treatments-xml/data/4E/56/37/4E56376DFFA6FF9946CB64912D7DFEAA.xml

<document id="5E558EC2483DA8F80F508B9EE2808FCD" ID-DOI="10.11646/zootaxa.3856.2.2" ID-GBIF-Dataset="24c01b66-c0d2-458b-a7fe-0e3de779a8e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="250932" ID-ZooBank="ABC0E7D7-2E22-4605-A97B-0F1E632F3930" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460889728808" checkinUser="plazi" docAuthor="Bota, Cornelio Andrés" docDate="2014" docId="4E56376DFFA6FF9946CB64912D7DFEAA" docLanguage="en" docName="zt03856p210.pdf" docOrigin="Zootaxa 3856 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxyallagma colombianum Bota, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="199" masterDocId="B26F4F15FFA3FF9E465C607B2E0DFFB6" masterDocTitle="A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)" masterLastPageNumber="210" masterPageNumber="192" pageNumber="197" updateTime="1698359463704" updateUser="plazi">
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<mods:title id="EC5645DEC765CB1DF71679973FF13D46">A brief look at the Odonata from the Páramo ecosystems in Colombia, with the descriptions of Oxyallagma colombianum sp. nov. and Rhionaeschna caligo sp. nov. (Odonata: Coenagrionidae, Aeshnidae, Libellulidae)</mods:title>
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<mods:namePart id="648C4234A362F443DC137A04FCC65CDF">Bota, Cornelio Andrés</mods:namePart>
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<mods:date id="09ED9193130F967AFCE66A39B5520B66">2014</mods:date>
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<mods:number id="D01C3C841E6617EFBE1BBB3C6D4B566B">3856</mods:number>
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<subSubSection id="8EE5D5F0FFA6FF9B46CB64912D6DFAFC" pageId="5" pageNumber="197" type="nomenclature">
<paragraph id="C640867BFFA6FF9B46CB64912C36FAB3" blockId="5.[151,1436,1258,1354]" box="[151,571,1258,1285]" pageId="5" pageNumber="197">
<heading id="9D083117FFA6FF9B46CB64912C36FAB3" bold="true" box="[151,571,1258,1285]" fontSize="11" level="1" pageId="5" pageNumber="197" reason="1">
<emphasis id="F48B5A69FFA6FF9B46CB64912C36FAB3" bold="true" box="[151,571,1258,1285]" pageId="5" pageNumber="197">
<taxonomicName id="01FFFDF8FFA6FF9B46CB64912FDAFAB2" ID-CoL="9X3X4" box="[151,471,1258,1284]" class="Insecta" family="Coenagrionidae" genus="Oxyallagma" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="colombianum" status="sp. nov.">
<emphasis id="F48B5A69FFA6FF9B46CB64912FDAFAB2" bold="true" box="[151,471,1258,1284]" italics="true" pageId="5" pageNumber="197">Oxyallagma colombianum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFB8E712FFA6FF9B478264902C36FAB3" box="[478,571,1259,1285]" pageId="5" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C640867BFFA6FF9B46CB65762D6DFAFC" blockId="5.[151,1436,1258,1354]" pageId="5" pageNumber="197">
<figureCitation id="5EC49AFEFFA6FF9B46CB65762EEEFA90" box="[151,227,1293,1318]" captionStart="FIGURE 4" captionStartId="13.[151,250,1954,1976]" captionTargetBox="[202,1384,193,1931]" captionTargetId="figure@13.[202,1385,193,1932]" captionTargetPageId="13" captionText="FIGURE 4. (a) Ischnura ramburii female (Páramo El Morro). (b) Oxyallagma colombianum, teneral male (Antioquia, Labores). Oxyallagma dissidens: (c) Female (Nariño, La Cocha); (d) Male (Cauca, Silvia)." httpUri="https://zenodo.org/record/250936/files/figure.png" pageId="5" pageNumber="197">Figs. 4</figureCitation>
b (teneral habitus ♂), 5a (pair in copula), 5b (pair in tandem, dorsal view), 5c (ovipositing pair), 5d (pair in tandem), 6a 
<emphasis id="F48B5A69FFA6FF9B474765492F29FAFF" box="[283,292,1330,1353]" italics="true" pageId="5" pageNumber="197">-</emphasis>
d (diagnostic traits), 7 (map), Table 1 (coordinates)
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA6FF9B46CB65022A87FA24" box="[151,1162,1401,1426]" pageId="5" pageNumber="197" type="etymology">
<paragraph id="C640867BFFA6FF9B46CB65022A87FA24" blockId="5.[151,1437,1401,2038]" box="[151,1162,1401,1426]" pageId="5" pageNumber="197">
<emphasis id="F48B5A69FFA6FF9B46CB65022F13FA24" bold="true" box="[151,286,1401,1426]" pageId="5" pageNumber="197">Etymology.</emphasis>
In reference to 
<collectingCountry id="BEE8C6EBFFA6FF9B479265022C32FA24" box="[462,575,1401,1426]" name="Colombia" pageId="5" pageNumber="197">Colombia</collectingCountry>
, the beautiful country where this species was found.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA6FF9B469B65E62B87F8AB" pageId="5" pageNumber="197" type="materials_examined">
<paragraph id="C640867BFFA6FF9B469B65E62B87F8AB" blockId="5.[151,1437,1401,2038]" pageId="5" pageNumber="197">
<emphasis id="F48B5A69FFA6FF9B469B65E62FBAFA00" bold="true" box="[199,439,1437,1462]" pageId="5" pageNumber="197">Examined material.</emphasis>
<specimenCount id="D0F94DF2FFA6FF9B479D65E52FF1FA03" box="[449,508,1438,1462]" pageId="5" pageNumber="197" type="generic">21 ♂</specimenCount>
&amp; 8 F 
<collectingCountry id="BEE8C6EBFFA6FF9B440465E62CC1FA00" box="[600,716,1437,1462]" name="Colombia" pageId="5" pageNumber="197">
<emphasis id="F48B5A69FFA6FF9B440465E62CC1FA00" bold="true" box="[600,716,1437,1462]" pageId="5" pageNumber="197">Colombia</emphasis>
</collectingCountry>
: Antioquia: 
<taxonomicName id="01FFFDF8FFA6FF9B453D65E52DBBFA00" box="[865,950,1438,1462]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
Las Baldías: 
<specimenCount id="D0F94DF2FFA6FF9B420565E52A84FA03" box="[1113,1161,1438,1462]" pageId="5" pageNumber="197" type="generic">1 ♂</specimenCount>
, 
<date id="B241A0BBFFA6FF9B42C465E62B17FA00" box="[1176,1306,1437,1462]" pageId="5" pageNumber="197" value="2013-07-16">16.vii.2013</date>
, Leg: J. R. Albertino &amp; A. L. Montoya. 
<taxonomicName id="01FFFDF8FFA6FF9B478765B92C3CFA6C" box="[475,561,1474,1498]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
El Morro: 
<specimenCount id="D0F94DF2FFA6FF9B44F265B92CDFFA6C" box="[686,722,1474,1498]" pageId="5" pageNumber="197" type="generic">1♂</specimenCount>
, 
<date id="B241A0BBFFA6FF9B448365B92D4DFA6C" box="[735,832,1474,1498]" pageId="5" pageNumber="197" value="2011-05-09">9.v.2011</date>
, Leg: C. Bota &amp; J. D. Castaño. 
<taxonomicName id="01FFFDF8FFA6FF9B42F865B92AF4FA6C" box="[1188,1273,1474,1498]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="paramo">Páramo</taxonomicName>
Sabanas: 
<specimenCount id="D0F94DF2FFA6FF9B433765BA2B9AFA6F" box="[1387,1431,1473,1498]" pageId="5" pageNumber="197" type="generic">5 ♂</specimenCount>
, 
<date id="B241A0BBFFA6FF9B46CB659D2F1BFA48" box="[151,278,1510,1534]" pageId="5" pageNumber="197" value="2011-10-01" valueMax="2011-10-03" valueMin="2011-10-01">
1 
<emphasis id="F48B5A69FFA6FF9B46F9659D2EBEFA4B" box="[165,179,1510,1533]" italics="true" pageId="5" pageNumber="197">–</emphasis>
3.x.2011
</date>
, Leg: C. Bota &amp; J.D. Castaño. 
<specimenCount id="D0F94DF2FFA6FF9B442E659D2C93FA4B" box="[626,670,1510,1534]" pageId="5" pageNumber="197" type="generic">1 ♂</specimenCount>
, path from Belmira town toward El Morro, 
<geoCoordinate id="A3CBE0BCFFA6FF9B42D4659E2ADAFA48" box="[1160,1239,1509,1534]" direction="north" orientation="latitude" pageId="5" pageNumber="197" precision="925" value="6.616667">6°37'N</geoCoordinate>
<geoCoordinate id="A3CBE0BCFFA6FF9B4281659E2B4DFA48" box="[1245,1344,1509,1534]" direction="west" orientation="longitude" pageId="5" pageNumber="197" precision="925" value="-75.666664">75°40'W</geoCoordinate>
<quantity id="01072B9EFFA6FF9B431B659E2B91FA4B" box="[1351,1436,1509,1534]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="5" pageNumber="197" unit="m" value="2950.0">2950 m</quantity>
approx., 
<date id="B241A0BBFFA6FF9B46A566722F76F994" box="[249,379,1545,1570]" pageId="5" pageNumber="197" value="2012-07-16">16.vii.2012</date>
. 
<specimenCount id="D0F94DF2FFA6FF9B47DA66722FA2F997" box="[390,431,1545,1570]" pageId="5" pageNumber="197" type="generic">5 ♂</specimenCount>
&amp; 3♀, Belmira, Corregimiento Labores, Finca El Paraíso, 
<geoCoordinate id="A3CBE0BCFFA6FF9B426366712A98F994" box="[1087,1173,1545,1570]" direction="north" orientation="latitude" pageId="5" pageNumber="197" precision="925" value="6.6666665">N 6°40'</geoCoordinate>
<geoCoordinate id="A3CBE0BCFFA6FF9B42C766712B0AF994" box="[1179,1287,1545,1570]" direction="west" orientation="longitude" pageId="5" pageNumber="197" precision="925" value="-75.63333">W 75°38'</geoCoordinate>
, 
<date id="B241A0BBFFA6FF9B434D66722B9AF994" box="[1297,1431,1545,1570]" pageId="5" pageNumber="197" value="2012-04-05" valueMax="2012-04-07" valueMin="2012-04-05">
5 
<emphasis id="F48B5A69FFA6FF9B434366712B20F997" box="[1311,1325,1546,1569]" italics="true" pageId="5" pageNumber="197">–</emphasis>
7.iv.2012
</date>
, Leg: C. Moreno, S. Bota, T. Bota &amp; C. Bota. 
