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<document id="768313C780E8DE9A5932B2D01EA66810" ID-DOI="10.5281/zenodo.187388" ID-GBIF-Dataset="566515ff-3e08-4daa-bc0d-8bb08381d5d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="187388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461069229596" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Hadiaty, Renny K." docDate="2009" docId="186B87BAFF8CFFEDFF59D056890E6AA3" docLanguage="en" docName="zt02085p068.pdf" docOrigin="Zootaxa 2085" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Glyptothorax ketambe Ng &amp; Hadiaty, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="E452FFC2FF8DFFE9FFCED10F8C6C6D28" masterDocTitle="Glyptothorax ketambe, a new catfish (Teleostei: Sisoridae) from northern Sumatra" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698584028730" updateUser="plazi">
<mods:mods id="9F246B5F6051352E1DA2BE083918F216" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="20DF1F41C600FAAF9AC5BD5E71AE9F63">
<mods:title id="EC5F38B2737D7DB68057BB8284D27494">Glyptothorax ketambe, a new catfish (Teleostei: Sisoridae) from northern Sumatra</mods:title>
</mods:titleInfo>
<mods:name id="369C705A2F21E41B5D3FAEC0F4C1EF0A" type="personal">
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</mods:role>
<mods:namePart id="275524EC55F3EC6EF1EF428702F66973">Ng, Heok Hee</mods:namePart>
</mods:name>
<mods:name id="BF68195BC00190FB04B4D935DB085CE5" type="personal">
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<mods:namePart id="947BB5A2AE2765103068348E3FCE3CF9">Hadiaty, Renny K.</mods:namePart>
</mods:name>
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<mods:title id="5CB224AA82BC4919EE0AF8F2959E486A">Zootaxa</mods:title>
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<mods:date id="B20842A88FEF9DFEE7B7BA877CF4A5E8">2009</mods:date>
<mods:detail id="8409E43D67B99797B7B329F9176044DE" type="volume">
<mods:number id="62D2FC683E5BFCEB83C481B8AF0FECD8">2085</mods:number>
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<mods:classification id="9ED4A4ACB7EB847B94F4848F990A08ED">journal article</mods:classification>
<mods:identifier id="03A4A3868548EA2D9BD90F6E6E68C07F" type="DOI">10.5281/zenodo.187388</mods:identifier>
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<mods:identifier id="AB89D2059B77D47E0AC771E45119647E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="55FB0899DFC7610860325CD8C990BFC6" type="Zenodo-Dep">187388</mods:identifier>
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<treatment id="186B87BAFF8CFFEDFF59D056890E6AA3" ID-DOI="http://doi.org/10.5281/zenodo.5674291" ID-GBIF-Taxon="119615687" ID-Zenodo-Dep="5674291" LSID="urn:lsid:plazi:treatment:186B87BAFF8CFFEDFF59D056890E6AA3" httpUri="http://treatment.plazi.org/id/186B87BAFF8CFFEDFF59D056890E6AA3" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62">
<subSubSection id="D8D86527FF8CFFE8FF59D0568D436CBE" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="907D36ACFF8CFFE8FF59D0568E6B6C5A" blockId="1.[151,519,344,406]" box="[151,519,344,371]" pageId="1" pageNumber="62">
<heading id="CB3581C0FF8CFFE8FF59D0568E6B6C5A" bold="true" box="[151,519,344,371]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="A2B6EABEFF8CFFE8FF59D0568E6B6C5A" bold="true" box="[151,519,344,371]" pageId="1" pageNumber="62">
<taxonomicName id="57C24D2FFF8CFFE8FF59D0568DCF6C5B" ID-CoL="3GJD5" box="[151,419,345,371]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="ketambe" status="sp. nov.">
<emphasis id="A2B6EABEFF8CFFE8FF59D0568DCF6C5B" bold="true" box="[151,419,345,371]" italics="true" pageId="1" pageNumber="62">Glyptothorax ketambe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B98557C5FF8CFFE8FE67D0578E6B6C5A" box="[425,519,344,370]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="907D36ACFF8CFFE8FF59D0738D436CBE" blockId="1.[151,519,344,406]" box="[151,303,380,406]" pageId="1" pageNumber="62">
(
<figureCitation id="08F92A29FF8CFFE8FF51D0738C9D6CBE" box="[159,241,380,406]" captionStart="FIGURE 1" captionStartId="2.[151,255,1798,1822]" captionTargetBox="[151,1436,197,1772]" captionTargetId="figure@2.[151,1436,197,1773]" captionTargetPageId="2" captionText="FIGURE 1. Glyptothorax ketambe, MZB 8694, holotype, 100.0 mm SL; Sumatra: Aceh, Alas River drainage." httpUri="https://zenodo.org/record/187389/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
&amp;
<figureCitation id="08F92A29FF8CFFE8FED8D0738D496CBE" box="[278,293,380,406]" captionStart="FIGURE 3" captionStartId="5.[151,255,777,801]" captionTargetBox="[151,1436,198,747]" captionTargetId="figure@5.[151,1436,197,752]" captionTargetPageId="5" captionText="FIGURE 3. Collection localities of G. ketambe." httpUri="https://zenodo.org/record/187391/files/figure.png" pageId="1" pageNumber="62">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D8D86527FF8CFFE8FF59D0C7890B6F7F" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="907D36ACFF8CFFE8FF59D0C788766F22" blockId="1.[151,1437,456,2030]" pageId="1" pageNumber="62">
<emphasis id="A2B6EABEFF8CFFE8FF59D0C78D226CCA" bold="true" box="[151,334,456,482]" pageId="1" pageNumber="62">
<typeStatus id="4F79880EFF8CFFE8FF59D0C78CB96CCA" box="[151,213,456,482]" pageId="1" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4F79880EFF8CFFE8FE98D0C68DA46CCB" box="[342,456,457,483]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
:
<collectionCode id="F6D3AE69FF8CFFE8FE1AD0C68E786CCB" box="[468,532,457,483]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="62">MZB</collectionCode>
8694, 100.0 mm
<collectionCode id="F6D3AE69FF8CFFE8FD29D0C68F616CCB" box="[743,781,457,483]" pageId="1" pageNumber="62">SL</collectionCode>
; Sumatra: Nangroe Aceh Darussalam province, Sungai
<collectingCountry id="E8D5763CFF8CFFE8FF59D0FF8CA16F22" box="[151,205,496,522]" name="Kazakhstan" pageId="1" pageNumber="62">Alas</collectingCountry>
at
<taxonomicName id="57C24D2FFF8CFFE8FF3ED0FF8D366F22" box="[240,346,496,522]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="66" phylum="Chordata" rank="species" species="ketambe">Ketambe</taxonomicName>
research station; R.K. Hadiaty &amp; A. Mun'im,
<date id="E47C106CFF8CFFE8FCB8D0FF887A6F22" box="[886,1046,496,522]" pageId="1" pageNumber="62" value="1998-06-16">16 June 1998</date>
.
