182 lines
22 KiB
XML
182 lines
22 KiB
XML
<document id="7FA5A8ECAD988383BBCE3B08DC975A71" ID-CLB-Dataset="35149" ID-DOI="http://dx.doi.org/10.3897/zookeys.536.5982" ID-GBIF-Dataset="f3546db1-d96a-40d8-ac8c-fe806547f201" ID-PMC="PMC4714036" ID-Pensoft-Pub="1313-2970-536-1" ID-PubMed="26798233" ID-ZooBank="A8B92CDD0B8C4384AAC559648BB45AA5" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-536-1" ModsDocOrigin="ZooKeys 536" ModsDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" checkinTime="1451243741663" checkinUser="pensoft" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2015" docId="1882882DAB1B715EFEEE5ED38E5CDAC6" docLanguage="en" docName="ZooKeys 536: 1-92" docOrigin="ZooKeys 536" docSource="http://dx.doi.org/10.3897/zookeys.536.5982" docTitle="Pterostichus (Circinatus) ailaoicus Shi & Liang, 2015, sp. n." docType="treatment" docUuid="D7ECDAEE-BB43-4C4A-B668-875E6B2540F2" docUuidSource="ZooBank" docVersion="8" lastPageNumber="11" masterDocId="FFEAFF976442467ED413FFC17531FF81" masterDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="9" updateTime="1731888672137" updateUser="admin">
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<mods:titleInfo id="7E5647F8BBF946F3B65486E6EB4E907A">
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<mods:title id="325C0F0EA45D9EC1F8A7DD82C4593B97">The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)</mods:title>
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<mods:namePart id="13B269957140D3F31A73B4F0BB7BBB09">Shi, Hongliang</mods:namePart>
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<mods:roleTerm id="BD15D9F0752967741655E70FA06402CC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D211D41D55F1D0A75ACC50FFAFA69F93">Liang, Hongbin</mods:namePart>
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<mods:typeOfResource id="731694AE78FDFFF08695192649B0A929">text</mods:typeOfResource>
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<mods:title id="A8D5AE082E093C38BFDCDB0AABED5355">ZooKeys</mods:title>
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<mods:part id="6C6197C304138437D7503E25D8EFA103">
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<mods:date id="257838AC8BAB71E54A6005E962057554">2015</mods:date>
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<mods:detail id="BE4072A8667504BC66F77EFDE5C1078F" type="volume">
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<mods:number id="F03900E4EF1DD8565DFC18D7E75F24CD">536</mods:number>
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<mods:extent id="08B444149A429FEC94507BB9D7031312" unit="page">
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<mods:start id="2CB9B9127D6E3E9D339625706EBE923B">1</mods:start>
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<mods:end id="16EE90406FDADEC4B621C76019B00E27">92</mods:end>
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<mods:location id="3583865135F9BEB78881AE99ABE46DA9">
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<mods:url id="2135D14D8DFC05B9E27C15012AC50A0F">http://dx.doi.org/10.3897/zookeys.536.5982</mods:url>
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</mods:location>
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<mods:classification id="A1E3AF0AC0D5D803A88358C8C7324356">journal article</mods:classification>
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<mods:identifier id="D856E46190650D157DF0BDB273ADD559" type="DOI">http://dx.doi.org/10.3897/zookeys.536.5982</mods:identifier>
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<mods:identifier id="B02E46DFF8DB5FC92F8089F7B2178A98" type="Pensoft-Pub">1313-2970-536-1</mods:identifier>
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<mods:identifier id="700F3CAD407AABD644F806B54DFA294F" type="ZooBank">A8B92CDD0B8C4384AAC559648BB45AA5</mods:identifier>
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</mods:mods>
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<treatment id="1882882DAB1B715EFEEE5ED38E5CDAC6" ID-GBIF-Taxon="152063178" LSID="urn:lsid:zoobank.org:act:D7ECDAEE-BB43-4C4A-B668-875E6B2540F2" httpUri="http://treatment.plazi.org/id/1882882DAB1B715EFEEE5ED38E5CDAC6" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="6F1F9A8ABDA655B093821484AFA0F187" pageId="8" pageNumber="9" type="multiple">
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<paragraph id="649684BEEE3B73A4C4122676D42DF920" pageId="8" pageNumber="9">
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<pageBreakToken id="3A0483D7212CC34B2C3A032A37C1D4DB" pageId="8" pageNumber="9" start="start">Taxon</pageBreakToken>
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classification Animalia
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<taxonomicName id="F6ED2EE60F45EE54D86F9FE80EC91223" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
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<taxonomicName id="CFBBDC6706EE75CFEF14D977AB28C728" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="676B01258AE30AF879075D3E9CCECAAF" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="A7965E05F3B2307E4F13BCFD9429ECEF" pageId="8" pageNumber="9">
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<taxonomicName id="05B17A73E4A69E8934DDD33F58358686" ID-CoL="8MNB8" LSID="http://zoobank.org/D7ECDAEE-BB43-4C4A-B668-875E6B2540F2" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus) ailaoicus" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ailaoicus" status="sp. nov." subGenus="Circinatus">Pterostichus (Circinatus) ailaoicus</taxonomicName>
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<taxonomicNameLabel id="26A5CEBDDD5EC42987EF12EC0FAB0C53" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
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Figures 11, 29, 97
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</paragraph>
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</subSubSection>
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<subSubSection id="079825241A7887AC9D91FD3A86D423D0" pageId="9" pageNumber="10" type="type locality">
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<paragraph id="DD6E50224645D20B70ECEBC00120579F" pageId="9" pageNumber="10">
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<pageBreakToken id="ABB5F66EDF6B45C9B1B3B5975020586F" pageId="9" pageNumber="10" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph id="C871CFF426049FC3336039515C7B6654" pageId="9" pageNumber="10">
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Yunnan: Xinping County, Jinshan pass in Ailao mountain (= Ailaoshan) (
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<geoCoordinate id="23EDE3AF0282DA7B213BD74D7D714C1A" direction="north" orientation="latitude" precision="55" value="23.947">N23.947°</geoCoordinate>
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,
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<geoCoordinate id="79CC0E43E97B4620EEBD1038B359EC26" direction="east" orientation="longitude" precision="55" value="101.499">E101.499°</geoCoordinate>
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), altitude 2351 m.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8D495A85478A5D19738804F4C290610" pageId="9" pageNumber="10" type="type material">
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<paragraph id="29992E6500FB8607A1846A2D59937916" pageId="9" pageNumber="10">Type material.</paragraph>
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<paragraph id="656AF9BB7BCA1D32C2462F73BE449F5B" pageId="9" pageNumber="10">
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Holotype (IZAS): male, body length = 10.9 mm, board mounted, genitalia dissected and glued on plastic film pinned under specimen, "CHINA, Yunnan, Xinping / county, Jinshan pass in Ailao / mount range, 2010.IX.14 / YANG Xiaodong lgt. / B10y5006, C.C.C.C."; "
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<geoCoordinate id="7756D92E3CC08A4B8DA97449C732FDB8" direction="north" orientation="latitude" precision="1" value="23.947483">N23°56.849'</geoCoordinate>
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/
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<geoCoordinate id="B87CBB42B218288AC1518F4D5D92F9A1" direction="east" orientation="longitude" precision="1" value="101.499886">E101°29.993'</geoCoordinate>
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2351 m / daytime, mixed forest, in or on dead log"; "HOLOTYPE ♂/
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<taxonomicName id="5D565D18D371C7B2C45E3DC620979A78" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Pterostichus (Circinatus)</taxonomicName>
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/
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<taxonomicName id="78C196A4A0D64801A01763B61ED4EA1C" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="ailaoicus" status="sp. nov.">ailaoicus</taxonomicName>
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<taxonomicNameLabel id="BD5958F0D5D15927B0497B108194422D" rank="species">new species</taxonomicNameLabel>
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/ des. SHI H.L. 2015" [red label].
