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<document id="B8979D3D5A3249967AF2FBCCC7B4EF2E" ID-DOI="10.5281/zenodo.12586003" ID-ISSN="2064-2474" ID-Zenodo-Dep="12586003" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719595525160" checkinUser="felipe" docAuthor="Nabozhenko, M." docDate="2005" docId="03DB7E6FFFE6EF53E55971AEFE390831" docLanguage="en" docName="ActaZoolAcadSciH.51.4.349-355.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 51 (4)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Hedyphanes semnanicus Nabozhenko, 2005, sp. n." docType="treatment" docVersion="2" lastPageNumber="354" masterDocId="FFE20617FFE5EF56E7047019FFC00E32" masterDocTitle="New Synonymies And New Species Of The Genus Hedyphanes Fischer Von Waldheim, 1822 (Coleoptera, Tenebrionidae: Helopini)" masterLastPageNumber="355" masterPageNumber="349" pageNumber="352" updateTime="1719599216648" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="9223E83F921EDCAD53174BDD2B58C586">New Synonymies And New Species Of The Genus Hedyphanes Fischer Von Waldheim, 1822 (Coleoptera, Tenebrionidae: Helopini)</mods:title>
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<mods:namePart id="02CF0B3A4DCD852FF85C0D2565209A96">Nabozhenko, M.</mods:namePart>
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<mods:date id="C1AE01F40084BFA31A224572FE84C64E">2005</mods:date>
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<treatment id="03DB7E6FFFE6EF53E55971AEFE390831" LSID="urn:lsid:plazi:treatment:03DB7E6FFFE6EF53E55971AEFE390831" httpUri="http://treatment.plazi.org/id/03DB7E6FFFE6EF53E55971AEFE390831" lastPageId="5" lastPageNumber="354" pageId="3" pageNumber="352">
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<heading id="D0857815FFE6EF55E55971AEFC160FE3" box="[605,982,439,465]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="352" reason="2">
<taxonomicName id="4C72B4FAFFE6EF55E55971AEFC500FE3" authorityName="Nabozhenko" authorityYear="2005" box="[605,912,439,465]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="semnanicus" status="sp. nov.">
<emphasis id="B906136BFFE6EF55E55971AEFC500FE3" bold="true" box="[605,912,439,465]" pageId="3" pageNumber="352">Hedyphanes semnanicus</emphasis>
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<taxonomicNameLabel id="A235AE10FFE6EF55E49371AEFC160FE3" box="[919,982,439,465]" pageId="3" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
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(
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)
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Type material
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(m) is with the following labels: “
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, prov.
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”; “leg.
<collectorName id="2687AAAFFFE6EF55E4C77222FB890C62" box="[963,1097,571,592]" pageId="3" pageNumber="352">Gy. Fábián, L</collectorName>
. Nádai,
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&amp;
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” (
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)
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.
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(f) is with the following label: “
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,
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leg.
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” (
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). (
<collectorName id="2687AAAFFFE6EF55E40F726DFCF20CBB" box="[779,818,628,649]" pageId="3" pageNumber="352">The</collectorName>
names [
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refer to the same person.)
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</paragraph>
<caption id="DF0D9FF1FFE6EF55E63976D7FBB60932" ID-DOI="http://doi.org/10.5281/zenodo.12586005" ID-Zenodo-Dep="12586005" httpUri="https://zenodo.org/record/12586005/files/figure.png" pageId="3" pageNumber="352" startId="3.[317,359,1742,1763]" targetBox="[444,1145,738,1703]" targetPageId="3" targetType="figure">
<paragraph id="8BCDCF79FFE6EF55E63976D7FBB60932" blockId="3.[317,1269,1742,1792]" pageId="3" pageNumber="352">
<emphasis id="B906136BFFE6EF55E63976D7FE5408D1" bold="true" box="[317,404,1742,1763]" pageId="3" pageNumber="352">Figs 14.</emphasis>
<taxonomicName id="4C72B4FAFFE6EF55E69976D7FD6208D1" authorityName="Nabozhenko" authorityYear="2005" box="[413,674,1742,1763]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="mesopotamicus" status="sp. nov.">
<emphasis id="B906136BFFE6EF55E69976D7FD6208D1" box="[413,674,1742,1763]" italics="true" pageId="3" pageNumber="352">Hedyphanes mesopotamicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235AE10FFE6EF55E5A376D7FD1A08D1" box="[679,730,1742,1763]" pageId="3" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
: 1 = habitus of male, 2 = aedeagus, ventral view, 3 = aedeagus, lateral view, 4 = spiculum gastrale, ventral view. Scale bars = 1 mm
</paragraph>
</caption>
<paragraph id="8BCDCF79FFE1EF52E67A71ADFE4F0FD4" blockId="4.[317,1270,436,630]" pageId="4" pageNumber="353">
Description Male (
<figureCitation id="1349D3FCFFE1EF52E54C71ADFD440FFB" box="[584,644,436,457]" captionStart="Figs 510" captionStartId="4.[317,360,1732,1753]" captionTargetBox="[436,1150,693,1691]" captionTargetId="figure-112@4.[434,1153,690,1703]" captionTargetPageId="4" captionText="Figs 510. Hedyphanes semnanicus sp. n.: 5 = habitus of male, 6 = spiculum gastrale, ventral view, 7 = spiculum gastrale, lateral view, 8 = parameres, lateral view, 9 = parameres, ventral view, 10 = pe-" figureDoi="http://doi.org/10.5281/zenodo.12586007" httpUri="https://zenodo.org/record/12586007/files/figure.png" pageId="4" pageNumber="353">Fig. 5</figureCitation>
). Body slender, black, with greasy sheen. Length
<quantity id="4C8A629CFFE1EF52E36971ADFB740FFB" box="[1133,1204,436,457]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="353" unit="mm" value="16.0">16 mm</quantity>
, width
<quantity id="4C8A629CFFE1EF52E63971C8FE4B0FD4" box="[317,395,465,486]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="353" unit="mm" value="6.3">6.3 mm</quantity>
.
</paragraph>
<paragraph id="8BCDCF79FFE1EF52E67A71F7FC080921" blockId="4.[317,1270,436,630]" lastBlockId="4.[317,1270,1732,1811]" pageId="4" pageNumber="353">
Head widest at level of eyes. Eyes large, convex. Ratio of head width at level of eyes to distance between eyes 1.6. Anterior margin of clypeus straight. Lateral margin of head at junction of gena and clypeus with weak emargination. Genae weakly rounded. Frons weakly convex. Head without transverse depression along frontal suture. Punctation of head coarse, moderately dense (intervals between punctures on frons subequal to puncture diameter). Antennae long, with 3 apical segments nis, ventral view. Scale bars =
<quantity id="4C8A629CFFE1EF52E49576E7FC080921" box="[913,968,1790,1811]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="353" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
<caption id="DF0D9FF1FFE1EF52E63976DDFB3608C4" ID-DOI="http://doi.org/10.5281/zenodo.12586007" ID-Zenodo-Dep="12586007" httpUri="https://zenodo.org/record/12586007/files/figure.png" pageId="4" pageNumber="353" startId="4.[317,360,1732,1753]" targetBox="[436,1150,693,1691]" targetPageId="4" targetType="figure">
<paragraph id="8BCDCF79FFE1EF52E63976DDFB3608C4" blockId="4.[317,1270,1732,1811]" pageId="4" pageNumber="353">
<emphasis id="B906136BFFE1EF52E63976DDFE6308EB" bold="true" box="[317,419,1732,1753]" pageId="4" pageNumber="353">Figs 510.</emphasis>
<taxonomicName id="4C72B4FAFFE1EF52E6AD76DDFD5608EB" authorityName="Nabozhenko" authorityYear="2005" box="[425,662,1732,1753]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="semnanicus" status="sp. nov.">
<emphasis id="B906136BFFE1EF52E6AD76DDFD5608EB" box="[425,662,1732,1753]" italics="true" pageId="4" pageNumber="353">Hedyphanes semnanicus</emphasis>
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<taxonomicNameLabel id="A235AE10FFE1EF52E59F76DDFD1108EB" box="[667,721,1732,1753]" pageId="4" pageNumber="353" rank="species">sp. n.</taxonomicNameLabel>
: 5 = habitus of male, 6 = spiculum gastrale, ventral view, 7 = spiculum gastrale, lateral view, 8 = parameres, lateral view, 9 = parameres, ventral view, 10 = pe-
</paragraph>
</caption>
<paragraph id="8BCDCF79FFE0EF53E63971ADFCEE0C31" blockId="5.[317,1271,436,1033]" pageId="5" pageNumber="354">extending beyond base of pronotum. Segments 911 filiform, flattened, asymmetric. Length to width ratio of segments 211: 1, 4.9, 2.4, 2.3, 2.3, 2.4, 2.2, 1.6, 1.3, 1.7. 3rd segment of antennae 2 times as long as 4th. Segment 11 1.3 times as long as 10th.</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A7212FD820CFF" blockId="5.[317,1271,436,1033]" pageId="5" pageNumber="354">Pronotum weakly transverse, widest at middle. Disc of pronotum strongly convex. Outer margins of pronotum emarginated only near angles. All margins of pronotum weakly rounded. Lateral sides of pronotum finely margined near angles; anterior margin and base very finely margined. Punctation of pronotum moderately coarse, diameter of punctures on the middle of disc about equal to distance between them; diameter of punctures on sides larger than distance between them. Propleura with coarse, not rasp-formed punctation; punctation on lateral portion sparser. Prosternal process weakly projecting.</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A72CDFDC00D11" blockId="5.[317,1271,436,1033]" pageId="5" pageNumber="354">Elytra oblong-oval. Disc of elytra moderately convex, only narrowly depressed near suture. Striae of punctures weak; interstrial punctation almost the same as punctures in striae. Elytra with transverse wrinkles.</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A7332FE0B0D6F" blockId="5.[317,1271,436,1033]" pageId="5" pageNumber="354">
Abdominal ventrites with fine punctation. Spiculum gastrale:
<figureCitation id="1349D3FCFFE0EF53E4C97332FBE30D72" box="[973,1059,811,832]" captionStart="Figs 510" captionStartId="4.[317,360,1732,1753]" captionTargetBox="[436,1150,693,1691]" captionTargetId="figure-112@4.[434,1153,690,1703]" captionTargetPageId="4" captionText="Figs 510. Hedyphanes semnanicus sp. n.: 5 = habitus of male, 6 = spiculum gastrale, ventral view, 7 = spiculum gastrale, lateral view, 8 = parameres, lateral view, 9 = parameres, ventral view, 10 = pe-" figureDoi="http://doi.org/10.5281/zenodo.12586007" httpUri="https://zenodo.org/record/12586007/files/figure.png" pageId="5" pageNumber="354">Figs 67</figureCitation>
. Parameres:
<figureCitation id="1349D3FCFFE0EF53E3987332FB330D72" box="[1180,1267,811,832]" captionStart="Figs 510" captionStartId="4.[317,360,1732,1753]" captionTargetBox="[436,1150,693,1691]" captionTargetId="figure-112@4.[434,1153,690,1703]" captionTargetPageId="4" captionText="Figs 510. Hedyphanes semnanicus sp. n.: 5 = habitus of male, 6 = spiculum gastrale, ventral view, 7 = spiculum gastrale, lateral view, 8 = parameres, lateral view, 9 = parameres, ventral view, 10 = pe-" figureDoi="http://doi.org/10.5281/zenodo.12586007" httpUri="https://zenodo.org/record/12586007/files/figure.png" pageId="5" pageNumber="354">Figs 89</figureCitation>
. Penis:
<figureCitation id="1349D3FCFFE0EF53E67B7351FE080D6F" box="[383,456,840,861]" captionStart="Figs 510" captionStartId="4.[317,360,1732,1753]" captionTargetBox="[436,1150,693,1691]" captionTargetId="figure-112@4.[434,1153,690,1703]" captionTargetPageId="4" captionText="Figs 510. Hedyphanes semnanicus sp. n.: 5 = habitus of male, 6 = spiculum gastrale, ventral view, 7 = spiculum gastrale, lateral view, 8 = parameres, lateral view, 9 = parameres, ventral view, 10 = pe-" figureDoi="http://doi.org/10.5281/zenodo.12586007" httpUri="https://zenodo.org/record/12586007/files/figure.png" pageId="5" pageNumber="354">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A737DFC8F0D81" blockId="5.[317,1271,436,1033]" pageId="5" pageNumber="354">Legs long; anterior and middle tibiae straight, posterior tibiae weakly curved inside in basal third. Anterior tarsi of male widened, its segments 12 transverse. Middle tarsi weakly widened, with transverse segments. All tarsi with dense rufous hairs.</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A73A2FCC50DE2" blockId="5.[317,1271,436,1033]" box="[382,773,955,976]" pageId="5" pageNumber="354">
Body length
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, body width
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.
</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A73C1FB200A3B" blockId="5.[317,1271,436,1033]" pageId="5" pageNumber="354">Female. Body wider and more robust. Antennae shorter, when posteriorly extended reaching beyond base of pronotum with one segment. Elytra without transverse wrinkles. Tarsi not widened.</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A7428FC820B2C" blockId="5.[317,1270,1073,1310]" pageId="5" pageNumber="354">
Diagnosis The new species is closely related to
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<emphasis id="B906136BFFE0EF53E3077428FB350A79" box="[1027,1269,1073,1099]" italics="true" pageId="5" pageNumber="354">Hedyphanes besseri</emphasis>
<bibRefCitation id="EFE3B288FFE0EF53E639744DFDF10A5C" author="FALDERMANN, F." box="[317,561,1108,1134]" pageId="5" pageNumber="354" refId="ref2246" refString="FALDERMANN, F. (1837) Fauna entomologica Trans-Caucasica. Coleoptera. Pars II. Auguste Semen, Moscou, 433 pp." type="book" year="1837">
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, 1837
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, but differs from it in the more widened body and the absence of emarginations on the outer margins of pronotum. Except
<taxonomicName id="4C72B4FAFFE0EF53E4FF7461FBB10AA0" authorityName="FALDERMANN" authorityYear="1837" box="[1019,1137,1144,1170]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="354" phylum="Arthropoda" rank="species" species="besseri">
<emphasis id="B906136BFFE0EF53E4FF7461FBB10AA0" box="[1019,1137,1144,1170]" italics="true" pageId="5" pageNumber="354">H. besseri</emphasis>
</taxonomicName>
, which has rectangular and transversal tarsomeres 14, only
<taxonomicName id="4C72B4FAFFE0EF53E4807482FBF20A87" authorityName="Nabozhenko" authorityYear="2005" box="[900,1074,1179,1205]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="354" phylum="Arthropoda" rank="species" species="semnanicus" status="sp. nov.">
<emphasis id="B906136BFFE0EF53E4807482FBF20A87" box="[900,1074,1179,1205]" italics="true" pageId="5" pageNumber="354">H. semnanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235AE10FFE0EF53E33F7482FBBA0A87" box="[1083,1146,1179,1205]" pageId="5" pageNumber="354" rank="species">sp. n.</taxonomicNameLabel>
has transversal head-shaped tarsomeres 13. But
<taxonomicName id="4C72B4FAFFE0EF53E41274A7FC020AEA" authorityName="Nabozhenko" authorityYear="2005" box="[790,962,1214,1240]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="354" phylum="Arthropoda" rank="species" species="semnanicus" status="sp. nov.">
<emphasis id="B906136BFFE0EF53E41274A7FC020AEA" box="[790,962,1214,1240]" italics="true" pageId="5" pageNumber="354">H. semnanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235AE10FFE0EF53E4CD74A7FBC80AEA" box="[969,1032,1214,1240]" pageId="5" pageNumber="354" rank="species">sp. n.</taxonomicNameLabel>
also differs from
<taxonomicName id="4C72B4FAFFE0EF53E3DD74A7FE500AC9" authorityName="FALDERMANN" authorityYear="1837" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="354" phylum="Arthropoda" rank="species" species="besseri">
<emphasis id="B906136BFFE0EF53E3DD74A7FE500AC9" italics="true" pageId="5" pageNumber="354">H. besseri</emphasis>
</taxonomicName>
by the structure of 11th antennal segment, the later having more elongate and less asymmetric 11th antennal segment.
</paragraph>
<paragraph id="8BCDCF79FFE0EF53E4177549FCDF0B57" blockId="5.[787,799,1360,1381]" box="[787,799,1360,1381]" pageId="5" pageNumber="354">*</paragraph>
<paragraph id="8BCDCF79FFE0EF53E67A7562FE390831" blockId="5.[317,1271,1403,1539]" pageId="5" pageNumber="354">
<emphasis id="B906136BFFE0EF53E67A7562FDF60BA2" box="[382,566,1403,1424]" italics="true" pageId="5" pageNumber="354">Acknowledgements</emphasis>
The author is much obliged to Prof. G. S.
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(ZIN), Dr. L.
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(DEI), Dr. M. Y.
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(Institute of Zoology of the Academy of Sciences of
<collectingCountry id="F3658FE9FFE0EF53E3987581FB330B9F" box="[1180,1267,1432,1453]" name="Armenia" pageId="5" pageNumber="354">Armenia</collectingCountry>
,
<collectingRegion id="49B6019BFFE0EF53E63975ADFE430BFB" box="[317,387,1460,1481]" country="Armenia" name="Yerevan" pageId="5" pageNumber="354">Erevan</collectingRegion>
) and Dr. O.
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(HNHM) for providing the material. The author also thanks Dr. A.
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.
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(Department of Zoology, Field Museum of Natural History, Chicago,
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) for reviewing this paper.
</paragraph>
</subSubSection>
</treatment>
</document>