<specimenCount id="D0F94DF2FFA6FF9B44EC66552CD2F9F3" box="[688,735,1582,1606]" pageId="5" pageNumber="197" type="generic">1 ♂</specimenCount>
, Municipality of La Ceja: Township Las Lomitas, 
<geoCoordinate id="A3CBE0BCFFA6FF9B437166552B74F9F0" box="[1325,1401,1581,1606]" direction="north" orientation="latitude" pageId="5" pageNumber="197" precision="925" value="6.0">N 6°0'</geoCoordinate>
<geoCoordinate id="A3CBE0BCFFA6FF9B43DE66552EEDF9DC" direction="west" orientation="longitude" pageId="5" pageNumber="197" precision="925" value="-75.38333">W 75°23'</geoCoordinate>
<quantity id="01072B9EFFA6FF9B46B666292F35F9DC" box="[234,312,1618,1642]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="197" unit="m" value="2300.0">2300m</quantity>
approx., 
<date id="B241A0BBFFA6FF9B47F5662A2C0CF9DC" box="[425,513,1617,1642]" pageId="5" pageNumber="197" value="2009-11">xi.2009</date>
, Leg: J. Campuzano. Municipality of Medellín, Corregimiento San Antonio de Prado: 1 ♀, Township Astilleros: Truchera Los Muñoz: 
<geoCoordinate id="A3CBE0BCFFA6FF9B4577660D2DBAF938" box="[811,951,1653,1678]" direction="north" orientation="latitude" pageId="5" pageNumber="197" precision="1" value="6.2476335">N 6°14,858'</geoCoordinate>
<geoCoordinate id="A3CBE0BCFFA6FF9B459C660D2A6DF938" box="[960,1120,1653,1678]" direction="west" orientation="longitude" pageId="5" pageNumber="197" precision="1" value="-75.665">W 75°39,900'</geoCoordinate>
<quantity id="01072B9EFFA6FF9B4236660E2AB6F938" box="[1130,1211,1653,1678]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="197" unit="m" value="2600.0">2600m</quantity>
, 
<date id="B241A0BBFFA6FF9B4296660E2B3BF938" box="[1226,1334,1653,1678]" pageId="5" pageNumber="197" value="2010-06-03">3.vi.2010</date>
, Leg: G. Valencia &amp; J. Cardona 
<emphasis id="F48B5A69FFA6FF9B47C066E12FB8F907" box="[412,437,1690,1713]" italics="true" pageId="5" pageNumber="197">—</emphasis>
D. Township Potrerito: 
<geoCoordinate id="A3CBE0BCFFA6FF9B449A66E12D57F904" box="[710,858,1689,1714]" direction="north" orientation="latitude" pageId="5" pageNumber="197" precision="1" value="6.174">N 6°10'26,4&quot;</geoCoordinate>
<geoCoordinate id="A3CBE0BCFFA6FF9B453966E12A07F904" box="[869,1034,1689,1714]" direction="west" orientation="longitude" pageId="5" pageNumber="197" precision="1" value="-75.6828">W 75°40'58,1&quot;</geoCoordinate>
<quantity id="01072B9EFFA6FF9B425566E22A65F907" box="[1033,1128,1689,1714]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="197" unit="m" value="2700.0">2700 m</quantity>
, 1 ♀, 
<date id="B241A0BBFFA6FF9B42E966E22B3DF904" box="[1205,1328,1689,1714]" pageId="5" pageNumber="197" value="2001-08-09">9.viii.2001</date>
, Leg: M. Castaño. 
<specimenCount id="D0F94DF2FFA6FF9B46A366C52F26F963" box="[255,299,1726,1750]" pageId="5" pageNumber="197" type="generic">1 ♂</specimenCount>
, 
<date id="B241A0BBFFA6FF9B476B66C62FAEF960" box="[311,419,1725,1750]" pageId="5" pageNumber="197" value="2001-09-09">9.ix.2001</date>
, Leg: M. C. Vélez. Leg: A. Clavijo &amp; Y. Correa, 
<specimenCount id="D0F94DF2FFA6FF9B459766C52DFAF963" box="[971,1015,1726,1750]" pageId="5" pageNumber="197" type="generic">2 ♂</specimenCount>
, 
<date id="B241A0BBFFA6FF9B425F66C62A7DF960" box="[1027,1136,1725,1750]" pageId="5" pageNumber="197" value="2011-03-05">5.iii.2011</date>
. 
<specimenCount id="D0F94DF2FFA6FF9B422766C52AA8F963" box="[1147,1189,1726,1750]" pageId="5" pageNumber="197" type="generic">1 ♂</specimenCount>
&amp; 2 ♀, 
<date id="B241A0BBFFA6FF9B435D66C62B8EF960" box="[1281,1411,1725,1750]" pageId="5" pageNumber="197" value="2010-07-12">12.vii.2010</date>
. 1 ♀, 
<date id="B241A0BBFFA6FF9B469D669A2F37F94C" box="[193,314,1761,1786]" pageId="5" pageNumber="197" value="2011-11-14">14.xi.2011</date>
, Leg: J. Ortiz. 
<specimenCount id="D0F94DF2FFA6FF9B47A066992C22F94F" box="[508,559,1762,1786]" pageId="5" pageNumber="197" type="generic">2 ♂</specimenCount>
, 
<date id="B241A0BBFFA6FF9B441E669A2CA3F94C" box="[578,686,1761,1786]" pageId="5" pageNumber="197" value="2013-01-18">18.i.2013</date>
, Leg: N. Uribe. 
<specimenCount id="D0F94DF2FFA6FF9B45DC66992DBEF94F" box="[896,947,1762,1786]" pageId="5" pageNumber="197" type="generic">1 ♂</specimenCount>
, Santander: Municipality of Piedecuesta: Corregimiento Sevilla: Finca El Razgón: 
<geoCoordinate id="A3CBE0BCFFA6FF9B443B677D2CDBF8A8" box="[615,726,1798,1822]" direction="north" orientation="latitude" pageId="5" pageNumber="197" precision="15" value="7.0416665">N 7°2'30&quot;</geoCoordinate>
<geoCoordinate id="A3CBE0BCFFA6FF9B4481677D2D63F8A8" box="[733,878,1797,1822]" direction="west" orientation="longitude" pageId="5" pageNumber="197" precision="15" value="-72.989716">W 72°59'23&quot;</geoCoordinate>
<quantity id="01072B9EFFA6FF9B4529677E2DCBF8A8" box="[885,966,1797,1822]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="197" unit="m" value="2250.0">2250m</quantity>
, 
<date id="B241A0BBFFA6FF9B458D677E2A5EF8A8" box="[977,1107,1797,1822]" pageId="5" pageNumber="197" value="2001-07-28">28.vii.2001</date>
, Leg: A. Veléz &amp; P. Duque.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA6FF98469B67522A63F9BE" lastPageId="6" lastPageNumber="198" pageId="5" pageNumber="197" type="description">
<paragraph id="C640867BFFA6FF98469B67522AB9FCA2" blockId="5.[151,1437,1401,2038]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="198" pageId="5" pageNumber="197">
<emphasis id="F48B5A69FFA6FF9B469B67522FDAF8F4" bold="true" box="[199,471,1833,1858]" pageId="5" pageNumber="197">
Description. 
<typeStatus id="194438D9FFA6FF9B473F67522FDFF8F4" box="[355,466,1833,1858]" pageId="5" pageNumber="197" type="holotype">Holotype</typeStatus>
.
</emphasis>
Body covered with golden hairs. 
<emphasis id="F48B5A69FFA6FF9B453767522DBDF8F4" bold="true" box="[875,944,1833,1858]" italics="true" pageId="5" pageNumber="197">Head.</emphasis>
Labium, labrum, base of mandible, gena, anteclypeus, and rear of head ivory white. Labrum with small black medio 
<emphasis id="F48B5A69FFA6FF9B45A067352A08F8D3" box="[1020,1029,1870,1893]" italics="true" pageId="5" pageNumber="197">-</emphasis>
basal spot. Anteclypeus with round brown spot on each side. Postclypeus, antenna, and frons reddish brown, postocular spots absent, with some green iridescence. Rear of head dorsally ivory white and ventrally pale blue. Occipital bar absent. Frons rounded. Postocular lobes protruding backwards beyond level of hind margin of compound eyes (
<figureCitation id="5EC49AFEFFA6FF9B42F167C22AF2F864" box="[1197,1279,1977,2002]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="5" pageNumber="197">Figs. 5</figureCitation>
a 
<emphasis id="F48B5A69FFA6FF9B435067C12B18F867" box="[1292,1301,1978,2001]" italics="true" pageId="5" pageNumber="197">-</emphasis>
d). 
<emphasis id="F48B5A69FFA6FF9B436367C22B96F864" bold="true" box="[1343,1435,1977,2002]" italics="true" pageId="5" pageNumber="197">Thorax.</emphasis>
Prothorax reddish brown with some iridescent green spots on pleural and dorsal sides, medial lobe of posterior prothoracic lobe developed into caudally projected, rounded plate with smoothly rounded margins and posterior edge slightly concave. Pterothorax reddish brown with mesepisternal and metepimeral iridescent green stripes, venter, and metepimeron mostly ivory white. Legs and coxae ivory white. Tarsi, spurs, and claws black (
<figureCitation id="5EC49AFEFFA5FF98436460A42B88FF4E" box="[1336,1413,223,248]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="6" pageNumber="198">Figs. 5</figureCitation>
a 
<emphasis id="F48B5A69FFA5FF9843CF609A2B91FF4E" box="[1427,1436,225,248]" italics="true" pageId="6" pageNumber="198">-</emphasis>
d). Seven external spurs on right metafemur and eight on left metafemur, as long as space between them or shorter, gradually increasing in size toward apex. Seven external spurs on right metatibia and eight on left metatibia, as long as space between them or shorter, gradually decreasing in size toward apex. Tarsal claws with well developed supplementary tooth. Wings hyaline. Pt red, ratio between distal and proximal length about 1:1.5. CuP reaching CuPAA slightly distal to confluence of CuPAA with hind margin of wing. Px 
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FW, 
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HW. 