</paragraph>
<paragraph id="907D36ACFF8CFFE8FF0BD319890B6F7F" blockId="1.[151,1437,456,2030]" pageId="1" pageNumber="62">
<typeStatus id="4F79880EFF8CFFE8FF0BD3198D2D6F18" box="[197,321,534,560]" pageId="1" pageNumber="62" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="F6D3AE69FF8CFFE8FE83D3198DE26F18" box="[333,398,534,560]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="62">MZB</collectionCode>
8695 (5), 49.0–95.0 mm
<collectionCode id="F6D3AE69FF8CFFE8FD04D3198E9D6F18" box="[714,753,534,560]" pageId="1" pageNumber="62">SL</collectionCode>
;
<collectionCode id="F6D3AE69FF8CFFE8FD31D3198F566F18" box="[767,826,534,560]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="62">ZRC</collectionCode>
46421 (1),
<quantity id="573A9B49FF8CFFE8FC04D31988546F18" box="[970,1080,534,560]" metricMagnitude="-2" metricUnit="m" metricValue="6.909999999999999" pageId="1" pageNumber="62" unit="mm" value="69.1">69.1 mm</quantity>
<collectionCode id="F6D3AE69FF8CFFE8FB8ED319880B6F18" box="[1088,1127,534,560]" pageId="1" pageNumber="62">SL</collectionCode>
; Sumatra: Nangroe Aceh Darussalam province, Kutacane fish market,
<geoCoordinate id="F5F6506BFF8CFFE8FD68D3328E976F7F" box="[678,763,573,599]" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="925" value="3.5">3°30'N</geoCoordinate>
,
<geoCoordinate id="F5F6506BFF8CFFE8FCC9D3328F0B6F7F" box="[775,871,573,599]" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="925" value="97.8">97°48'E</geoCoordinate>
; R.K. Hadiaty &amp; A. Mun'im,
<date id="E47C106CFF8CFFE8FB0DD332890F6F7F" box="[1219,1379,573,599]" pageId="1" pageNumber="62" value="1998-06-22">22 June 1998</date>
.
</paragraph>
</subSubSection>
<subSubSection id="D8D86527FF8CFFE8FF0BD36C8D8468F8" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="907D36ACFF8CFFE8FF0BD36C8D8468F8" blockId="1.[151,1437,456,2030]" pageId="1" pageNumber="62">
<emphasis id="A2B6EABEFF8CFFE8FF0BD36C8D206F55" bold="true" box="[197,332,611,637]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="57C24D2FFF8CFFE8FE9BD36B8E076F55" box="[341,619,612,637]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="A2B6EABEFF8CFFE8FE9BD36B8E076F55" box="[341,619,612,637]" italics="true" pageId="1" pageNumber="62">Glyptothorax ketambe</emphasis>
</taxonomicName>
can be distinguished from all other Sundaic congeners except
<taxonomicName id="57C24D2FFF8CFFE8FA4CD36B8C966F8B" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="A2B6EABEFF8CFFE8FA4CD36B8C966F8B" italics="true" pageId="1" pageNumber="62">G. schmidti</emphasis>
</taxonomicName>
and
<taxonomicName id="57C24D2FFF8CFFE8FEFDD3858DA96F8B" box="[307,453,650,675]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="A2B6EABEFF8CFFE8FEFDD3858DA96F8B" box="[307,453,650,675]" italics="true" pageId="1" pageNumber="62">G. siamensis</emphasis>
</taxonomicName>
in having a color pattern consisting of a dark brown body with yellowish midlateral and mid-dorsal stripes. It differs from
<taxonomicName id="57C24D2FFF8CFFE8FDBAD3BE8E926FE2" box="[628,766,689,714]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="A2B6EABEFF8CFFE8FDBAD3BE8E926FE2" box="[628,766,689,714]" italics="true" pageId="1" pageNumber="62">G. schmidti</emphasis>
</taxonomicName>
in having a deeper caudal peduncle (7.6–8.9% SL vs. 5.8–7.3) and a less posteriorly-extended thoracic adhesive apparatus (not reaching vs. reaching to the level of the base of the last pectoral-fin ray) and from
<taxonomicName id="57C24D2FFF8CFFE8FD09D3F08F326E30" box="[711,862,767,792]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="A2B6EABEFF8CFFE8FD09D3F08F326E30" box="[711,862,767,792]" italics="true" pageId="1" pageNumber="62">G. siamensis</emphasis>
</taxonomicName>
in having a narrower head (16.1–17.9% SL vs. 17.2–20.1) and an almost straight (vs. noticeably curved) lateral margin of the thoracic adhesive apparatus (
<figureCitation id="08F92A29FF8CFFE8FF51D2438C846E4E" box="[159,232,844,870]" captionStart="FIGURE 2. T" captionStartId="4.[151,259,1336,1360]" captionTargetBox="[273,1312,199,1307]" captionTargetId="figure@4.[273,1312,197,1308]" captionTargetPageId="4" captionText="FIGURE 2. T horacic adhesive apparatus of: a. Glyptothorax ketambe, holotype, MZB 8694, 100.0 mm SL; b. G. plectilis, paratype, ZRC 46420, 44.3 mm SL; c. G. siamensis, CMK 10829, 71.8 mm SL. Illustrations not to scale." httpUri="https://zenodo.org/record/187390/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
). There are eight other congeners from mainland Southeast Asia that share a similar color pattern of a dark-colored body (with or without distinct pale midlateral and mid-dorsal stripes) with
<taxonomicName id="57C24D2FFF8CFFE8FB29D27D891C6EA3" box="[1255,1392,882,907]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="A2B6EABEFF8CFFE8FB29D27D891C6EA3" box="[1255,1392,882,907]" italics="true" pageId="1" pageNumber="62">G. ketambe</emphasis>
</taxonomicName>
:
<taxonomicName id="57C24D2FFF8CFFE8FA4CD27D8D676E9A" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="coracinus">
<emphasis id="A2B6EABEFF8CFFE8FA4CD27D8D676E9A" italics="true" pageId="1" pageNumber="62">G. coracinus</emphasis>
</taxonomicName>
,
<taxonomicName id="57C24D2FFF8CFFE8FED4D2968DB86E9A" box="[282,468,921,946]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="interspinalus">
<emphasis id="A2B6EABEFF8CFFE8FED4D2968DB86E9A" box="[282,468,921,946]" italics="true" pageId="1" pageNumber="62">G. interspinalus</emphasis>
</taxonomicName>
,
<taxonomicName id="57C24D2FFF8CFFE8FE2AD2968E1D6E9A" box="[484,625,921,946]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="laosensis">
<emphasis id="A2B6EABEFF8CFFE8FE2AD2968E1D6E9A" box="[484,625,921,946]" italics="true" pageId="1" pageNumber="62">G. laosensis</emphasis>
</taxonomicName>
,
<taxonomicName id="57C24D2FFF8CFFE8FD4ED2968F496E9A" box="[640,805,921,946]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="A2B6EABEFF8CFFE8FD4ED2968F496E9A" box="[640,805,921,946]" italics="true" pageId="1" pageNumber="62">G. longicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="57C24D2FFF8CFFE8FCFBD296886D6E9A" box="[821,1025,921,946]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="longjiangensis">
<emphasis id="A2B6EABEFF8CFFE8FCFBD296886D6E9A" box="[821,1025,921,946]" italics="true" pageId="1" pageNumber="62">G. longjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName id="57C24D2FFF8CFFE8FBDFD29688F36E9A" box="[1041,1183,921,946]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="strabonis">
<emphasis id="A2B6EABEFF8CFFE8FBDFD29688F36E9A" box="[1041,1183,921,946]" italics="true" pageId="1" pageNumber="62">G. strabonis</emphasis>
</taxonomicName>
,
<taxonomicName id="57C24D2FFF8CFFE8FB61D29689256E9A" box="[1199,1353,921,946]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="trilineatus">
<emphasis id="A2B6EABEFF8CFFE8FB61D29689256E9A" box="[1199,1353,921,946]" italics="true" pageId="1" pageNumber="62">G. trilineatus</emphasis>
</taxonomicName>
and
<emphasis id="A2B6EABEFF8CFFE8FA4DD2968E496EF1" italics="true" pageId="1" pageNumber="62">
<taxonomicName id="57C24D2FFF8CFFE8FA4DD2968D636EF1" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="zanaensis">G. zanaensis</taxonomicName>
.
