174 lines
12 KiB
XML
174 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.773.24878" ID-GBIF-Dataset="3b67ab3e-bd75-4194-a90c-c12b3dc8332b" ID-Pensoft-Pub="1313-2970-773-1" ID-ZBK="900F9307844A4B9AB428DA97FF3B4B5C" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-773-1" ModsDocOrigin="ZooKeys 773" ModsDocTitle="A molecular and conchological dissection of the “scaly” Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1531209184653" checkinUser="pensoft" docAuthor="Khalik, Mohd Zacaery, Hendriks, Kasper, Vermeulen, Jaap J. & Schilthuizen, Menno" docDate="2018" docId="79A2CF435F54680FFC2B699CC1416DE5" docLanguage="en" docName="ZooKeys 773: 1-55" docOrigin="ZooKeys 773" docSource="http://dx.doi.org/10.3897/zookeys.773.24878" docTitle="Georissa muluensis Khalik, Hendriks, Vermeulen & Schilthuizen, 2018, sp. n." docType="treatment" docUuid="8CAE7706-39F4-47ED-9275-7606DDD5FC26" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFACFFA96149FF8D3477FFCCD227FF8E" masterDocTitle="A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="1" updateTime="1643514872515" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khalik, Mohd Zacaery</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendriks, Kasper</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>773</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.773.24878</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.773.24878</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-773-1</mods:identifier>
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<mods:identifier type="ZBK">900F9307844A4B9AB428DA97FF3B4B5C</mods:identifier>
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<mods:identifier type="ZooBank">900F9307844A4B9AB428DA97FF3B4B5C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="146053687" LSID="urn:lsid:zoobank.org:act:8CAE7706-39F4-47ED-9275-7606DDD5FC26" httpUri="http://treatment.plazi.org/id/79A2CF435F54680FFC2B699CC1416DE5" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/8CAE7706-39F4-47ED-9275-7606DDD5FC26" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa muluensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muluensis">Georissa muluensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Lagang Cave, Mulu National Park, Mulu, Sarawak, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="4.051">04°03.06'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="114.82283">114°49.37'E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype. Lagang Cave, Mulu National Park, Mulu, Sarawak, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="4.051">04°03.06'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="114.82283">114°49.37'E</geoCoordinate>
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): MZU/MOL 17.86 (leg. MZ Khalik and SK Reduan). Paratypes. Lagang Cave, Mulu National Park, Mulu, Sarawak (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="4.051">04°03.06'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="114.82283">114°49.37'E</geoCoordinate>
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): MZU/MOL 17.30 (13), MZU/MOL 17.31 (9).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Deer Cave, Mulu National Park, Mulu, Sarawak: JJV 10533 (this sample, approximately 120 individuals, also contains specimens of
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<taxonomicName genus="G." lsidName="G. hadra" pageId="0" pageNumber="1" rank="species" species="hadra">G. hadra</taxonomicName>
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), JJV 10554 (this sample contains 5 individual of
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<taxonomicName genus="G." lsidName="G. muluensis" pageId="0" pageNumber="1" rank="species" species="muluensis">G. muluensis</taxonomicName>
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, 1 individual
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<taxonomicName genus="G." lsidName="G. hadra" pageId="0" pageNumber="1" rank="species" species="hadra">G. hadra</taxonomicName>
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), JJV 10533 (this sample, approximately 150 individuals, also contains specimens of
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<taxonomicName genus="G." lsidName="G. hadra" pageId="0" pageNumber="1" rank="species" species="hadra">G. hadra</taxonomicName>
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and
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<taxonomicName genus="G." lsidName="G. kobelti" pageId="0" pageNumber="1" rank="species" species="kobelti">G. kobelti</taxonomicName>
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). Mulu N.P., Mulu, Sarawak: JJV 10527.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Named after Mulu National Park, Sarawak, Malaysia, the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Protoconch. Color: yellow. Sculpture pattern: meshed - ellipsoidal mesh shape. Mesh width: 16-26
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. Teleoconch. Color: yellow. First whorl: shouldered, above the shoulder flat, nearly horizontal; below the shoulder flat, cylindrical, but abruptly withdrawn into the deeply incised suture. Subsequent whorls: convex to rounded. SH: 1.67-2.05 mm, SW: 1.57-1.79 mm, SI: 1.08-1.18. Total number of whorls: 2
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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-3. Shell sculpture. Radial sculpture: absent, only weak to strong growth lines are visible. Spiral sculpture: present, consisting of thin, but strongly sculpted and continuous ribs. Scales: two to three series of tall and diagonal scales, regularly spaced, the upper scale series always stronger than the lower ones, weaker series appear later at the spire or consist only of randomly spaced arrays of acute nodules, widely spaced between the first and second scale series, more densely spaced later. Aperture. Shape: rounded to slightly oval. Basal side: rounded, angular at the columellar region. Parietal side: straight to slightly curved. AH: 0.82-0.98 mm, AW: 1.03-1.18 mm, AI: 0.77-0.83. Holotype dimension. SH: 1.67 mm, SW: 1.53 mm, AH: 0.82 mm, AW: 1.07 mm.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 9.
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa muluensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="muluensis">Georissa muluensis</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Holotype: MZU/MOL 17.86
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<normalizedToken originalValue="D–K">D-K</normalizedToken>
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Paratypes: MZU/MOL 17.30. A, D Shell apertural view B Shell side view C Shell rear view
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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Shell cross-section from 3D model
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<normalizedToken originalValue="G–H">G-H</normalizedToken>
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Operculum frontal and ventral view I Shell top view J Protoconch side view. K. Close up of protoconch from top at 1000
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken originalValue="A–I">A-I</normalizedToken>
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); 200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(J); 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(K).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="cross diagnosis">
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<paragraph pageId="0" pageNumber="1">Cross diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The wide spacing of the major spiral scale series of
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<taxonomicName genus="G." lsidName="G. muluensis" pageId="0" pageNumber="1" rank="species" species="muluensis">G. muluensis</taxonomicName>
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is similar to
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<taxonomicName genus="G." lsidName="G. kinabatanganensis" pageId="0" pageNumber="1" rank="species" species="kinabatanganensis">G. kinabatanganensis</taxonomicName>
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, but
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<taxonomicName genus="G." lsidName="G. muluensis" pageId="0" pageNumber="1" rank="species" species="muluensis">G. muluensis</taxonomicName>
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has a more elongated shell shape, rather than the more flattened habitus of
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<taxonomicName genus="G." lsidName="G. kinabatanganensis" pageId="0" pageNumber="1" rank="species" species="kinabatanganensis">G. kinabatanganensis</taxonomicName>
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. In general shell shape and sculpture
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<taxonomicName genus="G." lsidName="G. muluensis" pageId="0" pageNumber="1" rank="species" species="muluensis">G. muluensis</taxonomicName>
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also resembles
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<taxonomicName genus="G." lsidName="G. hadra" pageId="0" pageNumber="1" rank="species" species="hadra">G. hadra</taxonomicName>
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, which, however, is larger and more elongated.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known only from the small area of Lagang Cave, Mulu National Park, Mulu, Sarawak, Malaysia.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="molecular analysis">
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<paragraph pageId="0" pageNumber="1">Molecular analysis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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ML and Bayesian analyses show that the individuals of
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<taxonomicName genus="G." lsidName="G. muluensis" pageId="0" pageNumber="1" rank="species" species="muluensis">G. muluensis</taxonomicName>
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(16S: n = 4; CO1: n = 4) form a monophyletic group with 100% BS and 100% PP, which is the sister group of
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<taxonomicName genus="G." lsidName="G. hadra" pageId="0" pageNumber="1" rank="species" species="hadra">G. hadra</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |