237 lines
31 KiB
XML
237 lines
31 KiB
XML
<document id="84B845D4AE859E128738E35E8184941A" ID-DOI="10.5281/zenodo.203341" ID-GBIF-Dataset="5d057188-bd12-44f5-903f-74f9a88b68d3" ID-ISSN="1175-5326" ID-Zenodo-Dep="203341" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460253477643" checkinUser="plazi" docAuthor="Ruiz, Gustavo Rodrigo Sanches" docDate="2011" docId="4E4F69489A5CFFA85DD4F974A0A2FB88" docLanguage="en" docName="zt02785p060.pdf" docOrigin="Zootaxa 2785" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Macutula Ruiz, 2011, gen. nov." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="B27611309A5CFFA95D43FFFEA444FF98" masterDocTitle="Description of Macutula, a new genus of jumping spiders from Northeastern Brazil (Araneae: Salticidae: Amycoida)" masterLastPageNumber="60" masterPageNumber="53" pageNumber="53" updateTime="1698243053875" updateUser="plazi">
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<mods:titleInfo id="E77FF16B6E6404E194FC015D003946DC">
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<mods:title id="F7EE2B4E86C4EBC455D75BC52394DFD2">Description of Macutula, a new genus of jumping spiders from Northeastern Brazil (Araneae: Salticidae: Amycoida)</mods:title>
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<mods:name id="E58132DD3A3B1B7141AE0E276E782F1E" type="personal">
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<mods:roleTerm id="40270AC199D36459AB554BDDBF932A4F">Author</mods:roleTerm>
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<mods:namePart id="B4FE5B4A50698EEAB03960EE1F4AEA9A">Ruiz, Gustavo Rodrigo Sanches</mods:namePart>
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<mods:typeOfResource id="1F3A47DE40F7CF401DA643777618370D">text</mods:typeOfResource>
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<mods:title id="2DB2F268F73D61AB94254E13B523C5D9">Zootaxa</mods:title>
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<mods:part id="A34FD9E8AFFD9AE98EE40BB56907D16C">
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<mods:date id="1FB9FD44F44D4DC08A882A44E5C4363D">2011</mods:date>
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<mods:number id="C085AF17FB755197CCF61F362A961AF2">2785</mods:number>
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<mods:classification id="FDEC01F08262933C97DC7B39407CE6BA">journal article</mods:classification>
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<mods:identifier id="BE2DE8FD9BDD5F77FF1558BF6348B6DA" type="DOI">10.5281/zenodo.203341</mods:identifier>
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<mods:identifier id="756EBDC97A218D8BE896130FA75578F6" type="Zenodo-Dep">203341</mods:identifier>
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<treatment id="4E4F69489A5CFFA85DD4F974A0A2FB88" ID-DOI="http://doi.org/10.5281/zenodo.6192080" ID-GBIF-Taxon="119392105" ID-Zenodo-Dep="6192080" LSID="urn:lsid:plazi:treatment:4E4F69489A5CFFA85DD4F974A0A2FB88" httpUri="http://treatment.plazi.org/id/4E4F69489A5CFFA85DD4F974A0A2FB88" lastPageId="1" lastPageNumber="54" pageId="0" pageNumber="53">
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<subSubSection id="8EFC8BD59A5CFFA95DD4F974A5C5F93C" box="[151,385,1674,1700]" pageId="0" pageNumber="53" type="nomenclature">
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<paragraph id="C659D85E9A5CFFA95DD4F974A5C5F93C" blockId="0.[151,385,1674,1700]" box="[151,385,1674,1700]" pageId="0" pageNumber="53">
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<heading id="9D116F329A5CFFA95DD4F974A5C5F93C" bold="true" box="[151,385,1674,1700]" fontSize="11" level="1" pageId="0" pageNumber="53" reason="1">
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<emphasis id="F492044C9A5CFFA95DD4F974A5C5F93C" bold="true" box="[151,385,1674,1700]" pageId="0" pageNumber="53">
