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<mods:title id="765E7DE6371DFC0A16522455E72B6D2D">Revision of the Mesobuthus caucasicus complex from Central Asia with descriptions of six new species (Scorpiones Buthidae)</mods:title>
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<mods:namePart id="C4A45A9A57AFC66C1F909A6B87AE226B">Kaiser, Ronald C.</mods:namePart>
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<mods:namePart id="C6F8734759529C0FDA2CB1DDA4983C11">Stewart, Alexander K.</mods:namePart>
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<mods:namePart id="BA3DA65BC3E4E54B4729C59B301C73A0">Graham, Matthew R.</mods:namePart>
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<taxonomicName id="4C0B3AC77A576A7FFF485932FD2F4038" authority="(Nordmann, 1840)" baseAuthorityName="Nordmann" baseAuthorityYear="1840" box="[203,719,861,888]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" sensu="stricto" species="caucasicus">
<emphasis id="B97F9D567A576A7FFF485932FE0A4037" bold="true" box="[203,490,861,888]" italics="true" pageId="10" pageNumber="9">Mesobuthus caucasicus</emphasis>
(
<bibRefCitation id="EF9A3CB57A576A7FFE7A5931FD274038" author="NORDMANN" box="[505,711,862,888]" pageId="10" pageNumber="9" refId="ref34768" refString="NORDMANN, A. 1840. Notice sur les Scorpions de la faune pontique. Pp. 751 - 752 in Voyage dans la Russie meridionale et la Crimee, par la Hongrie, la Valachie et la Moldavie, execute an 1837, sous la direction de M. Anatole de Demidoff, par Mm. de- Sainson, Le-Play, Huot, Leleveille, Raflet, Rousseau, de Nordmann et du Ponceau; dedie a S. M. Nicolas 1 - er, Empereur de toutes les Russies. Paris, vol. 3." type="journal volume" year="1840">Nordmann, 1840</bibRefCitation>
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<paragraph id="8BB441447A576A7FFF565910FDD140F5" blockId="10.[203,791,861,949]" pageId="10" pageNumber="9">
(
<figureCitation id="13305DC17A576A7FFF5E5910FEBE40D6" box="[221,350,895,918]" captionStart-0="Figures 2125" captionStart-1="Figures 2629" captionStart-2="Figures 3035" captionStart-3="Figures 3641" captionStartId-0="9.[192,278,1781,1805]" captionStartId-1="11.[192,278,1773,1797]" captionStartId-2="12.[192,278,1353,1377]" captionStartId-3="13.[192,278,1448,1472]" captionTargetBox-0="[198,1439,224,1743]" captionTargetBox-1="[209,1414,223,1739]" captionTargetBox-2="[191,1437,191,1324]" captionTargetBox-3="[207,1425,198,1394]" captionTargetId-0="figure-6@9.[192,1439,211,1752]" captionTargetId-1="figure-6@11.[192,1439,216,1744]" captionTargetId-2="figure-11@12.[192,1437,192,1324]" captionTargetId-3="figure-6@13.[192,1437,192,1405]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="Figures 2125: Mesobuthus caucasicus, holotype female. Dorsal (21) and ventral (22) views. Metasoma V and telson lateral (23), and metasoma and telson, lateral (24) and dorsal (25) views. The original labels are also included in the plate. Scale bar: 10 mm (2122)." captionText-1="Figures 2629: Mesobuthus caucasicus. Figures 2627. Male from Melekli, Turkey, dorsal (26) and ventral (27) views. Figures 2829. Female from Melekli, Turkey, dorsal (28) and ventral (29) views. Scale bar: 10 mm." captionText-2="Figures 3035: Mesobuthus caucasicus. Figures 30, 32, 34. Male from Melekli, Turkey, chelicerae, carapace and tergites IIII (30), sternopectinal region and sternites IIIIV (32), metasoma V and telson lateral (34). Figures 31, 33, 35. Female from Melekli, Turkey, chelicerae, carapace and tergites IIII (31), sternopectinal region and sternites IIIIV (33), metasoma V and telson lateral (35)." captionText-3="Figures 3641: Mesobuthus caucasicus. Figures 3638. Male from Melekli, Turkey, metasoma and telson, lateral (36), ventral (37), and dorsal (38) views. Figures 3941.Female from Melekli, Turkey, metasoma and telson, lateral (39), ventral (40), and dorsal (41) views." pageId="10" pageNumber="9">Figs. 2141</figureCitation>
,
<figureCitation id="13305DC17A576A7FFEEA5910FE7440D6" box="[361,404,895,918]" captionStart="Figures 265273" captionStartId="56.[192,278,1569,1593]" captionTargetBox="[208,1427,176,1539]" captionTargetId="figure-49@56.[192,1439,168,1540]" captionTargetPageId="56" captionText="Figures 265273: Mesobuthus, metasoma and telson ventral of males. Figure 265. M. brutus sp. n., holotype. Figure 266. M. caucasicus, Turkey. Figure 267. M. gorelovi sp. n., holotype. Figure 268. M. fuscus, Tajikistan. Figure 269. M. kaznakovi, Tajikistan. Figure 270. M. kreuzbergi sp. n., holotype. Figure 271. M. mischi sp. n., holotype. Figure 272. M. nenilini sp. n., holotype. Figure 273. M. parthorum, Afghanistan." pageId="10" pageNumber="9">266</figureCitation>
,
<figureCitation id="13305DC17A576A7FFE1D5910FE2940D6" box="[414,457,895,918]" captionStart="Figures 274284" captionStartId="57.[192,278,1468,1492]" captionTargetBox="[196,1428,206,1431]" captionTargetId="figure-6@57.[192,1440,192,1439]" captionTargetPageId="57" captionText="Figures 274284: Mesobuthus, tarsomeres of third leg. Figure 274. M. brutus sp. n., male holotype. Figure 275. M. caucasicus, female from Turkey. Figure 276. M. elenae sp. n., female holotype. Figure 277. M. gorelovi sp. n., female paratype from type locality. Figure 278. M. fuscus, female from Tajikistan. Figure 279. M. kaznakovi, male from Tajikistan. Figure 280. M. kreuzbergi sp. n., female paratype. Figure 281. M. mischi sp. n., male holotype. Figure 282. M. nenilini sp. n., female paratype. Figure 283284. M. parthorum, male from Ghazni, Afghanistan (283) and female from Kholm, Afghanistan (284)." pageId="10" pageNumber="9">275</figureCitation>
,
<figureCitation id="13305DC17A576A7FFE505910FDD440D6" box="[467,564,895,918]" captionStart="Figures 285303" captionStartId="58.[192,278,1281,1305]" captionTargetBox="[198,1431,202,1246]" captionTargetId="figure-11@58.[192,1440,192,1252]" captionTargetPageId="58" captionText="Figures 285303: Mesobuthus, pedipalp chela dorsal. Figure 285286. M. brutus sp. n., male holotype (285) and female paratype (286). Figure 287288. M. caucasicus, male (287) and female (288) from Turkey. Figure 289. M. elenae sp. n., female holotype. Figure 290291. M. fuscus, differently colored male (290) and female (291) from Tajikistan. Figure 292293. M. gorelovi sp. n., male holotype (292) and female paratype (293). Figure 294295. M. kaznakovi, male (294) and female (295) from Tajikistan. Figure 296297. M. kreuzbergi sp. n., male holotype (296) and female paratype (297). Figure 298299. M. mischi sp. n., male holotype (298) and female paratype (299). Figure 300301. M. nenilini sp. n., male holotype (300) and female paratype (301). Figure 302303. M. parthorum, male (302) and female (303) from Afghanistan." pageId="10" pageNumber="9">287288</figureCitation>
,
<figureCitation id="13305DC17A576A7FFDBD5910FD8940D6" box="[574,617,895,918]" captionStart="Figures 304315" captionStartId="59.[192,278,953,977]" captionTargetBox="[208,1427,193,918]" captionTargetId="figure-6@59.[192,1437,192,924]" captionTargetPageId="59" captionText="Figures 304315: Mesobuthus, pedipalp movable (304313) and fixed (314315) fingers. Figure 304. M. brutus sp. n., male paratype. Figure 305. M. caucasicus, male from Turkey. Figure 306. M. elenae sp. n., female holotype. Figure 307. M. gorelovi sp. n., male holotype. Figure 308. M. fuscus, female from Tajikistan. Figure 309. M. kaznakovi, male from Tajikistan. Figure 310, 314. M. kreuzbergi sp. n., male holotype. Figure 311. M. mischi sp. n., male holotype. Figure 312, 315. M. nenilini sp. n., male holotype. Figure 313. M. parthorum, male from Afghanistan." pageId="10" pageNumber="9">305</figureCitation>
,
<figureCitation id="13305DC17A576A7FFDF05910FD7D40D6" box="[627,669,895,918]" captionStart="Figures 316321" captionStartId="60.[192,278,1445,1469]" captionTargetBox="[191,1437,204,1417]" captionTargetId="figure-304@60.[192,1437,205,1416]" captionTargetPageId="60" captionText="Figures 316321: Mesobuthus, live specimens. Figure 316. M. kaznakovi, female from Uzbekistan. Figure 317. M. kaznakovi, male from Tajikistan. Figure 318. M. caucasicus, female from Iran. Figure 319. M. elenae sp. n., female. Figure 320. M. kreuzbergi sp. n., female paratype. Figure 321. M. parthorum, juvenile from Afghanistan." pageId="10" pageNumber="9">318</figureCitation>
,
<figureCitation id="13305DC17A576A7FFD2B5910FCE840D6" box="[680,776,895,918]" captionStart="Figures 322327" captionStartId="61.[192,278,1432,1456]" captionTargetBox="[191,1437,191,1404]" captionTargetId="figure-45@61.[192,1437,192,1403]" captionTargetPageId="61" captionText="Figures 322327: Mesobuthus, localities. Figures 322323. M. caucasicus, Turkey, Iğdir Province, Melekli Village. Figures 324325: M. fuscus, Tajikistan, Khatlon Province, Khuroson District, Ganjina. Figure 326. M. gorelovi, Uzbekistan, Fargona Province, Yazyavan. Figure 327. M. kreuzbergi, Uzbekistan, type locality." pageId="10" pageNumber="9">322323</figureCitation>
,
<tableCitation id="C68974FF7A576A7FFE3259F1FDC940F5" box="[433,553,926,949]" captionStart-0="Table 4" captionStart-1="Table 5" captionStart-2="Table 6" captionStartId-0="53.[192,257,1057,1081]" captionStartId-1="54.[192,257,1090,1114]" captionStartId-2="62.[192,257,967,991]" captionText-0="Table 4: Comparison among Mesobuthus species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." captionText-1="Table 5: Comparison among Mesobuhus species (specimens), based upon selected morphometric ratios of adult females. Abbreviations: length (L), width (W), depth (D)." captionText-2="Table 6: Comparison among Mesobuhus species, based upon pectinal teeth number. Data from selected specimens studied in this revision; additional data for M. caucasicus (Caucasus) from ZISP collection (Fet, 1989) scored by VF in 1987-88." pageId="10" pageNumber="9">Tables 46</tableCitation>
)
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<taxonomicName id="4C0B3AC77A576A7FFF4359BAFD5740AC" authority="Nordmann, 1840: 731" authorityName="Nordmann" authorityPageNumber="731" authorityYear="1840" box="[192,695,981,1004]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="caucasicus">
<emphasis id="B97F9D567A576A7FFF4359BAFE2440AC" box="[192,452,981,1004]" italics="true" pageId="10" pageNumber="9">Androctonus caucasicus</emphasis>
<bibRefCitation id="EF9A3CB57A576A7FFE4859BAFD5740AC" author="NORDMANN" box="[459,695,981,1004]" pageId="10" pageNumber="9" refId="ref34768" refString="NORDMANN, A. 1840. Notice sur les Scorpions de la faune pontique. Pp. 751 - 752 in Voyage dans la Russie meridionale et la Crimee, par la Hongrie, la Valachie et la Moldavie, execute an 1837, sous la direction de M. Anatole de Demidoff, par Mm. de- Sainson, Le-Play, Huot, Leleveille, Raflet, Rousseau, de Nordmann et du Ponceau; dedie a S. M. Nicolas 1 - er, Empereur de toutes les Russies. Paris, vol. 3." type="journal volume" year="1840">Nordmann, 1840: 731</bibRefCitation>
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, pl. 1, fig. 2.
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