="distribution">
|
|
<paragraph id="F605C99E7265D1052493902D06E6A83C" pageId="0" pageNumber="69">Distribution and ecology.</paragraph>
|
|
<paragraph id="9CFBEBEF720B17CC56B7655D44C6B3D8" pageId="0" pageNumber="69">
|
|
Known only from the Aleutian Islands (Fig.
|
|
<figureCitation id="22E8DD3A13888ABBB8FD8195EF7B96D2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Pseudomohnia gen. nov. in the North Pacific. Squares, Pseudomohnia kurilana (Dall, 1913) comb. nov.; circles, Pseudomohnia rogerclarki sp. nov.; stars, Pseudomohnia sp. Symbols with black fill are type localities. Scale in kilometers." figureDoi="10.3897/zookeys.1055.68911.figure2" httpUri="https://binary.pensoft.net/fig/574833" pageId="0" pageNumber="69">2</figureCitation>
|
|
) in 325-384 m, feeding on sponges.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3169914DF744C7690328ACCE5D6D46B9" pageId="0" pageNumber="69" type="remarks">
|
|
<paragraph id="68A9EDCDDBD9E8C11704AFDF4F48DB3F" pageId="0" pageNumber="69">Remarks.</paragraph>
|
|
<paragraph id="C48EA52D9BEA083782F2D551AED28D35" pageId="0" pageNumber="69">
|
|
Given the absence of accessory sperm storage pouches in the pallial gonoduct (
|
|
<bibRefCitation id="2ABB8D2A8FA2FF1253342D03D4E55FBF" DOI="https://doi.org/10.1017/S0025315400052784" author="Fretter, V" journalOrPublisher="Journal of the Marine Biological Association of Great Britain" pageId="0" pageNumber="69" pagination="567 - 586" refId="B10" refString="Fretter, V, 1951. Observations on the life histories and functional morphology of Cerithiopsis tubercularis (Montagu) and Triphora perversa (L). Journal of the Marine Biological Association of Great Britain 29: 567 - 586, DOI: https://doi.org/10.1017/S0025315400052784" title="Observations on the life histories and functional morphology of Cerithiopsis tubercularis (Montagu) and Triphora perversa (L)." url="https://doi.org/10.1017/S0025315400052784" volume="29" year="1951">Fretter 1951</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="B77F8367339E06C6B1D5DE614AEEC0FC" author="Houston, RS" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="69" pagination="183 - 189" refId="B16" refString="Houston, RS, 1985. Genital ducts of the Cerithiacea (Gastropoda: Mesogastropoda) from the Gulf of California. Journal of Molluscan Studies 51: 183 - 189" title="Genital ducts of the Cerithiacea (Gastropoda: Mesogastropoda) from the Gulf of California." volume="51" year="1985">Houston 1985</bibRefCitation>
|
|
), the dissected individual is inferred to be male.
|
|
</paragraph>
|
|
<paragraph id="2D48D723A654F593362CBD455FAA316D" pageId="0" pageNumber="69">
|
|
The disposition of the remnants of the shell from LACM 1997-168.10 on which the anatomical observations were made is unknown and there is no known photograph (RN Clark, L Groves, pers. comm.). The anatomy and radula morphology show several differences compared to
|
|
<taxonomicName id="0D30B803AA36EE045F129C1988165947" authorityName="Strong & Sirenko & McLean" authorityYear="2021" baseAuthorityName="Dall" baseAuthorityYear="1913" class="Gastropoda" family="Newtoniellidae" genus="Pseudomohnia" kingdom="Animalia" lsidName="Pseudomohnia kurilana" pageId="0" pageNumber="69" phylum="Mollusca" rank="species" species="kurilana">
|
|
<emphasis id="1F445B40E4397B060E7805DD71FECBD2" italics="true" pageId="0" pageNumber="69">Pseudomohnia kurilana</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="2D568B38ACE886394C538CA1C49B0220" lsidName="P. rogerclarki" pageId="0" pageNumber="69" rank="species" species="rogerclarki">
|
|
<emphasis id="D45ED6CA1E4FB73908BBA3E6E2309F77" italics="true" pageId="0" pageNumber="69">P. rogerclarki</emphasis>
|
|
</taxonomicName>
|
|
(see below). The two specimens in LACM 1997-156.7 (Fig.
|
|
<figureCitation id="71FD4FC8832FA1AC76A4326A325A7D24" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shell morphology of Pseudomohnia gen. nov. A, B Pseudomohnia kurilana (Dall, 1913) comb. nov A lectotype, USNM 205224. Kuril Islands B ZIN 57494 / 1. Kuril Islands C, D Pseudomohnia sp. LACM 1997 - 156.7 Aleutian Islands E-I Pseudomohnia rogerclarki sp. nov. Aleutian Islands E holotype, LACM 3776 F LACM 1997 - 174 G LACM 1997 - 174 H LACM 1997 - 163.21 I LACM 1997 - 165.20. Scale bar: 1 cm." figureDoi="10.3897/zookeys.1055.68911.figure1" httpUri="https://binary.pensoft.net/fig/574832" pageId="0" pageNumber="69">1C, D</figureCitation>
|
|
) have a broader spire angle than
|
|
<taxonomicName id="83C1331D8C1AA54DE055783DB6B83174" lsidName="P. rogerclarki" pageId="0" pageNumber="69" rank="species" species="rogerclarki">
|
|
<emphasis id="3AA334DBAC30D652A79CA38EA2599959" italics="true" pageId="0" pageNumber="69">P. rogerclarki</emphasis>
|
|
</taxonomicName>
|
|
, but share the constricted, elevated early whorls. The spiral sculpture of the teleoconch is less distinct and more irregular than in
|
|
<taxonomicName id="9820DA36327FB9AA35DEC4204C69AB08" lsidName="P. kurilana" pageId="0" pageNumber="69" rank="species" species="kurilana">
|
|
<emphasis id="122DE54D2C2B72E1955497C77CFCE276" italics="true" pageId="0" pageNumber="69">P. kurilana</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="5CCBEB4D2D63B01BBB70DB19F95DF4CC" lsidName="P. rogerclarki" pageId="0" pageNumber="69" rank="species" species="rogerclarki">
|
|
<emphasis id="C4E091EDF2614577C17A2DFFD85DA298" italics="true" pageId="0" pageNumber="69">P. rogerclarki</emphasis>
|
|
</taxonomicName>
|
|
. Fragmentary soft parts from one of the two specimens in LACM 1997-156.7 (Fig.
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<figureCitation id="42BEF8D0848E049944838581C1EE569C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Shell morphology of Pseudomohnia gen. nov. A, B Pseudomohnia kurilana (Dall, 1913) comb. nov A lectotype, USNM 205224. Kuril Islands B ZIN 57494 / 1. Kuril Islands C, D Pseudomohnia sp. LACM 1997 - 156.7 Aleutian Islands E-I Pseudomohnia rogerclarki sp. nov. Aleutian Islands E holotype, LACM 3776 F LACM 1997 - 174 G LACM 1997 - 174 H LACM 1997 - 163.21 I LACM 1997 - 165.20. Scale bar: 1 cm." figureDoi="10.3897/zookeys.1055.68911.figure1" httpUri="https://binary.pensoft.net/fig/574832" pageId="0" pageNumber="69">1D</figureCitation>
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) produced a radula that we infer to be teratological, lacking a rachidian and bearing stunted marginal teeth with weakly lobed tips. The two available lots were collected in deeper waters (325-384 m) than
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<taxonomicName id="06095E85E8C56BAE4B0FE149CC999794" lsidName="P. rogerclarki" pageId="0" pageNumber="69" rank="species" species="rogerclarki">
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<emphasis id="9C1062826297A4268E87894BEE3B26E0" italics="true" pageId="0" pageNumber="69">P. rogerclarki</emphasis>
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</taxonomicName>
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(114-166 m). Given the fragmentary and incomplete information available, we cautiously conclude that the broad morph represented by the two specimens in LACM 1997-156.7 is conspecific with LACM 1997-168.10 and represents a third and undescribed species of
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<taxonomicName id="498F7CA617519778E1CFDE4F14A3C25A" class="Gastropoda" family="Newtoniellidae" genus="Pseudomohnia" kingdom="Animalia" lsidName="Pseudomohnia" pageId="0" pageNumber="69" phylum="Mollusca" rank="genus">
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<emphasis id="E45E5FB210E314F58EA86A61F5D057F3" italics="true" pageId="0" pageNumber="69">Pseudomohnia</emphasis>
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</taxonomicName>
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. It is possible that the shells and the soft parts are not conspecific, and that the broad morph represents population variation or sexual dimorphism within the range of
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<taxonomicName id="AAC77063B816AD98ED81A0A293B767A5" lsidName="P. rogerclarki" pageId="0" pageNumber="69" rank="species" species="rogerclarki">
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<emphasis id="41BFB65998879294DF73BE88D8E4F83A" italics="true" pageId="0" pageNumber="69">P. rogerclarki</emphasis>
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</taxonomicName>
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. Thus, we refrain from describing another species until additional comparative material becomes available.
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</paragraph>
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<paragraph id="094E23E5D1F14573B834C50F13E2FB6F" pageId="0" pageNumber="69">
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To the extent that comparisons are possible, the anatomy of
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<taxonomicName id="C68281501D85BC03AD8D2F91BC15F301" class="Gastropoda" family="Newtoniellidae" genus="Pseudomohnia" kingdom="Animalia" lsidName="Pseudomohnia" pageId="0" pageNumber="69" phylum="Mollusca" rank="genus">
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<emphasis id="326FBABACE0A5648D4027D02C004A1E8" italics="true" pageId="0" pageNumber="69">Pseudomohnia</emphasis>
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</taxonomicName>
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sp. agrees well with that of
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<taxonomicName id="B9108969C0B30EAA9C769836BFF1BB65" lsidName="P. kurilana" pageId="0" pageNumber="69" rank="species" species="kurilana">
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<emphasis id="025947E6FF243BEEE640A78926191900" italics="true" pageId="0" pageNumber="69">P. kurilana</emphasis>
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</taxonomicName>
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. The most conspicuous differences between the two concern details of the anterior alimentary system; specifically, the salivary glands appeared more irregular, the snout longer, the radular sac shorter, the mid-esophageal gland less developed, and the length of the introvert shorter in
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<taxonomicName id="686FB8A6C527110F219ADCC39B80A6DA" lsidName="P. kurilana" pageId="0" pageNumber="69" rank="species" species="kurilana">
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<emphasis id="0FA5E65D7BBD508F881ADD96992DDB79" italics="true" pageId="0" pageNumber="69">P. kurilana</emphasis>
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</taxonomicName>
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. The length of the introvert is known to vary within the family (
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<bibRefCitation id="7F6BF7288D4CD7E79ED2AB5E09D776EB" DOI="https://doi.org/10.5962/bhl.part.17411" author="Houbrick, RS" journalOrPublisher="The Nautilus" pageId="0" pageNumber="69" pagination="155 - 165" refId="B15" refString="Houbrick, RS, 1987. Description of a new, giant Ataxocerithium species from Australia, with remarks on the systematic placement of the genus (Prosobranchia: Cerithiopsidae). The Nautilus 101: 155 - 165, DOI: https://doi.org/10.5962/bhl.part.17411" title="Description of a new, giant Ataxocerithium species from Australia, with remarks on the systematic placement of the genus (Prosobranchia: Cerithiopsidae)." url="https://doi.org/10.5962/bhl.part.17411" volume="101" year="1987">Houbrick 1987</bibRefCitation>
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;
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<bibRefCitation id="8CD58417F3A7BEB7A2BE0212B030C381" DOI="https://doi.org/10.1080/00222930903219954" author="Golding, RE" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="69" pagination="2723 - 2763" refId="B12" refString="Golding, RE, Ponder, WF, Byrne, M, 2009. The evolutionary and biomechanical implications of snout and proboscis morphology in Caenogastropoda (Mollusca: Gastropoda). Journal of Natural History 43: 2723 - 2763, DOI: https://doi.org/10.1080/00222930903219954" title="The evolutionary and biomechanical implications of snout and proboscis morphology in Caenogastropoda (Mollusca: Gastropoda)." url="https://doi.org/10.1080/00222930903219954" volume="43" year="2009">Golding et al. 2009</bibRefCitation>
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), but that of
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<taxonomicName id="F33CC5C2DEB3D999C6272BF1B33C7280" lsidName="P. kurilana" pageId="0" pageNumber="69" rank="species" species="kurilana">
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<emphasis id="919C34EDCF212664CAF425BA0A82F3CB" italics="true" pageId="0" pageNumber="69">P. kurilana</emphasis>
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</taxonomicName>
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also may have been incompletely retracted which would also explain the appearance of the snout.
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<taxonomicName id="1E7752F12F013C90C9872B9D8A290135" class="Gastropoda" family="Newtoniellidae" genus="Pseudomohnia" kingdom="Animalia" lsidName="Pseudomohnia" pageId="0" pageNumber="69" phylum="Mollusca" rank="genus">
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<emphasis id="1401919E9DF89FE00076DB05EDD331BB" italics="true" pageId="0" pageNumber="69">Pseudomohnia</emphasis>
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</taxonomicName>
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sp. is more similar to
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<taxonomicName id="EF5DEAE2D5A7C9B9E9B26D28989673FF" lsidName="P. kurilana" pageId="0" pageNumber="69" rank="species" species="kurilana">
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<emphasis id="3C5B362562E76C07F8A6C679E131B0EE" italics="true" pageId="0" pageNumber="69">P. kurilana</emphasis>
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</taxonomicName>
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in morphology of the radula, but differs in the broader, more triangular and finely serrated central cusp of the rachidian and in the morphology of the marginal teeth which bear slightly fewer (two to four versus three to five), shorter, more smoothly conical cusps in
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<taxonomicName id="DFC4D7EFBBF2E8D7E35950CAAF600184" lsidName="P. kurilana" pageId="0" pageNumber="69" rank="species" species="kurilana">
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<emphasis id="BCF6FCD476603D2C6DBB567521C9DDF4" italics="true" pageId="0" pageNumber="69">P. kurilana</emphasis>
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</taxonomicName>
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. However, the range of values of marginal cusp counts overlaps in the two species and its significance could diminish with greater sampling. Like other triphoroideans, the nervous system is epiathroid with a long supra-esophageal connective, but differs in the presence of numerous, tiny statoconia in the statocysts rather than a single statolith (
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<bibRefCitation id="ED8362BFAC6AB411FAA8B623D462C070" author="Risbec, J" journalOrPublisher="Annales des Sciences Naturelles, Zoologie" pageId="0" pageNumber="69" pagination="89 - 107" refId="B37" refString="Risbec, J, 1943. Recherches anatomiques sur les Prosobranches de Nouvelle-Caledonie. (Quatrieme partie). Annales des Sciences Naturelles, Zoologie 11: 89 - 107" title="Recherches anatomiques sur les Prosobranches de Nouvelle-Caledonie. (Quatrieme partie)." volume="11" year="1943">Risbec 1943</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |