173 lines
20 KiB
XML
173 lines
20 KiB
XML
<document id="8EC487397184C579DE822901103B1527" ID-CLB-Dataset="31828" ID-DOI="10.11646/zootaxa.4337.1.8" ID-GBIF-Dataset="b001f995-9e4f-45cf-bb6f-a67eeb29045b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1014111" ID-ZooBank="269E2AA7-7E1C-49D6-99A0-306DF6F4864B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1508235745325" checkinUser="plazi" docAuthor="Alekseev, Vitalii I." docDate="2017" docId="84693408FF91FFD9FF63FECAFAAEFB40" docLanguage="en" docName="zootaxa.4337.1.8.pdf" docOrigin="Zootaxa 4337 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clambus helheimricus Alekseev 2017, sp. nov." docType="treatment" docUuid="99D6B24C-2EDA-4FD9-8F0D-F1E58F35FF1B" docUuidSource="ZooBank" docVersion="9" lastPageNumber="143" masterDocId="78504C70FF90FFDBFFF4FFADFE43FFA1" masterDocTitle="A new species of minute beetle (Coleoptera: Clambidae) from Baltic amber (Paleogene, Eocene)" masterLastPageNumber="145" masterPageNumber="141" pageNumber="142" updateTime="1698469662158" updateUser="ExternalLinkService">
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<mods:title id="80623F81FD7EFDD7FD92F4EA5C016F7A">A new species of minute beetle (Coleoptera: Clambidae) from Baltic amber (Paleogene, Eocene)</mods:title>
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<mods:name id="34B82D9E8E30C52C224FF4AB3F114A6A" type="personal">
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<mods:roleTerm id="170E3E445A95606FF8D9718BEBA66FDE">Author</mods:roleTerm>
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<mods:namePart id="E7726975B45E0DF6690ED6448DB893F4">Alekseev, Vitalii I.</mods:namePart>
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<mods:title id="32FD2C881E18319CE8BCC9F2BF54DFE6">Zootaxa</mods:title>
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<mods:part id="68048B6858340A35BBDAFBBDC5CB2F43">
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<mods:date id="D8923044E66F36C3FF1FDD8F216E1C6E">2017</mods:date>
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<mods:number id="5B1DEA535D9E34048053D2ABB02E4A16">2017-10-17</mods:number>
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<mods:number id="A7925F917EFB6244CD6AFE794E6360A1">4337</mods:number>
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<mods:identifier id="CFBBF3CEB5102366B6EE382EF08A21EB" type="DOI">10.11646/zootaxa.4337.1.8</mods:identifier>
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<treatment id="84693408FF91FFD9FF63FECAFAAEFB40" ID-DOI="http://doi.org/10.5281/zenodo.6045801" ID-GBIF-Taxon="135450158" ID-Zenodo-Dep="6045801" LSID="urn:lsid:zoobank.org:act:99D6B24C-2EDA-4FD9-8F0D-F1E58F35FF1B" httpUri="http://treatment.plazi.org/id/84693408FF91FFD9FF63FECAFAAEFB40" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142">
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<subSubSection id="44DAD695FF91FFDAFF63FECAFCCCFE23" box="[151,655,359,386]" pageId="1" pageNumber="142" type="nomenclature">
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<paragraph id="0C7F851EFF91FFDAFF63FECAFCCCFE23" blockId="1.[151,1436,359,903]" box="[151,655,359,386]" pageId="1" pageNumber="142">
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<heading id="57373272FF91FFDAFF63FECAFCCCFE23" bold="true" box="[151,655,359,386]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
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<emphasis id="3EB4590CFF91FFDAFF63FECAFCCCFE23" bold="true" box="[151,655,359,386]" pageId="1" pageNumber="142">
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<taxonomicName id="CBC0FE9DFF91FFDAFF63FECAFFEBFE20" authority="Alekseev, 2017" authorityName="Alekseev" authorityYear="2017" box="[151,424,359,385]" class="Insecta" family="Clambidae" genus="Clambus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="helheimricus" status="sp. nov.">
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<emphasis id="3EB4590CFF91FFDAFF63FECAFFEBFE20" bold="true" box="[151,424,359,385]" italics="true" pageId="1" pageNumber="142">Clambus helheimricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2587E477FF91FFDAFE5BFEC5FC4FFE23" box="[431,524,360,386]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="94FB999BFF91FFDAFDE8FEC5FCC5FE23" box="[540,646,360,386]" captionStart="FIGURES 1 – 3" captionStartId="1.[151,258,2016,2038]" captionTargetBox="[151,1419,944,1997]" captionTargetId="figure@1.[151,1436,944,1999]" captionTargetPageId="1" captionText="FIGURES 1 – 3. Clambus helheimricus sp. nov., holotype, habitus, ventro-lateral view." httpUri="https://zenodo.org/record/1014113/files/figure.png" pageId="1" pageNumber="142">Figs 1–3</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="44DAD695FF91FFDAFF63FE27FEA6FDD0" pageId="1" pageNumber="142" type="materials_examined">
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<paragraph id="0C7F851EFF91FFDAFF63FE27FA4EFDA8" blockId="1.[151,1436,359,903]" pageId="1" pageNumber="142">
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<emphasis id="3EB4590CFF91FFDAFF63FE27FF03FE00" bold="true" box="[151,320,394,417]" pageId="1" pageNumber="142">Type material.</emphasis>
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Holotype No. 1470-2 [CCHH], adult, sex unknown, complete beetle, included in small-sized yellow amber piece embedded in a block of GTS polyester resin with approximate side-length dimensions of 13 × 10 ×
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<quantity id="CB3828FBFF91FFDAFAA2FE00FBDBFE65" box="[1366,1432,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="142" unit="mm" value="5.0">5 mm</quantity>
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. The specimen is clearly visible in lateral aspect. Syninclusions are represented by one ant specimens (?
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<taxonomicName id="CBC0FE9DFF91FFDAFB12FE7DFB68FE46" box="[1254,1323,464,487]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="genus">
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<emphasis id="3EB4590CFF91FFDAFB12FE7DFB68FE46" box="[1254,1323,464,487]" italics="true" pageId="1" pageNumber="142">Lasius</emphasis>
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</taxonomicName>
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sp.), three specimens of Acari, the head of an insect (? aphid) and minute particles of detritus.
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</paragraph>
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<paragraph id="0C7F851EFF91FFDAFF33FDB8FC9AFD8D" blockId="1.[151,1436,359,903]" box="[199,729,533,556]" pageId="1" pageNumber="142">
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<emphasis id="3EB4590CFF91FFDAFF33FDB8FF0EFD8D" bold="true" box="[199,333,533,556]" pageId="1" pageNumber="142">Type strata.</emphasis>
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Baltic amber, Upper or mid-Eocene.
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</paragraph>
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<paragraph id="0C7F851EFF91FFDAFF33FD95FEA6FDD0" blockId="1.[151,1436,359,903]" pageId="1" pageNumber="142">
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<materialsCitation id="BCA88F43FF91FFDAFF33FD95FEA6FDD0" ID-GBIF-Occurrence="1675163014" country="Russia" county="Peninsula" location="Yantarny" municipality="Samland" pageId="1" pageNumber="142" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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<emphasis id="3EB4590CFF91FFDAFF33FD95FF22FDEE" bold="true" box="[199,353,568,591]" pageId="1" pageNumber="142">
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<typeStatus id="D37B3BBCFF91FFDAFF33FD95FEBCFDEE" box="[199,255,568,591]" pageId="1" pageNumber="142">Type</typeStatus>
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locality.
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</emphasis>
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<location id="091FD3C5FF91FFDAFE9AFD95FF8DFDEE" LSID="urn:lsid:plazi:treatment:84693408FF91FFD9FF63FECAFAAEFB40:091FD3C5FF91FFDAFE9AFD95FF8DFDEE" box="[366,462,568,591]" country="Russia" county="Peninsula" municipality="Samland" name="Yantarny" pageId="1" pageNumber="142" stateProvince="Kaliningrad">Yantarny</location>
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settlement (formerly
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<location id="091FD3C5FF91FFDAFD32FD95FD0BFDEE" LSID="urn:lsid:plazi:treatment:84693408FF91FFD9FF63FECAFAAEFB40:091FD3C5FF91FFDAFD32FD95FD0BFDEE" box="[710,840,568,591]" country="Russia" county="Peninsula" municipality="Samland" name="Palmnicken" pageId="1" pageNumber="142" stateProvince="Kaliningrad">Palmnicken</location>
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),
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<location id="091FD3C5FF91FFDAFCABFD95FDFFFDEE" LSID="urn:lsid:plazi:treatment:84693408FF91FFD9FF63FECAFAAEFB40:091FD3C5FF91FFDAFCABFD95FDFFFDEE" box="[863,956,568,591]" country="Russia" county="Peninsula" municipality="Samland" name="Sambian" pageId="1" pageNumber="142" stateProvince="Kaliningrad">Sambian</location>
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(
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<collectingMunicipality id="EC1B1F64FF91FFDAFC3BFD95FA6CFDEE" box="[975,1071,568,591]" pageId="1" pageNumber="142">Samland</collectingMunicipality>
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)
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<collectingCounty id="E51EFD92FF91FFDAFBB7FD95FAEEFDEE" box="[1091,1197,568,591]" pageId="1" pageNumber="142">Peninsula</collectingCounty>
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,
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<collectingRegion id="CE044BFCFF91FFDAFB49FD95FB7FFDEE" box="[1213,1340,568,591]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="142">Kaliningrad</collectingRegion>
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Region,
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<collectingCountry id="74D7C58EFF91FFDAFF63FDF7FEA1FDD0" box="[151,226,602,625]" name="Russia" pageId="1" pageNumber="142">Russia</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="44DAD695FF91FFDAFF33FDD0FD62FD78" pageId="1" pageNumber="142" type="etymology">
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<paragraph id="0C7F851EFF91FFDAFF33FDD0FD62FD78" blockId="1.[151,1436,359,903]" pageId="1" pageNumber="142">
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<emphasis id="3EB4590CFF91FFDAFF33FDD0FF03FD35" bold="true" box="[199,320,637,660]" pageId="1" pageNumber="142">Etymology</emphasis>
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. The specific epithet “
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<taxonomicName id="CBC0FE9DFF91FFDAFDC7FDD0FC82FD35" authorityName="Alekseev" authorityYear="2017" box="[563,705,637,660]" class="Insecta" family="Clambidae" genus="Clambus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="helheimricus">helheimricus</taxonomicName>
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” is formed after the word Helheimr (Helheim, or Hel)—“concealed place, the underworld”—realm of the dead, existing beneath one of the three roots of the world ash-tree Yggdrasil and ruled by the goddess Hel in the old Scandinavian mythology.
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</paragraph>
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</subSubSection>
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<subSubSection id="44DAD695FF91FFDAFF33FD48FB1FFC26" pageId="1" pageNumber="142" type="diagnosis">
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<paragraph id="0C7F851EFF91FFDAFF33FD48FB1FFC26" blockId="1.[151,1436,359,903]" pageId="1" pageNumber="142">
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<emphasis id="3EB4590CFF91FFDAFF33FD48FF87FD5D" bold="true" box="[199,452,741,764]" pageId="1" pageNumber="142">Differential diagnosis.</emphasis>
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<taxonomicName id="CBC0FE9DFF91FFDAFE39FD48FC83FD5D" authority="Alekseev, 2017" authorityName="Alekseev" authorityYear="2017" box="[461,704,741,764]" class="Insecta" family="Clambidae" genus="Clambus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="helheimricus" status="sp. nov.">
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<emphasis id="3EB4590CFF91FFDAFE39FD48FC83FD5D" box="[461,704,741,764]" italics="true" pageId="1" pageNumber="142">Clambus helheimricus</emphasis>
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</taxonomicName>
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<emphasis id="3EB4590CFF91FFDAFD3DFD48FD61FD5D" bold="true" box="[713,802,741,764]" pageId="1" pageNumber="142">
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<taxonomicNameLabel id="2587E477FF91FFDAFD3DFD48FD61FD5D" box="[713,802,741,764]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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differs from extant species of the genus in the following combination of characters: dorsal surface apparently bare, impunctate, and shiny; temporal angles rounded, situated behind posterior margin of eyes; eyes relatively small, longitudinally oval; metaventrite and coxal plates pubescent, not shagreened; posterior horizontal plate of metaventrite moderately reduced in middle, approximately 3× shorter medially than laterally; metacoxal plates sparsely punctate; abdominal ventrites 2–5 pubescent, finely and densely punctate.
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</paragraph>
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</subSubSection>
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<caption id="58BFD596FF91FFDAFF63F84DFD9AF854" box="[151,985,2016,2038]" httpUri="https://zenodo.org/record/1014113/files/figure.png" pageId="1" pageNumber="142" targetBox="[151,1419,944,1997]" targetPageId="1">
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<paragraph id="0C7F851EFF91FFDAFF63F84DFD9AF854" blockId="1.[151,985,2016,2038]" box="[151,985,2016,2038]" pageId="1" pageNumber="142">
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<emphasis id="3EB4590CFF91FFDAFF63F84DFF71F857" bold="true" box="[151,306,2016,2038]" pageId="1" pageNumber="142">FIGURES 1–3.</emphasis>
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<taxonomicName id="CBC0FE9DFF91FFDAFEC3F84DFC4DF854" authority="Alekseev, 2017" authorityName="Alekseev" authorityYear="2017" box="[311,526,2016,2037]" class="Insecta" family="Clambidae" genus="Clambus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="helheimricus" status="sp. nov.">
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<emphasis id="3EB4590CFF91FFDAFEC3F84DFC4DF854" box="[311,526,2016,2037]" italics="true" pageId="1" pageNumber="142">Clambus helheimricus</emphasis>
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</taxonomicName>
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<emphasis id="3EB4590CFF91FFDAFDE0F84DFC22F857" bold="true" box="[532,609,2016,2038]" pageId="1" pageNumber="142">
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<taxonomicNameLabel id="2587E477FF91FFDAFDE0F84DFC22F857" box="[532,609,2016,2038]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype, habitus, ventro-lateral view.
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</paragraph>
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</caption>
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<subSubSection id="44DAD695FF92FFD9FF33FF35FA7CFB95" pageId="2" pageNumber="143" type="description">
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<paragraph id="0C7F851EFF92FFD9FF33FF35FD6BFF70" blockId="2.[151,1436,152,1249]" pageId="2" pageNumber="143">
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<emphasis id="3EB4590CFF92FFD9FF33FF35FF08FF0E" bold="true" box="[199,331,152,175]" pageId="2" pageNumber="143">Description</emphasis>
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.
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<emphasis id="3EB4590CFF92FFD9FEA8FF35FFF3FF0E" box="[348,432,152,175]" italics="true" pageId="2" pageNumber="143">Habitus</emphasis>
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. Body length
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<quantity id="CB3828FBFF92FFD9FDA6FF35FCFAFF0E" box="[594,697,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="2" pageNumber="143" unit="mm" value="1.01">1.01 mm</quantity>
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; maximum width
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<quantity id="CB3828FBFF92FFD9FC71FF35FDA8FF0E" box="[901,1003,152,175]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="143" unit="mm" value="0.78">0.78 mm</quantity>
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; rounded, conglobate; convex, smooth, impunctate and shiny dorsally. Body dark brown, legs rufous.
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</paragraph>
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<paragraph id="0C7F851EFF92FFD9FF33FF70FFF6FD8D" blockId="2.[151,1436,152,1249]" pageId="2" pageNumber="143">
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<emphasis id="3EB4590CFF92FFD9FF33FF70FF42FF55" box="[199,257,221,244]" italics="true" pageId="2" pageNumber="143">Head</emphasis>
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. Hypognathous, transverse (approximately 1.4× as wide as long), convex, widest across temporal angles behind eyes, narrower than pronotum. Clypeal margin arcuate, with sparse short setation. Eyes from dorsal view relatively small, longitudinally oval, very finely facetted, consisting of approximately 50 facets. Interfacetal setae absent. Lateral margin of eye diverging from temporal margin. Temporal angles rounded, situated behind posterior margin of eyes. Antennal insertions exposed in dorsal view, widely separated. Antennae distinctly clubbed, relatively short (when posteriorly extended reaching middle of pronotum), consisting of 10 antennomeres. Scape and pedicel enlarged; antennomere 3 long and cylindrical; antennomeres 4–7 elongate; antennomere 8 rounded, as long as wide; antennomeres 9–10 distinctly enlarged and clubbed. Antennomere 9 the largest, triangular with rounded angles, slightly longer than broad. Ratio of antennomere lengths: 6.0: 4.0: 8.0: 4.0: 2.0: 2.0: 2.0: 3.0: 11.0: 5.0. Antennal club covered with long, thick, flexible, white hairs.
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</paragraph>
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<paragraph id="0C7F851EFF92FFD9FF33FD95FCBFFCBE" blockId="2.[151,1436,152,1249]" pageId="2" pageNumber="143">
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<emphasis id="3EB4590CFF92FFD9FF33FD95FF57FDEE" box="[199,276,568,591]" italics="true" pageId="2" pageNumber="143">Thorax</emphasis>
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. Pronotum transverse (in dorsal view about 0.41× as long as wide), convex. Lateral pronotal carina rounded. Anterior pronotal angles rectangular, rounded; posterior angles obtuse, broadly rounded. Posterior pronotal margin concave; anterior pronotal margin broadly arcuate. Scutellar shield triangular, as wide as long. Elytra strongly convex, rounded, almost as broad as pronotum basally, approximately 0.7× as long as their maximal combined width and 2.7× as long as pronotum. Elytral punctation or setation not apparent. Epipleura incomplete, narrow, present only anteriorly. Pro- and mesocoxal cavities open. Horizontal plate of metaventrite narrow in middle (approximately 0.3× as long as it lateral length), densely pubescent. Pubescence long, decumbent.
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</paragraph>
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<paragraph id="0C7F851EFF92FFD9FF33FC87FD53FC6D" blockId="2.[151,1436,152,1249]" pageId="2" pageNumber="143">
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||
<emphasis id="3EB4590CFF92FFD9FF33FC87FEBAFCE0" box="[199,249,810,833]" italics="true" pageId="2" pageNumber="143">Legs</emphasis>
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. Long and slender. Metaxocal plates large, covering metacoxae and abdominal ventrite 1, punctate, pubescent like horizontal plate of metaventrite. Punctation large, irregular; distance between punctures equal to 1.0–4.0× diameter of one puncture. Coxae cylindrical, projecting. Femora and tibiae slender. Tibiae pubescent. Tarsi tetramerous with tarsomeres simple, cylindrical, pubescent. Tarsomeres 1 and 4 long, almost equal in length; tarsomeres 2 and 3 shorter than tarsomeres 1 and 4. Claws simple, acute, symmetrical.
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</paragraph>
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<paragraph id="0C7F851EFF92FFD9FF33FC75FA7CFB95" blockId="2.[151,1436,152,1249]" pageId="2" pageNumber="143">
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<emphasis id="3EB4590CFF92FFD9FF33FC75FF6FFC4E" box="[199,300,984,1007]" italics="true" pageId="2" pageNumber="143">Abdomen</emphasis>
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. With five ventrites. Ventrite 1 almost concealed by metacoxal plates; ratio of ventrite lengths 2–5: 2.0: 1.0: 1.0: 2.0. Ventrites 2–5 pubescent, finely and densely punctate. Distance between punctures equal to 0.8–1.0× diameter of one puncture. Ventrite 5 simple, without apparent long pubescence apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="44DAD695FF92FFD9FF33FBEDFAAEFB40" pageId="2" pageNumber="143" type="discussion">
|
||
<paragraph id="0C7F851EFF92FFD9FF33FBEDFAD2FBF6" blockId="2.[151,1436,152,1249]" box="[199,1169,1088,1111]" pageId="2" pageNumber="143">
|
||
<emphasis id="3EB4590CFF92FFD9FF33FBEDFF67FBF6" bold="true" box="[199,292,1088,1111]" pageId="2" pageNumber="143">Remark</emphasis>
|
||
. Genitalia internal and obscured in this specimen. Metathoracic wings not visible.
|
||
</paragraph>
|
||
<paragraph id="0C7F851EFF92FFD9FF33FBCFFAAEFB40" blockId="2.[151,1436,152,1249]" pageId="2" pageNumber="143">
|
||
<emphasis id="3EB4590CFF92FFD9FF33FBCFFFDBFBD8" bold="true" box="[199,408,1122,1145]" pageId="2" pageNumber="143">Assumed bionomy</emphasis>
|
||
. Extant representatives of the genus
|
||
<taxonomicName id="CBC0FE9DFF92FFD9FCD0FBCFFDC1FBD8" authorityName="Fischer von Waldheim" authorityYear="1821" box="[804,898,1122,1145]" class="Insecta" family="Clambidae" genus="Clambus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EB4590CFF92FFD9FCD0FBCFFDC1FBD8" box="[804,898,1122,1145]" italics="true" pageId="2" pageNumber="143">Clambus</emphasis>
|
||
</taxonomicName>
|
||
are collected from decaying vegetation, leaf litter and rotten wood. Records indicate that the family is probably feeding on microfungi, though they may be found on Ascomycetes, Myxomycetes and Basidiomycetes, where they feed mainly on spores and hyphae (
|
||
<bibRefCitation id="6851F8EFFF92FFD9FB57FB05FB7DFB1E" author="Leschen" box="[1187,1342,1192,1215]" pageId="2" pageNumber="143" refString="Leschen, R. A. B. (2016) 10.3. Clambidae Fischer von Waldheim, 1821. In: Beutel, R. G. & Leschen, R. A. B. (Eds.), Handbook of Zoology - Coleoptera, Beetles. Vol. 1. Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter de Gruyter GmbH & Co KG, Berlin, pp. 211 - 215." type="book chapter" year="2016">Leschen 2016</bibRefCitation>
|
||
). For the fossil species, a similar biology and relation with saproxylic environment and soil litter can be assumed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |