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<document id="5FC2E354C23CAF3915846BDA4197673C" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="1175­5326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FFB5390E4D2DFB87FCFAFB9B" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Geostiba (Sibiota) crepusculigena Gusarov, sp. n." docType="treatment" docVersion="14" lastPageNumber="46" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="44" updateTime="1698188143631" updateUser="plazi">
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<mods:title id="43D33BC3F52EAF294F486D95E9E96DB0">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="D12B95370260063C4CE2F53A39B0EF73">Gusarov, Vladimir I.</mods:namePart>
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<mods:date id="B9096FF2437264BEEF5BAEFB63EC5E9D">2002</mods:date>
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<treatment id="5B50E916FFB5390E4D2DFB87FCFAFB9B" ID-DOI="http://doi.org/10.5281/zenodo.6277573" ID-Zenodo-Dep="6277573" LSID="urn:lsid:plazi:treatment:5B50E916FFB5390E4D2DFB87FCFAFB9B" httpUri="http://treatment.plazi.org/id/5B50E916FFB5390E4D2DFB87FCFAFB9B" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
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<paragraph id="D3465800FFB539084D2DFB87FBA8FB8F" blockId="43.[264,1088,1083,1110]" box="[264,1088,1083,1110]" pageId="43" pageNumber="44">
<heading id="880EEF6CFFB539084D2DFB87FBA8FB8F" bold="true" box="[264,1088,1083,1110]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<keyLead id="6808FB35FFB539084D2DFB87FBA8FB8F" box="[264,1088,1083,1110]" pageId="43" pageNumber="44">
<emphasis id="E18D8412FFB539084D2DFB87FBA8FB8F" bold="true" box="[264,1088,1083,1110]" pageId="43" pageNumber="44">
<keyStep id="680D40A5FFB539084D2DFB87FBA8FB8F" box="[264,1088,1083,1110]" pageId="43" pageNumber="44">
<key id="A7680F31FFB539084D2DFB87FBA8FB8F" box="[264,1088,1083,1110]" pageId="43" pageNumber="44">
7.
<taxonomicName id="14F92383FFB539084D03FB80FCD9FB8F" ID-CoL="6KD6S" authority="Gusarov" authorityName="Gusarov" box="[294,817,1083,1110]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="crepusculigena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FFB539084D03FB80FE78FB8C" bold="true" box="[294,400,1084,1110]" italics="true" pageId="43" pageNumber="44">Geostiba</emphasis>
(
<emphasis id="E18D8412FFB539084D85FB80FE1DFB8C" bold="true" box="[416,501,1084,1110]" italics="true" pageId="43" pageNumber="44">Sibiota</emphasis>
)
<emphasis id="E18D8412FFB539084E23FB80FD56FB8C" bold="true" box="[518,702,1084,1110]" italics="true" pageId="43" pageNumber="44">crepusculigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FFB539084F1BFB87FC68FB8F" box="[830,896,1083,1109]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="4BC24485FFB539084FB5FB87FBD0FB8F" box="[912,1080,1083,1109]" captionStart-0="FIGURES 169 ­ 173" captionStart-1="FIGURES 174 ­ 177" captionStart-2="FIGURES 178 ­ 186" captionStartId-0="44.[264,383,1756,1780]" captionStartId-1="45.[264,383,1766,1790]" captionStartId-2="46.[264,382,1225,1249]" captionTargetBox-0="[314,1272,742,1729]" captionTargetBox-1="[317,1274,1164,1735]" captionTargetBox-2="[393,1195,318,1201]" captionTargetId-0="figure@44.[306,1280,729,1738]" captionTargetId-1="figure@45.[305,1283,1154,1742]" captionTargetId-2="figure@46.[384,1202,310,1211]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionText-0="FIGURES 169 ­ 173. Abdominal segment 8 of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 169 male tergum 8; 170 male sternum 8; 171 female tergum 8; 172 female sternum 8, 173 apex of female sternum 8. Scale bar 0.2 mm (169 ­ 172), 0.1 mm (173)." captionText-1="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." captionText-2="FIGURES 178 ­ 186. Genitalia of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 178 details of internal sac retracted into median lobe, ventral view; 179 distal diverticula of everted internal sac, ventral view; 180 everted internal sac, ventral view; 181 spermatheca; 182 apex of left paramere, side facing median lobe; 183 ­ 184 details of internal sac retracted into median lobe, lateral view; 185 right distal diverticulum of everted internal sac, lateral view; 186 everted internal sac, lateral view. Scale bar 0.2 mm (178, 180, 183 ­ 184, 186), 0.1 mm (179, 181 ­ 182, 185)." httpUri-0="https://zenodo.org/record/155725/files/figure.png" httpUri-1="https://zenodo.org/record/155726/files/figure.png" httpUri-2="https://zenodo.org/record/155727/files/figure.png" pageId="43" pageNumber="44">Figs. 169­186</figureCitation>
)
</key>
</keyStep>
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB539084D2DFB3AFC51FAB2" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="D3465800FFB539084D2DFB3AFC85FB2B" blockId="43.[264,1324,1158,1904]" pageId="43" pageNumber="44">
<emphasis id="E18D8412FFB539084D2DFB3AFE56FB7A" bold="true" box="[264,446,1158,1184]" pageId="43" pageNumber="44">
<typeStatus id="0C42E6A2FFB539084D2DFB3AFEABFB7A" box="[264,323,1158,1184]" pageId="43" pageNumber="44">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0C42E6A2FFB539084DF7FB3BFDAAFB7B" box="[466,578,1159,1185]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
,,
<emphasis id="E18D8412FFB539084E51FB3AFC16FB7A" bold="true" box="[628,1022,1158,1184]" pageId="43" pageNumber="44">
<collectingCountry id="ABEE1890FFB539084E51FB3AFC89FB7A" box="[628,865,1158,1184]" name="United States of America" pageId="43" pageNumber="44">UNITED STATES</collectingCountry>
: Tennessee:
</emphasis>
Cocke Co.: Snake Den Ridge Trail,
<quantity id="1401F5E5FFB539084DBEFB13FE3EFB13" box="[411,470,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="43" pageNumber="44" unit="km" value="9.0">9 km</quantity>
S Cosby,
<geoCoordinate id="B6CD3EC7FFB539084E69FB13FD39FB13" box="[588,721,1199,1225]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="9" value="35.736332">35°44.18'N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFB539084EFCFB13FC8CFB13" box="[729,868,1199,1225]" direction="west" orientation="longitude" pageId="43" pageNumber="44" precision="9" value="-83.24333">83°14.60'W</geoCoordinate>
,
<quantity id="1401F5E5FFB539084F55FB13FC24FB13" box="[880,972,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="43" pageNumber="44" unit="m" value="1520.0">1520 m</quantity>
, in forest litter,
<taxonomicName id="14F92383FFB5390848A6FB13FB2EFB12" box="[1155,1222,1199,1224]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFB5390848A6FB13FB2EFB12" box="[1155,1222,1199,1224]" italics="true" pageId="43" pageNumber="44">Picea</emphasis>
</taxonomicName>
,
<emphasis id="E18D8412FFB5390848F1FB13FE98FB2A" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="14F92383FFB5390848F1FB13FE84FB2A" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
,
</emphasis>
<taxonomicName id="14F92383FFB539084D52FB6BFD87FB2B" authority="(V.I.Gusarov)" authorityName="V.I.Gusarov" baseAuthorityName="V. I. Gusarov" box="[375,623,1239,1265]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFB539084D52FB6BFE2CFB2A" box="[375,452,1239,1264]" italics="true" pageId="43" pageNumber="44">Betula</emphasis>
(V.I.Gusarov)
</taxonomicName>
,
<date id="A7477EC0FFB539084E5EFB6BFD11FB2B" box="[635,761,1239,1265]" pageId="43" pageNumber="44" value="2001-09-23">23.ix.2001</date>
(
<collectionCode id="B5E8C0C5FFB539084F2DFB6BFC88FB2B" box="[776,864,1239,1265]" pageId="43" pageNumber="44">KSEM</collectionCode>
).
</paragraph>
<paragraph id="D3465800FFB539084D1DFB43FC51FAB2" blockId="43.[264,1324,1158,1904]" pageId="43" pageNumber="44">
<typeStatus id="0C42E6A2FFB539084D1DFB43FE47FAC3" box="[312,431,1279,1305]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
<emphasis id="E18D8412FFB539084DE4FB42FCADFAC2" bold="true" box="[449,837,1278,1304]" pageId="43" pageNumber="44">
<collectingCountry id="ABEE1890FFB539084DE4FB42FD44FAC2" box="[449,684,1278,1304]" name="United States of America" pageId="43" pageNumber="44">UNITED STATES</collectingCountry>
: Tennessee:
</emphasis>
Cocke Co.:
<specimenCount id="C5FF9389FFB539084FCDFB43FB62FAC3" box="[1000,1162,1279,1305]" pageId="43" pageNumber="44" type="generic">20 specimens</specimenCount>
, same data as the
<typeStatus id="0C42E6A2FFB539084D12FA9BFE49FA9B" box="[311,417,1319,1345]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
;
<specimenCount id="C5FF9389FFB539084D94FA9BFDBFFA9B" box="[433,599,1319,1345]" pageId="43" pageNumber="44" type="generic">10 specimens</specimenCount>
, ditto but
<quantity id="1401F5E5FFB539084EFDFA9BFCF0FA9B" box="[728,792,1319,1345]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="43" pageNumber="44" unit="km" value="8.0">8 km</quantity>
S Cosby,
<geoCoordinate id="B6CD3EC7FFB539084FBFFA9BFBC8FA9B" box="[922,1056,1319,1345]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="9" value="35.7435">35°44.61'N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFB53908480EFA9BFB5FFA9B" box="[1067,1207,1319,1345]" direction="west" orientation="longitude" pageId="43" pageNumber="44" precision="9" value="-83.23333">83°14.00'W</geoCoordinate>
,
<quantity id="1401F5E5FFB5390848E2FA9BFACFFA9B" box="[1223,1319,1319,1345]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="43" pageNumber="44" unit="m" value="1470.0">1470 m</quantity>
,
<taxonomicName id="14F92383FFB539084D2DFAF3FEA2FAB2" box="[264,330,1359,1384]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFB539084D2DFAF3FEA2FAB2" box="[264,330,1359,1384]" italics="true" pageId="43" pageNumber="44">Tsuga</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFB539084D79FAF3FE77FAB2" box="[348,415,1359,1384]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFB539084D79FAF3FE77FAB2" box="[348,415,1359,1384]" italics="true" pageId="43" pageNumber="44">Picea</emphasis>
</taxonomicName>
,
<emphasis id="E18D8412FFB539084D88FAF3FD5DFAB2" box="[429,693,1359,1384]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="14F92383FFB539084D88FAF3FDB5FAB2" box="[429,605,1359,1384]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
,
<taxonomicName id="14F92383FFB539084E4CFAF3FD5DFAB2" box="[617,693,1359,1384]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
</emphasis>
(all
<collectionCode id="B5E8C0C5FFB539084F24FAF2FCBEFAB2" box="[769,854,1358,1384]" pageId="43" pageNumber="44">KSEM</collectionCode>
,
<collectionCode id="B5E8C0C5FFB539084F44FAF2FC45FAB2" box="[865,941,1358,1384]" pageId="43" pageNumber="44">SPSU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB539084D1DFACAFC89F90A" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="D3465800FFB539084D1DFACAFD5AF982" blockId="43.[264,1324,1158,1904]" pageId="43" pageNumber="44">
<emphasis id="E18D8412FFB539084D1DFACAFE50FA4A" bold="true" box="[312,440,1398,1424]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
<taxonomicName id="14F92383FFB539084DE3FACBFD02FA4A" box="[454,746,1399,1424]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="crepusculigena">
<emphasis id="E18D8412FFB539084DE3FACBFD02FA4A" box="[454,746,1399,1424]" italics="true" pageId="43" pageNumber="44">Geostiba crepusculigena</emphasis>
</taxonomicName>
can be distinguished from other Nearctic species of
<taxonomicName id="14F92383FFB539084D09FA23FE7EFA62" box="[300,406,1439,1464]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FFB539084D09FA23FE7EFA62" box="[300,406,1439,1464]" italics="true" pageId="43" pageNumber="44">Geostiba</emphasis>
</taxonomicName>
by having small eyes (temple length to eye length ratio 3.0­3.1), pronotal pubescence of
<typeStatus id="0C42E6A2FFB539084D93FA7AFE00FA3A" box="[438,488,1478,1504]" pageId="43" pageNumber="44">type</typeStatus>
V, reduced wings, short elytra (pronotum length to elytron length ratio 1.3), the presence of two short parallel carinae in the middle of abdominal tergum
<quantity id="1401F5E5FFB5390848DDFA52FAC1F9D2" box="[1272,1321,1518,1544]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="43" pageNumber="44" unit="in" value="7.0">7 in</quantity>
front of posterior margin, the shape of the aedeagus (
<figureCitation id="4BC24485FFB539084FB9F9AAFB52F9EA" box="[924,1210,1558,1584]" captionStart-0="FIGURES 174 ­ 177" captionStart-1="FIGURES 178 ­ 186" captionStartId-0="45.[264,383,1766,1790]" captionStartId-1="46.[264,382,1225,1249]" captionTargetBox-0="[317,1274,1164,1735]" captionTargetBox-1="[393,1195,318,1201]" captionTargetId-0="figure@45.[305,1283,1154,1742]" captionTargetId-1="figure@46.[384,1202,310,1211]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." captionText-1="FIGURES 178 ­ 186. Genitalia of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 178 details of internal sac retracted into median lobe, ventral view; 179 distal diverticula of everted internal sac, ventral view; 180 everted internal sac, ventral view; 181 spermatheca; 182 apex of left paramere, side facing median lobe; 183 ­ 184 details of internal sac retracted into median lobe, lateral view; 185 right distal diverticulum of everted internal sac, lateral view; 186 everted internal sac, lateral view. Scale bar 0.2 mm (178, 180, 183 ­ 184, 186), 0.1 mm (179, 181 ­ 182, 185)." httpUri-0="https://zenodo.org/record/155726/files/figure.png" httpUri-1="https://zenodo.org/record/155727/files/figure.png" pageId="43" pageNumber="44">Figs. 174­180, 182­186</figureCitation>
) and the shape of the spermatheca (
<figureCitation id="4BC24485FFB539084E1BF982FD4CF982" box="[574,676,1598,1624]" captionStart="FIGURES 178 ­ 186" captionStartId="46.[264,382,1225,1249]" captionTargetBox="[393,1195,318,1201]" captionTargetId="figure@46.[384,1202,310,1211]" captionTargetPageId="46" captionText="FIGURES 178 ­ 186. Genitalia of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 178 details of internal sac retracted into median lobe, ventral view; 179 distal diverticula of everted internal sac, ventral view; 180 everted internal sac, ventral view; 181 spermatheca; 182 apex of left paramere, side facing median lobe; 183 ­ 184 details of internal sac retracted into median lobe, lateral view; 185 right distal diverticulum of everted internal sac, lateral view; 186 everted internal sac, lateral view. Scale bar 0.2 mm (178, 180, 183 ­ 184, 186), 0.1 mm (179, 181 ­ 182, 185)." httpUri="https://zenodo.org/record/155727/files/figure.png" pageId="43" pageNumber="44">Fig. 181</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB539084D1DF9DBFC89F90A" blockId="43.[264,1324,1158,1904]" pageId="43" pageNumber="44">
<taxonomicName id="14F92383FFB539084D1DF9DBFDB5F95A" box="[312,605,1639,1664]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="crepusculigena">
<emphasis id="E18D8412FFB539084D1DF9DBFDB5F95A" box="[312,605,1639,1664]" italics="true" pageId="43" pageNumber="44">Geostiba crepusculigena</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="14F92383FFB539084F97F9DBFBA5F95A" box="[946,1101,1639,1664]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFB539084F97F9DBFBA5F95A" box="[946,1101,1639,1664]" italics="true" pageId="43" pageNumber="44">G. nimbicola</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFB53908487BF9DBFB1FF95A" box="[1118,1271,1639,1664]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pluvigena">
<emphasis id="E18D8412FFB53908487BF9DBFB1FF95A" box="[1118,1271,1639,1664]" italics="true" pageId="43" pageNumber="44">G. pluvigena</emphasis>
</taxonomicName>
and
<taxonomicName id="14F92383FFB539084D2DF933FE5CF972" box="[264,436,1679,1704]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nebuligena">
<emphasis id="E18D8412FFB539084D2DF933FE5CF972" box="[264,436,1679,1704]" italics="true" pageId="43" pageNumber="44">G. nebuligena</emphasis>
</taxonomicName>
in having apex of median lobe with obtuse apical projection (in lateral view) (
<figureCitation id="4BC24485FFB539084D7EF90AFDEAF90A" box="[347,514,1718,1744]" captionStart="FIGURES 174 ­ 177" captionStartId="45.[264,383,1766,1790]" captionTargetBox="[317,1274,1164,1735]" captionTargetId="figure@45.[305,1283,1154,1742]" captionTargetPageId="45" captionText="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." httpUri="https://zenodo.org/record/155726/files/figure.png" pageId="43" pageNumber="44">Figs. 176­177</figureCitation>
,
<figureCitation id="4BC24485FFB539084E28F90AFD9AF90A" box="[525,626,1718,1744]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="43" pageNumber="44">102­103</figureCitation>
,
<figureCitation id="4BC24485FFB539084E5BF90AFD0AF90A" box="[638,738,1718,1744]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="43" pageNumber="44">128­129</figureCitation>
,
<figureCitation id="4BC24485FFB539084ECBF90AFCBCF90A" box="[750,852,1718,1744]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 median lobe, ventral view; 153 ­ 154 apex of median lobe, ventral view; 155 median lobe, lateral view; 156 apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="43" pageNumber="44">155­156</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB5390E4D1DF962FD96FCCB" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44" type="description">
<paragraph id="D3465800FFB539084D1DF962FC64F8FA" blockId="43.[264,1324,1158,1904]" pageId="43" pageNumber="44">
<emphasis id="E18D8412FFB539084D1DF962FE38F922" bold="true" box="[312,464,1758,1784]" pageId="43" pageNumber="44">Description.</emphasis>
Length
<quantity id="1401F5E5FFB539084E18F962FD22F922" box="[573,714,1758,1784]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="43" pageNumber="44" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8­2.3 mm</quantity>
. Body brown to dark brown, antennae brown, legs and mouthparts brownish yellow. Body parallel­sided.
</paragraph>
<paragraph id="D3465800FFB5390F4D1DF892FD6DFE5C" blockId="43.[264,1324,1158,1904]" lastBlockId="44.[264,1324,284,670]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
Head as wide as long, surface on disk with fine isodiametric microsculpture, puncturation very fine, distance between punctures equal to 3­4 times their diameter. Temple length to eye length ratio 3.0­3.1. Antennal article 2 longer than article 3, article 4 transverse (width to length ratio 1.6), articles 5­10 strongly transverse, last article as long as 9 and 10 combined (as in
<figureCitation id="4BC24485FFB2390F4E04FED0FD9FFE5C" box="[545,631,364,390]" captionStart="FIGURES 8 ­ 17" captionStartId="6.[264,383,1709,1733]" captionTargetBox="[307,1277,321,1687]" captionTargetId="figure@6.[297,1289,310,1697]" captionTargetPageId="6" captionText="FIGURES 8 ­ 17. Mouthparts of Geostiba circellaris (Gravenhorst) (8, Moscow Region, Russia), G. infirma (Weise) (9, Ivano­Frankovsk Region, Ukraine), G. carteriensis Pace (10 ­ 15, Tennessee), G. appalachigena Gusarov, sp. n. (16, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (17, paratype from North Carolina). 8 ­ 10, prementum; 11 hypopharynx; 12 left maxilla, ventral view; 13 left galea, dorsal view; 14 left lacinia, dorsal view; 15 mentum; 16 ­ 17 right antenna. Scale bar 0.1 mm (8 ­ 15), 0.2 mm (16 ­ 17). L Length of antennal article." httpUri="https://zenodo.org/record/155703/files/figure.png" pageId="44" pageNumber="45">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB2390F4D1DFE28FDC3FD44" blockId="44.[264,1324,284,670]" pageId="44" pageNumber="45">
Pronotum as wide as long, width
<quantity id="1401F5E5FFB2390F4EEBFE28FC91FE74" box="[718,889,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.0" metricValueMin="3.4" pageId="44" pageNumber="45" unit="mm" value="0.37" valueMax="0.4" valueMin="0.34">0.34­0.40 mm</quantity>
, wider than head (pronotal width to head width ratio 1.1); microsculpture and puncturation as on head. Pronotal pubescence of
<typeStatus id="0C42E6A2FFB2390F4D2DFE58FED2FE24" box="[264,314,484,510]" pageId="44" pageNumber="45">type</typeStatus>
V. Elytra measured from humeral angle shorter than pronotum (pronotal length to elytral length ratio 1.3), wider than long (1.5), with fine isodiametric microsculpture and fine asperate puncturation, distance between punctures equals 1­3 times their diameter. Elytral suture behind scutellum slightly raised in both sexes. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<caption id="87860888FFB2390F4D2DF960FE91F88E" httpUri="https://zenodo.org/record/155725/files/figure.png" pageId="44" pageNumber="45" targetBox="[314,1272,742,1729]" targetPageId="44">
<paragraph id="D3465800FFB2390F4D2DF960FE91F88E" blockId="44.[264,1323,1756,1876]" pageId="44" pageNumber="45">
<emphasis id="E18D8412FFB2390F4D2DF960FE36F92E" bold="true" box="[264,478,1756,1780]" pageId="44" pageNumber="45">FIGURES 169­173</emphasis>
. Abdominal segment 8 of
<taxonomicName id="14F92383FFB2390F4F26F961FB86F92E" authority="Gusarov" authorityName="Gusarov" box="[771,1134,1757,1780]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="crepusculigena" status="sp. nov.">
<emphasis id="E18D8412FFB2390F4F26F961FBE4F92E" box="[771,1036,1757,1780]" italics="true" pageId="44" pageNumber="45">Geostiba crepusculigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FFB2390F485DF960FB5DF92E" box="[1144,1205,1756,1780]" pageId="44" pageNumber="45" rank="species">
<emphasis id="E18D8412FFB2390F485DF960FB5DF92E" bold="true" box="[1144,1205,1756,1780]" pageId="44" pageNumber="45">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Snake Ridge Trail, Tennessee). 169 male tergum 8; 170 male sternum 8; 171 female tergum 8; 172 female sternum 8, 173 apex of female sternum 8. Scale bar 0.2 mm (169­172), 0.1 mm (173).
</paragraph>
</caption>
<paragraph id="D3465800FFB3390E4D1DFEA0FB22FE5C" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">Abdominal terga with fine microsculpture of transverse meshes, with fine and sparse puncturation, puncturation becoming finer towards abdomen apex, on terga 3­5 distance between punctures equals 2­6 times their diameter. Tergum 7 without white edge.</paragraph>
<paragraph id="D3465800FFB3390E4D1DFE28FB8BFE24" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">
Male tergum 7 with two medial carinae in front of posterior margin. Male tergum 8 with two or four weak and short carinae in front of posterior margin, posterior margin convex (
<figureCitation id="4BC24485FFB3390E4D66FE58FE41FE24" box="[323,425,484,510]" captionStart="FIGURES 169 ­ 173" captionStartId="44.[264,383,1756,1780]" captionTargetBox="[314,1272,742,1729]" captionTargetId="figure@44.[306,1280,729,1738]" captionTargetPageId="44" captionText="FIGURES 169 ­ 173. Abdominal segment 8 of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 169 male tergum 8; 170 male sternum 8; 171 female tergum 8; 172 female sternum 8, 173 apex of female sternum 8. Scale bar 0.2 mm (169 ­ 172), 0.1 mm (173)." httpUri="https://zenodo.org/record/155725/files/figure.png" pageId="45" pageNumber="46">Fig. 169</figureCitation>
). Male sternum 8 with convex posterior margin (
<figureCitation id="4BC24485FFB3390E4FCAFE58FBBEFE24" box="[1007,1110,484,510]" captionStart="FIGURES 169 ­ 173" captionStartId="44.[264,383,1756,1780]" captionTargetBox="[314,1272,742,1729]" captionTargetId="figure@44.[306,1280,729,1738]" captionTargetPageId="44" captionText="FIGURES 169 ­ 173. Abdominal segment 8 of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 169 male tergum 8; 170 male sternum 8; 171 female tergum 8; 172 female sternum 8, 173 apex of female sternum 8. Scale bar 0.2 mm (169 ­ 172), 0.1 mm (173)." httpUri="https://zenodo.org/record/155725/files/figure.png" pageId="45" pageNumber="46">Fig. 170</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB3390E4D1DFDB0FCF0FD91" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">
Female tergum 8 with convex posterior margin (
<figureCitation id="4BC24485FFB3390E4FA2FDB0FC18FDFC" box="[903,1008,524,550]" captionStart="FIGURES 169 ­ 173" captionStartId="44.[264,383,1756,1780]" captionTargetBox="[314,1272,742,1729]" captionTargetId="figure@44.[306,1280,729,1738]" captionTargetPageId="44" captionText="FIGURES 169 ­ 173. Abdominal segment 8 of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 169 male tergum 8; 170 male sternum 8; 171 female tergum 8; 172 female sternum 8, 173 apex of female sternum 8. Scale bar 0.2 mm (169 ­ 172), 0.1 mm (173)." httpUri="https://zenodo.org/record/155725/files/figure.png" pageId="45" pageNumber="46">Fig. 171</figureCitation>
), sternum 8 with strongly emarginate posterior margin (
<figureCitation id="4BC24485FFB3390E4E46FD8DFCE4FD91" box="[611,780,561,587]" captionStart="FIGURES 169 ­ 173" captionStartId="44.[264,383,1756,1780]" captionTargetBox="[314,1272,742,1729]" captionTargetId="figure@44.[306,1280,729,1738]" captionTargetPageId="44" captionText="FIGURES 169 ­ 173. Abdominal segment 8 of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 169 male tergum 8; 170 male sternum 8; 171 female tergum 8; 172 female sternum 8, 173 apex of female sternum 8. Scale bar 0.2 mm (169 ­ 172), 0.1 mm (173)." httpUri="https://zenodo.org/record/155725/files/figure.png" pageId="45" pageNumber="46">Figs. 172­173</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB3390E4D1DFDEBFE2EFD33" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">
Aedeagus as in
<figureCitation id="4BC24485FFB3390E4DD4FDEBFCE1FDAB" box="[497,777,599,625]" captionStart-0="FIGURES 174 ­ 177" captionStart-1="FIGURES 178 ­ 186" captionStartId-0="45.[264,383,1766,1790]" captionStartId-1="46.[264,382,1225,1249]" captionTargetBox-0="[317,1274,1164,1735]" captionTargetBox-1="[393,1195,318,1201]" captionTargetId-0="figure@45.[305,1283,1154,1742]" captionTargetId-1="figure@46.[384,1202,310,1211]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." captionText-1="FIGURES 178 ­ 186. Genitalia of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 178 details of internal sac retracted into median lobe, ventral view; 179 distal diverticula of everted internal sac, ventral view; 180 everted internal sac, ventral view; 181 spermatheca; 182 apex of left paramere, side facing median lobe; 183 ­ 184 details of internal sac retracted into median lobe, lateral view; 185 right distal diverticulum of everted internal sac, lateral view; 186 everted internal sac, lateral view. Scale bar 0.2 mm (178, 180, 183 ­ 184, 186), 0.1 mm (179, 181 ­ 182, 185)." httpUri-0="https://zenodo.org/record/155726/files/figure.png" httpUri-1="https://zenodo.org/record/155727/files/figure.png" pageId="45" pageNumber="46">Figs. 174­180, 182­186</figureCitation>
. Apex of median lobe in ventral view narrow, its outline convex apically (
<figureCitation id="4BC24485FFB3390E4E6BFDC3FD1EFD43" box="[590,758,639,665]" captionStart="FIGURES 174 ­ 177" captionStartId="45.[264,383,1766,1790]" captionTargetBox="[317,1274,1164,1735]" captionTargetId="figure@45.[305,1283,1154,1742]" captionTargetPageId="45" captionText="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." httpUri="https://zenodo.org/record/155726/files/figure.png" pageId="45" pageNumber="46">Figs. 174­175</figureCitation>
), in lateral view strait, with strong obtuse apical projection ventrally (
<figureCitation id="4BC24485FFB3390E4DDBFD1BFD4DFD1B" box="[510,677,679,705]" captionStart="FIGURES 174 ­ 177" captionStartId="45.[264,383,1766,1790]" captionTargetBox="[317,1274,1164,1735]" captionTargetId="figure@45.[305,1283,1154,1742]" captionTargetPageId="45" captionText="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." httpUri="https://zenodo.org/record/155726/files/figure.png" pageId="45" pageNumber="46">Figs. 176­177</figureCitation>
), distal diverticula of internal sac in ventral view broad (
<figureCitation id="4BC24485FFB3390E4D34FD73FE51FD33" box="[273,441,719,745]" captionStart="FIGURES 178 ­ 186" captionStartId="46.[264,382,1225,1249]" captionTargetBox="[393,1195,318,1201]" captionTargetId="figure@46.[384,1202,310,1211]" captionTargetPageId="46" captionText="FIGURES 178 ­ 186. Genitalia of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 178 details of internal sac retracted into median lobe, ventral view; 179 distal diverticula of everted internal sac, ventral view; 180 everted internal sac, ventral view; 181 spermatheca; 182 apex of left paramere, side facing median lobe; 183 ­ 184 details of internal sac retracted into median lobe, lateral view; 185 right distal diverticulum of everted internal sac, lateral view; 186 everted internal sac, lateral view. Scale bar 0.2 mm (178, 180, 183 ­ 184, 186), 0.1 mm (179, 181 ­ 182, 185)." httpUri="https://zenodo.org/record/155727/files/figure.png" pageId="45" pageNumber="46">Figs. 179­180</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB3390E4D1DFD4BFD96FCCB" blockId="45.[264,1325,284,1089]" box="[312,638,759,785]" pageId="45" pageNumber="46">
Spermatheca as in
<figureCitation id="4BC24485FFB3390E4E30FD4BFD92FCCB" box="[533,634,759,785]" captionStart="FIGURES 178 ­ 186" captionStartId="46.[264,382,1225,1249]" captionTargetBox="[393,1195,318,1201]" captionTargetId="figure@46.[384,1202,310,1211]" captionTargetPageId="46" captionText="FIGURES 178 ­ 186. Genitalia of Geostiba crepusculigena Gusarov, sp. n. (paratypes from Snake Ridge Trail, Tennessee). 178 details of internal sac retracted into median lobe, ventral view; 179 distal diverticula of everted internal sac, ventral view; 180 everted internal sac, ventral view; 181 spermatheca; 182 apex of left paramere, side facing median lobe; 183 ­ 184 details of internal sac retracted into median lobe, lateral view; 185 right distal diverticulum of everted internal sac, lateral view; 186 everted internal sac, lateral view. Scale bar 0.2 mm (178, 180, 183 ­ 184, 186), 0.1 mm (179, 181 ­ 182, 185)." httpUri="https://zenodo.org/record/155727/files/figure.png" pageId="45" pageNumber="46">Fig. 181</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB3390E4D1DFCA7FD6EFC0B" pageId="45" pageNumber="46" type="distribution">
<paragraph id="D3465800FFB3390E4D1DFCA7FD6BFC84" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">
<emphasis id="E18D8412FFB3390E4D1DFCA7FE30FCEF" bold="true" box="[312,472,795,821]" pageId="45" pageNumber="46">Distribution.</emphasis>
Known from Snake Den Mountain in the Great Smoky Mountains massif (Tennessee) (
<figureCitation id="4BC24485FFB3390E4DEDFCF8FDD1FC84" box="[456,569,836,862]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="45" pageNumber="46">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FFB3390E4E60FCF8FD9EFC84" box="[581,630,836,862]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="45" pageNumber="46">340</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB3390E4D1DFCD7FD6EFC0B" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">
<emphasis id="E18D8412FFB3390E4D1DFCD7FDEFFC5F" bold="true" box="[312,519,875,901]" pageId="45" pageNumber="46">Natural History.</emphasis>
<taxonomicName id="14F92383FFB3390E4E3DFCD0FCE5FC5F" box="[536,781,876,901]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nebuligena">
<emphasis id="E18D8412FFB3390E4E3DFCD0FCE5FC5F" box="[536,781,876,901]" italics="true" pageId="45" pageNumber="46">Geostiba nebuligena</emphasis>
</taxonomicName>
was collected at altitudes of
<quantity id="1401F5E5FFB3390E4849FCD0FAE4FC5C" box="[1132,1292,876,902]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="45" pageNumber="46" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">1400­1600 m</quantity>
in leaf litter in mixed forest with red spruce (
<taxonomicName id="14F92383FFB3390E4F24FC2EFC75FC71" box="[769,925,914,939]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="rubens">
<emphasis id="E18D8412FFB3390E4F24FC2EFC75FC71" box="[769,925,914,939]" italics="true" pageId="45" pageNumber="46">Picea rubens</emphasis>
</taxonomicName>
), birch, purple laurel (
<taxonomicName id="14F92383FFB3390E488FFC2EFE3FFC0A" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="catawbiense">
<emphasis id="E18D8412FFB3390E488FFC2EFE3FFC0A" italics="true" pageId="45" pageNumber="46">Rhododendron catawbiense</emphasis>
</taxonomicName>
) and hemlock.
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB3390E4D1DFC67FCFAFB9B" pageId="45" pageNumber="46" type="etymology">
<paragraph id="D3465800FFB3390E4D1DFC67FCFAFB9B" blockId="45.[264,1325,284,1089]" pageId="45" pageNumber="46">
<emphasis id="E18D8412FFB3390E4D1DFC67FE2DFC2F" bold="true" box="[312,453,987,1013]" pageId="45" pageNumber="46">Etymology.</emphasis>
The specific name is derived from the Latin noun
<emphasis id="E18D8412FFB3390E4866FC60FB30FC2F" box="[1091,1240,988,1013]" italics="true" pageId="45" pageNumber="46">crepusculum</emphasis>
(dusk, twilight) and the verb
<emphasis id="E18D8412FFB3390E4E2BFBBEFDB9FBC1" box="[526,593,1026,1051]" italics="true" pageId="45" pageNumber="46">gigno</emphasis>
(to be born, to arise). It refers to the dark forests in the Great Smoky Mountains where the species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>