257 lines
37 KiB
XML
257 lines
37 KiB
XML
<document id="343B90D959909DE1A91BEABBEED1B60F" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="11755326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FFA639194D2DFC7FFE0CFCCB" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Geostiba (Sibiota) coeligena Gusarov, sp. n." docType="treatment" docVersion="14" lastPageNumber="59" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="57" updateTime="1698188143631" updateUser="plazi">
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<mods:titleInfo id="AD2C404D83AC302D8F3DFE7A0CA6A17F">
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<mods:title id="55140719E0BCA17177A8C4D5FF65BDB7">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="6BFA117141D99AC6A549655847690112" type="personal">
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<mods:namePart id="C4C228381A0F3BA541335980F0211567">Gusarov, Vladimir I.</mods:namePart>
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<mods:title id="D4A44B9D1DEA8FC7DDBF30158EAD2C78">Zootaxa</mods:title>
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<mods:part id="88527F990E9F50B198A1CA7529F0FFC3">
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<mods:date id="FA7E5063503FCC24B71A22E32C11592F">2002</mods:date>
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<mods:number id="F6B63EAEFF97808B06E25C0CD6E21580">81</mods:number>
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<mods:start id="04707F3279FED08085D7971D3441C077">1</mods:start>
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<mods:classification id="197EC6CBD104DFEC985680E381EFA9E1">journal article</mods:classification>
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<mods:identifier id="BD73C6A0ACA7ACFDABE076947E6CDE9C" type="DOI">10.5281/zenodo.155701</mods:identifier>
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<treatment id="5B50E916FFA639194D2DFC7FFE0CFCCB" ID-DOI="http://doi.org/10.5281/zenodo.6277579" ID-Zenodo-Dep="6277579" LSID="urn:lsid:plazi:treatment:5B50E916FFA639194D2DFC7FFE0CFCCB" httpUri="http://treatment.plazi.org/id/5B50E916FFA639194D2DFC7FFE0CFCCB" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57">
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<subSubSection id="9BE30B8BFFA6391B4D2DFC7FFBE1FC07" box="[264,1033,963,990]" pageId="56" pageNumber="57" type="key">
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<paragraph id="D3465800FFA6391B4D2DFC7FFBE1FC07" blockId="56.[264,1033,963,990]" box="[264,1033,963,990]" pageId="56" pageNumber="57">
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<heading id="880EEF6CFFA6391B4D2DFC7FFBE1FC07" bold="true" box="[264,1033,963,990]" fontSize="11" level="1" pageId="56" pageNumber="57" reason="1">
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<keyStep id="680D40A5FFA6391B4D2DFC7FFBE1FC07" box="[264,1033,963,990]" pageId="56" pageNumber="57">
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<key id="A7680F31FFA6391B4D2DFC7FFBE1FC07" box="[264,1033,963,990]" pageId="56" pageNumber="57">
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<keyLead id="6808FB35FFA6391B4D2DFC7FFBE1FC07" box="[264,1033,963,990]" pageId="56" pageNumber="57">
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<emphasis id="E18D8412FFA6391B4D2DFC7FFBE1FC07" bold="true" box="[264,1033,963,990]" pageId="56" pageNumber="57">
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10.
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||
<taxonomicName id="14F92383FFA6391B4D11FC78FD13FC07" ID-CoL="3FSLB" authority="Gusarov" authorityName="Gusarov" box="[308,763,963,990]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="coeligena" status="sp. nov." subGenus="Sibiota">
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<emphasis id="E18D8412FFA6391B4D11FC78FE76FC04" bold="true" box="[308,414,964,990]" italics="true" pageId="56" pageNumber="57">Geostiba</emphasis>
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(
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<emphasis id="E18D8412FFA6391B4D8AFC78FDEBFC04" bold="true" box="[431,515,964,990]" italics="true" pageId="56" pageNumber="57">Sibiota</emphasis>
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||
)
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<emphasis id="E18D8412FFA6391B4E30FC78FD61FC04" bold="true" box="[533,649,964,990]" italics="true" pageId="56" pageNumber="57">coeligena</emphasis>
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Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="FABE3969FFA6391B4F2DFC7FFCA2FC07" box="[776,842,963,989]" pageId="56" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="4BC24485FFA6391B4F7FFC7FFBE9FC07" box="[858,1025,963,989]" captionStart-0="FIGURES 247 252" captionStart-1="FIGURES 253 261" captionStartId-0="57.[264,382,1726,1750]" captionStartId-1="58.[264,383,1544,1568]" captionTargetBox-0="[282,1305,745,1687]" captionTargetBox-1="[361,1227,879,1516]" captionTargetId-0="figure@57.[273,1313,729,1700]" captionTargetId-1="figure@58.[355,1231,873,1521]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionText-0="FIGURES 247 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 249, 251), 0.1 mm (250, 252)." captionText-1="FIGURES 253 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 258, 260 261 – paratype, 254 255, 259 holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 257, 259 261)." httpUri-0="https://zenodo.org/record/155735/files/figure.png" httpUri-1="https://zenodo.org/record/155736/files/figure.png" pageId="56" pageNumber="57">Figs. 247261</figureCitation>
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)
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</emphasis>
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</keyLead>
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</key>
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</keyStep>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="9BE30B8BFFA6391B4D2DFBB2FD7EFB2B" pageId="56" pageNumber="57" type="materials_examined">
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||
<paragraph id="D3465800FFA6391B4D2DFBB2FD41FB7B" blockId="56.[264,1324,1038,1864]" pageId="56" pageNumber="57">
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||
<emphasis id="E18D8412FFA6391B4D2DFBB2FE56FBF2" bold="true" box="[264,446,1038,1064]" pageId="56" pageNumber="57">
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<typeStatus id="0C42E6A2FFA6391B4D2DFBB2FEADFBF2" box="[264,325,1038,1064]" pageId="56" pageNumber="57">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="0C42E6A2FFA6391B4DEAFBB3FDD7FBF3" box="[463,575,1039,1065]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
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,,
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<emphasis id="E18D8412FFA6391B4E4AFBB2FB27FBF2" bold="true" box="[623,1231,1038,1064]" pageId="56" pageNumber="57">
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<collectingCountry id="ABEE1890FFA6391B4E4AFBB2FCB2FBF2" box="[623,858,1038,1064]" name="United States of America" pageId="56" pageNumber="57">UNITED STATES</collectingCountry>
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: Tennessee / North Carolina:
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</emphasis>
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Sevier Co. / Swain Co.: Appalachian National Scenic Trail, W of Clingmans Dome,
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<quantity id="1401F5E5FFA6391B48E6FB8BFAF8FB8B" box="[1219,1296,1079,1105]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="56" pageNumber="57" unit="km" value="16.0">16 km</quantity>
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S Gatlinburg,
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<geoCoordinate id="B6CD3EC7FFA6391B4DB2FBE3FDF5FBA3" box="[407,541,1119,1145]" direction="north" orientation="latitude" pageId="56" pageNumber="57" precision="9" value="35.564667">35°33.88'N</geoCoordinate>
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<geoCoordinate id="B6CD3EC7FFA6391B4E00FBE3FD59FBA3" box="[549,689,1119,1145]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="9" value="-83.5235">83°31.41'W</geoCoordinate>
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,
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||
<quantity id="1401F5E5FFA6391B4E9AFBE3FCF4FBA3" box="[703,796,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="56" pageNumber="57" unit="m" value="1930.0">1930 m</quantity>
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, in forest litter,
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<taxonomicName id="14F92383FFA6391B4FF8FBE3FBF7FBA2" box="[989,1055,1119,1144]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
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<emphasis id="E18D8412FFA6391B4FF8FBE3FBF7FBA2" box="[989,1055,1119,1144]" italics="true" pageId="56" pageNumber="57">Picea</emphasis>
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</taxonomicName>
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,
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||
<taxonomicName id="14F92383FFA6391B4815FBE3FB99FBA2" box="[1072,1137,1119,1144]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="E18D8412FFA6391B4815FBE3FB99FBA2" box="[1072,1137,1119,1144]" italics="true" pageId="56" pageNumber="57">Abies</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicName id="14F92383FFA6391B48A4FBE3FB25FBA2" box="[1153,1229,1119,1144]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="E18D8412FFA6391B48A4FBE3FB25FBA2" box="[1153,1229,1119,1144]" italics="true" pageId="56" pageNumber="57">Betula</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="14F92383FFA6391B48FBFBE3FE43FB7B" authority="V.I.Gusarov" authorityName="V.I.Gusarov" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
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<emphasis id="E18D8412FFA6391B48FBFBE3FAC2FBA2" box="[1246,1322,1119,1144]" italics="true" pageId="56" pageNumber="57">Oxalis</emphasis>
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||
(V.I.Gusarov)
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</taxonomicName>
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||
,
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||
<date id="A7477EC0FFA6391B4D92FB3BFDDCFB7B" box="[439,564,1159,1185]" pageId="56" pageNumber="57" value="2001-06-22">22.vi.2001</date>
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||
(
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||
<collectionCode id="B5E8C0C5FFA6391B4E61FB3BFD74FB7B" box="[580,668,1159,1185]" pageId="56" pageNumber="57">KSEM</collectionCode>
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||
).
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||
</paragraph>
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||
<paragraph id="D3465800FFA6391B4D1DFB13FD7EFB2B" blockId="56.[264,1324,1038,1864]" pageId="56" pageNumber="57">
|
||
<typeStatus id="0C42E6A2FFA6391B4D1DFB13FE4BFB13" box="[312,419,1199,1225]" pageId="56" pageNumber="57" type="paratype">Paratype</typeStatus>
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||
:, as the
|
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<typeStatus id="0C42E6A2FFA6391B4E1FFB13FD48FB13" box="[570,672,1199,1225]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
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but
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||
<geoCoordinate id="B6CD3EC7FFA6391B4EF8FB13FC8BFB13" box="[733,867,1199,1225]" direction="north" orientation="latitude" pageId="56" pageNumber="57" precision="9" value="35.562332">35°33.74'N</geoCoordinate>
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||
<geoCoordinate id="B6CD3EC7FFA6391B4F4AFB13FC13FB13" box="[879,1019,1199,1225]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="9" value="-83.514">83°30.84'W</geoCoordinate>
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||
,
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||
<quantity id="1401F5E5FFA6391B4829FB13FB84FB13" box="[1036,1132,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="56" pageNumber="57" unit="m" value="1950.0">1950 m</quantity>
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, in forest litter,
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||
<taxonomicName id="14F92383FFA6391B4D2DFB6BFEA1FB2A" box="[264,329,1239,1264]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E18D8412FFA6391B4D2DFB6BFEA1FB2A" box="[264,329,1239,1264]" italics="true" pageId="56" pageNumber="57">Abies</emphasis>
|
||
</taxonomicName>
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||
,
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||
<taxonomicName id="14F92383FFA6391B4D7DFB6BFE3AFB2A" box="[344,466,1239,1264]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="E18D8412FFA6391B4D7DFB6BFE3AFB2A" box="[344,466,1239,1264]" italics="true" pageId="56" pageNumber="57">Vaccinium</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicName id="14F92383FFA6391B4DC4FB6BFDC5FB2A" box="[481,557,1239,1264]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="56" pageNumber="57" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E18D8412FFA6391B4DC4FB6BFDC5FB2A" box="[481,557,1239,1264]" italics="true" pageId="56" pageNumber="57">Oxalis</emphasis>
|
||
</taxonomicName>
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||
(
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||
<collectionCode id="B5E8C0C5FFA6391B4E18FB6BFD61FB2B" box="[573,649,1239,1265]" pageId="56" pageNumber="57">SPSU</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFA6391B4D1DFB42FC35F982" pageId="56" pageNumber="57" type="diagnosis">
|
||
<paragraph id="D3465800FFA6391B4D1DFB42FEADFA63" blockId="56.[264,1324,1038,1864]" pageId="56" pageNumber="57">
|
||
<emphasis id="E18D8412FFA6391B4D1DFB42FE5FFAC2" bold="true" box="[312,439,1278,1304]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
|
||
<taxonomicName id="14F92383FFA6391B4DE9FB43FD5AFAC2" box="[460,690,1279,1304]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="coeligena">
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<emphasis id="E18D8412FFA6391B4DE9FB43FD5AFAC2" box="[460,690,1279,1304]" italics="true" pageId="56" pageNumber="57">Geostiba coeligena</emphasis>
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</taxonomicName>
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can be distinguished from other Nearctic species of
|
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<taxonomicName id="14F92383FFA6391B4D2DFA9BFE9AFA9A" box="[264,370,1319,1344]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FFA6391B4D2DFA9BFE9AFA9A" box="[264,370,1319,1344]" italics="true" pageId="56" pageNumber="57">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
by having small eyes (temple length to eye length ratio 4.05.6), pronotal pubescence of
|
||
<typeStatus id="0C42E6A2FFA6391B4D56FAF3FE4DFAB3" box="[371,421,1359,1385]" pageId="56" pageNumber="57">type</typeStatus>
|
||
V, reduced wings, short elytra (pronotum length to elytron length ratio 1.4), the absence of carinae on abdominal tergum 7 and the shape of the aedeagus (
|
||
<figureCitation id="4BC24485FFA6391B4897FACBFED0FA63" captionStart-0="FIGURES 247 252" captionStart-1="FIGURES 253 261" captionStartId-0="57.[264,382,1726,1750]" captionStartId-1="58.[264,383,1544,1568]" captionTargetBox-0="[282,1305,745,1687]" captionTargetBox-1="[361,1227,879,1516]" captionTargetId-0="figure@57.[273,1313,729,1700]" captionTargetId-1="figure@58.[355,1231,873,1521]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionText-0="FIGURES 247 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 249, 251), 0.1 mm (250, 252)." captionText-1="FIGURES 253 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 258, 260 261 – paratype, 254 255, 259 holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 257, 259 261)." httpUri-0="https://zenodo.org/record/155735/files/figure.png" httpUri-1="https://zenodo.org/record/155736/files/figure.png" pageId="56" pageNumber="57">Figs. 249 261</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFA6391B4D1DFA7BFC35F982" blockId="56.[264,1324,1038,1864]" pageId="56" pageNumber="57">
|
||
<taxonomicName id="14F92383FFA6391B4D1DFA7BFDF3FA3A" box="[312,539,1479,1504]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="coeligena">
|
||
<emphasis id="E18D8412FFA6391B4D1DFA7BFDF3FA3A" box="[312,539,1479,1504]" italics="true" pageId="56" pageNumber="57">Geostiba coeligena</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="14F92383FFA6391B4E93FA7BFC81FA3A" box="[694,873,1479,1504]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="balsamensis">
|
||
<emphasis id="E18D8412FFA6391B4E93FA7BFC81FA3A" box="[694,873,1479,1504]" italics="true" pageId="56" pageNumber="57">G. balsamensis</emphasis>
|
||
</taxonomicName>
|
||
in lacking the carinae on male tergum 7; from
|
||
<taxonomicName id="14F92383FFA6391B4D4EFA53FE16F9D2" box="[363,510,1519,1544]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="nubigena">
|
||
<emphasis id="E18D8412FFA6391B4D4EFA53FE16F9D2" box="[363,510,1519,1544]" italics="true" pageId="56" pageNumber="57">G. nubigena</emphasis>
|
||
</taxonomicName>
|
||
in having broader median lobe of aedeagus (in ventral view) (
|
||
<figureCitation id="4BC24485FFA6391B48CBFA52FE85F9EA" captionStart="FIGURES 247 252" captionStartId="57.[264,382,1726,1750]" captionTargetBox="[282,1305,745,1687]" captionTargetId="figure@57.[273,1313,729,1700]" captionTargetPageId="57" captionText="FIGURES 247 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 249, 251), 0.1 mm (250, 252)." httpUri="https://zenodo.org/record/155735/files/figure.png" pageId="56" pageNumber="57">Figs. 249250</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FFA6391B4D5EF9AAFDBBF9EA" box="[379,595,1558,1584]" captionStart="FIGURES 219 229" captionStartId="54.[264,383,1720,1744]" captionTargetBox="[333,1253,320,1706]" captionTargetId="figure@54.[323,1267,310,1709]" captionTargetPageId="54" captionText="FIGURES 219 229. Aedeagus of Geostiba nubigena Lohse & Smetana (Richland Balsam Mountain (219 222), Blue Ridge Parkway (223), Graveyard Fields (224 227) and Grassy Ridge Mine Overlook (228 229), North Carolina). 219, 224, 226, 228 – median lobe, ventral view; 220, 223, 225, 227, 229 – apex of median lobe, ventral view; 221 – median lobe, lateral view; 222 – apex of median lobe, lateral view. Scale bar 0.2 mm (219, 221, 224, 226, 228), 0.1 mm (220, 222 223, 225, 227, 229)." httpUri="https://zenodo.org/record/155733/files/figure.png" pageId="56" pageNumber="57">219220, 223229</figureCitation>
|
||
), short distal diverticula (
|
||
<figureCitation id="4BC24485FFA6391B4FA2F9AAFBDFF9EA" box="[903,1079,1558,1584]" captionStart="FIGURES 253 261" captionStartId="58.[264,383,1544,1568]" captionTargetBox="[361,1227,879,1516]" captionTargetId="figure@58.[355,1231,873,1521]" captionTargetPageId="58" captionText="FIGURES 253 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 258, 260 261 – paratype, 254 255, 259 holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 257, 259 261)." httpUri="https://zenodo.org/record/155736/files/figure.png" pageId="56" pageNumber="57">Figs. 253, 256</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FFA6391B4860F9AAFB40F9EA" box="[1093,1192,1558,1584]" captionStart="FIGURES 230 246" captionStartId="55.[264,382,1503,1527]" captionTargetBox="[316,1271,321,1479]" captionTargetId="figure@55.[305,1282,310,1487]" captionTargetPageId="55" captionText="FIGURES 230 246. Genitalia of Geostiba nubigena Lohse & Smetana (Richland Balsam Mountain (230 231, 234 235, 237 238, 242, 246) and Blue Ridge Parkway (232 233, 236, 239 241, 243 245), North Carolina). 230 – details of internal sac retracted into median lobe, ventral view; 231 – copulatory piece of internal sac, dorsal view; 232 – medial lamellae of internal sac, ventral view; 233 – distal diverticula of everted internal sac, ventral view; 234 – everted internal sac, ventral view; 235 236 – spermatheca; 237 238 – details of internal sac retracted into median lobe, lateral view; 239 – partially everted internal sac, lateral view; 240 – details of internal sac inside partially everted internal sac, lateral view; 241 – everted internal sac, lateral view; 242 – completely everted internal sac, lateral view; 243 – copulatory piece of internal sac, lateral view; 244 – medial lamella of internal sac, lateral view; 245 – right distal diverticulum of everted internal sac, lateral view; 246 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (230, 234, 237 242), 0.1 mm (231 233, 235 236, 243 246)." httpUri="https://zenodo.org/record/155734/files/figure.png" pageId="56" pageNumber="57">233234</figureCitation>
|
||
) and large proximal diverticula of internal sac (
|
||
<figureCitation id="4BC24485FFA6391B4E96F982FC7CF982" box="[691,916,1598,1624]" captionStart="FIGURES 253 261" captionStartId="58.[264,383,1544,1568]" captionTargetBox="[361,1227,879,1516]" captionTargetId="figure@58.[355,1231,873,1521]" captionTargetPageId="58" captionText="FIGURES 253 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 258, 260 261 – paratype, 254 255, 259 holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 257, 259 261)." httpUri="https://zenodo.org/record/155736/files/figure.png" pageId="56" pageNumber="57">Figs. 253, 257258</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FFA6391B4F85F982FC39F982" box="[928,977,1598,1624]" captionStart="FIGURES 230 246" captionStartId="55.[264,382,1503,1527]" captionTargetBox="[316,1271,321,1479]" captionTargetId="figure@55.[305,1282,310,1487]" captionTargetPageId="55" captionText="FIGURES 230 246. Genitalia of Geostiba nubigena Lohse & Smetana (Richland Balsam Mountain (230 231, 234 235, 237 238, 242, 246) and Blue Ridge Parkway (232 233, 236, 239 241, 243 245), North Carolina). 230 – details of internal sac retracted into median lobe, ventral view; 231 – copulatory piece of internal sac, dorsal view; 232 – medial lamellae of internal sac, ventral view; 233 – distal diverticula of everted internal sac, ventral view; 234 – everted internal sac, ventral view; 235 236 – spermatheca; 237 238 – details of internal sac retracted into median lobe, lateral view; 239 – partially everted internal sac, lateral view; 240 – details of internal sac inside partially everted internal sac, lateral view; 241 – everted internal sac, lateral view; 242 – completely everted internal sac, lateral view; 243 – copulatory piece of internal sac, lateral view; 244 – medial lamella of internal sac, lateral view; 245 – right distal diverticulum of everted internal sac, lateral view; 246 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (230, 234, 237 242), 0.1 mm (231 233, 235 236, 243 246)." httpUri="https://zenodo.org/record/155734/files/figure.png" pageId="56" pageNumber="57">234</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFA639194D1DF9DAFDE3FE0C" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57" type="description">
|
||
<paragraph id="D3465800FFA6391B4D1DF9DAFBF4F972" blockId="56.[264,1324,1038,1864]" pageId="56" pageNumber="57">
|
||
<emphasis id="E18D8412FFA6391B4D1DF9DAFE38F95A" bold="true" box="[312,464,1638,1664]" pageId="56" pageNumber="57">Description.</emphasis>
|
||
Length
|
||
<quantity id="1401F5E5FFA6391B4E67F9DAFD3AF95A" box="[578,722,1638,1664]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="56" pageNumber="57" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9 mm</quantity>
|
||
. Body brownish yellow to brown, antennae light brown, mouthparts and legs brownish yellow. Body parallelsided.
|
||
</paragraph>
|
||
<paragraph id="D3465800FFA6391B4D1DF90AFB9DF892" blockId="56.[264,1324,1038,1864]" pageId="56" pageNumber="57">Head as wide as long, surface on disk with fine isodiametric microsculpture, puncturation very fine, distance between punctures equal to 34 times their diameter. Temple length to eye length ratio 4.05.6. Antennal article 2 longer than article 3, article 410 transverse to strongly transverse, last article as long as 9 and 10 combined.</paragraph>
|
||
<paragraph id="D3465800FFA7391A4D1DFEA0FEBAFDFC" blockId="57.[264,1324,284,670]" pageId="57" pageNumber="58">
|
||
Pronotum as wide as long, width
|
||
<quantity id="1401F5E5FFA7391A4EE8FEA0FCD3FEEC" box="[717,827,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="57" pageNumber="58" unit="mm" value="0.34">0.34 mm</quantity>
|
||
, wider than head (pronotal width to head width ratio 1.1); microsculpture and puncturation as on head. Pronotal pubescence of
|
||
<typeStatus id="0C42E6A2FFA7391A48DCFEF8FAC4FE84" box="[1273,1324,324,350]" pageId="57" pageNumber="58">type</typeStatus>
|
||
V. Elytra measured from humeral angle shorter than pronotum (pronotal length to elytral length ratio 1.4), wider than long (1.5), with fine isodiametric microsculpture and fine asperate puncturation, distance between punctures equals 23 times their diameter. Elytral suture behind scutellum very slightly raised. Wings reduced to short vestiges, shorter than elytra.
|
||
</paragraph>
|
||
<paragraph id="D3465800FFA7391A4D1DFD88FB22FD44" blockId="57.[264,1324,284,670]" pageId="57" pageNumber="58">Abdominal terga with fine microsculpture of transverse meshes, with fine and sparse puncturation, puncturation becoming finer towards abdomen apex, on terga 35 distance between punctures equals 36 times their diameter. Tergum 7 without white edge.</paragraph>
|
||
<caption id="87860888FFA7391A4D2DF902FC29F8E4" httpUri="https://zenodo.org/record/155735/files/figure.png" pageId="57" pageNumber="58" targetBox="[282,1305,745,1687]" targetPageId="57">
|
||
<paragraph id="D3465800FFA7391A4D2DF902FC29F8E4" blockId="57.[264,1323,1726,1854]" pageId="57" pageNumber="58">
|
||
<emphasis id="E18D8412FFA7391A4D2DF902FE08F90C" bold="true" box="[264,480,1726,1750]" pageId="57" pageNumber="58">FIGURES 247252</emphasis>
|
||
. Abdominal segment 8 and aedeagus of
|
||
<taxonomicName id="14F92383FFA7391A4F8EF903FB08F90C" authority="Gusarov" authorityName="Gusarov" box="[939,1248,1727,1750]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="coeligena" status="sp. nov.">
|
||
<emphasis id="E18D8412FFA7391A4F8EF903FB94F90C" box="[939,1148,1727,1750]" italics="true" pageId="57" pageNumber="58">Geostiba coeligena</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FABE3969FFA7391A48C9F902FAC3F90C" box="[1260,1323,1726,1750]" pageId="57" pageNumber="58" rank="species">
|
||
<emphasis id="E18D8412FFA7391A48C9F902FAC3F90C" bold="true" box="[1260,1323,1726,1750]" pageId="57" pageNumber="58">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247249, 251), 0.1 mm (250, 252).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D3465800FFA439194D1DFEA0FE10FE84" blockId="58.[264,1324,284,785]" pageId="58" pageNumber="59">
|
||
Male tergum 7 without carinae. Posterior margin of male tergum 8 and sternum 8 convex (
|
||
<figureCitation id="4BC24485FFA439194D67FEF8FE03FE84" box="[322,491,324,350]" captionStart="FIGURES 247 252" captionStartId="57.[264,382,1726,1750]" captionTargetBox="[282,1305,745,1687]" captionTargetId="figure@57.[273,1313,729,1700]" captionTargetPageId="57" captionText="FIGURES 247 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 249, 251), 0.1 mm (250, 252)." httpUri="https://zenodo.org/record/155735/files/figure.png" pageId="58" pageNumber="59">Figs. 247248</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFA439194D1DFED0FEADFE74" blockId="58.[264,1324,284,785]" pageId="58" pageNumber="59">
|
||
Aedeagus as in
|
||
<figureCitation id="4BC24485FFA439194DD1FED0FD75FE5C" box="[500,669,364,390]" captionStart-0="FIGURES 247 252" captionStart-1="FIGURES 253 261" captionStartId-0="57.[264,382,1726,1750]" captionStartId-1="58.[264,383,1544,1568]" captionTargetBox-0="[282,1305,745,1687]" captionTargetBox-1="[361,1227,879,1516]" captionTargetId-0="figure@57.[273,1313,729,1700]" captionTargetId-1="figure@58.[355,1231,873,1521]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionText-0="FIGURES 247 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 249, 251), 0.1 mm (250, 252)." captionText-1="FIGURES 253 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 258, 260 261 – paratype, 254 255, 259 holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 257, 259 261)." httpUri-0="https://zenodo.org/record/155735/files/figure.png" httpUri-1="https://zenodo.org/record/155736/files/figure.png" pageId="58" pageNumber="59">Figs. 249261</figureCitation>
|
||
. Apex of median lobe in lateral view strait (
|
||
<figureCitation id="4BC24485FFA439194897FED0FED1FE74" captionStart="FIGURES 247 252" captionStartId="57.[264,382,1726,1750]" captionTargetBox="[282,1305,745,1687]" captionTargetId="figure@57.[273,1313,729,1700]" captionTargetPageId="57" captionText="FIGURES 247 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 249, 251), 0.1 mm (250, 252)." httpUri="https://zenodo.org/record/155735/files/figure.png" pageId="58" pageNumber="59">Figs. 251 252</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFA439194D1DFE00FDE3FE0C" blockId="58.[264,1324,284,785]" box="[312,523,444,470]" pageId="58" pageNumber="59">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFA439194D1DFE5FFDF9FD44" pageId="58" pageNumber="59" type="distribution">
|
||
<paragraph id="D3465800FFA439194D1DFE5FFDF2FDFC" blockId="58.[264,1324,284,785]" pageId="58" pageNumber="59">
|
||
<emphasis id="E18D8412FFA439194D1DFE5FFE30FE27" bold="true" box="[312,472,483,509]" pageId="58" pageNumber="59">Distribution.</emphasis>
|
||
Known from the Great Smoky Mountains massif west of Clingmans Dome (
|
||
<figureCitation id="4BC24485FFA439194D45FDB0FE39FDFC" box="[352,465,524,550]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="58" pageNumber="59">Figs. 338</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FFA439194DF8FDB0FDE6FDFC" box="[477,526,524,550]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse & Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse & Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse & Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="58" pageNumber="59">340</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFA439194D1DFD8FFDF9FD44" blockId="58.[264,1324,284,785]" pageId="58" pageNumber="59">
|
||
<emphasis id="E18D8412FFA439194D1DFD8FFDEFFD97" bold="true" box="[312,519,563,589]" pageId="58" pageNumber="59">Natural History.</emphasis>
|
||
<taxonomicName id="14F92383FFA439194E3DFD88FD15FD97" box="[536,765,564,589]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="coeligena">
|
||
<emphasis id="E18D8412FFA439194E3DFD88FD15FD97" box="[536,765,564,589]" italics="true" pageId="58" pageNumber="59">Geostiba coeligena</emphasis>
|
||
</taxonomicName>
|
||
was collected in forest litter at altitudes above
|
||
<quantity id="1401F5E5FFA439194D2DFDE0FE8BFDAC" box="[264,355,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="58" pageNumber="59" unit="m" value="1900.0">1900 m</quantity>
|
||
in leaf litter mostly in coniferous forest with Fraser’s fir (
|
||
<taxonomicName id="14F92383FFA439194800FDE0FB56FDAF" box="[1061,1214,604,629]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="fraseri">
|
||
<emphasis id="E18D8412FFA439194800FDE0FB56FDAF" box="[1061,1214,604,629]" italics="true" pageId="58" pageNumber="59">Abies fraseri</emphasis>
|
||
</taxonomicName>
|
||
) and red spruce (
|
||
<taxonomicName id="14F92383FFA439194D43FD38FDE8FD47" box="[358,512,644,669]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="rubens">
|
||
<emphasis id="E18D8412FFA439194D43FD38FDE8FD47" box="[358,512,644,669]" italics="true" pageId="58" pageNumber="59">Picea rubens</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFA439194D1DFD17FE0CFCCB" pageId="58" pageNumber="59" type="etymology">
|
||
<paragraph id="D3465800FFA439194D1DFD17FE0CFCCB" blockId="58.[264,1324,284,785]" pageId="58" pageNumber="59">
|
||
<emphasis id="E18D8412FFA439194D1DFD17FE2DFD1F" bold="true" box="[312,453,683,709]" pageId="58" pageNumber="59">Etymology.</emphasis>
|
||
The specific name is derived from the Latin noun
|
||
<emphasis id="E18D8412FFA43919481CFD10FB66FD1F" box="[1081,1166,684,709]" italics="true" pageId="58" pageNumber="59">coelum</emphasis>
|
||
(the sky, the heavens) and the verb
|
||
<emphasis id="E18D8412FFA439194E33FD6EFDB1FD31" box="[534,601,722,747]" italics="true" pageId="58" pageNumber="59">gigno</emphasis>
|
||
(to be born, to arise). It refers to the high mountains where the species occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |