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<document id="3829F5A638DE2A64E5F3F5A3793BA630" ID-DOI="10.1515/bot-2023-0037" ID-ISSN="1437-4323" ID-Zenodo-Dep="11000291" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1713580552975" checkinUser="felipe" docAuthor="Hernández-Becerril, David U., Pichardo-Velarde, Jorge G., Alonso-Rodríguez, Rosalba, Maciel-Baltazar, Ebodio, Morquecho, Lourdes, Esqueda-Lara, Karina, Barón-Campis, Sofía A. &amp; Quiroz-González, Nataly" docDate="2023" docId="03E50944FFB42A1B4E98E730FA3DFCF7" docLanguage="en" docName="BotMar.66.6.539-557.pdf" docOrigin="Botanica Marina (Warsaw, Poland) 66 (6)" docSource="http://dx.doi.org/10.1515/bot-2023-0037" docStyle="DocumentStyle:852F6101A3F2630EFBDAB42A7703A325.1:BotMar.2015-.journal_article.open" docStyleId="852F6101A3F2630EFBDAB42A7703A325" docStyleName="BotMar.2015-.journal_article.open" docStyleVersion="1" docTitle="Alexandrium Halim 1960" docType="treatment" docVersion="4" lastPageNumber="542" masterDocId="FFDC713CFFB72A184E3EE248FF90FD3F" masterDocTitle="Diversity and distribution of species of the planktonic dinoflagellate genus Alexandrium (Dinophyta) from the tropical and subtropical Mexican Pacific Ocean" masterLastPageNumber="557" masterPageNumber="539" pageNumber="542" updateTime="1714506209649" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="26819292BEB24FD60CA31B5EF98EB7B8">Diversity and distribution of species of the planktonic dinoflagellate genus Alexandrium (Dinophyta) from the tropical and subtropical Mexican Pacific Ocean</mods:title>
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<paragraph id="8BF3B852FFB42A1B4E98E730FEFEF88D" blockId="3.[166,701,1400,1458]" pageId="3" pageNumber="542">
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3.1.1
<emphasis id="B9386440FFB42A1B4EDDE730FEE8F8B0" box="[227,376,1400,1423]" italics="true" pageId="3" pageNumber="542">
<taxonomicName id="4C4CC3D1FFB42A1B4EDDE730FEF5F8B0" authorityName="Halim" authorityYear="1960" box="[227,357,1400,1423]" class="Dinophyceae" family="Ostreopsidaceae" genus="Alexandrium" higherTaxonomySource="GBIF" kingdom="Chromista" order="Gonyaulacales" pageId="3" pageNumber="542" phylum="Myzozoa" rank="genus">Alexandrium</taxonomicName>
a
</emphasis>
</emphasis>
ffi
<emphasis id="B9386440FFB42A1B4FB3E730FEFEF88D" bold="true" pageId="3" pageNumber="542">
ne (Inoue
<emphasis id="B9386440FFB42A1B4FC5E730FD80F8B0" bold="true" box="[507,528,1400,1423]" italics="true" pageId="3" pageNumber="542">et</emphasis>
Fukuyo) Balech (
<figureCitation id="1377A4D7FFB42A1B4ED7E7D3FEF8F88D" box="[233,360,1434,1458]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figures 28</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356EBD9FFB42A1B4E98E7A8FA3DFCF7" pageId="3" pageNumber="542" type="description">
<paragraph id="8BF3B852FFB42A1B4E98E7A8FD3DFB04" blockId="3.[166,794,1503,1941]" pageId="3" pageNumber="542">
References:
<bibRefCitation id="EFDDC5A3FFB42A1B4F16E7A8FE36F8C8" author="Balech, E." box="[296,422,1504,1527]" pageId="3" pageNumber="542" pagination="151" refId="ref11445" refString="Balech, E. (1995). The genus Alexandrium Halim (Dino fl agellata). Sherkin Island Marine Station, Sherkin Island, p. 151." type="book chapter" year="1995">Balech 1995</bibRefCitation>
, 55, pl. XIII, figs 2036;
<bibRefCitation id="EFDDC5A3FFB42A1B4C99E7A9FF47FB27" author="Usup, G. &amp; Leaw, C. P. &amp; Ahmad, A. &amp; Lim, P. T." pageId="3" pageNumber="542" pagination="265 - 275" refId="ref15102" refString="Usup, G., Leaw, C. P., Ahmad, A., and Lim, P. T. (2002). Alexandrium (Dinophyceae) species in Malaysian waters. Harmful Algae 1: 265 - 275." type="journal article" year="2002">Usup et al. 2002</bibRefCitation>
, 267, figs 2 AD;
<bibRefCitation id="EFDDC5A3FFB42A1B4F43E44BFD94FB27" author="Gu, H. &amp; Zeng, N. &amp; Liu, T. &amp; Yang, W. &amp; Muller, A. &amp; Krock, B." box="[381,516,1539,1561]" pageId="3" pageNumber="542" pagination="68 - 81" refId="ref12162" refString="Gu, H., Zeng, N., Liu, T., Yang, W., Muller, A., and Krock, B. (2013). Morphology, toxicity, and phylogeny of Alexandrium (Dinophyceae) species along the coast of China. Harmful Algae 27: 68 - 81." type="journal article" year="2013">Gu et al. 2013</bibRefCitation>
, 72, figs 2 AD;
<bibRefCitation id="EFDDC5A3FFB42A1B4CA0E44BFF46FB04" author="Lassus, P. &amp; Chomerat, N. &amp; Hess, P. &amp; Nezan, E." pageId="3" pageNumber="542" pagination="1 - 523" refId="ref13215" refString="Lassus, P., Chomerat, N., Hess, P., and Nezan, E. (2016). Toxic and harmful microalgae of the World Ocean. Micro-algues toxiques et nuisibles de l' Ocean Mondial, IOC Manuals and guides, 68 (Bilingual English / French). International Society for the Study of Harmful Algae / Intergovernmental Oceanographic Commission of UNESCO, Copenhagen, pp. 1 - 523, 54 pls." type="book chapter" year="2016">Lassus et al. 2016</bibRefCitation>
, 42, pl. 2 AE;
<bibRefCitation id="EFDDC5A3FFB42A1B4F52E46DFD97FB05" author="Kim, E. S. &amp; Li, Z. &amp; Oh, S. J. &amp; Yoon, Y. H. &amp; Shin, H. H." box="[364,519,1573,1595]" pageId="3" pageNumber="542" pagination="427 - 437" refId="ref12879" refString="Kim, E. S., Li, Z., Oh, S. J., Yoon, Y. H., and Shin, H. H. (2017). Morphological identification of Alexandrium species (Dinophyceae) from Jinhae- Masan Bay, Korea. Ocean Sci. J. 52: 427 - 437." type="journal article" year="2017">Kim et al. 2017</bibRefCitation>
, 428, figs 1 AL.
</paragraph>
<paragraph id="8BF3B852FFB42A1B4EEDE400FD6DFB61" blockId="3.[166,794,1503,1941]" box="[211,765,1607,1630]" pageId="3" pageNumber="542">
Synonyms =
<taxonomicName id="4C4CC3D1FFB42A1B4F67E40FFD6DFB61" authority="Inouye et Fukuyo" authorityName="Inouye et Fukuyo" box="[345,765,1607,1630]" class="Dinophyceae" family="Ostreopsidaceae" genus="Protogonyaulax" higherTaxonomySource="GBIF" kingdom="Chromista" order="Gonyaulacales" pageId="3" pageNumber="542" phylum="Myzozoa" rank="species" species="affine">
<emphasis id="B9386440FFB42A1B4F67E40FFDAEFB62" bold="true" box="[345,574,1607,1630]" italics="true" pageId="3" pageNumber="542">Protogonyaulax affine</emphasis>
Inouye
<emphasis id="B9386440FFB42A1B4CAAE400FD37FB62" bold="true" box="[660,679,1608,1629]" italics="true" pageId="3" pageNumber="542">et</emphasis>
Fukuyo
</taxonomicName>
</paragraph>
<paragraph id="8BF3B852FFB42A1B4EEDE422FA93FDC6" blockId="3.[166,794,1503,1941]" lastBlockId="3.[824,1453,193,456]" pageId="3" pageNumber="542">
Chain-forming species, with chains of 38 cells observed (
<figureCitation id="1377A4D7FFB42A1B4F2BE4C5FE26FB9C" box="[277,438,1677,1699]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figures 2 and 3</figureCitation>
). Individual cells of medium-size, slightly longer than wide (
<figureCitation id="1377A4D7FFB42A1B4FFFE4E7FDF6FBF9" box="[449,614,1711,1734]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figures 2 and 3</figureCitation>
). The epitheca is domed to conical, cingulum excavated and the hypotheca is concave (
<figureCitation id="1377A4D7FFB42A1B4F34E4BDFEF5FA34" box="[266,357,1781,1803]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figure 4</figureCitation>
). The first apical plate (1) is rhomboidal, in contact with Po, and with a small ventral pore on its right margin (
<figureCitation id="1377A4D7FFB42A1B4F3DE572FE3DFA6F" box="[259,429,1849,1872]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figures 5 and 6</figureCitation>
). The apical complex pore is conspicuous, with a large connecting pore and a relatively small foramen, both aligned in the same axis (
<figureCitation id="1377A4D7FFB42A1B4CB5E537FCD5FDE9" box="[651,837,193,1941]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figures 6 and 7</figureCitation>
). The posterior sulcal plate (Sp) has a small pore and the two anterior ends are projected (
<figureCitation id="1377A4D7FFB42A1B4AA2E2ABFB66FDC6" box="[1180,1270,227,249]" captionStart="Figures 28" captionStartId="4.[1018,1085,1697,1716]" captionTargetBox="[138,985,1298,1944]" captionTargetId="figure-451@4.[136,988,1295,1945]" captionTargetPageId="4" captionText="Figures 28: Alexandrium affine, LM. (2, 3) Chains of 8 and 3 cells, respectively. (4) An empty cell showing only the theca in ventral view. (5) Epitheca in ventral view showing the ventral pore (arrow) in the first apical plate (1). (6) Epitheca with Po and 1 showing the ventral pore (arrow). (7, 8) Po and posterior sulcal plate (Sp) (with a connecting pore), respectively." figureDoi="http://doi.org/10.5281/zenodo.11000295" httpUri="https://zenodo.org/record/11000295/files/figure.png" pageId="3" pageNumber="542">Figure 8</figureCitation>
).
</paragraph>
<paragraph id="8BF3B852FFB42A1B4D5BE34EFC54FC01" blockId="3.[824,1453,193,456]" pageId="3" pageNumber="542">Measurements: 3043 µm length (L), 3343 µm width (W) (Table 1).</paragraph>
<paragraph id="8BF3B852FFB42A1B4D5BE303FBC5FC99" blockId="3.[824,1453,193,456]" pageId="3" pageNumber="542">
Toxicity: this is a species that produces saxitoxins at very low concentrations, so they do not represent a risk of poisoning (
<bibRefCitation id="EFDDC5A3FFB42A1B4D92E3D8FBDBFC99" author="Kim, H. - S. &amp; Park, H. &amp; Wang, H. &amp; Kim, T. &amp; Ki, J. - S." box="[940,1099,400,422]" pageId="3" pageNumber="542" pagination="105874" refId="ref12942" refString="Kim, H. - S., Park, H., Wang, H., Kim, T., and Ki, J. - S. (2023). Saxitoxinsproducing potential of the marine dinoflagellate Alexandrium a ffi ne and its environmental implications revealed by toxins and transcriptome profiling. Mar. Environ. Res. 185: 105874." type="journal article" year="2023">Kim et al. 2023</bibRefCitation>
).
</paragraph>
<paragraph id="8BF3B852FFB42A1B4D5BE3FAFA3DFCF7" blockId="3.[824,1453,193,456]" box="[869,1453,434,456]" pageId="3" pageNumber="542">
Distribution: in the southern part of the Gulf of
<collectingRegion id="498876B0FFB42A1B4B7EE3FAFA39FCF7" box="[1344,1449,434,456]" country="United States of America" name="California" pageId="3" pageNumber="542">California</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
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