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<emphasis id="B913EADDFFAAFFFFCC8B00EBFE71FF3D" bold="true" box="[151,401,151,178]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
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(
<figureCitation id="135C2A4AFFAAFFFFCC8300C7FEE9FF59" box="[159,265,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support &lt;80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="44" pageNumber="45">Figs. 18</figureCitation>
)
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http://zoobank.org/NomenclaturalActs/
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, figs. 14.
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<emphasis id="B913EADDFFAAFFFFCC8B013EFE8AFED6" box="[151,362,322,346]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
(Abrams and Harvey)
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:
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Abrams
<emphasis id="B913EADDFFAAFFFFCEAC013FFD3CFED6" box="[688,732,322,346]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
. 2019
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MPE 106532: 8, fig. 2.
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<emphasis id="B913EADDFFAAFFFFCC8B01F4FE63FE2E" bold="true" box="[151,387,392,418]" pageId="44" pageNumber="45">Material examined.</emphasis>
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male.
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,
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(
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).
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</paragraph>
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<emphasis id="B913EADDFFAAFFFFCCDB01ADFED9FE66" box="[199,313,465,490]" italics="true" pageId="44" pageNumber="45">
<typeStatus id="54DC886DFFAAFFFFCCDB01ADFED9FE66" box="[199,313,465,490]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="F370765FFFAAFFFFCD5701ADFE0EFE67" box="[331,494,465,491]" name="Australia" pageId="44" pageNumber="45">AUSTRALIA</collectingCountry>
:
<emphasis id="B913EADDFFAAFFFFCDE301ADFD2DFE66" box="[511,717,464,490]" italics="true" pageId="44" pageNumber="45">
<collectingRegion id="49A3F82DFFAAFFFFCDE301ADFD2DFE66" box="[511,717,464,490]" country="Australia" name="Western Australia" pageId="44" pageNumber="45">Western Australia</collectingRegion>
</emphasis>
:
<specimenCount id="9D61FD46FFAAFFFFCEFE01ADFCF3FE67" box="[738,787,465,491]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="6BBCACB5FFAAFFFFCF3B01ADFC78FE67" box="[807,920,465,491]" pageId="44" pageNumber="45">Bungaroo</collectingMunicipality>
,
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SE. of
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, 215607”S, 1162654”E,
<date id="FFD9100FFFAAFFFFCD320188FE2EFD83" box="[302,462,500,527]" pageId="44" pageNumber="45" value="2011-04-15">
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,
<collectorName id="26925319FFAAFFFFCE820189FCCCFD82" box="[670,812,500,527]" pageId="44" pageNumber="45">J. Alexander</collectorName>
, and
<collectorName id="26925319FFAAFFFFCF750189FC39FD83" box="[873,985,501,527]" pageId="44" pageNumber="45">S. Werner</collectorName>
(
<collectionCode id="ED76AE0AFFAAFFFFCFF40189FBCFFD83" box="[1000,1071,501,527]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="45" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFAAFFFFC82A0189FB40FD83" box="[1078,1184,501,527]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="44" pageNumber="45">T114972</specimenCode>
)
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;
<materialsCitation id="3B0F3C92FFAAFFFFC8A80189FEE8FDDB" ID-GBIF-Occurrence="3013449360" collectingDate="2011-04-15" collectionCode="WAM" collectorName="J. Alexander &amp; S. Werner" country="Australia" latitude="-21.943611" location="Pannawonica" longLatPrecision="21" longitude="116.44417" municipality="Bungaroo" pageId="44" pageNumber="45" specimenCode="T114969" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
<specimenCount id="9D61FD46FFAAFFFFC8A80189FB01FD83" box="[1204,1249,501,527]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="6BBCACB5FFAAFFFFC8F20189FABFFD83" box="[1262,1375,501,527]" pageId="44" pageNumber="45">Bungaroo</collectingMunicipality>
,
<quantity id="4C9F9B2AFFAAFFFFC9770188FF5BFDBE" metricMagnitude="4" metricUnit="m" metricValue="3.53" pageId="44" pageNumber="45" unit="km" value="35.3">35.3 km</quantity>
SE. of
<location id="8EB86014FFAAFFFFCD100264FE44FDBE" LSID="urn:lsid:plazi:treatment:03CE87D9FFAAFFFFCC8B00EBFCE1FB1B:8EB86014FFAAFFFFCD100264FE44FDBE" box="[268,420,536,562]" country="Australia" latitude="-21.943611" longLatPrecision="21" longitude="116.44417" municipality="Bungaroo" name="Pannawonica" pageId="44" pageNumber="45" stateProvince="Western Australia">Pannawonica</location>
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,
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<collectorName id="26925319FFAAFFFFC8580265FB30FDBE" box="[1092,1232,536,563]" pageId="44" pageNumber="45">J. Alexander</collectorName>
,
<collectorName id="26925319FFAAFFFFC8C70265FAA9FDBF" box="[1243,1353,537,563]" pageId="44" pageNumber="45">S. Werner</collectorName>
(
<collectionCode id="ED76AE0AFFAAFFFFC94A0265FA7DFDBF" box="[1366,1437,537,563]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="45" type="Museum">WAM</collectionCode>
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)
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.
</paragraph>
<paragraph id="8BD836CFFFAAFFFFCCDB021CFB87FDF7" blockId="44.[151,1437,392,1175]" box="[199,1127,608,635]" pageId="44" pageNumber="45">
<emphasis id="B913EADDFFAAFFFFCCDB021CFE96FDF6" box="[199,374,608,634]" italics="true" pageId="44" pageNumber="45">Other material.</emphasis>
<collectingCountry id="F370765FFFAAFFFFCD67021DFDFEFDF7" box="[379,542,609,635]" name="Australia" pageId="44" pageNumber="45">AUSTRALIA</collectingCountry>
:
<emphasis id="B913EADDFFAAFFFFCE34021DFD10FDF6" box="[552,752,608,634]" italics="true" pageId="44" pageNumber="45">
<collectingRegion id="49A3F82DFFAAFFFFCE34021DFD10FDF6" box="[552,752,608,634]" country="Australia" name="Western Australia" pageId="44" pageNumber="45">Western Australia</collectingRegion>
</emphasis>
: See
<bibRefCitation id="EFF64B3EFFAAFFFFCF31021CFB83FDF7" author="Abrams, K. M. &amp; Harvey, M. S." box="[813,1123,608,635]" pageId="44" pageNumber="45" pagination="132 - 136" refId="ref46052" refString="Abrams, K. M. &amp; Harvey, M. S. (2015) A new troglobitic schizomid (Hubbardiidae: Paradraculoides) from the Pilbara region, Western Australia. Records of the Western Australian Museum, 30 (2), 132 - 136. https: // doi. org / 10.18195 / issn. 0312 - 3162.30 (2). 2015.132 - 136" type="journal article" year="2015">Abrams and Harvey (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFAAFFFFCCDB02F8FB8AFBA6" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8BD836CFFFAAFFFFCCDB02F8FB8AFBA6" blockId="44.[151,1437,392,1175]" pageId="44" pageNumber="45">
<emphasis id="B913EADDFFAAFFFFCCDB02F8FEA2FD12" bold="true" box="[199,322,644,670]" pageId="44" pageNumber="45">Diagnosis.</emphasis>
The shape of the male flagellum of
<taxonomicName id="4C674D4CFFAAFFFFCEEE02F8FC3FFD12" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[754,991,644,670]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFAAFFFFCEEE02F8FC3FFD12" box="[754,991,644,670]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C674D4CFFAAFFFFC8E202F9FA89FD12" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1278,1385,644,670]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B913EADDFFAAFFFFC8E202F9FA89FD12" box="[1278,1385,644,670]" italics="true" pageId="44" pageNumber="45">D. affinis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFAAFFFFCC8B02D5FEDAFD4E" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,314,680,706]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFAAFFFFCC8B02D5FEDAFD4E" box="[151,314,680,706]" italics="true" pageId="44" pageNumber="45">D. gnophicola</emphasis>
</taxonomicName>
, particularly in the distally rounded end of the flagellum, which is tapering in most other similar described
<taxonomicName id="4C674D4CFFAAFFFFCD1702B0FE75FD6A" box="[267,405,716,742]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFAAFFFFCD1702B0FE75FD6A" box="[267,405,716,742]" italics="true" pageId="44" pageNumber="45">Draculoides</emphasis>
</taxonomicName>
species (
<taxonomicName id="4C674D4CFFAAFFFFCE1E02B1FD4FFD6A" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[514,687,716,742]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFAAFFFFCE1E02B1FD4FFD6A" box="[514,687,716,742]" italics="true" pageId="44" pageNumber="45">D. anachoretus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCEDC02B1FCB9FD6A" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[704,857,716,742]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="cochranus">
<emphasis id="B913EADDFFAAFFFFCEDC02B1FCB9FD6A" box="[704,857,716,742]" italics="true" pageId="44" pageNumber="45">D. cochranus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCF7502B1FC3FFD6A" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[873,991,716,742]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFAAFFFFCF7502B1FC3FFD6A" box="[873,991,716,742]" italics="true" pageId="44" pageNumber="45">D. kryptus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCFF302B1FB24FD6A" authorityName="Abrams and Harvey" authorityYear="2020" box="[1007,1220,716,742]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFAAFFFFCFF302B1FB24FD6A" box="[1007,1220,716,742]" italics="true" pageId="44" pageNumber="45">D. mckechnieorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFC8C902B1FA7CFD6A" box="[1237,1436,717,742]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="noctigrassator">
<emphasis id="B913EADDFFAAFFFFC8C902B1FA7CFD6A" box="[1237,1436,717,742]" italics="true" pageId="44" pageNumber="45">D. noctigrassator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFAAFFFFCCD6028DFE92FC86" authorityName="Abrams and Harvey" authorityYear="2020" box="[202,370,752,778]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFAAFFFFCCD6028DFE92FC86" box="[202,370,752,778]" italics="true" pageId="44" pageNumber="45">D. warramboo</emphasis>
</taxonomicName>
).
<taxonomicName id="4C674D4CFFAAFFFFCD90028CFD98FC86" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[396,632,752,778]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFAAFFFFCD90028CFD98FC86" box="[396,632,752,778]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C674D4CFFAAFFFFCF09028DFC6BFC86" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[789,907,752,778]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFAAFFFFCF09028DFC6BFC86" box="[789,907,752,778]" italics="true" pageId="44" pageNumber="45">D. bythius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFAAFFFFCFD4028DFB8BFC86" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[968,1131,752,778]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFAAFFFFCFD4028DFB8BFC86" box="[968,1131,752,778]" italics="true" pageId="44" pageNumber="45">D. gnophicola</emphasis>
</taxonomicName>
in the distal placement of dm4 on the margin of the flagellum (dm4 is sub-distal and anterior to dl
<quantity id="4C9F9B2AFFAAFFFFCFA70368FC04FCA2" box="[955,996,788,814]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="44" pageNumber="45" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C674D4CFFAAFFFFCFF70369FBB3FCA2" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1003,1107,788,814]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B913EADDFFAAFFFFCFF70369FBB3FCA2" box="[1003,1107,788,814]" italics="true" pageId="44" pageNumber="45">D. affinis</emphasis>
</taxonomicName>
and slightly sub-distal in and
<taxonomicName id="4C674D4CFFAAFFFFCC8B0345FED9FCDE" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,313,824,850]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFAAFFFFCC8B0345FED9FCDE" box="[151,313,824,850]" italics="true" pageId="44" pageNumber="45">D. gnophicola</emphasis>
</taxonomicName>
). Females of this species differ from all other females by the rectangular-shaped backward folding genital gonopod.
<taxonomicName id="4C674D4CFFAAFFFFCD420320FDAAFCFA" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[350,586,860,886]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFAAFFFFCD420320FDAAFCFA" box="[350,586,860,886]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFAAFFFFCFD40320FBB2FCFA" box="[968,1106,860,886]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFAAFFFFCFD40320FBB2FCFA" box="[968,1106,860,886]" italics="true" pageId="44" pageNumber="45">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at COI by the 50bp mini-barcode shown in
<figureCitation id="135C2A4AFFAAFFFFCE9E03FCFD02FC17" box="[642,738,896,923]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="44" pageNumber="45">Figure 3</figureCitation>
.
<taxonomicName id="4C674D4CFFAAFFFFCEF303FCFC39FC16" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[751,985,896,922]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFAAFFFFCEF303FCFC39FC16" box="[751,985,896,922]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFAAFFFFC95703FDFF3AFC32" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFAAFFFFC95703FDFF3AFC32" italics="true" pageId="44" pageNumber="45">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at 12S (
<figureCitation id="135C2A4AFFAAFFFFCE6503D8FD5BFC33" box="[633,699,932,959]" captionStart="FIGURE 5" captionStartId="20.[151,250,1216,1240]" captionTargetBox="[158,1424,183,1189]" captionTargetId="figure-20@20.[151,1436,181,1192]" captionTargetPageId="20" captionText="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416863" httpUri="https://zenodo.org/record/4416863/files/figure.png" pageId="44" pageNumber="45">Fig. 5</figureCitation>
) except for
<taxonomicName id="4C674D4CFFAAFFFFCF5F03D9FC3EFC32" authorityName="Abrams and Harvey" authorityYear="2020" box="[835,990,932,958]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="piscivultus">
<emphasis id="B913EADDFFAAFFFFCF5F03D9FC3EFC32" box="[835,990,932,958]" italics="true" pageId="44" pageNumber="45">D. piscivultus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C674D4CFFAAFFFFCFF703D8FB35FC32" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[1003,1237,932,958]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFAAFFFFCFF703D8FB35FC32" box="[1003,1237,932,958]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFAAFFFFCD2603B4FE24FC6E" box="[314,452,968,994]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFAAFFFFCD2603B4FE24FC6E" box="[314,452,968,994]" italics="true" pageId="44" pageNumber="45">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at ITS2 (
<figureCitation id="135C2A4AFFAAFFFFCF6A03B4FC5AFC6F" box="[886,954,968,995]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="44" pageNumber="45">Fig. 7</figureCitation>
) except for
<taxonomicName id="4C674D4CFFAAFFFFC85F03B5FB0FFC6E" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1091,1263,968,994]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFAAFFFFC85F03B5FB0FFC6E" box="[1091,1263,968,994]" italics="true" pageId="44" pageNumber="45">D. anachoretus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFC8E203B5FA92FC6E" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1278,1394,968,994]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFAAFFFFC8E203B5FA92FC6E" box="[1278,1394,968,994]" italics="true" pageId="44" pageNumber="45">D. bythius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFC99D03B5FEF5FB8A" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFAAFFFFC99D03B5FEF5FB8A" italics="true" pageId="44" pageNumber="45">D. gnophicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCD390391FE7BFB8A" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[293,411,1004,1030]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFAAFFFFCD390391FE7BFB8A" box="[293,411,1004,1030]" italics="true" pageId="44" pageNumber="45">D. kryptus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCDB70391FD60FB8A" authorityName="Abrams and Harvey" authorityYear="2020" box="[427,640,1004,1030]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFAAFFFFCDB70391FD60FB8A" box="[427,640,1004,1030]" italics="true" pageId="44" pageNumber="45">D. mckechnieorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCE8C0391FCD7FB8A" authorityName="Abrams and Harvey" authorityYear="2020" box="[656,823,1004,1030]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFAAFFFFCE8C0391FCD7FB8A" box="[656,823,1004,1030]" italics="true" pageId="44" pageNumber="45">D. warramboo</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCF5A0391FC03FB8A" authorityName="Abrams and Harvey" authorityYear="2020" box="[838,995,1004,1030]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="immortalis">
<emphasis id="B913EADDFFAAFFFFCF5A0391FC03FB8A" box="[838,995,1004,1030]" italics="true" pageId="44" pageNumber="45">D. immortalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFCFE80391FB74FB8A" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[1012,1172,1004,1030]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="belalugosii">
<emphasis id="B913EADDFFAAFFFFCFE80391FB74FB8A" box="[1012,1172,1004,1030]" italics="true" pageId="44" pageNumber="45">D. belalugosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFC8B80391FA92FB8A" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[1188,1394,1004,1030]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFAAFFFFC8B80391FA92FB8A" box="[1188,1394,1004,1030]" italics="true" pageId="44" pageNumber="45">D. christopherleei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFAAFFFFC99D0391FEEFFBA6" authorityName="Abrams and Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="piscivultus">
<emphasis id="B913EADDFFAAFFFFC99D0391FEEFFBA6" italics="true" pageId="44" pageNumber="45">D. piscivultus</emphasis>
</taxonomicName>
and
<emphasis id="B913EADDFFAAFFFFCD59046DFE2AFBA6" box="[325,458,1040,1066]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="4C674D4CFFAAFFFFCD59046DFE27FBA6" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[325,455,1040,1066]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="akashae">D. akashae</taxonomicName>
,
</emphasis>
which are not distinguishable using the ITS2 mini-barcode.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFAAFFFFCCDB0448FD28FBC3" box="[199,712,1076,1103]" pageId="44" pageNumber="45" type="description">
<paragraph id="8BD836CFFFAAFFFFCCDB0448FD28FBC3" blockId="44.[151,1437,392,1175]" box="[199,712,1076,1103]" pageId="44" pageNumber="45">
<emphasis id="B913EADDFFAAFFFFCCDB0448FEB9FBC2" bold="true" box="[199,345,1076,1102]" pageId="44" pageNumber="45">Description.</emphasis>
See
<bibRefCitation id="EFF64B3EFFAAFFFFCD920448FD24FBC3" author="Abrams, K. M. &amp; Harvey, M. S." box="[398,708,1076,1103]" pageId="44" pageNumber="45" pagination="132 - 136" refId="ref46052" refString="Abrams, K. M. &amp; Harvey, M. S. (2015) A new troglobitic schizomid (Hubbardiidae: Paradraculoides) from the Pilbara region, Western Australia. Records of the Western Australian Museum, 30 (2), 132 - 136. https: // doi. org / 10.18195 / issn. 0312 - 3162.30 (2). 2015.132 - 136" type="journal article" year="2015">Abrams and Harvey (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFAAFFFFCCDB0425FCE1FB1B" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8BD836CFFFAAFFFFCCDB0425FCE1FB1B" blockId="44.[151,1437,392,1175]" pageId="44" pageNumber="45">
<emphasis id="B913EADDFFAAFFFFCCDB0425FEDBFBFF" bold="true" box="[199,315,1113,1139]" pageId="44" pageNumber="45">Remarks.</emphasis>
<taxonomicName id="4C674D4CFFAAFFFFCD5E0424FDCCFBFE" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[322,556,1112,1138]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFAAFFFFCD5E0424FDCCFBFE" box="[322,556,1112,1138]" italics="true" pageId="44" pageNumber="45">Draculoides eremius</emphasis>
</taxonomicName>
has only been found in the Bungaroo valley, situated south-east of Pannawonica in the Pilbara region of
<collectingRegion id="49A3F82DFFAAFFFFCDDB0401FD6FFB1A" box="[455,655,1148,1175]" country="Australia" name="Western Australia" pageId="44" pageNumber="45">Western Australia</collectingRegion>
(
<figureCitation id="135C2A4AFFAAFFFFCE820400FD13FB1B" box="[670,755,1148,1175]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="44" pageNumber="45">Fig. 1E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>