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<taxonomicName id="01E6A3DD9A5CFFA95DD4F974A549F93C" ID-CoL="62XVY" box="[151,269,1674,1700]" class="Arachnida" family="Salticidae" genus="Macutula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="F492044C9A5CFFA95DD4F974A549F93C" bold="true" box="[151,269,1674,1700]" italics="true" pageId="0" pageNumber="53">Macutula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EFA1B9379A5CFFA95C50F974A5C5F93C" box="[275,385,1674,1700]" pageId="0" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8EFC8BD59A5CFFA95DD4F92CA615F97F" box="[151,593,1745,1768]" pageId="0" pageNumber="53" type="materials_examined">
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<paragraph id="C659D85E9A5CFFA95DD4F92CA615F97F" blockId="0.[151,593,1745,1768]" box="[151,593,1745,1768]" pageId="0" pageNumber="53">
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<emphasis id="F492044C9A5CFFA95DD4F92CA561F970" bold="true" box="[151,293,1745,1768]" pageId="0" pageNumber="53">
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<typeStatus id="195D66FC9A5CFFA95DD4F92CA488F97F" box="[151,204,1746,1767]" pageId="0" pageNumber="53">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="01E6A3DD9A5CFFA95C68F92FA5BFF97F" box="[299,507,1745,1767]" class="Arachnida" family="Salticidae" genus="Macutula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="aracoiaba" status="sp. nov.">
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<emphasis id="F492044C9A5CFFA95C68F92FA5BFF97F" box="[299,507,1745,1767]" italics="true" pageId="0" pageNumber="53">Macutula aracoiaba</emphasis>
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</taxonomicName>
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<emphasis id="F492044C9A5CFFA95F42F92CA615F97F" bold="true" box="[513,593,1746,1767]" pageId="0" pageNumber="53">
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<taxonomicNameLabel id="EFA1B9379A5CFFA95F42F92CA615F97F" box="[513,593,1746,1767]" pageId="0" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="8EFC8BD59A5CFFA85DD4F8EAA611FEFC" lastPageId="1" lastPageNumber="54" pageId="0" pageNumber="53" type="etymology">
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<paragraph id="C659D85E9A5CFFA95DD4F8EAA011F8B4" blockId="0.[151,1436,1812,2017]" box="[151,1109,1812,1836]" pageId="0" pageNumber="53">
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<emphasis id="F492044C9A5CFFA95DD4F8EAA55AF8B4" bold="true" box="[151,286,1812,1836]" pageId="0" pageNumber="53">Etymology.</emphasis>
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The name is an arbitrary combination of letters to be treated as feminine.
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</paragraph>
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<paragraph id="C659D85E9A5CFFA85D86F8C9A611FEFC" blockId="0.[151,1436,1812,2017]" lastBlockId="1.[151,1436,151,1040]" lastPageId="1" lastPageNumber="54" pageId="0" pageNumber="53">
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<emphasis id="F492044C9A5CFFA95D86F8C9A5BFF8C8" bold="true" box="[197,507,1847,1872]" pageId="0" pageNumber="53">Phylogenetic relationship.</emphasis>
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Although DNA data consistently support the monophyly of amycoids (e.g.,
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<bibRefCitation id="A277A5AF9A5CFFA9581CF8C9A5C6F8EC" author="Maddison" pageId="0" pageNumber="53" refString="Maddison, W. P. & Hedin, M. C. (2003) Jumping spider phylogeny (Araneae: Salticidae). Invertebrate Systematics, 17, 529 - 549." type="journal article" year="2003">Maddison & Hedin 2003</bibRefCitation>
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;
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<bibRefCitation id="A277A5AF9A5CFFA95CCCF8A2A6C2F8EC" author="Maddison" box="[399,646,1884,1908]" pageId="0" pageNumber="53" refString="Maddison, W. P., Bodner, M. & Needham, K. (2008) Salticid spider phylogeny revisited, with the discovery of a large Australasian clade (Araneae: Salticidae). Zootaxa, 1893, 49 - 64." type="journal article" year="2008">