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</paragraph>
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</subSubSection>
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<subSubSection id="4077998DE0DA56DBDD49B49510424256" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="83D50599B0926C6B093BF67ABE969F51" pageId="9" pageNumber="10">Diagnosis.</paragraph>
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<paragraph id="3A7D803BA8A9583005F826C9D799987F" pageId="9" pageNumber="10">
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Pronotum with single mid-lateral seta; posterior seta located almost at hind angle; hind angle forming indistinct obtuse angle, lateral margin straight before hind angle; elytral microsculpture almost invisible; elytral shoulder angle distinct; males with two large tubercles on terminal sternum; tubercles close to each other, distance between tubercles less than distance between setae on penultimate sternum; fifth tarsomeres glabrous beneath. Apical lamella of aedeagus long, its length approx 2.1 times basal width, apex widened, with a small tooth on right margin. Comparisons between similar species are given in Table 1 under
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<taxonomicName id="89720901E9CA90269EF4314F558A785E" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus maitreya" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maitreya" status="sp. nov.">Pterostichus maitreya</taxonomicName>
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<taxonomicNameLabel id="FA0D8E8F3558445C3C4E29DFA7081514" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<caption id="1BA44D0A5BE6CEA47CD04AF3E6589FF3" ID-Table-UUID="1BA44D0A5BE6CEA47CD04AF3E6589FF3" httpUri="http://table.plazi.org/id/1BA44D0A5BE6CEA47CD04AF3E6589FF3" pageId="9" pageNumber="10">
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<paragraph id="D06E1B6860A9DB93874B8C7AB340F05D" pageId="9" pageNumber="10">
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Table 1. Comparisons for
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<taxonomicName id="13ABE07B423834F09C8F1D30AE89494D" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus maitreya" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maitreya">Pterostichus maitreya</taxonomicName>
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,
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<taxonomicName id="671FAF21D39137A8E2A35962532FEE4C" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="baenningeri">Pterostichus baenningeri</taxonomicName>
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,
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<taxonomicName id="D4AF860DC3D74DF26326F584451B15D1" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus yan" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yan">Pterostichus yan</taxonomicName>
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,
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<taxonomicName id="7C1964AD29D04FD10FF85CC74864EEE4" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus ailaoicus" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ailaoicus">Pterostichus ailaoicus</taxonomicName>
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and
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<taxonomicName id="47DD41210A45711F3D2B6F959C6AE294" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus miao" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="miao">Pterostichus miao</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph id="795243C24B61F27AC1A339647EAA6500" pageId="9" pageNumber="10">
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<table id="E4714C22D52C6E2F3F5EF4F1B3E06985" pageId="9" pageNumber="10">
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<tr id="884C126520EAA933C9D178B3ECE107E4" pageId="9" pageNumber="10">
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<th id="9280C8D8F5451C9D49DF853CB5B599E4" colspan="1" pageId="9" pageNumber="10" rowspan="1">Characters / Species</th>
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<th id="929C19786779BF659427DDC1ED9924B6" colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<taxonomicName id="001A1E32F81DB26DD7C031121966E196" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="baenningeri">Pterostichus baenningeri</taxonomicName>
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</th>
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<th id="14F387FDD9C367374AB50C072265C439" colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<taxonomicName id="FEFB1B1041899DC3B72AA7BF82CC2A84" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus maitreya" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="maitreya">Pterostichus maitreya</taxonomicName>
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</th>
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<th id="1C81C723447322FB4894EA0080D3167D" colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<taxonomicName id="CDFCBF5EE1E3169A43F094328BE36AA4" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus miao" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="miao">Pterostichus miao</taxonomicName>
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</th>
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<th id="B742AE91007A47DF3AFDC343E9961BE1" colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<taxonomicName id="D36098780E91139C6DDF098D7ACC563E" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus ailaoicus" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ailaoicus">Pterostichus ailaoicus</taxonomicName>
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</th>
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<th id="7E1B767125994603E6BC67397D02E177" colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<taxonomicName id="54592393C953B352D20E28E90BCE0F4A" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus yan" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yan">Pterostichus yan</taxonomicName>
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</th>
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</tr>
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</table>
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</paragraph>
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<paragraph id="BC4A20983206E3047A9A4A17DBD0DE78" pageId="9" pageNumber="10">
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This new species is similar to
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<taxonomicName id="48B5E1AE5EE72690AB1FE148B2B9AD3D" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="baenningeri">Pterostichus baenningeri</taxonomicName>
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in body size and form, pronotal shape, and male terminal sternum. Despite the significant differences in male genitalia (shape of apical lamella, see Figs 29B and 40B), they also differ in: (1)
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<taxonomicName id="64AB0A9C5D2C3BD08D53E412024AB049" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus ailaoicus" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ailaoicus" status="sp. nov.">Pterostichus ailaoicus</taxonomicName>
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<taxonomicNameLabel id="D6FEBB93EC1F58C2F86A767920726780" rank="species">sp. n.</taxonomicNameLabel>
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have the fifth tarsomeres glabrous beneath, but these are setose in
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<taxonomicName id="A7801AE982918A06DD7F066A13B9BDDD" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="baenningeri">Pterostichus baenningeri</taxonomicName>
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; (2)
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<taxonomicName id="E0332D8A8CE144A58036F7D43FB49AC8" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus ailaoicus" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ailaoicus" status="sp. nov.">Pterostichus ailaoicus</taxonomicName>
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<taxonomicNameLabel id="60BEE9FA43031183F391033C92D0FDA7" rank="species">sp. n.</taxonomicNameLabel>
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have the two tubercles on the male terminal sternum close to each other, distance between tubercles less than the distance between setae on penultimate sternum; but in
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<taxonomicName id="FA06C58AB50F5F70D96A900541C5F8A8" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="baenningeri">Pterostichus baenningeri</taxonomicName>
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, distance between tubercles on terminal sternum is greater than the distance between setae on penultimate sternum.
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</paragraph>
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</subSubSection>
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<subSubSection id="5D19855D379FAFB6CA95B178E9307841" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
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<paragraph id="110DCBBFA31DFD3378C312889C0EDCF8" pageId="9" pageNumber="10">Description.</paragraph>
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<paragraph id="0D4659024FED80618D31E95B3D090534" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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Body form fairly elongate, body length 10.9 mm; dorsal side almost black, moderately shining, elytron with faint iridescent shine; mouthparts, antenna, tarsus, tibia, and apex of femur reddish brown; ventral side blackish. Elytral microsculpture weak, barely visible, linear. Head. Frons without punctures; antenna reaching elytron basal sixth; gena shorter than length of eye, briefly tumid behind eye. Pronotum round, lateral margin curved in middle, nearly straight before hind angle, widest at approx anterior two fifths; posterior margin a little narrower than anterior margin; PW/PL = 1.10; one mid-lateral seta present, located a little before greatest width; posterior seta close to but a little anterior to hind angle; hind angle forming indistinct obtuse angle; basal fovea shallow, faintly defined; inner groove subparallel to median line, not reaching posterior margin; outer groove completely vanished; outer area of inner groove flat; basal foveal area finely punctate along inner groove. Elytron oviform, with basal ridge almost straight; elytral shoulder moderately widened, shoulder angle between basal ridge and lateral margin forming obtuse angle, humeral tooth very
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<pageBreakToken id="B89AFBEBCBE64A8438F8C4333F19134B" pageId="10" pageNumber="11" start="start">small</pageBreakToken>
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, not jutting out; intervals feebly convex; striae moderately deep, without punctures; scutellar stria short, apex free; third interval with two setigerous pores adjacent to second stria; umbilical pore series on ninth interval sparse in middle, composed of 16-17 pores (6, 1, 9-10). Ventral side. Proepisternum without punctures, mesepisternum and metepisternum finely punctate; penultimate sternum not modified; terminal sternum with two large tubercles vaguely defined, approx one third length of terminal sternum; tubercles located a little before middle of sternum, distance between tubercles less than distance between two setae on penultimate sternum; region between tubercles not depressed. Legs. Fifth tarsomeres glabrous beneath; males with apical half of mesotibia slightly widened, inner margin crenulate; first metatarsomere with distinct carina on outer surface, such carina superficial on full length of second metatarsomere. Male genitalia. Median lobe of male genitalia bent approx 90 degrees, apex gradually bent ventrally (Fig. 29A); ventral margin slightly curved in middle, weakly sinuate before apex; dorsal margin gradually curved; apical orifice large, slightly turned to left side, opened on ventral side; in lateral view, apical lamella very long, laminate with base slightly thickened, sinuate before apex, apex pointing apical-ventrally, its length approx one third length of apical orifice; in dorsal view, apical lamella located on right side of aedeagal apex, its base not grooved on dorsal surface; apical lamella long with rounded apex, length approx 2.1 times basal width, distinctly widened at approx apical third, with a small tooth on right margin (Fig. 29B). Right paramere straight and stout, apical half fusiform, slightly enlarged and then narrowed to apex, inner margin evenly curved before apex; length approx three times greatest width; apex obtuse (Fig. 29C). Endophallus not studied. Female genitalia unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C8E95DF0BEE88F5D69A0D05A7AC24EE" pageId="10" pageNumber="11" type="distribution">
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<paragraph id="14DA765EA8AB2D510235ED3915AF7E49" pageId="10" pageNumber="11">Distribution.</paragraph>
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<paragraph id="9D8AC64EE11D4EC962ECE7C6D90BFCB2" pageId="10" pageNumber="11">This species is known only from the holotype collected from Ailaoshan Mt. in Yunnan Province (Map 1). The altitude is 2351 m.</paragraph>
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</subSubSection>
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<subSubSection id="84E6A01CC83CB17BDA7E669C1F92DDF8" pageId="10" pageNumber="11" type="etymology">
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<paragraph id="A606B1B3427C20E61014B85C7C652939" pageId="10" pageNumber="11">Etymology.</paragraph>
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<paragraph id="91148542B8498A6F8B493B774EC84654" pageId="10" pageNumber="11">This new species is named for its type locality, Ailao Mountain.</paragraph>
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</subSubSection>
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<subSubSection id="064FCDDAA27D04FDD457B387CD6E90ED" pageId="10" pageNumber="11" type="affinities">
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<paragraph id="68B854DF55C432A44E8C0616BB3BF405" pageId="10" pageNumber="11">Affinities.</paragraph>
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<paragraph id="7D79654DB1F1A966C1DEDBA40A76DB24" pageId="10" pageNumber="11">
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<taxonomicName id="0CF8D1D09A53500FC8C5E211F36D061C" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus ailaoicus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ailaoicus" status="sp. nov.">Pterostichus ailaoicus</taxonomicName>
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<taxonomicNameLabel id="02DCD85B9D1E4C2602235B4CFD3EB69A" rank="species">sp. n.</taxonomicNameLabel>
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is close to
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<taxonomicName id="BB68311AFEB606A05CFF20E306DAC765" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus baenningeri" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="baenningeri">Pterostichus baenningeri</taxonomicName>
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,
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<taxonomicName id="8A67AC0746F639E81ADEAD93B00659CC" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus maitreya" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="maitreya" status="sp. nov.">Pterostichus maitreya</taxonomicName>
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<taxonomicNameLabel id="DCC5E57EC26D4C96D010B950A989AD6F" rank="species">sp. n.</taxonomicNameLabel>
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,
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<taxonomicName id="79834B9581894A4F53E3AA5AC68E3D77" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus miao" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="miao" status="sp. nov.">Pterostichus miao</taxonomicName>
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<taxonomicNameLabel id="CFD6BEE6ED09E5C4233B3BE1FFBF92F4" rank="species">sp. n.</taxonomicNameLabel>
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, and
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<taxonomicName id="872C92A06FB8C4452A675A0540EC0328" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus yan" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="yan" status="sp. nov.">Pterostichus yan</taxonomicName>
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<taxonomicNameLabel id="24243A070D8E4103BFE4223B0F26ADDD" rank="species">sp. n.</taxonomicNameLabel>
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in their similarities of male terminal sternum and male genitalia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |