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<document id="A5DB805D2A2338B07BEE23F80191680B" ID-CLB-Dataset="270512" ID-DOI="10.11646/zootaxa.5351.4.1" ID-GBIF-Dataset="5c01a707-b7ab-4497-8980-9752bfab74e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391904" ID-ZooBank="01FF28C0-58F7-47FC-B2A5-03BA8DAF2662" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1695991637813" checkinUser="plazi" docAuthor="Kury, Adriano B. &amp; Bernabé, Tiago" docDate="2023" docId="44270927FFBD575F6C8EFAEDFDD09DBC" docLanguage="en" docName="zootaxa.5351.4.1.pdf" docOrigin="Zootaxa 5351 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Titanoleptes eros Kury &amp; Bernabé 2023, sp. nov." docType="treatment" docUuid="EB41BA5F-6C2C-4E04-97CD-DBD02ABADBAA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="427" masterDocId="B81E715FFFA957456C19FFE6FFE89A74" masterDocTitle="Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae)" masterLastPageNumber="434" masterPageNumber="401" pageNumber="421" updateTime="1699693093697" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="EF4031A1E408459572FBD6AC3947F665" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F99485CF0D71EF3CB08708019B52AF18">Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae)</mods:title>
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<mods:name id="3352488208881BAC58B4C93BF7C46F3C" type="personal">
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<mods:namePart id="562C08F9BB0F8B79B3F285B0B0C3890C">Kury, Adriano B.</mods:namePart>
<mods:nameIdentifier id="448E0569CAAF5E7B4FBA0AFFCB4F5524" type="ORCID">0000-0002-8334-6204</mods:nameIdentifier>
<mods:affiliation id="D1C45975EC0232E1FCA9A8DCDD106802">Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20.940 - 040 Rio de Janeiro (RJ), Brazil. adrianok @ gmail. com; https: // orcid. org / 0000 - 0002 - 8334 - 6204</mods:affiliation>
<mods:nameIdentifier id="11D0D115A57771F32F5E9631C22F737E" type="email">adrianok@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4498E1EBE3F3248F9A17337637774D22">Bernabé, Tiago</mods:namePart>
<mods:nameIdentifier id="D925F7657DC5E721BE9EFC23C0F83448" type="ORCID">0000-0001-5820-2076</mods:nameIdentifier>
<mods:affiliation id="02BBA9415F79D2CFC5A664E952499481">Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia e Botânica, IBILCE, Universidade Estadual Paulista, UNESP, Rua Cristóvão Colombo 2265, CEP 15054 - 000, São José do Rio Preto, SP, Brazil.</mods:affiliation>
<mods:nameIdentifier id="87854A81301975A2533D6FDD1AEE921F" type="email">bernabe.t@gmail.com</mods:nameIdentifier>
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<mods:title id="88E53F89D5E21566F05A0AFB9B4017EA">Zootaxa</mods:title>
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<mods:date id="B839EBE022409569B15515FAF5D8278A">2023</mods:date>
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<mods:number id="AA15CAFDC71A0F8CE98028BAC8B6A9FD">2023-09-28</mods:number>
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<mods:number id="34B3C5D5186FA852324CBB9CBEBEC2F6">5351</mods:number>
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<treatment id="44270927FFBD575F6C8EFAEDFDD09DBC" ID-DOI="http://doi.org/10.5281/zenodo.8400104" ID-GBIF-Taxon="213635629" ID-Zenodo-Dep="8400104" LSID="urn:lsid:zoobank.org:act:EB41BA5F-6C2C-4E04-97CD-DBD02ABADBAA" httpUri="http://treatment.plazi.org/id/44270927FFBD575F6C8EFAEDFDD09DBC" lastPageId="26" lastPageNumber="427" pageId="20" pageNumber="421">
<subSubSection id="8494EBBAFFBD57516C8EFAEDFE219F52" box="[151,457,1291,1318]" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="CC31B831FFBD57516C8EFAEDFE219F52" blockId="20.[151,1025,1291,1391]" box="[151,457,1291,1318]" pageId="20" pageNumber="421">
<heading id="97790F5DFFBD57516C8EFAEDFE219F52" bold="true" box="[151,457,1291,1318]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<emphasis id="FEFA6423FFBD57516C8EFAEDFE219F52" bold="true" box="[151,457,1291,1318]" pageId="20" pageNumber="421">
<taxonomicName id="0B8EC3B2FFBD57516C8EFAEDFE8D9F52" ID-CoL="BTLX6" authority="Kury &amp; Bernabé, 2023" authorityName="Kury &amp; Bernabé" authorityYear="2023" box="[151,357,1291,1318]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="eros" status="sp. nov.">
<emphasis id="FEFA6423FFBD57516C8EFAEDFE8D9F52" bold="true" box="[151,357,1291,1318]" italics="true" pageId="20" pageNumber="421">Titanoleptes eros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5C9D958FFBD57516D75FAEAFE219F52" box="[364,457,1292,1318]" pageId="20" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="8494EBBAFFBD57516C8EFAD6FBE99F1B" pageId="20" pageNumber="421" type="description">
<paragraph id="CC31B831FFBD57516C8EFAD6FECF9F3E" blockId="20.[151,1025,1291,1391]" box="[151,295,1328,1354]" pageId="20" pageNumber="421">
(
<figureCitation id="54B5A4B4FFBD57516C86FAD6FEC89F3E" box="[159,288,1328,1354]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStart-3="FIGURE 26" captionStartId-0="25.[151,250,1919,1943]" captionStartId-1="26.[151,250,1463,1487]" captionStartId-2="27.[151,250,1125,1149]" captionStartId-3="28.[151,250,1943,1967]" captionTargetBox-0="[151,1435,417,1895]" captionTargetBox-1="[151,1436,181,1439]" captionTargetBox-2="[151,1436,181,1101]" captionTargetBox-3="[197,1390,181,1919]" captionTargetId-0="figure-123@25.[151,1435,417,1895]" captionTargetId-1="figure-20@26.[151,1436,181,1439]" captionTargetId-2="figure-17@27.[151,1436,181,1101]" captionTargetId-3="figure-20@28.[197,1390,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionText-0="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." captionText-1="FIGURE 24. Titanoleptes eros sp. nov., male paratype (MNRJ 7077) from ES, Santa Teresa, free tergites and proximal left leg IV: a. Dorsal view; b. Prolateral view; c. Ventral view; d. Retrolateral view. Scale bars = 1 mm." captionText-2="FIGURE 25. Titanoleptes eros sp. nov. male holotype (MNRJ 18440) from Santa Teresa. Femur IV (each figure is rotated 90° along the proximal-distal axis relative to each adjacent one). From the top: retrolateral view. ventral view, prolateral view and dorsal view. Scale bar = 5 mm." captionText-3="FIGURE 26. Titanoleptes eros sp. nov., male paratype (MNRJ 18927), ES, Santa Teresa, male genitalia, distal part: a. Dorsal view; b. Ventral view; c. Dextrolateral view; d. Detail of glans, dextrolateral view. Scale bars = 100 μm (a, c), 50 μm (b, d)." figureDoi-0="http://doi.org/10.5281/zenodo.8391957" figureDoi-1="http://doi.org/10.5281/zenodo.8391959" figureDoi-2="http://doi.org/10.5281/zenodo.8391961" figureDoi-3="http://doi.org/10.5281/zenodo.8391965" httpUri-0="https://zenodo.org/record/8391957/files/figure.png" httpUri-1="https://zenodo.org/record/8391959/files/figure.png" httpUri-2="https://zenodo.org/record/8391961/files/figure.png" httpUri-3="https://zenodo.org/record/8391965/files/figure.png" pageId="20" pageNumber="421">Figs 23–26</figureCitation>
)
</paragraph>
<paragraph id="CC31B831FFBD57516C8EFAB2FBE99F1B" blockId="20.[151,1025,1291,1391]" box="[151,1025,1364,1391]" pageId="20" pageNumber="421">
<uri id="B81FB433FFBD57516C8EFAB2FBE99F1B" box="[151,1025,1364,1391]" pageId="20" pageNumber="421">
urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="8494EBBAFFBD57516C8EFA7FFD859FC4" box="[151,621,1432,1457]" pageId="20" pageNumber="421" type="reference_group">
<paragraph id="CC31B831FFBD57516C8EFA7FFD859FC4" blockId="20.[151,621,1432,1457]" box="[151,621,1432,1457]" pageId="20" pageNumber="421">
“SOD-01 sp. D”—
<bibRefCitation id="A81FC5C0FFBD57516D4FFA7FFDCE9FC5" author="Kury, A. B. &amp; Bernabe, T. N. &amp; de Azara, L. N. &amp; Araujo, D. C. &amp; Benedetti, A. R." box="[342,550,1432,1457]" pageId="20" pageNumber="421" pagination="70 - 122" refId="ref16599" refString="Kury, A. B., Bernabe, T. N., de Azara, L. N., Araujo, D. C. &amp; Benedetti, A. R. (2022) Phylogeny of the clade K 92 (Opiliones, Laniatores, Gonyleptidae) with description of a new subfamily and discussion on the evolution of caelopygine facies and sexual dimorphism. Zoologischer Anzeiger, 298, 70 - 122. https: // doi. org / 10.1016 / j. jcz. 2022.03.004" type="journal article" year="2022">
Kury
<emphasis id="FEFA6423FFBD57516D8BFA7FFE2D9FC4" box="[402,453,1432,1456]" italics="true" pageId="20" pageNumber="421">et al.</emphasis>
2022: 81
</bibRefCitation>
, fig. 7.
</paragraph>
</subSubSection>
<subSubSection id="8494EBBAFFBD57516C8EFA39FC0B9C69" pageId="20" pageNumber="421" type="etymology">
<paragraph id="CC31B831FFBD57516C8EFA39FC0B9C69" blockId="20.[151,1437,1503,2033]" pageId="20" pageNumber="421">
<emphasis id="FEFA6423FFBD57516C8EFA39FEF69F8D" bold="true" box="[151,286,1503,1529]" pageId="20" pageNumber="421">Etymology.</emphasis>
Noun in apposition. In Greek mythology, Eros was the god of love, whose Roman counterpart was Cupid. Thus, this name mirrors another species of the genus,
<taxonomicName id="0B8EC3B2FFBD57516F5CF9E5FC349C69" authorityName=": Kury" authorityYear="2003" box="[837,988,1539,1565]" class="Arachnida" family="Gonyleptidae" genus="Metagonyleptes" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="cupidensis">
<emphasis id="FEFA6423FFBD57516F5CF9E5FC349C69" box="[837,988,1539,1565]" italics="true" pageId="20" pageNumber="421">C. cupidensis</emphasis>
</taxonomicName>
.
</paragraph>
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<subSubSection id="8494EBBAFFBD57516CDEF9C1FACC9D49" pageId="20" pageNumber="421" type="materials_examined">
<paragraph id="CC31B831FFBD57516CDEF9C1FACC9D49" blockId="20.[151,1437,1503,2033]" pageId="20" pageNumber="421">
<emphasis id="FEFA6423FFBD57516CDEF9C1FEB99C34" bold="true" box="[199,337,1574,1601]" pageId="20" pageNumber="421">Type series.</emphasis>
<materialsCitation id="7CE6B26CFFBD57516D4FF9C1FC529C11" ID-GBIF-Occurrence="4418826343" collectingDate="2006-08-11" collectionCode="REBIO" collectorName="Cachoeira II &amp; D. Mota" country="Brazil" latitude="-19.9066" location="REBIO Augusto Ruschi" longLatPrecision="1" longitude="-40.562035" municipality="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ 18440" specimenCount="1" specimenCount-female="2" stateProvince="Espirito Santo State" typeStatus="holotype">
<collectingCountry id="B499F8A1FFBD57516D4FF9C1FE569C35" box="[342,446,1575,1601]" name="Brazil" pageId="20" pageNumber="421">BRAZIL</collectingCountry>
,
<collectingRegion id="0E4A76D3FFBD57516DD1F9C1FD4F9C35" box="[456,679,1575,1601]" country="Brazil" name="Espirito Santo" pageId="20" pageNumber="421">Espirito Santo State</collectingRegion>
:
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(
<specimenCode id="9C28104AFFBD57516F20F9C1FC3D9C35" box="[825,981,1575,1601]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20" pageNumber="421">MNRJ 18440</specimenCode>
),,
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,
<collectorName id="617BDDE7FFBD57516C8EF9ADFECE9C11" box="[151,294,1611,1637]" pageId="20" pageNumber="421">Cachoeira II</collectorName>
, (
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,
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),
<date id="B8309EF1FFBD57516E92F9ADFCFC9C11" box="[651,788,1611,1637]" pageId="20" pageNumber="421" value="2006-08-11">
<collectingDate id="A8746719FFBD57516E92F9ADFCFC9C11" box="[651,788,1611,1637]" pageId="20" pageNumber="421" value="2006-08-11">11.viii.2006</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516F38F9ADFC689C11" box="[801,896,1611,1637]" pageId="20" pageNumber="421">D. Mota</collectorName>
leg.;
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<materialsCitation id="7CE6B26CFFBD57516FDAF9ADFC309CFD" ID-GBIF-Occurrence="4418826363" collectingDate="2006-08-11" collectionCode="REBIO" collectorName="Cachoeira II &amp; D. Mota" country="Brazil" latitude="-19.9066" location="Augusto Ruschi" longLatPrecision="1" longitude="-40.562035" municipality="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ 18440" specimenCount="11" specimenCount-female="2" specimenCount-male="2" stateProvince="Espirito Santo" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516FDAF9ADFC199C13" box="[963,1009,1611,1639]" pageId="20" pageNumber="421" type="male">2 ♂</specimenCount>
,
<specimenCount id="DA8873B8FFBD57516FE6F9ADFBC39C11" box="[1023,1067,1611,1637]" pageId="20" pageNumber="421" type="female">2 ♀</specimenCount>
,
<specimenCount id="DA8873B8FFBD57516823F9ADFB939C11" box="[1082,1147,1611,1637]" pageId="20" pageNumber="421" type="generic">7 juv.</specimenCount>
<typeStatus id="13350693FFBD5751689AF9ADFB069C11" box="[1155,1262,1611,1637]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
(
<specimenCode id="9C28104AFFBD575168E6F9ADFA789C11" box="[1279,1424,1611,1637]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20" pageNumber="421">MNRJ 7077</specimenCode>
),
<collectingMunicipality id="2C55224BFFBD57516C8EF989FEC39CFD" box="[151,299,1647,1673]" pageId="20" pageNumber="421">Santa Teresa</collectingMunicipality>
,
<location id="C951EEEAFFBD57516D21F989FD569CFD" LSID="urn:lsid:plazi:treatment:44270927FFBD575F6C8EFAEDFDD09DBC:C951EEEAFFBD57516D21F989FD569CFD" box="[312,702,1647,1673]" country="Brazil" municipality="Santa Teresa" name="Estacao Biologica de Santa Lucia" pageId="20" pageNumber="421" stateProvince="Espirito Santo State">Estação Biológica de Santa Lúcia</location>
,
<date id="B8309EF1FFBD57516ED2F989FCCB9CFD" box="[715,803,1647,1673]" pageId="20" pageNumber="421" value="2006-11">
<collectingDate id="A8746719FFBD57516ED2F989FCCB9CFD" box="[715,803,1647,1673]" pageId="20" pageNumber="421" value="2006-11">xi.2006</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516F28F989FC769CFD" box="[817,926,1647,1673]" pageId="20" pageNumber="421">J. Wesley</collectorName>
leg.;
</materialsCitation>
<materialsCitation id="7CE6B26CFFBD57516FFBF989FD069CD9" ID-GBIF-Occurrence="4418826346" collectingDate="2006-11" collectionCode="MNRJ" collectorName="J. Wesley" country="Brazil" location="Estacao Biologica de Santa Lucia" municipality="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ 7077" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Espirito Santo State" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516FFBF989FBF89CFF" box="[994,1040,1647,1675]" pageId="20" pageNumber="421" type="male">2 ♂</specimenCount>
,
<specimenCount id="DA8873B8FFBD57516806F989FBA49CFD" box="[1055,1100,1647,1673]" pageId="20" pageNumber="421" type="female">1 ♀</specimenCount>
<typeStatus id="13350693FFBD5751684CF989FB289CFD" box="[1109,1216,1647,1673]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
(
<specimenCode id="9C28104AFFBD575168CBF989FAAF9CFD" box="[1234,1351,1647,1673]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20" pageNumber="421">MNRJ 22</specimenCode>
),
<location id="C951EEEAFFBD57516947F989FF0B9CD9" LSID="urn:lsid:plazi:treatment:44270927FFBD575F6C8EFAEDFDD09DBC:C951EEEAFFBD57516947F989FF0B9CD9" country="Brazil" name="Santa Teresa" pageId="20" pageNumber="421" stateProvince="Espirito Santo State">Santa Teresa</location>
,
<date id="B8309EF1FFBD57516CF5F975FE869CD9" box="[236,366,1683,1709]" pageId="20" pageNumber="421" value="1980-12-22">
<collectingDate id="A8746719FFBD57516CF5F975FE869CD9" box="[236,366,1683,1709]" pageId="20" pageNumber="421" value="1980-12-22">22.xii.1980</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516D60F975FDE09CD9" box="[377,520,1683,1709]" pageId="20" pageNumber="421">E. Izecksohn</collectorName>
&amp;
<collectorName id="617BDDE7FFBD57516E30F975FD509CD9" box="[553,696,1683,1709]" pageId="20" pageNumber="421">O.L. Peixoto</collectorName>
leg.;
</materialsCitation>
<materialsCitation id="7CE6B26CFFBD57516EEDF975FC969CA5" ID-GBIF-Occurrence="4418826372" collectingDate="1980-12-22" collectingDateMax="1981-04-05" collectingDateMin="1981-04-03" collectionCode="MNRJ" collectorName="E. Izecksohn &amp; O. L. Peixoto" country="Brazil" location="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ 22" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Espirito Santo State" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516EEDF975FCF69CDB" box="[756,798,1683,1711]" pageId="20" pageNumber="421" type="male">3 ♂</specimenCount>
,
<specimenCount id="DA8873B8FFBD57516F30F975FCB99CD9" box="[809,849,1683,1709]" pageId="20" pageNumber="421" type="female">1 ♀</specimenCount>
<typeStatus id="13350693FFBD57516F4EF975FC2A9CD9" box="[855,962,1683,1709]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
(
<specimenCode id="9C28104AFFBD57516FD6F975FB839CD9" box="[975,1131,1683,1709]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20" pageNumber="421">MNRJ 18927</specimenCode>
),
<location id="C951EEEAFFBD57516864F975FAE59CD9" LSID="urn:lsid:plazi:treatment:44270927FFBD575F6C8EFAEDFDD09DBC:C951EEEAFFBD57516864F975FAE59CD9" box="[1149,1293,1683,1709]" country="Brazil" name="Santa Teresa" pageId="20" pageNumber="421" stateProvince="Espirito Santo State">Santa Teresa</location>
,
<date id="B8309EF1FFBD5751690FF974FA7F9CD9" box="[1302,1431,1682,1709]" pageId="20" pageNumber="421" value="1981-04-03" valueMax="1981-04-05" valueMin="1981-04-03">
<collectingDate id="A8746719FFBD5751690FF974FA7F9CD9" box="[1302,1431,1682,1709]" pageId="20" pageNumber="421" value="1981-04-03" valueMax="1981-04-05" valueMin="1981-04-03">3-5.iv.1981</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516C8EF951FEC09CA5" box="[151,296,1719,1745]" pageId="20" pageNumber="421">C.A.G. Cruz</collectorName>
,
<collectorName id="617BDDE7FFBD57516D2DF951FE2F9CA5" box="[308,455,1719,1745]" pageId="20" pageNumber="421">O.L. Peixoto</collectorName>
,
<collectorName id="617BDDE7FFBD57516DCDF951FDC69CA5" box="[468,558,1719,1745]" pageId="20" pageNumber="421">S. Silva</collectorName>
,
<collectorName id="617BDDE7FFBD57516E20F951FD669CA5" box="[569,654,1719,1745]" pageId="20" pageNumber="421">A. Reis</collectorName>
&amp;
<collectorName id="617BDDE7FFBD57516EAAF951FCAD9CA5" box="[691,837,1719,1745]" pageId="20" pageNumber="421">E. Izecksohn</collectorName>
leg.;
</materialsCitation>
<materialsCitation id="7CE6B26CFFBD57516F9CF951FCDC9C81" ID-GBIF-Occurrence="4418826356" collectingDate="1984-09-27" collectingDateMax="1984-09-28" collectingDateMin="1984-09-27" collectionCode="MNRJ" collectorName="O. L. Peixoto &amp; C. A. G. Cruz &amp; E. Izecksohn" country="Brazil" location="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ 18941" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo State" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516F9CF951FC589CA7" box="[901,944,1719,1747]" pageId="20" pageNumber="421" type="male">1 ♂</specimenCount>
<typeStatus id="13350693FFBD57516FA1F951FBF09CA5" box="[952,1048,1719,1745]" pageId="20" pageNumber="421" type="paratype">paratype</typeStatus>
(
<specimenCode id="9C28104AFFBD5751683EF951FB2E9CA5" box="[1063,1222,1719,1745]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20" pageNumber="421">MNRJ 18941</specimenCode>
),
<location id="C951EEEAFFBD575168C3F951FA839CA5" LSID="urn:lsid:plazi:treatment:44270927FFBD575F6C8EFAEDFDD09DBC:C951EEEAFFBD575168C3F951FA839CA5" box="[1242,1387,1719,1745]" country="Brazil" name="Santa Teresa" pageId="20" pageNumber="421" stateProvince="Espirito Santo State">Santa Teresa</location>
,
<date id="B8309EF1FFBD5751696EF951FEF99C81" pageId="20" pageNumber="421" value="1984-09-27" valueMax="1984-09-28" valueMin="1984-09-27">
<collectingDate id="A8746719FFBD5751696EF951FEF99C81" pageId="20" pageNumber="421" value="1984-09-27" valueMax="1984-09-28" valueMin="1984-09-27">27- 28.ix.1984</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516D04F93DFE479C81" box="[285,431,1755,1781]" pageId="20" pageNumber="421">O.L. Peixoto</collectorName>
,
<collectorName id="617BDDE7FFBD57516DA2F93DFDA09C81" box="[443,584,1755,1781]" pageId="20" pageNumber="421">C.A.G. Cruz</collectorName>
&amp;
<collectorName id="617BDDE7FFBD57516E75F93DFD159C81" box="[620,765,1755,1781]" pageId="20" pageNumber="421">E. Izecksohn</collectorName>
leg.;
</materialsCitation>
<materialsCitation id="7CE6B26CFFBD57516F22F93DFD9F9D6D" ID-GBIF-Occurrence="4418826339" collectingDate="2006-01-16" collectingDateMax="2006-01-20" collectingDateMin="2006-01-16" collectionCode="MNRJ" collectorName="AracnoLab" country="Brazil" location="REBIO de Santa Lucia" municipality="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ 18951" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo State" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516F22F93DFC8D9C83" box="[827,869,1755,1783]" pageId="20" pageNumber="421" type="male">1 ♂</specimenCount>
<typeStatus id="13350693FFBD57516F75F93DFC249C81" box="[876,972,1755,1781]" pageId="20" pageNumber="421" type="paratype">paratype</typeStatus>
(
<specimenCode id="9C28104AFFBD57516FC2F93DFB909C81" box="[987,1144,1754,1781]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="20" pageNumber="421">MNRJ 18951</specimenCode>
),
<collectingMunicipality id="2C55224BFFBD57516895F93DFAF49C81" box="[1164,1308,1755,1781]" pageId="20" pageNumber="421">Santa Teresa</collectingMunicipality>
,
<location id="C951EEEAFFBD5751693EF93DFEF69D6D" LSID="urn:lsid:plazi:treatment:44270927FFBD575F6C8EFAEDFDD09DBC:C951EEEAFFBD5751693EF93DFEF69D6D" country="Brazil" municipality="Santa Teresa" name="REBIO de Santa Lucia" pageId="20" pageNumber="421" stateProvince="Espirito Santo State">REBIO de Santa Lúcia</location>
,
<date id="B8309EF1FFBD57516D31F919FE519D6D" box="[296,441,1791,1817]" pageId="20" pageNumber="421" value="2006-01-16" valueMax="2006-01-20" valueMin="2006-01-16">
<collectingDate id="A8746719FFBD57516D31F919FE519D6D" box="[296,441,1791,1817]" pageId="20" pageNumber="421" value="2006-01-16" valueMax="2006-01-20" valueMin="2006-01-16">16-20.i.2006</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516DDBF919FDA89D6D" box="[450,576,1791,1817]" pageId="20" pageNumber="421">AracnoLab</collectorName>
leg.;
</materialsCitation>
<materialsCitation id="7CE6B26CFFBD57516E64F919FF2A9D49" ID-GBIF-Occurrence="4418826360" collectingDate="1974-08-24" collectingDateMax="1984-09-28" collectingDateMin="1984-09-27" collectionCode="MNRJ" collectorName="A. L. Perachi" country="Brazil" location="Santa Teresa" municipality="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MNRJ-HS 872" specimenCount="2" specimenCount-female="1" specimenCount-male="2" stateProvince="Espirito Santo State" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516E64F919FD4D9D6F" box="[637,677,1791,1819]" pageId="20" pageNumber="421" type="male">2 ♂</specimenCount>
<typeStatus id="13350693FFBD57516EB2F919FCFE9D6D" box="[683,790,1791,1817]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
(
<specimenCode id="9C28104AFFBD57516F3DF919FC399D6D" box="[804,977,1791,1817]" collectionCode="MNRJ-HS" pageId="20" pageNumber="421">MNRJ-HS 872</specimenCode>
),
<collectingMunicipality id="2C55224BFFBD57516FFAF919FB9B9D6D" box="[995,1139,1791,1817]" pageId="20" pageNumber="421">Santa Teresa</collectingMunicipality>
,
<date id="B8309EF1FFBD57516865F919FAEF9D6D" box="[1148,1287,1791,1817]" pageId="20" pageNumber="421" value="1974-08-24">
<collectingDate id="A8746719FFBD57516865F919FAEF9D6D" box="[1148,1287,1791,1817]" pageId="20" pageNumber="421" value="1974-08-24">24.viii.1974</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516916F919FA749D6D" box="[1295,1436,1791,1817]" pageId="20" pageNumber="421">A.L. Perachi</collectorName>
leg;
</materialsCitation>
<materialsCitation id="7CE6B26CFFBD57516CD1F8C5FACC9D49" ID-GBIF-Occurrence="4418826312" box="[200,1316,1826,1853]" collectingDate="2008-05-24" collectingDateMax="2008-05-30" collectingDateMin="2008-05-24" collectionCode="MNRJ" collectorName="F. Carbayo" country="Brazil" location="REBIO Augusto Ruschi" municipality="Santa Teresa" pageId="20" pageNumber="421" specimenCode="MZSP 73450" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo State" typeStatus="paratype">
<specimenCount id="DA8873B8FFBD57516CD1F8C5FF1A9D49" box="[200,242,1827,1853]" pageId="20" pageNumber="421" type="female">1 ♀</specimenCount>
<typeStatus id="13350693FFBD57516CE0F8C5FE8C9D49" box="[249,356,1827,1853]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
(
<specimenCode id="9C28104AFFBD57516D6AF8C5FDE69D49" box="[371,526,1826,1853]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="20" pageNumber="421" type="Museum">MZSP 73450</specimenCode>
),
<collectingMunicipality id="2C55224BFFBD57516E38F8C5FD5A9D49" box="[545,690,1827,1853]" pageId="20" pageNumber="421">Santa Teresa</collectingMunicipality>
,
<location id="C951EEEAFFBD57516EA4F8C5FC259D49" LSID="urn:lsid:plazi:treatment:44270927FFBD575F6C8EFAEDFDD09DBC:C951EEEAFFBD57516EA4F8C5FC259D49" box="[701,973,1827,1853]" country="Brazil" municipality="Santa Teresa" name="REBIO Augusto Ruschi" pageId="20" pageNumber="421" stateProvince="Espirito Santo State">REBIO Augusto Ruschi</location>
,
<date id="B8309EF1FFBD57516FC1F8C5FB869D49" box="[984,1134,1827,1853]" pageId="20" pageNumber="421" value="2008-05-24" valueMax="2008-05-30" valueMin="2008-05-24">
<collectingDate id="A8746719FFBD57516FC1F8C5FB869D49" box="[984,1134,1827,1853]" pageId="20" pageNumber="421" value="2008-05-24" valueMax="2008-05-30" valueMin="2008-05-24">24-30.v.2008</collectingDate>
</date>
,
<collectorName id="617BDDE7FFBD57516860F8C5FB1C9D49" box="[1145,1268,1827,1853]" pageId="20" pageNumber="421">F. Carbayo</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8494EBBAFFBD57516CDEF8A0FCC39D85" pageId="20" pageNumber="421" type="diagnosis">
<paragraph id="CC31B831FFBD57516CDEF8A0FCC39D85" blockId="20.[151,1437,1503,2033]" pageId="20" pageNumber="421">
<emphasis id="FEFA6423FFBD57516CDEF8A0FEAA9D14" bold="true" box="[199,322,1862,1888]" pageId="20" pageNumber="421">Diagnosis.</emphasis>
Ocularium in both sexes with two high erect spines, concolorous with background (
<figureCitation id="54B5A4B4FFBD57516932F8A1FA679D15" box="[1323,1423,1863,1889]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="20" pageNumber="421">Fig. 23c</figureCitation>
). Scutal area III armed with small paramedian rounded tubercles (
<figureCitation id="54B5A4B4FFBD57516F70F88DFC189DF1" box="[873,1008,1899,1925]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="20" pageNumber="421">Figs. 23d, e</figureCitation>
). Free tergite III armed with a median apophysis in both sexes (
<figureCitation id="54B5A4B4FFBD57516DA1F869FDD79DDD" box="[440,575,1935,1961]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="20" pageNumber="421">Figs. 23a, f</figureCitation>
). Coxa IV of male without retrolateral apophysis (
<figureCitation id="54B5A4B4FFBD5751689AF869FB009DDD" box="[1155,1256,1935,1961]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="20" pageNumber="421">Fig. 23b</figureCitation>
). Trochanter IV of male with proximal prolateral conical apophysis forming short pincer with coxal apophysis (
<figureCitation id="54B5A4B4FFBD575168DEF855FAC09DB9" box="[1223,1320,1971,1997]" captionStart="FIGURE 24" captionStartId="26.[151,250,1463,1487]" captionTargetBox="[151,1436,181,1439]" captionTargetId="figure-20@26.[151,1436,181,1439]" captionTargetPageId="26" captionText="FIGURE 24. Titanoleptes eros sp. nov., male paratype (MNRJ 7077) from ES, Santa Teresa, free tergites and proximal left leg IV: a. Dorsal view; b. Prolateral view; c. Ventral view; d. Retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391959" httpUri="https://zenodo.org/record/8391959/files/figure.png" pageId="20" pageNumber="421">Fig. 24a</figureCitation>
). Glans of penis free, without a circular wall around stylus (
<figureCitation id="54B5A4B4FFBD57516EA5F831FCF69D85" box="[700,798,2007,2033]" captionStart="FIGURE 26" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[197,1390,181,1919]" captionTargetId="figure-20@28.[197,1390,181,1919]" captionTargetPageId="28" captionText="FIGURE 26. Titanoleptes eros sp. nov., male paratype (MNRJ 18927), ES, Santa Teresa, male genitalia, distal part: a. Dorsal view; b. Ventral view; c. Dextrolateral view; d. Detail of glans, dextrolateral view. Scale bars = 100 μm (a, c), 50 μm (b, d)." figureDoi="http://doi.org/10.5281/zenodo.8391965" httpUri="https://zenodo.org/record/8391965/files/figure.png" pageId="20" pageNumber="421">Fig. 26d</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="98F1E8B9FFBC57506C8EF871FAAF9D83" ID-DOI="http://doi.org/10.5281/zenodo.8391948" ID-Zenodo-Dep="8391948" httpUri="https://zenodo.org/record/8391948/files/figure.png" pageId="21" pageNumber="422" startId="21.[151,250,1943,1967]" targetBox="[181,1406,181,1919]" targetPageId="21" targetType="figure">
<paragraph id="CC31B831FFBC57506C8EF871FAAF9D83" blockId="21.[151,1437,1943,2040]" pageId="21" pageNumber="422">
<emphasis id="FEFA6423FFBC57506C8EF871FEF69DDB" bold="true" box="[151,286,1943,1967]" pageId="21" pageNumber="422">FIGURE 19.</emphasis>
<taxonomicName id="0B8EC3B2FFBC57506D3AF871FCE09DDB" authority="(Soares &amp; Soares, 1946)" authorityName="Kury &amp; Bernabé" authorityYear="2023" baseAuthorityName="Soares &amp; Soares" baseAuthorityYear="1946" box="[291,776,1943,1967]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="cupidensis">
<emphasis id="FEFA6423FFBC57506D3AF871FDE79DDB" box="[291,527,1943,1967]" italics="true" pageId="21" pageNumber="422">Titanoleptes cupidensis</emphasis>
(
<bibRefCitation id="A81FC5C0FFBC57506E03F871FCE99DDB" author="Soares, B. A. M. &amp; Soares, H. E. M." box="[538,769,1943,1967]" pageId="21" pageNumber="422" pagination="195 - 212" refId="ref16998" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1946) Novos opilioes do Espirito Santo e um novo opiliao do estado do Para. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, 7, 195 - 212." type="journal article" year="1946">Soares &amp; Soares, 1946</bibRefCitation>
)
</taxonomicName>
, (MNRJ 17869), male (A–F), and female (G, H), from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Same, ventral view; d. Same, dextrolateral view; e. Same, posterior view; f. Same, frontal view; g. Female, habitus, dextrolateral view; h. Same, dorsal view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="98F1E8B9FFBF57536C8EFB1AFE799F28" ID-DOI="http://doi.org/10.5281/zenodo.8391950" ID-Zenodo-Dep="8391950" httpUri="https://zenodo.org/record/8391950/files/figure.png" pageId="22" pageNumber="423" startId="22.[151,250,1276,1300]" targetBox="[151,1435,181,1252]" targetPageId="22" targetType="figure">
<paragraph id="CC31B831FFBF57536C8EFB1AFE799F28" blockId="22.[151,1436,1276,1372]" pageId="22" pageNumber="423">
<emphasis id="FEFA6423FFBF57536C8EFB1AFEF69F60" bold="true" box="[151,286,1276,1300]" pageId="22" pageNumber="423">FIGURE 20.</emphasis>
<taxonomicName id="0B8EC3B2FFBF57536D3BFB1AFCED9F60" authority="(Soares &amp; Soares, 1946)" authorityName="Kury &amp; Bernabé" authorityYear="2023" baseAuthorityName="Soares &amp; Soares" baseAuthorityYear="1946" box="[290,773,1276,1300]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="cupidensis">
<emphasis id="FEFA6423FFBF57536D3BFB1AFDE59F60" box="[290,525,1276,1300]" italics="true" pageId="22" pageNumber="423">Titanoleptes cupidensis</emphasis>
(
<bibRefCitation id="A81FC5C0FFBF57536E00FB1AFD169F60" author="Soares, B. A. M. &amp; Soares, H. E. M." box="[537,766,1276,1300]" pageId="22" pageNumber="423" pagination="195 - 212" refId="ref16998" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1946) Novos opilioes do Espirito Santo e um novo opiliao do estado do Para. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, 7, 195 - 212." type="journal article" year="1946">Soares &amp; Soares, 1946</bibRefCitation>
)
</taxonomicName>
, male (MNRJ 17869), from Sooretama: a. Left coxa and trochanter IV, dorsal view; b. Left patella IV, prolateral view; c. Left coxa to proximal femur IV, prolateral view; d. Left leg IV, ventral view. Scale bars = 1mm.
</paragraph>
</caption>
<subSubSection id="8494EBBAFFBF57536CDEFA6CFCD39FD1" box="[199,827,1418,1445]" pageId="22" pageNumber="423" type="distribution">
<paragraph id="CC31B831FFBF57536CDEFA6CFCD39FD1" blockId="22.[151,1437,1418,2021]" box="[199,827,1418,1445]" pageId="22" pageNumber="423">
<emphasis id="FEFA6423FFBF57536CDEFA6CFE899FD0" bold="true" box="[199,353,1418,1444]" pageId="22" pageNumber="423">Distribution.</emphasis>
<collectingCountry id="B499F8A1FFBF57536D7EFA6CFE469FD0" box="[359,430,1418,1444]" name="Brazil" pageId="22" pageNumber="423">Brazil</collectingCountry>
,
<collectingRegion id="0E4A76D3FFBF57536DA3FA6CFD739FD1" box="[442,667,1418,1445]" country="Brazil" name="Espirito Santo" pageId="22" pageNumber="423">Espírito Santo State</collectingRegion>
, Santa Teresa.
</paragraph>
</subSubSection>
<subSubSection id="8494EBBAFFBF575F6CDEFA48FDD09DBC" lastPageId="26" lastPageNumber="427" pageId="22" pageNumber="423" type="description">
<paragraph id="CC31B831FFBF57536CDEFA48FEE69C65" blockId="22.[151,1437,1418,2021]" pageId="22" pageNumber="423">
<emphasis id="FEFA6423FFBF57536CDEFA48FDCD9FBD" bold="true" box="[199,549,1454,1481]" pageId="22" pageNumber="423">
Description of male
<typeStatus id="13350693FFBF57536DA1FA49FDC99FBD" box="[440,545,1455,1481]" pageId="22" pageNumber="423" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="FEFA6423FFBF57536E35FA49FD309FBC" box="[556,728,1455,1480]" italics="true" pageId="22" pageNumber="423">Measurements:</emphasis>
CL = 2.9, CW = 3.6, AL = 3.9, AW = 6.3. Pp = 8.4. Legs, I: Fe = 4.8, Ti = 3.2, Mt = 5.1; II: Fe = 10.5, Ti = 8.0, Mt = 9.8; III: Fe = 8.7, Ti = 4.6, Mt = 7.3; IV: Fe = 12.1, Ti = 7.5, Mt = 12.1.
</paragraph>
<paragraph id="CC31B831FFBF57526CDEF9FDFD859A8D" blockId="22.[151,1437,1418,2021]" lastBlockId="23.[151,1437,151,393]" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">
<emphasis id="FEFA6423FFBF57536CDEF9FDFEC19C40" box="[199,297,1563,1588]" italics="true" pageId="22" pageNumber="423">Dorsum.</emphasis>
Dorsal scutum outline gamma pyriform in dorsal view, quite flattened, so that Cx IV is much thicker than scutum in lateral view. Carapace mid-area cordiform, with narrow and low ocularium. Ocularium armed with a pair of paramedian, strong, divergent spines. Frontal hump low, armed with a pair of small granules. Two ozopores, first slit-like, second rounded. Mesotergum divided into 4 scutal areas, although areas III and IV are extensively fused to each other, except for the lateral thirds, where faint grooves are visible. Area I long, divided into broadly joined halves. Area II projected into area I. Area III strongly arched frontwards and moderately encased into area II. Carapace smooth and unarmed, area I with a transverse row of five granules on each side, area II with a similar row of 10 granules, without any especially developed in the middle. Areas III-IV with two rows of a few granules plus a paramedian pair of larger rounded capitate tubercles, these forming a trapezoid with posterior tubercles much smaller than anterior ones, and more widely spaced. Area V (posterior border of scutum) with a transverse row of granules. Lateral margins of scutum with a row of tubercles stretching from mid-carapace to area V, and which increase in size toward the posterior. Each free tergite with a row of larger granules, free tergite III with an immense spiniform apophysis. Posterior border of dorsal scutum with arched concavity.
<emphasis id="FEFA6423FFBF5753681FF82DFBBA9D90" box="[1030,1106,1995,2020]" italics="true" pageId="22" pageNumber="423">Venter.</emphasis>
Cx I free, other connected by tubercular bridges. Cx IV much larger than the others combined, without any ventro-apical tubercle. Stigmatic area posterior border deeply concave. Free tergites each with a transverse row of granules. Stigmatic area with very long axis and large, elliptical, slanted stigmata.
</paragraph>
<paragraph id="CC31B831FFBE57526CDEFEE5FE559BFD" blockId="23.[151,1437,151,393]" pageId="23" pageNumber="424">
<emphasis id="FEFA6423FFBE57526CDEFEE5FED79B69" box="[199,319,259,285]" italics="true" pageId="23" pageNumber="424">Pedipalps.</emphasis>
Moderately elongate, as long as body. Tr with thin neck, 1 + 3 setiferous tubercles dorsal and ventral. Fe with a row of 3 extremely weak ventral setiferous tubercles and much stronger meso-distal spine. Pa strongly curved, unarmed. Ti and Ta flattened ventrally. Spination of Ti: mesal IiIi, ectal IiIi (last two geminated). Spination of Ta: mesal IiIi, ectal IiIi.
</paragraph>
<caption id="98F1E8B9FFBE57526C8EF8C8FEF69DFA" ID-DOI="http://doi.org/10.5281/zenodo.8391953" ID-Zenodo-Dep="8391953" httpUri="https://zenodo.org/record/8391953/files/figure.png" pageId="23" pageNumber="424" startId="23.[151,250,1838,1862]" targetBox="[151,1435,417,1815]" targetPageId="23" targetType="figure">
<paragraph id="CC31B831FFBE57526C8EF8C8FEF69DFA" blockId="23.[151,1437,1838,1934]" pageId="23" pageNumber="424">
<emphasis id="FEFA6423FFBE57526C8EF8C8FECA9D32" bold="true" box="[151,290,1838,1862]" pageId="23" pageNumber="424">FIGURE 21.</emphasis>
<taxonomicName id="0B8EC3B2FFBE57526D32F8C8FCCB9D32" authority="(Soares &amp; Soares, 1946)" authorityName="Kury &amp; Bernabé" authorityYear="2023" baseAuthorityName="Soares &amp; Soares" baseAuthorityYear="1946" box="[299,803,1838,1862]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="cupidensis">
<emphasis id="FEFA6423FFBE57526D32F8C8FDF29D32" box="[299,538,1838,1862]" italics="true" pageId="23" pageNumber="424">Titanoleptes cupidensis</emphasis>
(
<bibRefCitation id="A81FC5C0FFBE57526E30F8C8FCF49D32" author="Soares, B. A. M. &amp; Soares, H. E. M." box="[553,796,1838,1862]" pageId="23" pageNumber="424" pagination="195 - 212" refId="ref16998" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1946) Novos opilioes do Espirito Santo e um novo opiliao do estado do Para. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, 7, 195 - 212." type="journal article" year="1946">Soares &amp; Soares, 1946</bibRefCitation>
)
</taxonomicName>
, male (MNRJ 17869), from Sooretama, left Fe IV: a. Dorsal view; b. Retrolateral (external) view; c. Ventral view; d. Prolateral (internal) view. Anterior direction upwards in a to c. Scale bars = 5 mm.
</paragraph>
</caption>
<caption id="98F1E8B9FFB1575D6C8EF991FCD49CAC" ID-DOI="http://doi.org/10.5281/zenodo.8391955" ID-Zenodo-Dep="8391955" httpUri="https://zenodo.org/record/8391955/files/figure.png" pageId="24" pageNumber="425" startId="24.[151,250,1655,1679]" targetBox="[151,1436,181,1632]" targetPageId="24" targetType="figure">
<paragraph id="CC31B831FFB1575D6C8EF991FCD49CAC" blockId="24.[151,1436,1655,1752]" pageId="24" pageNumber="425">
<emphasis id="FEFA6423FFB1575D6C8EF991FEC89CFB" bold="true" box="[151,288,1655,1679]" pageId="24" pageNumber="425">FIGURE 22.</emphasis>
<taxonomicName id="0B8EC3B2FFB1575D6D3EF991FCFF9CFB" authority="(Soares &amp; Soares, 1946)" authorityName="Kury &amp; Bernabé" authorityYear="2023" baseAuthorityName="Soares &amp; Soares" baseAuthorityYear="1946" box="[295,791,1655,1680]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="cupidensis">
<emphasis id="FEFA6423FFB1575D6D3EF991FDFD9CFB" box="[295,533,1655,1679]" italics="true" pageId="24" pageNumber="425">Titanoleptes cupidensis</emphasis>
(
<bibRefCitation id="A81FC5C0FFB1575D6E3BF99EFCF89CFB" author="Soares, B. A. M. &amp; Soares, H. E. M." box="[546,784,1655,1680]" pageId="24" pageNumber="425" pagination="195 - 212" refId="ref16998" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1946) Novos opilioes do Espirito Santo e um novo opiliao do estado do Para. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, 7, 195 - 212." type="journal article" year="1946">Soares &amp; Soares, 1946</bibRefCitation>
)
</taxonomicName>
, (MNRJ 17869), from Sooretama, male genitalia, distal part: a. Dorsal view; b. Ventral view; c. Dextrolateral view; d. Same, panoramic; e. Latero-apical view; f. Dorso-apical view; g. Glans, dextrolateral view. Scale bars = 100 μm (a, b, d, e), 50 μm (c, f, g).
</paragraph>
</caption>
<paragraph id="CC31B831FFB1575C6CDEF8E1FAB29B69" blockId="24.[151,1437,1798,2040]" lastBlockId="25.[151,1437,151,393]" lastPageId="25" lastPageNumber="426" pageId="24" pageNumber="425">
<emphasis id="FEFA6423FFB1575D6CDEF8E1FEEC9D54" box="[199,260,1799,1824]" italics="true" pageId="24" pageNumber="425">Legs.</emphasis>
All legs slender, only leg IV with significant armature. Dorsal apophyses of Cx I and II: Cx I anterior single, triangular erect, posterior larger, conical, directed backwards. Cx II anterior single, triangular erect, posterior flat triangular with small accessory tubercle. Tr IV with prodorsal median conical apophysis, which connects with apophysis of Cx IV and retrolateral distal conical apophysis of the same size. Otherwise only with a few small and scattered ventral tubercles. Legs I to III entirely unarmed. Fe I–II straight, III slightly arched. Fe IV: elongate, curved to the median line at 1/2 length; neither curved to ventral nor to dorsal. Proventral: row of numerous, small, acuminate tubercles all along the femur length. Retroventral: row of numerous, small, acuminate tubercles + unarmed mid-region + 6 larger tubercles distally. Retrodorsal: row of tubercles, being the basalmost and distalmost ones larger, with RD9 the largest. Dorsal: row of small tubercles, of which Do6 to Do8 more outstanding. Prodorsal: row of numerous small acuminate tubercles all along the femur length. Pa IV with many rows of tubercles, Ti IV as well, but tubercles are coarser. Mt IV smooth, thin, and slender. Tarsal counts: 6(3)-6(3)/10(3)-10(3)/7-7/8-8.
</paragraph>
<paragraph id="CC31B831FFB0575C6CDEFEC1FD849BFD" blockId="25.[151,1437,151,393]" pageId="25" pageNumber="426">
<emphasis id="FEFA6423FFB0575C6CDEFEC1FE349B35" box="[199,476,295,321]" italics="true" pageId="25" pageNumber="426">Color (in 75% ethanol).</emphasis>
Body and appendages background Strong Greenish Yellow (#99) with some darker mottling, especially on carapace. Leg IV considerably darker starting on apical portion of Cx. Venter as dorsum, with unobtrusive honeycomb reticulation.
</paragraph>
<caption id="98F1E8B9FFB0575C6C8EF899FC8F9DAA" ID-DOI="http://doi.org/10.5281/zenodo.8391957" ID-Zenodo-Dep="8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="25" pageNumber="426" startId="25.[151,250,1919,1943]" targetBox="[151,1435,417,1895]" targetPageId="25" targetType="figure">
<paragraph id="CC31B831FFB0575C6C8EF899FC8F9DAA" blockId="25.[151,1436,1919,2015]" pageId="25" pageNumber="426">
<emphasis id="FEFA6423FFB0575C6C8EF899FEC99DE3" bold="true" box="[151,289,1919,1943]" pageId="25" pageNumber="426">FIGURE 23.</emphasis>
<taxonomicName id="0B8EC3B2FFB0575C6D3EF899FE3D9DE3" authority="Kury &amp; Bernabé, 2023" authorityName="Kury &amp; Bernabé" authorityYear="2023" box="[295,469,1919,1943]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="426" phylum="Arthropoda" rank="species" species="eros" status="sp. nov.">
<emphasis id="FEFA6423FFB0575C6D3EF899FE3D9DE3" box="[295,469,1919,1943]" italics="true" pageId="25" pageNumber="426">Titanoleptes eros</emphasis>
</taxonomicName>
<emphasis id="FEFA6423FFB0575C6DC5F899FDC59DE3" bold="true" box="[476,557,1919,1943]" pageId="25" pageNumber="426">
<taxonomicNameLabel id="E5C9D958FFB0575C6DC5F899FDC59DE3" box="[476,557,1919,1943]" pageId="25" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="98F1E8B9FFB3575F6C8EFA51FBB49F87" ID-DOI="http://doi.org/10.5281/zenodo.8391959" ID-Zenodo-Dep="8391959" httpUri="https://zenodo.org/record/8391959/files/figure.png" pageId="26" pageNumber="427" startId="26.[151,250,1463,1487]" targetBox="[151,1436,181,1439]" targetPageId="26" targetType="figure">
<paragraph id="CC31B831FFB3575F6C8EFA51FBB49F87" blockId="26.[151,1436,1463,1524]" pageId="26" pageNumber="427">
<emphasis id="FEFA6423FFB3575F6C8EFA51FEC89FBB" bold="true" box="[151,288,1463,1487]" pageId="26" pageNumber="427">FIGURE 24.</emphasis>
<taxonomicName id="0B8EC3B2FFB3575F6D3FFA51FE3B9FBB" authority="Kury &amp; Bernabé, 2023" authorityName="Kury &amp; Bernabé" authorityYear="2023" box="[294,467,1463,1487]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="26" pageNumber="427" phylum="Arthropoda" rank="species" species="eros" status="sp. nov.">
<emphasis id="FEFA6423FFB3575F6D3FFA51FE3B9FBB" box="[294,467,1463,1487]" italics="true" pageId="26" pageNumber="427">Titanoleptes eros</emphasis>
</taxonomicName>
<emphasis id="FEFA6423FFB3575F6DC1FA5EFDC19FA4" bold="true" box="[472,553,1464,1488]" pageId="26" pageNumber="427">
<taxonomicNameLabel id="E5C9D958FFB3575F6DC1FA5EFDC19FA4" box="[472,553,1464,1488]" pageId="26" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (MNRJ 7077) from ES, Santa Teresa, free tergites and proximal left leg IV: a. Dorsal view; b. Prolateral view; c. Ventral view; d. Retrolateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="CC31B831FFB3575F6CDEF9C4FE5B9D28" blockId="26.[151,1437,1570,1992]" pageId="26" pageNumber="427">
<emphasis id="FEFA6423FFB3575F6CDEF9C4FE859C48" box="[199,365,1570,1596]" italics="true" pageId="26" pageNumber="427">Male genitalia</emphasis>
(
<figureCitation id="54B5A4B4FFB3575F6D64F9C4FDEB9C48" box="[381,515,1570,1596]" captionStart="FIGURE 26" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[197,1390,181,1919]" captionTargetId="figure-20@28.[197,1390,181,1919]" captionTargetPageId="28" captionText="FIGURE 26. Titanoleptes eros sp. nov., male paratype (MNRJ 18927), ES, Santa Teresa, male genitalia, distal part: a. Dorsal view; b. Ventral view; c. Dextrolateral view; d. Detail of glans, dextrolateral view. Scale bars = 100 μm (a, c), 50 μm (b, d)." figureDoi="http://doi.org/10.5281/zenodo.8391965" httpUri="https://zenodo.org/record/8391965/files/figure.png" pageId="26" pageNumber="427">Figs. 26a–d</figureCitation>
). VP trapezoid, and distal border with a moderately deep parabolic cleft. Field of
<typeStatus id="13350693FFB3575F6C8EF9A0FF2F9C14" box="[151,199,1606,1632]" pageId="26" pageNumber="427">type</typeStatus>
1 microsetae occupying most of VP except distal corners and not extending dorsally into podium but rather to latero-basal lobe. Basal lobe as a large, rounded polygon projecting dorsally from VP. Macrosetae (MS) A and B moderately long, forming an arch on latero-basal lobe, B only slightly smaller than A1–A3. MS C1–C3 elongate, straight basally, but distally helical. MS D1 minute, midway between C and A, E1–E2 short, situated on flange. Flabellum diamond-shaped, with ventral half outline gently serrated. Glans sac short, multi-folded, forming apically a solea projected as a short and blunt dorsal process. Stylus stalk arising from ventralmost part of glans sac; stylus inserted on stalk in candelabrum-mode. Stylus distally provided with distal denticles, which more apically form a large spiny protuberance.
</paragraph>
<paragraph id="CC31B831FFB3575F6CDEF880FDD09DBC" blockId="26.[151,1437,1570,1992]" pageId="26" pageNumber="427">
<emphasis id="FEFA6423FFB3575F6CDEF880FE389DF4" bold="true" box="[199,464,1893,1920]" pageId="26" pageNumber="427">Female (MNRJ 7077).</emphasis>
The female is essentially similar to male in build, appendages, and armature of dorsal scutum. Main differences are in the apophysis of free tergite III and in the armature of leg IV, which are much reduced. Tarsal counts: 6(3)/9(3)/7/8.
</paragraph>
</subSubSection>
</treatment>
</document>
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