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2 branching between Px 5 and 
<quantity id="01072B9EFFA5FF98473F61CC2F80FE66" box="[355,397,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="198" unit="in" value="6.0">6 in</quantity>
FW, and between Px 4 and 
<quantity id="01072B9EFFA5FF98449F61CC2CE3FE66" box="[707,750,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="198" unit="in" value="5.0">5 in</quantity>
right HW and under Px 
<quantity id="01072B9EFFA5FF9845A361C32A27FE66" box="[1023,1066,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="198" unit="in" value="4.0">4 in</quantity>
left HW. 
<emphasis id="F48B5A69FFA5FF9842C961CC2B06FE66" bold="true" box="[1173,1291,439,464]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF9842C961CC2B0EFE66" bold="true" box="[1173,1283,439,464]" italics="true" pageId="6" pageNumber="198">Abdomen</emphasis>
.
</emphasis>
Vent er i vory white. S1 
<emphasis id="F48B5A69FFA5FF98475E61A62F1DFE42" box="[258,272,477,500]" italics="true" pageId="6" pageNumber="198">–</emphasis>
6 dorsum and lateral terga red except: distal margin of S2 
<emphasis id="F48B5A69FFA5FF9845C361A62DA0FE42" box="[927,941,477,500]" italics="true" pageId="6" pageNumber="198">–</emphasis>
3, distal fifth of S4 
<emphasis id="F48B5A69FFA5FF9842D961A62A9EFE42" box="[1157,1171,477,500]" italics="true" pageId="6" pageNumber="198">–</emphasis>
5, distal ¾ of S6 black. Dorsum and lateral terga of S7 
<emphasis id="F48B5A69FFA5FF9847A5627A2C0AFDAE" box="[505,519,513,536]" italics="true" pageId="6" pageNumber="198">–</emphasis>
10 blue except dorso-distal margin of S10 black. S7 
<emphasis id="F48B5A69FFA5FF984200627A2A67FDAE" box="[1116,1130,513,536]" italics="true" pageId="6" pageNumber="198">–</emphasis>
9 with distal dorsal row of denticles (
<figureCitation id="5EC49AFEFFA5FF984750625F2F56FD8B" box="[268,347,548,573]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="6" pageNumber="198">Figs. 5</figureCitation>
a 
<emphasis id="F48B5A69FFA5FF984734625E2F7BFD8A" box="[360,374,549,572]" italics="true" pageId="6" pageNumber="198">–</emphasis>
d). Genital ligula (as in figure 6b) with apex slightly concave and pair of latero-apical lobes ending in sharp process directed proximad, small latero-medial pointed lobes and an inner medial hooklike process. Cercus (
<figureCitation id="5EC49AFEFFA5FF9846A562172F41FD33" box="[249,332,620,645]" captionStart="FIGURE 6" captionStartId="15.[151,250,1831,1853]" captionTargetBox="[233,1352,206,1809]" captionTargetId="figure@15.[226,1360,193,1810]" captionTargetPageId="15" captionText="FIGURE 6. Oxyallagma colombianum (Páramo Sabanas): Female paratype: (a) Dorsal view of pronotum posterior lobe and mesostigmal plates. Male paratype (Páramo Las Baldías): (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio - dorsal view of caudal appendages. l-m. l.: Latero-medial lobes. m. p.: Mesostigmal plates. p. p. l.: Pronotum posterior lobe. e. v – ap. p.: Ventroapical external process. v – b t.: Ventrobasal tooth. i. d. p.: Inner dorsal process. p. d. b.: Paraproct dorsal branch. p. v. b.: Paraproct ventral branch. Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250938/files/figure.png" pageId="6" pageNumber="198">Figs. 6</figureCitation>
c 
<emphasis id="F48B5A69FFA5FF98470462162F6BFD32" box="[344,358,621,644]" italics="true" pageId="6" pageNumber="198">–</emphasis>
d) external side dark brown, internal light brown, approximately half as long as S10, with ventrobasal tooth, one external ventroapical, rounded process which is most distal point of the body, and rounded inner dorsal process. Paraproct bifid with ventral branch ivory white becoming dark brown through dorsal branch, both branches with rounded apices, distal point of ventral process reaching about 3/4 cercus length or slightly beyond. Total length 
<quantity id="01072B9EFFA5FF9847D962872FDBFCA2" box="[389,470,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="198" unit="mm" value="30.0">30 mm</quantity>
. Abdomen length 
<quantity id="01072B9EFFA5FF9844FF62872CFBFCA2" box="[675,758,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="198" unit="mm" value="23.0">23 mm</quantity>
. FW length 
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. HW length 
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.
</paragraph>
<paragraph id="C640867BFFA5FF98469B63642CD5FC71" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF98469B63642C22FC8E" bold="true" box="[199,559,799,824]" pageId="6" pageNumber="198">
Variations in male 
<typeStatus id="194438D9FFA5FF9847EE63642C27FC8E" box="[434,554,799,824]" pageId="6" pageNumber="198" type="paratype">paratypes</typeStatus>
.
</emphasis>
A black spot may be present anterior to median ocellus. Green iridescence varies from completely absent to covering most of frons. Pterothorax can be completely reddish brown. Seven to ten external metafemoral spurs. Six to eleven external metatibial spurs. 11 
<emphasis id="F48B5A69FFA5FF98425563122A1AFC36" box="[1033,1047,873,896]" italics="true" pageId="6" pageNumber="198">–</emphasis>
14 Px in FW, 
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8 
<emphasis id="F48B5A69FFA5FF98428863122AEFFC36" box="[1236,1250,873,896]" italics="true" pageId="6" pageNumber="198">–</emphasis>
10 in
</quantity>
HW. 
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2 branching between Px 5 and 
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FW, between Px 4 and 
<quantity id="01072B9EFFA5FF98455663F72D39FC13" box="[778,820,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="198" unit="in" value="5.0">5 in</quantity>
HW. Total length 
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28 
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32 mm
</quantity>
. Abdominal length 
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22 
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25 mm
</quantity>
. FW length 
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17 
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19 mm
</quantity>
. HW length 
<quantity id="01072B9EFFA5FF98440B63D42CDEFC71" box="[599,723,943,968]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="6" pageNumber="198" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">
16 
<emphasis id="F48B5A69FFA5FF98442863CA2C8FFC7E" box="[628,642,945,968]" italics="true" pageId="6" pageNumber="198">–</emphasis>
18 mm
</quantity>
.
</paragraph>
<paragraph id="C640867BFFA5FF98469B63AF2DAEFA2A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF98469B63AF2F3EFC5B" bold="true" box="[199,307,980,1005]" pageId="6" pageNumber="198">Allotype.</emphasis>
Color pattern as in 
<typeStatus id="194438D9FFA5FF98444963AF2C7AFC5B" box="[533,631,980,1005]" pageId="6" pageNumber="198" type="holotype">holotype</typeStatus>
but: 
<emphasis id="F48B5A69FFA5FF9844EF63AF2CF5FC5B" bold="true" box="[691,760,980,1005]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF9844EF63AF2CFCFC5B" bold="true" box="[691,753,980,1005]" italics="true" pageId="6" pageNumber="198">Head</emphasis>
.
</emphasis>
Olive green except rear and gena downwards from level of antenna base ivory white (
<figureCitation id="5EC49AFEFFA5FF98479C638C2C03FBA6" box="[448,526,1015,1040]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="6" pageNumber="198">Figs. 5</figureCitation>
a, c 
<emphasis id="F48B5A69FFA5FF98446163822C4BFBA6" box="[573,582,1017,1040]" italics="true" pageId="6" pageNumber="198">-</emphasis>
d). 
<emphasis id="F48B5A69FFA5FF984430638C2CCCFBA6" bold="true" box="[620,705,1015,1040]" italics="true" pageId="6" pageNumber="198">Thorax</emphasis>
. Brownish red areas in males olive green (
<figureCitation id="5EC49AFEFFA5FF9842FA638C2AF9FBA6" box="[1190,1268,1015,1040]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="6" pageNumber="198">Figs. 5</figureCitation>
a 
<emphasis id="F48B5A69FFA5FF98435B63822B1DFBA6" box="[1287,1296,1017,1040]" italics="true" pageId="6" pageNumber="198">-</emphasis>
d). Posterior prothoracic lobe directed dorsally, in dorsal view sinuous. Mesostigmal plates flat with latero-apical prominent process directed proximad (
<figureCitation id="5EC49AFEFFA5FF98478B64442C13FBEE" box="[471,542,1087,1112]" captionStart="FIGURE 6" captionStartId="15.[151,250,1831,1853]" captionTargetBox="[233,1352,206,1809]" captionTargetId="figure@15.[226,1360,193,1810]" captionTargetPageId="15" captionText="FIGURE 6. Oxyallagma colombianum (Páramo Sabanas): Female paratype: (a) Dorsal view of pronotum posterior lobe and mesostigmal plates. Male paratype (Páramo Las Baldías): (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio - dorsal view of caudal appendages. l-m. l.: Latero-medial lobes. m. p.: Mesostigmal plates. p. p. l.: Pronotum posterior lobe. e. v – ap. p.: Ventroapical external process. v – b t.: Ventrobasal tooth. i. d. p.: Inner dorsal process. p. d. b.: Paraproct dorsal branch. p. v. b.: Paraproct ventral branch. Illustrations by Natalia Uribe Macias." httpUri="https://zenodo.org/record/250938/files/figure.png" pageId="6" pageNumber="198">Fig. 6</figureCitation>
a). Eight external spurs on right metafemur and six on left metafemur; seven external spurs on right metatibia and eight on left metatibia. Px 
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FW, 
<quantity id="01072B9EFFA5FF984587641F2A19FBCB" box="[987,1044,1124,1149]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="198" unit="in" value="10.0">10 in</quantity>
HW. 
<collectingCountry id="BEE8C6EBFFA5FF98420A641F2A74FBCA" box="[1110,1145,1124,1148]" name="Poland" pageId="6" pageNumber="198">RP</collectingCountry>
2 branching between Px 5 and 
<quantity id="01072B9EFFA5FF98469464FC2EF8FB16" box="[200,245,1159,1184]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="198" unit="in" value="6.0">6 in</quantity>
FW, between Px 4 and 
<quantity id="01072B9EFFA5FF98445664FC2C3AFB16" box="[522,567,1159,1184]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="198" unit="in" value="5.0">5 in</quantity>
HW. 
<emphasis id="F48B5A69FFA5FF98442764FC2CFCFB16" bold="true" box="[635,753,1159,1184]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF98442764FC2CE4FB16" bold="true" box="[635,745,1159,1184]" italics="true" pageId="6" pageNumber="198">Abdomen</emphasis>
.
</emphasis>
S6 with middorsal black line starting at ¾ and widening in distal fourth, surrounded by greenish ivory white; S7–10 dorsum black and lateral terga greenish ivory white except for pair of bluish gray lateral spots on dorsum of S10 (
<figureCitation id="5EC49AFEFFA5FF98452664B42DC7FB5E" box="[890,970,1231,1256]" captionStart="FIGURE 5" captionStartId="14.[151,250,1953,1975]" captionTargetBox="[202,1383,193,1930]" captionTargetId="figure@14.[202,1384,193,1932]" captionTargetPageId="14" captionText="FIGURE 5. Oxyallagma colombianum: (Páramo Sabanas): (a) Pair in wheel position; (b) Tandem pair; (Antioquia, Labores): (c) Ovipositing pair with microparasitoid close to the oviposition place; (d) Tandem pair (photograph by Cintia Moreno)." httpUri="https://zenodo.org/record/250937/files/figure.png" pageId="6" pageNumber="198">Figs. 5</figureCitation>
a 
<emphasis id="F48B5A69FFA5FF98458464AA2DECFB5E" box="[984,993,1233,1256]" italics="true" pageId="6" pageNumber="198">-</emphasis>
d). Brown sub 
<emphasis id="F48B5A69FFA5FF9842D564AA2A9FFB5E" box="[1161,1170,1233,1256]" italics="true" pageId="6" pageNumber="198">-</emphasis>
conical cercus, slightly shorter than S10, apex of cercus most distal point of body. Paraproct ivory white with pruinescence. Welldeveloped vulvar spine, ivory white with brown tip. Sub 
<emphasis id="F48B5A69FFA5FF98457565622D3FFA86" box="[809,818,1305,1328]" italics="true" pageId="6" pageNumber="198">-</emphasis>
basal plate of ovipositor triangular. Teeth along outer valve of ovipositor arranged in row that begins at half its length and runs distally to tip. Ovipositor not including stylus reaching distal margin of S10. Ovipositor ivory white with edge of valve and stylus brown. Total length 
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. Abdomen length 
<quantity id="01072B9EFFA5FF9847CF65FF2FE8FA2A" box="[403,485,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="198" unit="mm" value="23.0">23 mm</quantity>
. FW length 
<quantity id="01072B9EFFA5FF98443265FF2CCDFA2A" box="[622,704,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="198" unit="mm" value="19.0">19 mm</quantity>
. HW length 
<quantity id="01072B9EFFA5FF98451165FF2D92FA2A" box="[845,927,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="198" unit="mm" value="17.0">17 mm</quantity>
.
</paragraph>
<paragraph id="C640867BFFA5FF98469B65DC2A63F9BE" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF98469B65DC2C30FA76" bold="true" box="[199,573,1447,1472]" pageId="6" pageNumber="198">
Variation in female 
<typeStatus id="194438D9FFA5FF98479C65DC2C35FA76" box="[448,568,1447,1472]" pageId="6" pageNumber="198" type="paratype">paratypes</typeStatus>
.
</emphasis>
Six to seven metafemoral external spurs. Six to eight metatibial external spurs. 11–12 Px in FW, 
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HW. 
<collectingCountry id="BEE8C6EBFFA5FF98441A65B72C64FA52" box="[582,617,1484,1508]" name="Poland" pageId="6" pageNumber="198">RP</collectingCountry>
2 branching between Px 4 and 
<quantity id="01072B9EFFA5FF98459E65B72DE0FA53" box="[962,1005,1484,1509]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="198" unit="in" value="6.0">6 in</quantity>
FW, between Px 4 and 
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HW. Total length 
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. Abdomen length 
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22 
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23 mm
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. FW 
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18 
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19 mm
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. HW length 
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16 
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17 mm
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.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA5FF98469B666F2C46F9C2" pageId="6" pageNumber="198" type="diagnosis">
<paragraph id="C640867BFFA5FF98469B666F2C46F9C2" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF98469B666F2F4CF99B" bold="true" box="[199,321,1556,1581]" pageId="6" pageNumber="198">Diagnosis.</emphasis>
The new species fits the generic diagnosis given by 
<bibRefCitation id="A26EFB8AFFA5FF9845C6666F2A73F99A" author="Garrison" box="[922,1150,1556,1581]" pageId="6" pageNumber="198" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World: An Illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, 528 pp." type="book" year="2010">
Garrison 
<emphasis id="F48B5A69FFA5FF984258666E2A33F99A" box="[1028,1086,1556,1580]" italics="true" pageId="6" pageNumber="198">et al.</emphasis>
2010
</bibRefCitation>
except for: genital ligula with small latero-medial pointed lobes; male cercus with only one ventro-apical process; and ventral branch of paraproct narrower than half of cercus.
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA5FF99469B66042DC1FF63" lastPageId="7" lastPageNumber="199" pageId="6" pageNumber="198" type="biology_ecology">
<paragraph id="C640867BFFA5FF99469B66042DC1FF63" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="199" pageId="6" pageNumber="198">
<emphasis id="F48B5A69FFA5FF98469B66042F2AF92E" bold="true" box="[199,295,1663,1688]" pageId="6" pageNumber="198">Biology.</emphasis>
This highland species is not exclusive to the 
<taxonomicName id="01FFFDF8FFA5FF98456866FB2D81F92E" box="[820,908,1664,1688]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="198" phylum="Arthropoda" rank="species" species="paramo">páramo</taxonomicName>
. Populations are numerous and conspicuous. I observed males perching on the emerging macrophytes in groups without apparent competition between them. In the evening, some males were seen in wooded areas close to the lakes; all populations occurred close to forest habitats, which can be a clue of how important this shelter is for this species. Pairs in tandem or in copula were seen ovipositing during midday. Two pairs were observed initiating the wheel position (
<figureCitation id="5EC49AFEFFA5FF98420D67742A9AF89E" box="[1105,1175,1807,1832]" captionStart="FIGURE 4" captionStartId="13.[151,250,1954,1976]" captionTargetBox="[202,1384,193,1931]" captionTargetId="figure@13.[202,1385,193,1932]" captionTargetPageId="13" captionText="FIGURE 4. (a) Ischnura ramburii female (Páramo El Morro). (b) Oxyallagma colombianum, teneral male (Antioquia, Labores). Oxyallagma dissidens: (c) Female (Nariño, La Cocha); (d) Male (Cauca, Silvia)." httpUri="https://zenodo.org/record/250936/files/figure.png" pageId="6" pageNumber="198">Fig. 4</figureCitation>
a) until end, the whole process taking 11 minutes 5 seconds (+/ 
<emphasis id="F48B5A69FFA5FF984400674E2C67F8FA" box="[604,618,1845,1868]" italics="true" pageId="6" pageNumber="198">–</emphasis>
1 minute) on average, which according to Córdoba 
<emphasis id="F48B5A69FFA5FF9842EE674E2AB6F8FA" box="[1202,1211,1845,1868]" italics="true" pageId="6" pageNumber="198">-</emphasis>
Aguilar &amp; Cordero 
<emphasis id="F48B5A69FFA5FF9843CF674E2B91F8FA" box="[1427,1436,1845,1868]" italics="true" pageId="6" pageNumber="198">-</emphasis>
Rivera (2008) could indicate female cryptic choice, or sperm displacement, or both. Numerous ovipositing pairs were observed, females laying their eggs on macrophytes by submerging their abdomen. On one occasion a microparasitoid (Hymenoptera) was seen close to where a female was ovipositing (
<figureCitation id="5EC49AFEFFA5FF98421F67DB2A85F80E" box="[1091,1160,1952,1977]" captionStart="FIGURE 4" captionStartId="13.[151,250,1954,1976]" captionTargetBox="[202,1384,193,1931]" captionTargetId="figure@13.[202,1385,193,1932]" captionTargetPageId="13" captionText="FIGURE 4. (a) Ischnura ramburii female (Páramo El Morro). (b) Oxyallagma colombianum, teneral male (Antioquia, Labores). Oxyallagma dissidens: (c) Female (Nariño, La Cocha); (d) Male (Cauca, Silvia)." httpUri="https://zenodo.org/record/250936/files/figure.png" pageId="6" pageNumber="198">Fig. 4</figureCitation>
c), but it did not seem to disturb female activity. One male teneral was observed perched on a macrophyte after hatching, but with its wings completely bent [according to 
<bibRefCitation id="A26EFB8AFFA4FF9947A860EC2C9EFF06" author="Corbet" box="[500,659,151,176]" pageId="7" pageNumber="199" refString="Corbet, P. S. (1999) Dragonflies: Behavior and ecology of Odonata. Comstock Publishing Associates, Cornell University Press. Ithaca, New York, 829 pp." type="book" year="1999">Corbet (1999)</bibRefCitation>
Stage 3]; it took 35 minutes to extend them completely and its first flight on that hot day was around noon. No exuviae were found (
<figureCitation id="5EC49AFEFFA4FF99453760C72DA3FF63" box="[875,942,188,213]" captionStart="FIGURE 3" captionStartId="12.[151,250,1621,1643]" captionTargetBox="[151,1436,193,1598]" captionTargetId="figure@12.[151,1436,193,1599]" captionTargetPageId="12" captionText="FIGURE 3. Sympetrum paramo: (a) Teneral female and exuvia (Páramo El Congo); (b) Male (Páramo El Morro). Erythrodiplax connata- group (Cauca, Timbio): (c) Mature male; (d) Immature male (same coloration as females)." httpUri="https://zenodo.org/record/250935/files/figure.png" pageId="7" pageNumber="199">Fig. 3</figureCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="8EE5D5F0FFA4FF99469B60A42D7DFEAA" pageId="7" pageNumber="199" type="distribution">
<paragraph id="C640867BFFA4FF99469B60A42D7DFEAA" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="199">
<emphasis id="F48B5A69FFA4FF99469B60A42F6DFF4E" bold="true" box="[199,352,223,248]" pageId="7" pageNumber="199">Distribution.</emphasis>
Known from the northern portion of Central and Oriental Mountain Ranges in Antioquia and Santander Departments (
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), between 2,400 and 
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.
</paragraph>
</subSubSection>
</treatment>
</document>