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Maddison
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<emphasis id="F492044C9A5CFFA95F4BF8A3A67FF8EC" box="[520,571,1884,1908]" italics="true" pageId="0" pageNumber="53">et al</emphasis>
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. 2008
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</bibRefCitation>
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), morphological synapomorphies remain unknown. The new genus is supposed to belong to Amycoida by similarities with several amycoid lineages. In general appearance, spiders of this group resemble those of
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<taxonomicName id="01E6A3DD9A5CFFA95CA3F85AA770F824" authority="Peckham and Peckham" authorityName="Peckham and Peckham" box="[480,820,1956,1980]" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="53" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5CFFA95CA3F85AA667F824" box="[480,547,1956,1980]" italics="true" pageId="0" pageNumber="53">Breda</emphasis>
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Peckham and Peckham
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</taxonomicName>
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, which could be an amycoid according to unpublished DNA data, by the dark, glabrous carapace. The third leg longest is also common in amycines, such as
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<taxonomicName id="01E6A3DD9A5CFFA95858F837A118F878" box="[1307,1372,1993,2016]" class="Arachnida" family="Salticidae" genus="Mago" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="53" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5CFFA95858F837A118F878" box="[1307,1372,1993,2016]" italics="true" pageId="0" pageNumber="53">Mago</emphasis>
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</taxonomicName>
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O.P.- Cambridge and
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<taxonomicName id="01E6A3DD9A5DFFA85C0FFF67A630FF37" authority="Simon. Cheliceral" authorityName="Simon. Cheliceral" box="[332,628,151,176]" class="Arachnida" family="Salticidae" genus="Noegus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA85C0FFF67A5E4FF28" box="[332,416,153,176]" italics="true" pageId="1" pageNumber="54">Noegus</emphasis>
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Simon. Cheliceral
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</taxonomicName>
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teeth are similar to the ones present in
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<taxonomicName id="01E6A3DD9A5DFFA8597BFF66A038FF28" box="[1080,1148,152,176]" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA8597BFF66A038FF28" box="[1080,1148,152,176]" italics="true" pageId="1" pageNumber="54">Breda</emphasis>
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</taxonomicName>
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and in the
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<taxonomicName id="01E6A3DD9A5DFFA859BEFF67A10FFF28" box="[1277,1355,153,176]" class="Arachnida" family="Salticidae" genus="Hurius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA859BEFF67A10FFF28" box="[1277,1355,153,176]" italics="true" pageId="1" pageNumber="54">Hurius</emphasis>
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</taxonomicName>
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group, with several teeth on promargin and one on retromargin (see
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<bibRefCitation id="A277A5AF9A5DFFA85E13FF42A054FF4C" author="Chickering" box="[848,1040,188,212]" pageId="1" pageNumber="54" refString="Chickering, A. M. (1946) The Salticidae of Panama. Bulletin of the Museum of Comparative Zoology, 97, 37 - 39." type="journal article" year="1946">Chickering 1946</bibRefCitation>