<taxonomicName id="57C24D2FFF8CFFE8FED5D2CF8E496EF1" box="[283,549,960,985]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="ketambe">Glyptothorax ketambe</taxonomicName>
</emphasis>
is distinguished from
<emphasis id="A2B6EABEFF8CFFE8FCFDD2CF8FAB6EF1" box="[819,967,960,985]" italics="true" pageId="1" pageNumber="62">G. c o r a c i n u s</emphasis>
in having a wider interorbital distance (29.0–31.2% HL vs. 26.2–26.7), from
<taxonomicName id="57C24D2FFF8CFFE8FDAED2E88F776928" box="[608,795,999,1024]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="interspinalus">
<emphasis id="A2B6EABEFF8CFFE8FDAED2E88F776928" box="[608,795,999,1024]" italics="true" pageId="1" pageNumber="62">G. interspinalus</emphasis>
</taxonomicName>
in having a deeper caudal peduncle (7.6–8.9% SL vs. 6.1–8.0) and a narrower head (16.1–17.9% SL vs. 17.2–22.1), from
<emphasis id="A2B6EABEFF8CFFE8FC04D502883A690E" box="[970,1110,1037,1062]" italics="true" pageId="1" pageNumber="62">G. l a o s e n s i s</emphasis>
in having a narrower head (16.1–17.9% SL vs. 19.2–21.1) and a less posteriorly-extended thoracic adhesive apparatus (not reaching vs. reaching beyond the level of the base of the last pectoral-fin ray), and from
<taxonomicName id="57C24D2FFF8CFFE8FBDAD55488D4695C" box="[1044,1208,1115,1140]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="A2B6EABEFF8CFFE8FBDAD55488D4695C" box="[1044,1208,1115,1140]" italics="true" pageId="1" pageNumber="62">G. longicauda</emphasis>
</taxonomicName>
in having a shorter caudal peduncle (20.1–22.9% SL vs. 23.6) and fewer vertebrae (37–38 vs. 42). It differs from
<taxonomicName id="57C24D2FFF8CFFE8FA4CD58E8D2469E9" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="longjiangensis">
<emphasis id="A2B6EABEFF8CFFE8FA4CD58E8D2469E9" italics="true" pageId="1" pageNumber="62">G. longjiangensis</emphasis>
</taxonomicName>
in having tubercles on the dorsal surface of the head with a single, median ridge (vs. with anastomosing tubercles with multiple ridges) and without any pale markings on the nuchal plate (vs. part or all of the nuchal plate pale-colored), from
<taxonomicName id="57C24D2FFF8CFFE8FD46D5FA8F736826" box="[648,799,1269,1294]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="strabonis">
<emphasis id="A2B6EABEFF8CFFE8FD46D5FA8F736826" box="[648,799,1269,1294]" italics="true" pageId="1" pageNumber="62">G. strabonis</emphasis>
</taxonomicName>
in having a larger eye (7.5–9.6% HL vs. 6.2–6.5), interorbital distance (29.0–31.2% HL vs. 26.0–27.2) and fewer serrations on the posterior margin of the pectoral spine (9–12 vs. 14–16), from
<taxonomicName id="57C24D2FFF8CFFE8FDAFD44C8E906874" box="[609,764,1347,1372]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="trilineatus">
<emphasis id="A2B6EABEFF8CFFE8FDAFD44C8E906874" box="[609,764,1347,1372]" italics="true" pageId="1" pageNumber="62">G. trilineatus</emphasis>
</taxonomicName>
in having a longer caudal peduncle (20.1–22.9% SL vs. 16.7–18.4), and from
<taxonomicName id="57C24D2FFF8CFFE8FE58D4668E4368AA" box="[406,559,1385,1410]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="zainaensis">
<emphasis id="A2B6EABEFF8CFFE8FE58D4668E4368AA" box="[406,559,1385,1410]" italics="true" pageId="1" pageNumber="62">G. zainaensis</emphasis>
</taxonomicName>
in having a shorter adipose-fin base (12.0–14.5% SL vs. 14.4–18.4), and a less posteriorly-extended thoracic adhesive apparatus (not reaching to the level of the base of the last pectoralfin ray vs. reaching beyond).
</paragraph>
</subSubSection>
<subSubSection id="D8D86527FF8CFFEDFF0BD4D28E9D6BE2" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62" type="description">
<paragraph id="907D36ACFF8CFFE8FF0BD4D28EA66B90" blockId="1.[151,1437,456,2030]" pageId="1" pageNumber="62">
<emphasis id="A2B6EABEFF8CFFE8FF0BD4D28D0E68DF" bold="true" box="[197,354,1501,1527]" pageId="1" pageNumber="62">Description.</emphasis>
Biometric data as in
<tableCitation id="DD400317FF8CFFE8FDA7D4D28EA868DF" box="[617,708,1501,1527]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetBox="[151,1275,330,1420]" captionTargetPageId="3" captionText="TABLE 1. Biometric data for Glyptothorax ketambe (n = 6)." httpUri="http://table.plazi.org/id/C4BD6624FF8EFFEAFF59D1988F7D6D87" pageId="1" pageNumber="62" tableUuid="C4BD6624FF8EFFEAFF59D1988F7D6D87">Table 1</tableCitation>
. Head depressed, body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile flat to anal-fin base, then sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin tuberculate, with small tubercles on sides of body. Lateral line complete and midlateral. Vertebrae 16+21=37* (2), 17+20=37 (3) or 17+21=38 (1).
</paragraph>
<paragraph id="907D36ACFF8CFFE8FF0BD7CA8FD26A7B" blockId="1.[151,1437,456,2030]" pageId="1" pageNumber="62">Head depressed and broad, triangular when viewed laterally. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill openings broad, extending from directly beneath post-temporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, horizontal axis longest; located entirely in dorsal half of head.</paragraph>
<paragraph id="907D36ACFF8CFFE8FF0BD66F88916AC6" blockId="1.[151,1437,456,2030]" pageId="1" pageNumber="62">Barbels in four pairs. Maxillary barbel long and slender, extending to middle of pectoral-fin base. Nasal barbel slender, extending to halfway between its base and anterior orbital margin. Inner mandibular barbel origin close to midline, extending to midway between its base and that of pectoral spine. Outer mandibular barbel originating posterolateral of inner mandibular barbel, extending to base of pectoral spine.</paragraph>
<caption id="C4BD6624FF8FFFEBFF59D609895E6A35" box="[151,1330,1798,1822]" httpUri="https://zenodo.org/record/187389/files/figure.png" pageId="2" pageNumber="63" targetBox="[151,1436,197,1772]" targetPageId="2">
<paragraph id="907D36ACFF8FFFEBFF59D609895E6A35" blockId="2.[151,1330,1798,1822]" box="[151,1330,1798,1822]" pageId="2" pageNumber="63">
<emphasis id="A2B6EABEFF8FFFEBFF59D6098D756A36" bold="true" box="[151,281,1798,1822]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
<taxonomicName id="57C24D2FFF8FFFEBFEEED6098E626A35" box="[288,526,1798,1821]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="63" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="A2B6EABEFF8FFFEBFEEED6098E626A35" box="[288,526,1798,1821]" italics="true" pageId="2" pageNumber="63">Glyptothorax ketambe</emphasis>
</taxonomicName>
, MZB 8694, holotype, 100.0 mm SL; Sumatra: Aceh, Alas River drainage.
</paragraph>
</caption>
<paragraph id="907D36ACFF8FFFEBFF0BD6618FC86AFD" blockId="2.[151,1436,1902,2005]" pageId="2" pageNumber="63">Mouth inferior, premaxillary tooth band partially exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth in single broad semilunate band. Dentary teeth in two narrow crescentic bands separated at midline.</paragraph>
<caption id="C4BD6624FF8EFFEAFF59D1988F7D6D87" ID-Table-UUID="C4BD6624FF8EFFEAFF59D1988F7D6D87" box="[151,785,151,175]" httpUri="http://table.plazi.org/id/C4BD6624FF8EFFEAFF59D1988F7D6D87" pageId="3" pageNumber="64" targetBox="[151,1275,330,1420]" targetIsTable="true" targetPageId="3">
<paragraph id="907D36ACFF8EFFEAFF59D1988F7D6D87" blockId="3.[151,1275,151,311]" box="[151,785,151,175]" pageId="3" pageNumber="64">
<emphasis id="A2B6EABEFF8EFFEAFF59D1988D646D87" bold="true" box="[151,264,151,175]" pageId="3" pageNumber="64">TABLE 1.</emphasis>
Biometric data for
<taxonomicName id="57C24D2FFF8EFFEAFE15D1978EA46D87" box="[475,712,152,175]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="A2B6EABEFF8EFFEAFE15D1978EA46D87" box="[475,712,152,175]" italics="true" pageId="3" pageNumber="64">Glyptothorax ketambe</emphasis>
</taxonomicName>
(n=6).
</paragraph>
</caption>
<paragraph id="907D36ACFF8EFFEAFD93D1C588976DC9" blockId="3.[151,1275,151,311]" box="[605,1275,202,225]" pageId="3" pageNumber="64">
<typeStatus id="4F79880EFF8EFFEAFD93D1C58EAC6DC9" box="[605,704,202,225]" pageId="3" pageNumber="64" type="holotype">Holotype</typeStatus>
MZB 8694 Range Mean±SD
</paragraph>
<paragraph id="907D36ACFF8EFFEAFF51D1FA8F826C24" blockId="3.[151,1275,151,311]" box="[159,1006,245,268]" pageId="3" pageNumber="64">Standard length (mm) 100.0 49.0–100.0</paragraph>
<paragraph id="907D36ACFF8EFFEAFF51D02F8CB96C1F" blockId="3.[151,1275,151,311]" box="[159,213,288,311]" pageId="3" pageNumber="64">%SL</paragraph>
<paragraph id="907D36ACFF8EFFEAFF51D045889568A4" pageId="3" pageNumber="64">
<table id="E2C2C40CFF8E0016FF59D045889768A4" box="[151,1275,330,1420]" gridcols="4" gridrows="26" pageId="3" pageNumber="64">
<tr id="2EF234EEFF8E0016FF59D04588976C49" box="[151,1275,330,353]" gridrow="0" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D0458DD06C49" box="[151,444,330,353]" gridcol="0" gridrow="0" pageId="3" pageNumber="64">Predorsal length</th>
<th id="6D235D92FF8E0016FD93D0458EE06C49" box="[605,652,330,353]" gridcol="1" gridrow="0" pageId="3" pageNumber="64">34.0</th>
<th id="6D235D92FF8E0016FCBBD0458F8C6C49" box="[885,992,330,353]" gridcol="2" gridrow="0" pageId="3" pageNumber="64">33.3–35.9</th>
<th id="6D235D92FF8E0016FB43D04588976C49" box="[1165,1275,330,353]" gridcol="3" gridrow="0" pageId="3" pageNumber="64">34.2±0.90</th>
</tr>
<tr id="2EF234EEFF8E0016FF59D07A88976CA4" box="[151,1275,373,396]" gridrow="1" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D07A8DD06CA4" box="[151,444,373,396]" gridcol="0" gridrow="1" pageId="3" pageNumber="64">Preanal length</th>
<td id="6D235D92FF8E0016FD93D07A8EE06CA4" box="[605,652,373,396]" gridcol="1" gridrow="1" pageId="3" pageNumber="64">65.0</td>
<td id="6D235D92FF8E0016FCBBD07A8F8C6CA4" box="[885,992,373,396]" gridcol="2" gridrow="1" pageId="3" pageNumber="64">64.3–67.5</td>
<td id="6D235D92FF8E0016FB43D07A88976CA4" box="[1165,1275,373,396]" gridcol="3" gridrow="1" pageId="3" pageNumber="64">66.0±1.24</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D0AF88976C9F" box="[151,1275,416,439]" gridrow="2" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D0AF8DD06C9F" box="[151,444,416,439]" gridcol="0" gridrow="2" pageId="3" pageNumber="64">Prepelvic length</th>
<td id="6D235D92FF8E0016FD93D0AF8EE06C9F" box="[605,652,416,439]" gridcol="1" gridrow="2" pageId="3" pageNumber="64">45.7</td>
<td id="6D235D92FF8E0016FCBBD0AF8F8C6C9F" box="[885,992,416,439]" gridcol="2" gridrow="2" pageId="3" pageNumber="64">45.4–47.8</td>
<td id="6D235D92FF8E0016FB43D0AF88976C9F" box="[1165,1275,416,439]" gridcol="3" gridrow="2" pageId="3" pageNumber="64">46.3±0.95</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D0C588976CC9" box="[151,1275,458,481]" gridrow="3" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D0C58DD06CC9" box="[151,444,458,481]" gridcol="0" gridrow="3" pageId="3" pageNumber="64">Prepectoral length</th>
<td id="6D235D92FF8E0016FD93D0C58EE06CC9" box="[605,652,458,481]" gridcol="1" gridrow="3" pageId="3" pageNumber="64">19.1</td>
<td id="6D235D92FF8E0016FCBBD0C58F8C6CC9" box="[885,992,458,481]" gridcol="2" gridrow="3" pageId="3" pageNumber="64">18.2–20.6</td>
<td id="6D235D92FF8E0016FB43D0C588976CC9" box="[1165,1275,458,481]" gridcol="3" gridrow="3" pageId="3" pageNumber="64">19.6±0.95</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D0FA88976F24" box="[151,1275,501,524]" gridrow="4" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D0FA8DD06F24" box="[151,444,501,524]" gridcol="0" gridrow="4" pageId="3" pageNumber="64">Length of dorsal-fin base</th>
<td id="6D235D92FF8E0016FD93D0FA8EE06F24" box="[605,652,501,524]" gridcol="1" gridrow="4" pageId="3" pageNumber="64">11.4</td>
<td id="6D235D92FF8E0016FCBBD0FA8F8C6F24" box="[885,992,501,524]" gridcol="2" gridrow="4" pageId="3" pageNumber="64">11.0–13.3</td>
<td id="6D235D92FF8E0016FB43D0FA88976F24" box="[1165,1275,501,524]" gridcol="3" gridrow="4" pageId="3" pageNumber="64">11.8±0.85</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D32F88976F1F" box="[151,1275,544,567]" gridrow="5" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D32F8DD06F1F" box="[151,444,544,567]" gridcol="0" gridrow="5" pageId="3" pageNumber="64">Dorsal-spine length</th>
<td id="6D235D92FF8E0016FD93D32F8EE06F1F" box="[605,652,544,567]" gridcol="1" gridrow="5" pageId="3" pageNumber="64">12.5</td>
<td id="6D235D92FF8E0016FCBBD32F8F8C6F1F" box="[885,992,544,567]" gridcol="2" gridrow="5" pageId="3" pageNumber="64">12.5–14.8</td>
<td id="6D235D92FF8E0016FB43D32F88976F1F" box="[1165,1275,544,567]" gridcol="3" gridrow="5" pageId="3" pageNumber="64">13.5±0.83</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D34588976F49" box="[151,1275,586,609]" gridrow="6" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D3458DD06F49" box="[151,444,586,609]" gridcol="0" gridrow="6" pageId="3" pageNumber="64">Length of anal-fin base</th>
<td id="6D235D92FF8E0016FD93D3458EE06F49" box="[605,652,586,609]" gridcol="1" gridrow="6" pageId="3" pageNumber="64">15.5</td>
<td id="6D235D92FF8E0016FCBBD3458F8C6F49" box="[885,992,586,609]" gridcol="2" gridrow="6" pageId="3" pageNumber="64">12.2–16.8</td>
<td id="6D235D92FF8E0016FB43D34588976F49" box="[1165,1275,586,609]" gridcol="3" gridrow="6" pageId="3" pageNumber="64">14.9±1.81</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D37A88976FA4" box="[151,1275,629,652]" gridrow="7" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D37A8DD06FA4" box="[151,444,629,652]" gridcol="0" gridrow="7" pageId="3" pageNumber="64">Pelvic-fin length</th>
<td id="6D235D92FF8E0016FD93D37A8EE06FA4" box="[605,652,629,652]" gridcol="1" gridrow="7" pageId="3" pageNumber="64">16.2</td>
<td id="6D235D92FF8E0016FCBBD37A8F8C6FA4" box="[885,992,629,652]" gridcol="2" gridrow="7" pageId="3" pageNumber="64">16.0–16.9</td>
<td id="6D235D92FF8E0016FB43D37A88976FA4" box="[1165,1275,629,652]" gridcol="3" gridrow="7" pageId="3" pageNumber="64">16.5±0.35</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D3AF88976F9F" box="[151,1275,672,695]" gridrow="8" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D3AF8DD06F9F" box="[151,444,672,695]" gridcol="0" gridrow="8" pageId="3" pageNumber="64">Pectoral-fin length</th>
<td id="6D235D92FF8E0016FD93D3AF8EE06F9F" box="[605,652,672,695]" gridcol="1" gridrow="8" pageId="3" pageNumber="64">20.3</td>
<td id="6D235D92FF8E0016FCBBD3AF8F8C6F9F" box="[885,992,672,695]" gridcol="2" gridrow="8" pageId="3" pageNumber="64">20.3–23.0</td>
<td id="6D235D92FF8E0016FB43D3AF88976F9F" box="[1165,1275,672,695]" gridcol="3" gridrow="8" pageId="3" pageNumber="64">22.2±1.01</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D3C588976FC9" box="[151,1275,714,737]" gridrow="9" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D3C58DD06FC9" box="[151,444,714,737]" gridcol="0" gridrow="9" pageId="3" pageNumber="64">Pectoral-spine length</th>
<td id="6D235D92FF8E0016FD93D3C58EE06FC9" box="[605,652,714,737]" gridcol="1" gridrow="9" pageId="3" pageNumber="64">14.3</td>
<td id="6D235D92FF8E0016FCBBD3C58F8C6FC9" box="[885,992,714,737]" gridcol="2" gridrow="9" pageId="3" pageNumber="64">14.3–16.6</td>
<td id="6D235D92FF8E0016FB43D3C588976FC9" box="[1165,1275,714,737]" gridcol="3" gridrow="9" pageId="3" pageNumber="64">15.3±0.96</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D3FA88976E24" box="[151,1275,757,780]" gridrow="10" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D3FA8DD06E24" box="[151,444,757,780]" gridcol="0" gridrow="10" pageId="3" pageNumber="64">Caudal-fin length</th>
<td id="6D235D92FF8E0016FD93D3FA8EE06E24" box="[605,652,757,780]" gridcol="1" gridrow="10" pageId="3" pageNumber="64">25.5</td>
<td id="6D235D92FF8E0016FCBBD3FA8F8C6E24" box="[885,992,757,780]" gridcol="2" gridrow="10" pageId="3" pageNumber="64">25.4–29.8</td>
<td id="6D235D92FF8E0016FB43D3FA88976E24" box="[1165,1275,757,780]" gridcol="3" gridrow="10" pageId="3" pageNumber="64">27.3±1.70</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D22F88976E1F" box="[151,1275,800,823]" gridrow="11" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D22F8DD06E1F" box="[151,444,800,823]" gridcol="0" gridrow="11" pageId="3" pageNumber="64">Length of adipose-fin base</th>