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, fig. 16;
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<bibRefCitation id="A277A5AF9A5DFFA85934FF42A157FF4C" author="Galiano" box="[1143,1299,188,212]" pageId="1" pageNumber="54" refString="Galiano, M. E. (1985) Revision del genero Hurius Simon, 1901 (Araneae, Salticidae). Journal of Arachnology, 13, 9 - 18." type="journal article" year="1985">Galiano 1985</bibRefCitation>
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, figs 4, 18). The reduced sperm loop immediately before the embolus is similar to that present in species of
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<taxonomicName id="01E6A3DD9A5DFFA85983FF1EA1DFFF6F" authority="Simon. The" authorityName="Simon. The" box="[1216,1435,223,247]" class="Arachnida" family="Salticidae" genus="Sitticus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA85983FF1EA156FF6F" box="[1216,1298,224,247]" italics="true" pageId="1" pageNumber="54">Sitticus</emphasis>
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Simon. The
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</taxonomicName>
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epigynal structure is also similar to that of many amycoids: the anterior atrium is found in
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<taxonomicName id="01E6A3DD9A5DFFA859E3FEFAA0A7FE84" box="[1184,1251,260,284]" class="Arachnida" family="Salticidae" genus="Breda" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA859E3FEFAA0A7FE84" box="[1184,1251,260,284]" italics="true" pageId="1" pageNumber="54">Breda</emphasis>
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</taxonomicName>
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and many amycines, such as
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<taxonomicName id="01E6A3DD9A5DFFA85C74FED7A53CFED8" box="[311,376,297,320]" class="Arachnida" family="Salticidae" genus="Mago" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA85C74FED7A53CFED8" box="[311,376,297,320]" italics="true" pageId="1" pageNumber="54">Mago</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="01E6A3DD9A5DFFA85CECFED7A647FED8" box="[431,515,297,320]" class="Arachnida" family="Salticidae" genus="Noegus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="F492044C9A5DFFA85CECFED7A647FED8" box="[431,515,297,320]" italics="true" pageId="1" pageNumber="54">Noegus</emphasis>
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</taxonomicName>
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. Moreover, copulation ducts coiled posteriorly and anterior spermathecae are also found in almost all groups of amycoids.
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</paragraph>
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</subSubSection>
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<subSubSection id="8EFC8BD59A5DFFA85D86FE91A140FE34" pageId="1" pageNumber="54" type="diagnosis">
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<paragraph id="C659D85E9A5DFFA85D86FE91A140FE34" blockId="1.[151,1436,151,1040]" pageId="1" pageNumber="54">
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<emphasis id="F492044C9A5DFFA85D86FE91A57BFE10" bold="true" box="[197,319,367,392]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
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<taxonomicName id="01E6A3DD9A5DFFA85C04FE8EA5F6FE10" box="[327,434,368,392]" class="Arachnida" family="Salticidae" genus="Macutula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="F492044C9A5DFFA85C04FE8EA5F6FE10" box="[327,434,368,392]" italics="true" pageId="1" pageNumber="54">Macutula</emphasis>
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</taxonomicName>
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<emphasis id="F492044C9A5DFFA85CF9FE8EA661FE10" bold="true" box="[442,549,368,392]" pageId="1" pageNumber="54">
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||
<taxonomicNameLabel id="EFA1B9379A5DFFA85CF9FE8EA661FE10" box="[442,549,368,392]" pageId="1" pageNumber="54" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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differs from the remaining genera of Amycoida by having a very deep prolateral excavation on the dorsal surface of the cymbium of the male palp, forming a dorsal keel (
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<figureCitation id="5EDDC4DB9A5DFFA859F2FE6AA0B2FE34" box="[1201,1270,404,428]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Fig. 7</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="8EFC8BD59A5DFFA85D86FE49A0A2FB88" pageId="1" pageNumber="54" type="description">
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<paragraph id="C659D85E9A5DFFA85D86FE49A03CFC74" blockId="1.[151,1436,151,1040]" pageId="1" pageNumber="54">
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<emphasis id="F492044C9A5DFFA85D86FE49A513FE48" bold="true" box="[197,343,439,464]" pageId="1" pageNumber="54">Description.</emphasis>
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Median sized jumping spiders (
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<quantity id="011E75BB9A5DFFA85F81FE49A75FFE57" box="[706,795,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="54" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7mm</quantity>
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||
) with slightly flattened bodies and dark brown, glabrous carapaces (
|
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<figureCitation id="5EDDC4DB9A5DFFA85C50FE22A53CFE6C" box="[275,376,476,500]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1913,1936]" captionTargetBox="[151,1436,1111,1872]" captionTargetId="figure@1.[151,1436,1095,1873]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Macutula aracoiaba sp. nov. 1 male habitus; 2 female habitus." httpUri="https://zenodo.org/record/203342/files/figure.png" pageId="1" pageNumber="54">Figs 1–2</figureCitation>
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||
); chelicera similar in males and females, with an anterior color pattern (
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<figureCitation id="5EDDC4DB9A5DFFA859E2FE22A0B6FE6C" box="[1185,1266,476,500]" captionStart="FIGURES 11 – 14" captionStartId="3.[151,244,1255,1278]" captionTargetBox="[168,1423,208,1217]" captionTargetId="figure@3.[151,1436,193,1234]" captionTargetPageId="3" captionText="FIGURES 11 – 14. Macutula aracoiaba sp. nov. 11 male face; 12 male chelicerae, retrolateral view; 13 male labium and endites, ventral view; 14 male sternum and coxae, ventral view." httpUri="https://zenodo.org/record/203344/files/figure.png" pageId="1" pageNumber="54">Fig. 11</figureCitation>
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), one big tooth on retromargin and five on promargin, the proximal two being the largest (
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<figureCitation id="5EDDC4DB9A5DFFA85EABFE01A079FD8F" box="[1000,1085,511,535]" captionStart="FIGURES 11 – 14" captionStartId="3.[151,244,1255,1278]" captionTargetBox="[168,1423,208,1217]" captionTargetId="figure@3.[151,1436,193,1234]" captionTargetPageId="3" captionText="FIGURES 11 – 14. Macutula aracoiaba sp. nov. 11 male face; 12 male chelicerae, retrolateral view; 13 male labium and endites, ventral view; 14 male sternum and coxae, ventral view." httpUri="https://zenodo.org/record/203344/files/figure.png" pageId="1" pageNumber="54">Fig. 12</figureCitation>
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); male endite similar to that of the female (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85C5FFDDAA52BFDA4" box="[284,367,548,572]" captionStart="FIGURES 11 – 14" captionStartId="3.[151,244,1255,1278]" captionTargetBox="[168,1423,208,1217]" captionTargetId="figure@3.[151,1436,193,1234]" captionTargetPageId="3" captionText="FIGURES 11 – 14. Macutula aracoiaba sp. nov. 11 male face; 12 male chelicerae, retrolateral view; 13 male labium and endites, ventral view; 14 male sternum and coxae, ventral view." httpUri="https://zenodo.org/record/203344/files/figure.png" pageId="1" pageNumber="54">Fig. 13</figureCitation>
|
||
); labium rounded with posterior ventral excavations (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85E88FDDAA058FDA4" box="[971,1052,548,572]" captionStart="FIGURES 11 – 14" captionStartId="3.[151,244,1255,1278]" captionTargetBox="[168,1423,208,1217]" captionTargetId="figure@3.[151,1436,193,1234]" captionTargetPageId="3" captionText="FIGURES 11 – 14. Macutula aracoiaba sp. nov. 11 male face; 12 male chelicerae, retrolateral view; 13 male labium and endites, ventral view; 14 male sternum and coxae, ventral view." httpUri="https://zenodo.org/record/203344/files/figure.png" pageId="1" pageNumber="54">Fig. 13</figureCitation>
|
||
) and sternum subpentagonal (
|
||
<figureCitation id="5EDDC4DB9A5DFFA8582CFDDAA4F2FDC7" captionStart="FIGURES 11 – 14" captionStartId="3.[151,244,1255,1278]" captionTargetBox="[168,1423,208,1217]" captionTargetId="figure@3.[151,1436,193,1234]" captionTargetPageId="3" captionText="FIGURES 11 – 14. Macutula aracoiaba sp. nov. 11 male face; 12 male chelicerae, retrolateral view; 13 male labium and endites, ventral view; 14 male sternum and coxae, ventral view." httpUri="https://zenodo.org/record/203344/files/figure.png" pageId="1" pageNumber="54">Fig. 14</figureCitation>
|
||
); male palp with a straight femur (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85F79FDB9A63AFDC7" box="[570,638,583,607]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Fig. 6</figureCitation>
|
||
), short tibia with a curved RTA and a poorly developed RvTA (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85804FDB9A4E3FD1C" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Figs 3– 5</figureCitation>
|
||
,
|
||
<figureCitation id="5EDDC4DB9A5DFFA85DF0FD92A4B8FD1C" box="[179,252,620,644]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,244,1415,1438]" captionTargetBox="[170,1425,210,1384]" captionTargetId="figure@6.[151,1436,193,1394]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Macutula santana sp. nov. 23 – 24 left male palp (23 ventral, 24 retrolateral). Macutula caruaru sp. nov. 25 – 26 left male palp (25 ventral, 26 retrolateral); 27 epigyne, ventral view; 28 vulva, dorsal view." httpUri="https://zenodo.org/record/203345/files/figure.png" pageId="1" pageNumber="54">24, 26</figureCitation>
|
||
), a triangular cymbium seen in anterior view, with a deep prolateral-dorsal excavation (
|
||
<figureCitation id="5EDDC4DB9A5DFFA8598DFD92A157FD1C" box="[1230,1299,620,644]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Fig. 7</figureCitation>
|
||
), small subtegulum and tegulum with a small loop of the sperm duct just before it enters the embolus (
|
||
<figureCitation id="5EDDC4DB9A5DFFA859EEFD71A157FD3F" box="[1197,1299,655,679]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Figs 3, 8</figureCitation>
|
||
) and a long curling embolus, whose tip lies in a ventral excavation on the distal cymbium (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85946FD4AA009FD54" box="[1029,1101,692,716]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="5EDDC4DB9A5DFFA8591AFD4AA0E6FD54" box="[1113,1186,692,716]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,244,1415,1438]" captionTargetBox="[170,1425,210,1384]" captionTargetId="figure@6.[151,1436,193,1394]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Macutula santana sp. nov. 23 – 24 left male palp (23 ventral, 24 retrolateral). Macutula caruaru sp. nov. 25 – 26 left male palp (25 ventral, 26 retrolateral); 27 epigyne, ventral view; 28 vulva, dorsal view." httpUri="https://zenodo.org/record/203345/files/figure.png" pageId="1" pageNumber="54">23, 25</figureCitation>