<td id="6D235D92FF8E0016FD93D22F8EE06E1F" box="[605,652,800,823]" gridcol="1" gridrow="11" pageId="3" pageNumber="64">13.8</td>
<td id="6D235D92FF8E0016FCBBD22F8F8C6E1F" box="[885,992,800,823]" gridcol="2" gridrow="11" pageId="3" pageNumber="64">12.0–14.5</td>
<td id="6D235D92FF8E0016FB43D22F88976E1F" box="[1165,1275,800,823]" gridcol="3" gridrow="11" pageId="3" pageNumber="64">13.5±0.89</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D24588976E49" box="[151,1275,842,865]" gridrow="12" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D2458DD06E49" box="[151,444,842,865]" gridcol="0" gridrow="12" pageId="3" pageNumber="64">Dorsal to adipose distance</th>
<td id="6D235D92FF8E0016FD93D2458EE06E49" box="[605,652,842,865]" gridcol="1" gridrow="12" pageId="3" pageNumber="64">23.1</td>
<td id="6D235D92FF8E0016FCBBD2458F8C6E49" box="[885,992,842,865]" gridcol="2" gridrow="12" pageId="3" pageNumber="64">21.5–25.6</td>
<td id="6D235D92FF8E0016FB43D24588976E49" box="[1165,1275,842,865]" gridcol="3" gridrow="12" pageId="3" pageNumber="64">23.3±1.38</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D27A88976EA4" box="[151,1275,885,908]" gridrow="13" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D27A8DD06EA4" box="[151,444,885,908]" gridcol="0" gridrow="13" pageId="3" pageNumber="64">Post-adipose distance</th>
<td id="6D235D92FF8E0016FD93D27A8EE06EA4" box="[605,652,885,908]" gridcol="1" gridrow="13" pageId="3" pageNumber="64">20.4</td>
<td id="6D235D92FF8E0016FCBBD27A8F8C6EA4" box="[885,992,885,908]" gridcol="2" gridrow="13" pageId="3" pageNumber="64">19.5–21.7</td>
<td id="6D235D92FF8E0016FB43D27A88976EA4" box="[1165,1275,885,908]" gridcol="3" gridrow="13" pageId="3" pageNumber="64">20.6±0.92</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D2AF88976E9F" box="[151,1275,928,951]" gridrow="14" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D2AF8DD06E9F" box="[151,444,928,951]" gridcol="0" gridrow="14" pageId="3" pageNumber="64">Length of caudal peduncle</th>
<td id="6D235D92FF8E0016FD93D2AF8EE06E9F" box="[605,652,928,951]" gridcol="1" gridrow="14" pageId="3" pageNumber="64">21.2</td>
<td id="6D235D92FF8E0016FCBBD2AF8F8C6E9F" box="[885,992,928,951]" gridcol="2" gridrow="14" pageId="3" pageNumber="64">20.1–22.9</td>
<td id="6D235D92FF8E0016FB43D2AF88976E9F" box="[1165,1275,928,951]" gridcol="3" gridrow="14" pageId="3" pageNumber="64">21.1±1.02</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D2C588976EC9" box="[151,1275,970,993]" gridrow="15" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D2C58DD06EC9" box="[151,444,970,993]" gridcol="0" gridrow="15" pageId="3" pageNumber="64">Depth of caudal peduncle</th>
<td id="6D235D92FF8E0016FD93D2C58EE06EC9" box="[605,652,970,993]" gridcol="1" gridrow="15" pageId="3" pageNumber="64">8.9</td>
<td id="6D235D92FF8E0016FCBBD2C58F8C6EC9" box="[885,992,970,993]" gridcol="2" gridrow="15" pageId="3" pageNumber="64">7.6–8.9</td>
<td id="6D235D92FF8E0016FB43D2C588976EC9" box="[1165,1275,970,993]" gridcol="3" gridrow="15" pageId="3" pageNumber="64">8.2±0.47</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D2FA88976924" box="[151,1275,1013,1036]" gridrow="16" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D2FA8DD06924" box="[151,444,1013,1036]" gridcol="0" gridrow="16" pageId="3" pageNumber="64">Body depth at anus</th>
<td id="6D235D92FF8E0016FD93D2FA8EE06924" box="[605,652,1013,1036]" gridcol="1" gridrow="16" pageId="3" pageNumber="64">15.3</td>
<td id="6D235D92FF8E0016FCBBD2FA8F8C6924" box="[885,992,1013,1036]" gridcol="2" gridrow="16" pageId="3" pageNumber="64">12.7–15.3</td>
<td id="6D235D92FF8E0016FB43D2FA88976924" box="[1165,1275,1013,1036]" gridcol="3" gridrow="16" pageId="3" pageNumber="64">14.4±1.04</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D52F8897691F" box="[151,1275,1056,1079]" gridrow="17" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D52F8DD0691F" box="[151,444,1056,1079]" gridcol="0" gridrow="17" pageId="3" pageNumber="64">Head length</th>
<td id="6D235D92FF8E0016FD93D52F8EE0691F" box="[605,652,1056,1079]" gridcol="1" gridrow="17" pageId="3" pageNumber="64">24.0</td>
<td id="6D235D92FF8E0016FCBBD52F8F8C691F" box="[885,992,1056,1079]" gridcol="2" gridrow="17" pageId="3" pageNumber="64">24.0–25.5</td>
<td id="6D235D92FF8E0016FB43D52F8897691F" box="[1165,1275,1056,1079]" gridcol="3" gridrow="17" pageId="3" pageNumber="64">25.0±0.58</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D54588976949" box="[151,1275,1098,1121]" gridrow="18" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D5458DD06949" box="[151,444,1098,1121]" gridcol="0" gridrow="18" pageId="3" pageNumber="64">Head width</th>
<td id="6D235D92FF8E0016FD93D5458EE06949" box="[605,652,1098,1121]" gridcol="1" gridrow="18" pageId="3" pageNumber="64">17.1</td>
<td id="6D235D92FF8E0016FCBBD5458F8C6949" box="[885,992,1098,1121]" gridcol="2" gridrow="18" pageId="3" pageNumber="64">16.1–17.9</td>
<td id="6D235D92FF8E0016FB43D54588976949" box="[1165,1275,1098,1121]" gridcol="3" gridrow="18" pageId="3" pageNumber="64">17.2±0.66</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D57A889769A4" box="[151,1275,1141,1164]" gridrow="19" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D57A8DD069A4" box="[151,444,1141,1164]" gridcol="0" gridrow="19" pageId="3" pageNumber="64">Head depth</th>
<td id="6D235D92FF8E0016FD93D57A8EE069A4" box="[605,652,1141,1164]" gridcol="1" gridrow="19" pageId="3" pageNumber="64">14.7</td>
<td id="6D235D92FF8E0016FCBBD57A8F8C69A4" box="[885,992,1141,1164]" gridcol="2" gridrow="19" pageId="3" pageNumber="64">11.8–14.7</td>
<td id="6D235D92FF8E0016FB43D57A889769A4" box="[1165,1275,1141,1164]" gridcol="3" gridrow="19" pageId="3" pageNumber="64">13.0±1.07</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D5AF8897699F" box="[151,1275,1184,1207]" gridrow="20" pageId="3" pageNumber="64" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="6D235D92FF8E0016FF59D5AF8DD0699F" box="[151,444,1184,1207]" gridcol="0" gridrow="20" pageId="3" pageNumber="64">%HL</th>
</tr>
<tr id="2EF234EEFF8E0016FF59D5C5889769C9" box="[151,1275,1226,1249]" gridrow="21" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D5C58DD069C9" box="[151,444,1226,1249]" gridcol="0" gridrow="21" pageId="3" pageNumber="64">Snout length</th>
<td id="6D235D92FF8E0016FD93D5C58EE069C9" box="[605,652,1226,1249]" gridcol="1" gridrow="21" pageId="3" pageNumber="64">50.4</td>
<td id="6D235D92FF8E0016FCBBD5C58F8C69C9" box="[885,992,1226,1249]" gridcol="2" gridrow="21" pageId="3" pageNumber="64">43.6–50.4</td>
<td id="6D235D92FF8E0016FB43D5C5889769C9" box="[1165,1275,1226,1249]" gridcol="3" gridrow="21" pageId="3" pageNumber="64">47.0±2.75</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D5FA88976824" box="[151,1275,1269,1292]" gridrow="22" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D5FA8DD06824" box="[151,444,1269,1292]" gridcol="0" gridrow="22" pageId="3" pageNumber="64">Interorbital distance</th>
<td id="6D235D92FF8E0016FD93D5FA8EE06824" box="[605,652,1269,1292]" gridcol="1" gridrow="22" pageId="3" pageNumber="64">30.4</td>
<td id="6D235D92FF8E0016FCBBD5FA8F8C6824" box="[885,992,1269,1292]" gridcol="2" gridrow="22" pageId="3" pageNumber="64">29.0–31.2</td>
<td id="6D235D92FF8E0016FB43D5FA88976824" box="[1165,1275,1269,1292]" gridcol="3" gridrow="22" pageId="3" pageNumber="64">30.1±0.99</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D42F8897681F" box="[151,1275,1312,1335]" gridrow="23" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D42F8DD0681F" box="[151,444,1312,1335]" gridcol="0" gridrow="23" pageId="3" pageNumber="64">Eye diameter</th>
<td id="6D235D92FF8E0016FD93D42F8EE0681F" box="[605,652,1312,1335]" gridcol="1" gridrow="23" pageId="3" pageNumber="64">7.5</td>
<td id="6D235D92FF8E0016FCBBD42F8F8C681F" box="[885,992,1312,1335]" gridcol="2" gridrow="23" pageId="3" pageNumber="64">7.5–9.6</td>
<td id="6D235D92FF8E0016FB43D42F8897681F" box="[1165,1275,1312,1335]" gridcol="3" gridrow="23" pageId="3" pageNumber="64">8.6±0.82</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D44588976849" box="[151,1275,1354,1377]" gridrow="24" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D4458DD06849" box="[151,444,1354,1377]" gridcol="0" gridrow="24" pageId="3" pageNumber="64">Nasal barbel length</th>
<td id="6D235D92FF8E0016FD93D4458EE06849" box="[605,652,1354,1377]" gridcol="1" gridrow="24" pageId="3" pageNumber="64">22.5</td>
<td id="6D235D92FF8E0016FCBBD4458F8C6849" box="[885,992,1354,1377]" gridcol="2" gridrow="24" pageId="3" pageNumber="64">8.0–25.2</td>
<td id="6D235D92FF8E0016FB43D44588976849" box="[1165,1275,1354,1377]" gridcol="3" gridrow="24" pageId="3" pageNumber="64">16.4±6.87</td>
</tr>
<tr id="2EF234EEFF8E0016FF59D47A889768A4" box="[151,1275,1397,1420]" gridrow="25" pageId="3" pageNumber="64">
<th id="6D235D92FF8E0016FF59D47A8DD068A4" box="[151,444,1397,1420]" gridcol="0" gridrow="25" pageId="3" pageNumber="64">Maxillary barbel length</th>
<td id="6D235D92FF8E0016FD93D47A8EE068A4" box="[605,652,1397,1420]" gridcol="1" gridrow="25" pageId="3" pageNumber="64">95.0</td>
<td id="6D235D92FF8E0016FCBBD47A8F8C68A4" box="[885,992,1397,1420]" gridcol="2" gridrow="25" pageId="3" pageNumber="64">74.6–98.8</td>
<td id="6D235D92FF8E0016FB43D47A889768A4" box="[1165,1275,1397,1420]" gridcol="3" gridrow="25" pageId="3" pageNumber="64">87.8±9.57</td>
</tr>
</table>
</paragraph>
<paragraph id="907D36ACFF8EFFEAFF51D4AF889568C9" blockId="3.[159,1274,1440,1505]" pageId="3" pageNumber="64">Inner mandibular barbel length 30.8 27.0–32.8 31.2±2.21 Outer mandibular barbel length 50.8 32.6–52.1 47.1±7.33</paragraph>
<paragraph id="907D36ACFF8EFFEAFF0BD72388F66A53" blockId="3.[151,1436,1580,2031]" pageId="3" pageNumber="64">Dorsal fin located above anterior third of body, with I,6 (6) rays; fin margin convex; spine short and straight, smooth on anterior and posterior margins. Adipose fin with anterior margin straight or very slightly concave and posterior margin angular. Caudal fin strongly forked, with lower lobe very slightly longer than upper lobe and i,7,8,i (6) principal rays. Procurrent rays symmetrical and extending only slightly anterior to fin base. Adipose-fin origin at vertical through base of first anal-fin ray. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with iv,8 (1) or iv,9,i* (5) rays. Pelvic-fin origin at vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (6) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,8,i (2) or I,9* (4) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 9–12* serrations.</paragraph>
<paragraph id="907D36ACFF8EFFEAFF0BD68789356AC7" blockId="3.[151,1436,1580,2031]" pageId="3" pageNumber="64">Thoracic adhesive apparatus present, consisting of ridges of skin in an elongate oval field extending from isthmus to just anterior to level of base of last pectoral-fin ray. Median depression present on posterior third of adhesive apparatus. Ridges uninterrupted and oriented anterodistally, radiating from median depression.</paragraph>
<caption id="C4BD6624FF89FFEDFF59D437893A685B" httpUri="https://zenodo.org/record/187390/files/figure.png" pageId="4" pageNumber="65" targetBox="[273,1312,199,1307]" targetPageId="4">
<paragraph id="907D36ACFF89FFEDFF59D437893A685B" blockId="4.[151,1434,1336,1395]" pageId="4" pageNumber="65">
<emphasis id="A2B6EABEFF89FFEDFF59D4378D556878" bold="true" box="[151,313,1336,1360]" pageId="4" pageNumber="65">FIGURE 2. T</emphasis>
horacic adhesive apparatus of: a.
<taxonomicName id="57C24D2FFF89FFEDFD78D4368FDD6878" box="[694,945,1337,1360]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="ketambe">
<emphasis id="A2B6EABEFF89FFEDFD78D4368FDD6878" box="[694,945,1337,1360]" italics="true" pageId="4" pageNumber="65">Glyptothorax ketambe</emphasis>
</taxonomicName>
, holotype, MZB 8694, 100.0 mm SL; b.
<taxonomicName id="57C24D2FFF89FFEDFA4BD4368C87685B" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="plectilis">
<emphasis id="A2B6EABEFF89FFEDFA4BD4368C87685B" italics="true" pageId="4" pageNumber="65">G. plectilis</emphasis>
</taxonomicName>
, paratype, ZRC 46420, 44.3 mm SL; c.
<taxonomicName id="57C24D2FFF89FFEDFD56D4538F70685B" box="[664,796,1372,1395]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="A2B6EABEFF89FFEDFD56D4538F70685B" box="[664,796,1372,1395]" italics="true" pageId="4" pageNumber="65">G. siamensis</emphasis>
</taxonomicName>
, CMK 10829, 71.8 mm SL. Illustrations not to scale.
</paragraph>
</caption>
<paragraph id="907D36ACFF89FFEDFF0BD4AF8E9D6BE2" blockId="4.[151,1437,1440,1931]" pageId="4" pageNumber="65">
<emphasis id="A2B6EABEFF89FFEDFF0BD4AF8D3F6892" bold="true" box="[197,339,1440,1466]" pageId="4" pageNumber="65">Coloration.</emphasis>
In 70% ethanol: Dorsal and lateral surfaces of head, and body dark brown, fading to beige on ventral surfaces. A faint thin, lighter brown mid-dorsal stripe extending from base of last dorsal-fin ray to origin of adipose fin; stripe very faint in some individuals. Laterosensory pores along lateral line rimmed in beige, imparting appearance of a diffuse lighter brown midlateral line. All fins with brown fin rays, and diffuse melanophores on fin membranes. Pectoral and pelvic fins with brown on bases of fin rays and hyaline posterior margin. Anal fin with brown base and hyaline posteroventral corner. Adipose fin brown, with hyaline distal margin. Caudal fin brown, with tips of lobes hyaline. Maxillary and nasal barbels brown dorsally, beige ventrally. Mandibular barbels beige.
</paragraph>
</subSubSection>
<subSubSection id="D8D86527FF89FFEDFF0BD7D98DF76A15" pageId="4" pageNumber="65" type="distribution">
<paragraph id="907D36ACFF89FFEDFF0BD7D98DF76A15" blockId="4.[151,1437,1440,1931]" pageId="4" pageNumber="65">
<emphasis id="A2B6EABEFF89FFEDFF0BD7D98D096BD8" bold="true" box="[197,357,1750,1776]" pageId="4" pageNumber="65">Distribution.</emphasis>
Known from the
<collectingCountry id="E8D5763CFF89FFEDFDF9D7D98E006BD8" box="[567,620,1750,1776]" name="Kazakhstan" pageId="4" pageNumber="65">Alas</collectingCountry>
River drainage in northern Sumatra, which drains the eastern face of the Leuser Mountain Range and the western face of the Central Gajo Mountain Range, debouching into the Indian Ocean (
<figureCitation id="08F92A29FF89FFEDFE8AD62C8DE76A15" box="[324,395,1827,1853]" captionStart="FIGURE 3" captionStartId="5.[151,255,777,801]" captionTargetBox="[151,1436,198,747]" captionTargetId="figure@5.[151,1436,197,752]" captionTargetPageId="5" captionText="FIGURE 3. Collection localities of G. ketambe." httpUri="https://zenodo.org/record/187391/files/figure.png" pageId="4" pageNumber="65">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D8D86527FF89FFEDFF0BD645890E6AA3" pageId="4" pageNumber="65" type="etymology">
<paragraph id="907D36ACFF89FFEDFF0BD645890E6AA3" blockId="4.[151,1437,1440,1931]" pageId="4" pageNumber="65">
<emphasis id="A2B6EABEFF89FFEDFF0BD6458D396A4C" bold="true" box="[197,341,1866,1892]" pageId="4" pageNumber="65">Etymology.</emphasis>
This species is named after its
<typeStatus id="4F79880EFF89FFEDFD1AD6458F646A4C" box="[724,776,1866,1892]" pageId="4" pageNumber="65">type</typeStatus>
locality, the
<taxonomicName id="57C24D2FFF89FFEDFC68D645887E6A4C" box="[934,1042,1866,1892]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="66" phylum="Chordata" rank="species" species="ketambe">Ketambe</taxonomicName>
research station, which is world renowned as a site for the study of the Sumatran orangutan (
<taxonomicName id="57C24D2FFF89FFEDFC94D67E8F9D6AA2" box="[858,1009,1905,1930]" class="Mammalia" family="Hominidae" genus="Pongo" kingdom="Animalia" order="Primates" pageId="4" pageNumber="65" phylum="Chordata" rank="species" species="abelii">
<emphasis id="A2B6EABEFF89FFEDFC94D67E8F9D6AA2" box="[858,1009,1905,1930]" italics="true" pageId="4" pageNumber="65">Pongo abelii</emphasis>
</taxonomicName>
). Used as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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