|
||
). Leg formula
|
||
<quantity id="011E75BB9A5DFFA8580BFD4AA1D8FD54" box="[1352,1436,692,716]" metricMagnitude="1" metricUnit="m" metricValue="3.4086799999999995" pageId="1" pageNumber="54" unit="in" value="1342.0">1342 in</quantity>
|
||
males (Figs 15–18) and
|
||
<quantity id="011E75BB9A5DFFA85CE5FD29A5BFFD77" box="[422,507,727,751]" metricMagnitude="1" metricUnit="m" metricValue="8.666479999999998" pageId="1" pageNumber="54" unit="in" value="3412.0">3412 in</quantity>
|
||
females (Figs 19–22). Spination: male: palpal femur 0; femur I d1-1-1 p1di, II d1- 1-1, p1di, III d1-1-1 p2di, IV d1-1-1; patella I=II 0, III=IV r1; tibia I
|
||
<date id="B258FE9E9A5DFFA85EE2FD02A793FC8C" box="[929,983,764,788]" pageId="1" pageNumber="54">v2-2</date>
|
||
-2, II
|
||
<date id="B258FE9E9A5DFFA85957FD02A06AFC8C" box="[1044,1070,764,788]" pageId="1" pageNumber="54">v1</date>
|
||
r-2-2, p1-0-1, III
|
||
<date id="B258FE9E9A5DFFA859B0FD02A149FC8C" box="[1267,1293,764,788]" pageId="1" pageNumber="54">v1</date>
|
||
p-0-1r, p1-1, r1-1-1, IV
|
||
<date id="B258FE9E9A5DFFA85C53FCE1A56EFCAF" box="[272,298,799,823]" pageId="1" pageNumber="54">v1</date>
|
||
p-0-1r, p1-0, r1-1-1; metatarsus I=II
|
||
<date id="B258FE9E9A5DFFA85F84FCE1A6B8FCAF" box="[711,764,799,823]" pageId="1" pageNumber="54">v2-2</date>
|
||
, III
|
||
<date id="B258FE9E9A5DFFA85E6FFCE1A725FCAF" box="[812,865,799,823]" pageId="1" pageNumber="54">v2-2</date>
|
||
, p1-2, r1-2, IV
|
||
<date id="B258FE9E9A5DFFA85953FCE1A06EFCAF" box="[1040,1066,799,823]" pageId="1" pageNumber="54">v1</date>
|
||
p-1r, p0-2, r1-1-2; female: femur I=II d1-1-1, p1di, III d1-1-1 p2di, IV d1-1-1; patella I-IV 0; tibia I
|
||
<date id="B258FE9E9A5DFFA85EC7FCBAA7FDFCC4" box="[900,953,836,860]" pageId="1" pageNumber="54">v2-2</date>
|
||
-2, II
|
||
<date id="B258FE9E9A5DFFA85EB5FCBAA054FCC4" box="[1014,1040,836,860]" pageId="1" pageNumber="54">v1</date>
|
||
r-2-2, III
|
||
<date id="B258FE9E9A5DFFA85939FCBAA0D0FCC4" box="[1146,1172,836,860]" pageId="1" pageNumber="54">v1</date>
|
||
p-0-1r, p0-1, r1-1-1, IV
|
||
<date id="B258FE9E9A5DFFA85DD4FC99A4F4FCE7" box="[151,176,871,895]" pageId="1" pageNumber="54">v1</date>
|
||
p-0-0, p0, r1-1; metatarsus I=II
|
||
<date id="B258FE9E9A5DFFA85F55FC99A608FCE7" box="[534,588,871,895]" pageId="1" pageNumber="54">v2-2</date>
|
||
, III
|
||
<date id="B258FE9E9A5DFFA85F39FC99A6F4FCE7" box="[634,688,871,895]" pageId="1" pageNumber="54">v2-2</date>
|
||
, p1-2, r1-2, IV
|
||
<date id="B258FE9E9A5DFFA85E1EFC99A73CFCE7" box="[861,888,871,895]" pageId="1" pageNumber="54">v1</date>
|
||
p-1r, p0-2, r1-2. Abdomen about the same length as the carapace with longitudinal dorsal color patterns (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85E41FC72A723FC3C" box="[770,871,908,932]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1913,1936]" captionTargetBox="[151,1436,1111,1872]" captionTargetId="figure@1.[151,1436,1095,1873]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Macutula aracoiaba sp. nov. 1 male habitus; 2 female habitus." httpUri="https://zenodo.org/record/203342/files/figure.png" pageId="1" pageNumber="54">Figs 1–2</figureCitation>
|
||
). Epigynum with an anterior atrium (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85844FC72A114FC3C" box="[1287,1360,908,932]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation id="5EDDC4DB9A5DFFA8581FFC72A13FFC3C" box="[1372,1403,908,932]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,244,1415,1438]" captionTargetBox="[170,1425,210,1384]" captionTargetId="figure@6.[151,1436,193,1394]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Macutula santana sp. nov. 23 – 24 left male palp (23 ventral, 24 retrolateral). Macutula caruaru sp. nov. 25 – 26 left male palp (25 ventral, 26 retrolateral); 27 epigyne, ventral view; 28 vulva, dorsal view." httpUri="https://zenodo.org/record/203345/files/figure.png" pageId="1" pageNumber="54">27</figureCitation>
|
||
), a membranous, spiral initial portion of the copulation duct coiling backwards around the second, sclerotized portion, which coils forward and becomes loose, reaching the anterior spermatheca (
|
||
<figureCitation id="5EDDC4DB9A5DFFA85EAAFC2AA07BFC74" box="[1001,1087,980,1004]" captionStart="FIGURES 3 – 10" captionStartId="2.[151,244,1929,1952]" captionTargetBox="[178,1426,218,1889]" captionTargetId="figure@2.[151,1436,193,1896]" captionTargetPageId="2" captionText="FIGURES 3 – 10. Macutula aracoiaba sp. nov. 3 – 8 left male palp, (3 ventral, 4 retrolateral, 5 dorsal, 6 prolateral, 7 anterior view, 8 clarified bulb, ventral view); 9 epigyne, ventral view; 10 vulva, dorsal view." httpUri="https://zenodo.org/record/203343/files/figure.png" pageId="1" pageNumber="54">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation id="5EDDC4DB9A5DFFA85908FC2AA02FFC74" box="[1099,1131,980,1004]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,244,1415,1438]" captionTargetBox="[170,1425,210,1384]" captionTargetId="figure@6.[151,1436,193,1394]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Macutula santana sp. nov. 23 – 24 left male palp (23 ventral, 24 retrolateral). Macutula caruaru sp. nov. 25 – 26 left male palp (25 ventral, 26 retrolateral); 27 epigyne, ventral view; 28 vulva, dorsal view." httpUri="https://zenodo.org/record/203345/files/figure.png" pageId="1" pageNumber="54">28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C659D85E9A5DFFA85D86FC09A0A2FB88" blockId="1.[151,1436,151,1040]" box="[197,1254,1015,1040]" pageId="1" pageNumber="54">
|
||
<emphasis id="F492044C9A5DFFA85D86FC09A520FB88" bold="true" box="[197,356,1015,1040]" pageId="1" pageNumber="54">Composition.</emphasis>
|
||
Three species:
|
||
<taxonomicName id="01E6A3DD9A5DFFA85F50FC07A6EFFB88" box="[531,683,1016,1040]" class="Arachnida" family="Salticidae" genus="Macutula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="aracoiaba" status="sp. nov.">
|
||
<emphasis id="F492044C9A5DFFA85F50FC07A6EFFB88" box="[531,683,1016,1040]" italics="true" pageId="1" pageNumber="54">M. aracoiaba</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F492044C9A5DFFA85FF0FC06A748FB88" bold="true" box="[691,780,1016,1040]" pageId="1" pageNumber="54">
|
||
<taxonomicNameLabel id="EFA1B9379A5DFFA85FF0FC06A748FB88" box="[691,780,1016,1040]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="01E6A3DD9A5DFFA85E59FC07A7DCFB88" box="[794,920,1017,1040]" class="Arachnida" family="Salticidae" genus="Macutula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
|
||
<emphasis id="F492044C9A5DFFA85E59FC07A7DCFB88" box="[794,920,1017,1040]" italics="true" pageId="1" pageNumber="54">M. santana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F492044C9A5DFFA85EDCFC06A7BCFB88" bold="true" box="[927,1016,1016,1040]" pageId="1" pageNumber="54">
|
||
<taxonomicNameLabel id="EFA1B9379A5DFFA85EDCFC06A7BCFB88" box="[927,1016,1016,1040]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="01E6A3DD9A5DFFA85945FC07A0C2FB88" box="[1030,1158,1017,1040]" class="Arachnida" family="Salticidae" genus="Macutula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="caruaru" status="sp. nov.">
|
||
<emphasis id="F492044C9A5DFFA85945FC07A0C2FB88" box="[1030,1158,1017,1040]" italics="true" pageId="1" pageNumber="54">M. caruaru</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F492044C9A5DFFA859CEFC06A0A2FB88" bold="true" box="[1165,1254,1016,1040]" pageId="1" pageNumber="54">
|
||
<taxonomicNameLabel id="EFA1B9379A5DFFA859CEFC06A0A2FB88" box="[1165,1254,1016,1040]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |