425 lines
55 KiB
XML
425 lines
55 KiB
XML
<document id="971DC5CE997D5CD043E4EACED876E1EB" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="11755326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FF8F39314D2DFE5FFB97FB2C" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Sibiota Casey 1906" docType="treatment" docVersion="14" lastPageNumber="19" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="18" updateTime="1698188143631" updateUser="plazi">
|
||
<mods:mods id="2CBF184D49837D9C5C8A917A09590A85" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E3AE3A291CE7792F967DEA5A2A5440D2">
|
||
<mods:title id="B669B118156B2E95F8EEEADBA88BC70F">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1837F6BFA1D6292C8EB8AE37FD4839ED" type="personal">
|
||
<mods:role id="B934DCCAD2334007172BE50AB53DCB2A">
|
||
<mods:roleTerm id="11D09D8E19E83F55BA9B98CF6FF8F6F1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8DA35CFAAC2720F5FE243236008B2956">Gusarov, Vladimir I.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8EB7C5FAE2461B896761DD842FBD6F54">text</mods:typeOfResource>
|
||
<mods:relatedItem id="CB8B209BA70299C4DE14DC759F427D30" type="host">
|
||
<mods:titleInfo id="BB86B57094C7FDB5DEEC114AF4CE0A2B">
|
||
<mods:title id="DE969329506CEE4FFFEF1300C306BFC8">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="AE34D351AD0DCF7C1C4266D09D9DC138">
|
||
<mods:date id="EF0C4B9750EEEF63BA420F8165281D98">2002</mods:date>
|
||
<mods:detail id="EBF2312AE690E5C6E4297F2A1E6A5614" type="volume">
|
||
<mods:number id="DBFA9401605D9C99B8BF1C5E98347F26">81</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="56D1C01F78A07B2C6ECCF7FCBA0B6A7E" unit="page">
|
||
<mods:start id="43A7832262AE3799634805039534B544">1</mods:start>
|
||
<mods:end id="204C0CAD955CE44A0F7B202F38E38CCF">88</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D75EA08FB24CE9BF7199793ED99E79F7">journal article</mods:classification>
|
||
<mods:identifier id="9D94999FD097BEF9877B7C2313138997" type="DOI">10.5281/zenodo.155701</mods:identifier>
|
||
<mods:identifier id="778E4CB7DCD95D23F8872C8DEB00B6FF" type="GBIF-Dataset">b947e6c9-d6cf-4988-a098-5e01dd69a888</mods:identifier>
|
||
<mods:identifier id="1ABD335353B99D68F0EF026DC2DB9350" type="ISSN">11755326</mods:identifier>
|
||
<mods:identifier id="D53E65A45763EA087DCCBA83955EC308" type="Zenodo-Dep">155701</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5B50E916FF8F39314D2DFE5FFB97FB2C" ID-DOI="http://doi.org/10.5281/zenodo.6277556" ID-GBIF-Taxon="119330229" ID-Zenodo-Dep="6277556" LSID="urn:lsid:plazi:treatment:5B50E916FF8F39314D2DFE5FFB97FB2C" httpUri="http://treatment.plazi.org/id/5B50E916FF8F39314D2DFE5FFB97FB2C" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
<subSubSection id="9BE30B8BFF8F39324D2DFE5FFD9FFE27" box="[264,631,483,510]" pageId="17" pageNumber="18" type="nomenclature">
|
||
<paragraph id="D3465800FF8F39324D2DFE5FFD9FFE27" blockId="17.[264,631,483,510]" box="[264,631,483,510]" pageId="17" pageNumber="18">
|
||
<heading id="880EEF6CFF8F39324D2DFE5FFD9FFE27" bold="true" box="[264,631,483,510]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
|
||
<emphasis id="E18D8412FF8F39324D2DFE5FFD9FFE27" bold="true" box="[264,631,483,510]" pageId="17" pageNumber="18">
|
||
Subgenus
|
||
<taxonomicName id="14F92383FF8F39324DADFE58FD9FFE27" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[392,631,483,510]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324DADFE58FE35FE24" bold="true" box="[392,477,484,510]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
<bibRefCitation id="B76825F1FF8F39324DC1FE5FFD9FFE27" author="Casey" box="[484,631,483,509]" pageId="17" pageNumber="87" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFF8F39324D2DFD93FB3FFC9C" pageId="17" pageNumber="18" type="reference_group">
|
||
<paragraph id="D3465800FF8F39324D2DFD93FCBBFDBC" blockId="17.[264,1324,559,870]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFD93FCBBFDBC" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFD93FDF4FD9C" authority="Casey, 1906: 350" authorityName="Casey" authorityPageNumber="350" authorityYear="1906" box="[264,540,559,582]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFD93FEBBFD9C" box="[264,339,559,582]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
<treatmentCitation id="52587E11FF8F39324D7EFD93FDF4FD9C" author="Casey" box="[347,540,559,582]" page="350" pageId="17" pageNumber="18" year="1906">
|
||
<bibRefCitation id="B76825F1FF8F39324D7EFD93FE09FD9C" author="Casey" box="[347,481,559,582]" pageId="17" pageNumber="87" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
|
||
: 350
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="0C42E6A2FF8F39324E0FFD93FD89FD9C" box="[554,609,559,582]" pageId="17" pageNumber="18">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="14F92383FF8F39324EE9FD93FBF7FD9C" ID-CoL="9K77Q" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[716,1055,559,582]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="impressula">
|
||
<emphasis id="E18D8412FF8F39324EE9FD93FC7AFD9C" box="[716,914,559,582]" italics="true" pageId="17" pageNumber="18">Sibiota impressula</emphasis>
|
||
<bibRefCitation id="B76825F1FF8F39324FBFFD93FBF7FD9C" author="Casey" box="[922,1055,559,582]" pageId="17" pageNumber="87" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
|
||
</taxonomicName>
|
||
, by original designation) (as a genus in tribe
|
||
<taxonomicName id="14F92383FF8F39324E2DFDF3FCAFFDBC" ID-CoL="8VWQ5" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[520,839,591,614]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Bolitocharini">Bolitocharini Thomson, 1859</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D2DFDD3FBC4FD5C" blockId="17.[264,1324,559,870]" box="[264,1068,623,646]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFDD3FBC4FD5C" box="[264,1068,623,646]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFDD3FDDBFD5C" authority="Fenyes, 1920: 249" authorityName="Fenyes" authorityPageNumber="249" authorityYear="1920" box="[264,563,623,646]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFDD3FEBCFD5C" box="[264,340,623,646]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
:
|
||
<treatmentCitation id="52587E11FF8F39324D4DFDD3FDDBFD5C" author="Fenyes" box="[360,563,623,646]" page="249" pageId="17" pageNumber="18" year="1920">
|
||
<bibRefCitation id="B76825F1FF8F39324D4DFDD3FE12FD5C" author="Fenyes" box="[360,506,623,646]" pageId="17" pageNumber="87" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. Bruxelles: L. Desmet-Verteneuil, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
|
||
: 249
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(as synonym of
|
||
<taxonomicName id="14F92383FF8F39324EC0FDD3FBC8FD5C" authority="Mulsant & Rey, 1853" authorityName="Mulsant & Rey" authorityYear="1853" box="[741,1056,623,646]" class="Insecta" family="Staphylinidae" genus="Sipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324EC0FDD3FCD8FD5C" box="[741,816,623,646]" italics="true" pageId="17" pageNumber="18">Sipalia</emphasis>
|
||
<bibRefCitation id="B76825F1FF8F39324F12FDD3FBC8FD5C" author="Mulsant" box="[823,1056,623,646]" pageId="17" pageNumber="88" refString="Mulsant, E. & Rey, C. (1853) Description de quelques Coleopteres nouveaux ou peu connus, de la tribu des Brachelytres. Annales de la Societe Linneenne de Lyon, ser. 2, 1, 22 - 72." type="journal article" year="1853">Mulsant & Rey, 1853</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D2DFD33FBEAFD7C" blockId="17.[264,1324,559,870]" box="[264,1026,655,678]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFD33FBEAFD7C" box="[264,1026,655,678]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFD33FD1EFD7C" authority="Bernhauer & Scheerpeltz, 1926: 599" authorityName="Bernhauer & Scheerpeltz" authorityPageNumber="599" authorityYear="1926" box="[264,758,655,678]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFD33FEBCFD7C" box="[264,340,655,678]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
:
|
||
<treatmentCitation id="52587E11FF8F39324D4DFD33FD1EFD7C" author="Bernhauer" box="[360,758,655,678]" page="599" pageId="17" pageNumber="18" year="1926">
|
||
<bibRefCitation id="B76825F1FF8F39324D4DFD33FD55FD7C" author="Bernhauer" box="[360,701,655,678]" pageId="17" pageNumber="87" refString="Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. Berlin: W. Junk, pp. 499 - 988." type="journal article" year="1926">Bernhauer & Scheerpeltz, 1926</bibRefCitation>
|
||
: 599
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(as synonym of
|
||
<taxonomicName id="14F92383FF8F39324F82FD33FC1BFD7C" box="[935,1011,655,678]" class="Insecta" family="Staphylinidae" genus="Sipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324F82FD33FC1BFD7C" box="[935,1011,655,678]" italics="true" pageId="17" pageNumber="18">Sipalia</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D2DFD13FCF3FD3C" blockId="17.[264,1324,559,870]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFD13FCF3FD3C" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFD13FD40FD1C" authority="Scheerpeltz, 1951: 172" authorityName="Scheerpeltz" authorityPageNumber="172" authorityYear="1951" box="[264,680,687,710]" class="Insecta" family="Staphylinidae" genus="Ditroposipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFD13FE4CFD1C" box="[264,420,687,710]" italics="true" pageId="17" pageNumber="18">Ditroposipalia</emphasis>
|
||
<treatmentCitation id="52587E11FF8F39324D8EFD13FD40FD1C" author="Scheerpeltz" box="[427,680,687,710]" page="172" pageId="17" pageNumber="18" year="1951">
|
||
<bibRefCitation id="B76825F1FF8F39324D8EFD13FD86FD1C" author="Scheerpeltz" box="[427,622,687,710]" pageId="17" pageNumber="88" refString="Scheerpeltz, O. (1951) Die neue Systematik der Gattung Sipalia Muls. Rey (Col. Staphylinidae). Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien, 92, 166 - 180." type="journal article" year="1951">Scheerpeltz, 1951</bibRefCitation>
|
||
: 172
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="0C42E6A2FF8F39324E93FD13FD05FD1C" box="[694,749,687,710]" pageId="17" pageNumber="18">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="14F92383FF8F39324F70FD13FB6AFD1C" authority="Baudi, 1869" authorityName="Baudi" authorityYear="1869" box="[853,1154,687,710]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bidens">
|
||
<emphasis id="E18D8412FF8F39324F70FD13FC1FFD1C" box="[853,1015,687,710]" italics="true" pageId="17" pageNumber="18">Leptusa bidens</emphasis>
|
||
Baudi, 1869
|
||
</taxonomicName>
|
||
, by original designation) (as subgenus of
|
||
<taxonomicName id="14F92383FF8F39324E73FD73FD49FD3C" box="[598,673,719,742]" class="Insecta" family="Staphylinidae" genus="Sipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324E73FD73FD49FD3C" box="[598,673,719,742]" italics="true" pageId="17" pageNumber="18">Sipalia</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<emphasis id="E18D8412FF8F39324E9DFD72FCF3FD3C" bold="true" box="[696,795,718,742]" pageId="17" pageNumber="18">syn. nov.</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D2DFD53FB6AFCDC" blockId="17.[264,1324,559,870]" box="[264,1154,751,774]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFD53FB6AFCDC" box="[264,1154,751,774]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFD53FDD3FCDC" authority="Seevers, 1978: 128" authorityName="Seevers" authorityPageNumber="128" authorityYear="1978" box="[264,571,751,774]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFD53FEBCFCDC" box="[264,340,751,774]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
:
|
||
<treatmentCitation id="52587E11FF8F39324D4DFD53FDD3FCDC" author="Seevers" box="[360,571,751,774]" page="128" pageId="17" pageNumber="18" year="1978">
|
||
<bibRefCitation id="B76825F1FF8F39324D4DFD53FDEAFCDC" author="Seevers" box="[360,514,751,774]" pageId="17" pageNumber="88" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
|
||
: 128
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(as valid genus in subtribe Geostibina
|
||
<bibRefCitation id="B76825F1FF8F39324FF9FD53FB9DFCDC" author="Seevers" box="[988,1141,751,774]" pageId="17" pageNumber="88" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D2DFCB3FC21FCFC" blockId="17.[264,1324,559,870]" box="[264,969,783,806]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFCB3FC21FCFC" box="[264,969,783,806]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFCB3FD4FFCFC" authority="Lohse & Smetana, 1988: 270" authorityName="Lohse & Smetana" authorityPageNumber="270" authorityYear="1988" box="[264,679,783,806]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFCB3FEBCFCFC" box="[264,340,783,806]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
:
|
||
<treatmentCitation id="52587E11FF8F39324D4DFCB3FD4FFCFC" author="Lohse" box="[360,679,783,806]" page="270" pageId="17" pageNumber="18" year="1988">
|
||
<bibRefCitation id="B76825F1FF8F39324D4DFCB3FD86FCFC" author="Lohse" box="[360,622,783,806]" pageId="17" pageNumber="88" refString="Lohse, G. A. & Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse & Smetana, 1988</bibRefCitation>
|
||
: 270
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(as synonym of
|
||
<taxonomicName id="14F92383FF8F39324F7DFCB3FC50FCFC" ID-CoL="8VRPP" box="[856,952,783,806]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324F7DFCB3FC50FCFC" box="[856,952,783,806]" italics="true" pageId="17" pageNumber="18">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D2DFC93FB3FFC9C" blockId="17.[264,1324,559,870]" box="[264,1239,815,838]" pageId="17" pageNumber="18">
|
||
<treatmentCitationGroup id="F3E97F2EFF8F39324D2DFC93FB3FFC9C" box="[264,1239,815,838]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="14F92383FF8F39324D2DFC93FE77FC9C" authority="Ashe" authorityName="Ashe" box="[264,415,815,838]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFC93FEBCFC9C" box="[264,340,815,838]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
: Ashe
|
||
</taxonomicName>
|
||
<emphasis id="E18D8412FF8F39324D83FC93FE53FC9C" box="[422,443,815,838]" italics="true" pageId="17" pageNumber="18">in</emphasis>
|
||
<treatmentCitation id="52587E11FF8F39324DE4FC93FC5BFC9C" author="Newton" box="[449,947,815,838]" page="371" pageId="17" pageNumber="18" year="2000">
|
||
<bibRefCitation id="B76825F1FF8F39324DE4FC93FC93FC9C" author="Newton" box="[449,891,815,838]" pageId="17" pageNumber="88" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. & Chandler, D. S. (2000). Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press, pp. 272 - 418." type="book chapter" year="2000">Newton, Thayer, Ashe & Chandler, 2000</bibRefCitation>
|
||
: 371
|
||
</treatmentCitation>
|
||
(as subgenus of
|
||
<taxonomicName id="14F92383FF8F39324842FC93FB2FFC9C" ID-CoL="8VRPP" box="[1127,1223,815,838]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324842FC93FB2FFC9C" box="[1127,1223,815,838]" italics="true" pageId="17" pageNumber="18">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFF8F39324D2DFCF3FC60FCBC" box="[264,904,847,870]" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="D3465800FF8F39324D2DFCF3FC60FCBC" blockId="17.[264,1324,559,870]" box="[264,904,847,870]" pageId="17" pageNumber="18">
|
||
(other references for Palaearctic
|
||
<taxonomicName id="14F92383FF8F39324E41FCF3FCE8FCBC" box="[612,768,847,870]" class="Insecta" family="Staphylinidae" genus="Ditroposipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324E41FCF3FCE8FCBC" box="[612,768,847,870]" italics="true" pageId="17" pageNumber="18">Ditroposipalia</emphasis>
|
||
</taxonomicName>
|
||
are omitted)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFF8F39324D2DFC2AFC14FBF3" pageId="17" pageNumber="18" type="diagnosis">
|
||
<paragraph id="D3465800FF8F39324D2DFC2AFB77FC03" blockId="17.[264,1324,918,1904]" pageId="17" pageNumber="18">
|
||
<emphasis id="E18D8412FF8F39324D2DFC2AFE60FC6A" bold="true" box="[264,392,918,944]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
|
||
<taxonomicName id="14F92383FF8F39324DB9FC2BFE07FC6A" box="[412,495,919,944]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324DB9FC2BFE07FC6A" box="[412,495,919,944]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
differs from other subgenera of
|
||
<taxonomicName id="14F92383FF8F39324F5CFC2BFC0BFC6A" box="[889,995,919,944]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324F5CFC2BFC0BFC6A" box="[889,995,919,944]" italics="true" pageId="17" pageNumber="18">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
in having two longitudinal carinae in the middle of male abdominal tergum
|
||
<quantity id="1401F5E5FF8F39324F67FC03FC87FC03" box="[834,879,959,985]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="17" pageNumber="18" unit="in" value="7.0">7 in</quantity>
|
||
front of posterior margin.
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D1DFC5AFC14FBF3" blockId="17.[264,1324,918,1904]" pageId="17" pageNumber="18">
|
||
<emphasis id="E18D8412FF8F39324D1DFC5AFE28FBDA" bold="true" box="[312,448,998,1024]" pageId="17" pageNumber="18">Synonyms.</emphasis>
|
||
<taxonomicName id="14F92383FF8F39324DF5FC5BFD94FBDA" box="[464,636,999,1024]" class="Insecta" family="Staphylinidae" genus="Ditroposipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324DF5FC5BFD94FBDA" box="[464,636,999,1024]" italics="true" pageId="17" pageNumber="18">Ditroposipalia</emphasis>
|
||
</taxonomicName>
|
||
is placed in synonymy with
|
||
<taxonomicName id="14F92383FF8F39324FF7FC5BFBCDFBDA" box="[978,1061,999,1024]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324FF7FC5BFBCDFBDA" box="[978,1061,999,1024]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
because the
|
||
<typeStatus id="0C42E6A2FF8F3932489AFC5BFB19FBDB" box="[1215,1265,999,1025]" pageId="17" pageNumber="18">type</typeStatus>
|
||
species of both have two longitudinal carinae on the male tergum 7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFF8F39314D1DFB8AFB97FB2C" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="D3465800FF8F39324D1DFB8AFE91FB2B" blockId="17.[264,1324,918,1904]" pageId="17" pageNumber="18">
|
||
<emphasis id="E18D8412FF8F39324D1DFB8AFE2BFB8A" bold="true" box="[312,451,1078,1104]" pageId="17" pageNumber="18">Discussion.</emphasis>
|
||
<bibRefCitation id="B76825F1FF8F39324DF4FB8BFCE3FB8B" author="Lohse" box="[465,779,1079,1105]" pageId="17" pageNumber="88" refString="Lohse, G. A. & Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse and Smetana (1988)</bibRefCitation>
|
||
did not assign any of their four species to subgenera. Pace assigned his three species to subgenera
|
||
<taxonomicName id="14F92383FF8F39324FB6FBE3FBD7FBA2" box="[915,1087,1119,1144]" class="Insecta" family="Staphylinidae" genus="Ditroposipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324FB6FBE3FBD7FBA2" box="[915,1087,1119,1144]" italics="true" pageId="17" pageNumber="18">Ditroposipalia</emphasis>
|
||
</taxonomicName>
|
||
(synonymized here with
|
||
<taxonomicName id="14F92383FF8F39324D61FB3BFE7FFB7A" box="[324,407,1159,1184]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D61FB3BFE7FFB7A" box="[324,407,1159,1184]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="14F92383FF8F39324DFEFB3BFC83FB7B" authority="Scheerpeltz, 1951" authorityName="Scheerpeltz" authorityYear="1951" box="[475,875,1159,1185]" class="Insecta" family="Staphylinidae" genus="Lioglutosipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324DFEFB3BFD66FB7A" box="[475,654,1159,1184]" italics="true" pageId="17" pageNumber="18">Lioglutosipalia</emphasis>
|
||
<bibRefCitation id="B76825F1FF8F39324EB3FB3BFC83FB7B" author="Scheerpeltz" box="[662,875,1159,1185]" pageId="17" pageNumber="88" refString="Scheerpeltz, O. (1951) Die neue Systematik der Gattung Sipalia Muls. Rey (Col. Staphylinidae). Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien, 92, 166 - 180." type="journal article" year="1951">Scheerpeltz, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
. The latter is now considered a synonym of
|
||
<taxonomicName id="14F92383FF8F39324D40FB13FC6BFB13" authority="Scheerpeltz, 1931 (Assing 1999)" authorityName="Scheerpeltz, 1931 (Assing" authorityYear="1999" box="[357,899,1199,1225]" class="Insecta" family="Staphylinidae" genus="Sipalotricha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D40FB13FE1EFB12" box="[357,502,1199,1224]" italics="true" pageId="17" pageNumber="18">Sipalotricha</emphasis>
|
||
Scheerpeltz, 1931 (
|
||
<bibRefCitation id="B76825F1FF8F39324EC5FB13FC92FB13" author="Assing" box="[736,890,1199,1225]" pageId="17" pageNumber="86" refString="Assing, V. (1999) A revision of the species of Geostiba Thomson 1858 from Greece and Cyprus (Coleoptera, Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 31 (2), 845 - 928." type="journal article" year="1999">Assing 1999</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which lacks modifications on male tergum 7.
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39324D1DFB43FD10F9D3" blockId="17.[264,1324,918,1904]" pageId="17" pageNumber="18">
|
||
The males of both western Nearctic species of
|
||
<taxonomicName id="14F92383FF8F39324F47FB43FC24FAC2" box="[866,972,1279,1304]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324F47FB43FC24FAC2" box="[866,972,1279,1304]" italics="true" pageId="17" pageNumber="18">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
have male abdominal tergum with two carinae and are placed in the subgenus
|
||
<taxonomicName id="14F92383FF8F39324F6AFA9BFC4AFA9A" box="[847,930,1319,1344]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324F6AFA9BFC4AFA9A" box="[847,930,1319,1344]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="14F92383FF8F39324FEAFA9BFB92FA9A" box="[975,1146,1319,1344]" class="Insecta" family="Staphylinidae" genus="Ditroposipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324FEAFA9BFB92FA9A" box="[975,1146,1319,1344]" italics="true" pageId="17" pageNumber="18">Ditroposipalia</emphasis>
|
||
</taxonomicName>
|
||
). Eight of the twelve Appalachian species have male secondary characters corresponding to
|
||
<taxonomicName id="14F92383FF8F39324880FAF3FB10FAB2" box="[1189,1272,1359,1384]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324880FAF3FB10FAB2" box="[1189,1272,1359,1384]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
and four species are consistent with
|
||
<taxonomicName id="14F92383FF8F39324EA3FACBFCFFFA4A" box="[646,791,1399,1424]" class="Insecta" family="Staphylinidae" genus="Sipalotricha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324EA3FACBFCFFFA4A" box="[646,791,1399,1424]" italics="true" pageId="17" pageNumber="18">Sipalotricha</emphasis>
|
||
</taxonomicName>
|
||
. However there are good reasons to believe that all sixteen native Nearctic species of
|
||
<taxonomicName id="14F92383FF8F39324ED3FA23FC88FA62" box="[758,864,1439,1464]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324ED3FA23FC88FA62" box="[758,864,1439,1464]" italics="true" pageId="17" pageNumber="18">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
(that is, excluding
|
||
<taxonomicName id="14F92383FF8F39324860FA23FB09FA62" box="[1093,1249,1439,1464]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="circellaris">
|
||
<emphasis id="E18D8412FF8F39324860FA23FB09FA62" box="[1093,1249,1439,1464]" italics="true" pageId="17" pageNumber="18">G. circellaris</emphasis>
|
||
</taxonomicName>
|
||
introduced to Newfoundland) form a monophyletic group in relation to Palaearctic species of
|
||
<taxonomicName id="14F92383FF8F39324D2DFA53FEB3F9D2" box="[264,347,1519,1544]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324D2DFA53FEB3F9D2" box="[264,347,1519,1544]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="14F92383FF8F39324DA1FA53FDD8F9D2" box="[388,560,1519,1544]" class="Insecta" family="Staphylinidae" genus="Ditroposipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324DA1FA53FDD8F9D2" box="[388,560,1519,1544]" italics="true" pageId="17" pageNumber="18">Ditroposipalia</emphasis>
|
||
</taxonomicName>
|
||
) or
|
||
<taxonomicName id="14F92383FF8F39324E45FA53FD19F9D2" box="[608,753,1519,1544]" class="Insecta" family="Staphylinidae" genus="Sipalotricha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324E45FA53FD19F9D2" box="[608,753,1519,1544]" italics="true" pageId="17" pageNumber="18">Sipalotricha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8F39314D1DF9ABFEA0FE74" blockId="17.[264,1324,918,1904]" lastBlockId="18.[264,1324,284,1270]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
There are some characters shared by all native Nearctic species of
|
||
<taxonomicName id="14F92383FF8F39324874F9ABFB53F9EA" box="[1105,1211,1559,1584]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8F39324874F9ABFB53F9EA" box="[1105,1211,1559,1584]" italics="true" pageId="17" pageNumber="18">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
that may be autapomorphies for this group of species. These include: slightly raised elytral suture behind scutellum (except
|
||
<taxonomicName id="14F92383FF8F39324E6BF9DBFD12F95A" box="[590,762,1639,1664]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="impressula">
|
||
<emphasis id="E18D8412FF8F39324E6BF9DBFD12F95A" box="[590,762,1639,1664]" italics="true" pageId="17" pageNumber="18">G. impressula</emphasis>
|
||
</taxonomicName>
|
||
); the proximal seta on the
|
||
<emphasis id="E18D8412FF8F39324841F9DBFB2CF95A" box="[1124,1220,1639,1664]" italics="true" pageId="17" pageNumber="18">external</emphasis>
|
||
side of paramere apex is the longest; internal sac of the aedeagus has two pairs of diverticula (
|
||
<figureCitation id="4BC24485FF8F39324D34F90BFE4AF90B" box="[273,418,1719,1745]" captionStart="FIGURES 38 50" captionStartId="22.[264,383,1403,1427]" captionTargetBox="[306,1285,460,1385]" captionTargetId="figure@22.[294,1294,449,1393]" captionTargetPageId="22" captionText="FIGURES 38 50. Genitalia of Geostiba appalachigena Gusarov, sp. n. (paratypes from Black Mountain (38, 40 45, 47 50) and Back Allegheny Mountain (39, 46), West Virginia). 38 – details of internal sac retracted into median lobe, ventral view; 39 – copulatory piece of internal sac, dorsal view; 40 – medial lamellae of internal sac, ventral view; 41 distal diverticula of everted internal sac, ventral view; 42 – everted internal sac, ventral view; 43 – spermatheca; 44 45 – details of internal sac retracted into median lobe, lateral view; 46 – copulatory piece of internal sac, lateral view; 47 – medial lamella of internal sac, lateral view; 48 – everted internal sac, lateral view; 49 – right distal diverticulum of everted internal sac, lateral view; 50 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (38, 42, 44 45, 48), 0.1 mm (39 41, 43, 46 47, 49 50)." httpUri="https://zenodo.org/record/155709/files/figure.png" pageId="17" pageNumber="18">Figs. 42, 48</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FF8F39324D95F90BFE36F90B" box="[432,478,1719,1745]" captionStart="FIGURES 197 203" captionStartId="49.[264,383,1650,1674]" captionTargetBox="[311,1273,594,1629]" captionTargetId="figure@49.[305,1283,588,1638]" captionTargetPageId="49" captionText="FIGURES 197 203. Genitalia of Geostiba balsamensis Pace (Cataloochee Balsam Mountain, North Carolina). 197 – details of internal sac retracted into median lobe, ventral view; 198 – everted internal sac, ventral view; 199 – completely everted internal sac, ventral view; 200 – copulatory piece of internal sac, dorsal view; 201 – medial lamellae of internal sac, ventral view; 202 – distal diverticula of everted internal sac, ventral view; 203 – spermatheca. Scale bar 0.2 mm (197 199), 0.1 mm (200 203). CP – copulatory piece, DD – distal diverticula of internal sac, LW – lateral wall of internal sac, ML – medial lamellae, PD – proximal diverticula of internal sac, S – suspensoria, UP – Ushaped plate." httpUri="https://zenodo.org/record/155730/files/figure.png" pageId="17" pageNumber="18">198</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FF8F39324DC9F90BFDF3F90B" box="[492,539,1719,1745]" captionStart="FIGURES 204 212" captionStartId="50.[264,383,1644,1668]" captionTargetBox="[311,1279,700,1618]" captionTargetId="figure@50.[300,1287,689,1625]" captionTargetPageId="50" captionText="FIGURES 204 212. Aedeagus of Geostiba balsamensis Pace (Cataloochee Balsam Mountain, North Carolina). 204 206 – details of internal sac retracted into median lobe, lateral view; 207 – copulatory piece of internal sac, lateral view; 208 – medial lamella of internal sac, lateral view; 209 – everted internal sac, lateral view; 210 – completely everted internal sac, lateral view; 211 – right distal diverticulum of everted internal sac, lateral view; 212 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (204 206, 209 210), 0.1 mm (207 208, 211 212)." httpUri="https://zenodo.org/record/155731/files/figure.png" pageId="17" pageNumber="18">210</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FF8F39324E0CF90BFDB1F90B" box="[553,601,1719,1745]" captionStart="FIGURES 316 320" captionStartId="71.[264,383,1481,1505]" captionTargetBox="[294,1301,904,1448]" captionTargetId="figure@71.[281,1305,889,1455]" captionTargetPageId="71" captionText="FIGURES 316 320. Genitalia of Geostiba impressula (Casey) (holotype (316) and specimens from Triangle Lake, Oregon (317 320 )). 316 317 – spermatheca; 318 – everted internal sac, ventral view; 319 – everted internal sac, lateral view; 320 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (316 317, 320), 0.4 mm (318 319)." httpUri="https://zenodo.org/record/155745/files/figure.png" pageId="17" pageNumber="18">318</figureCitation>
|
||
), medial lamellae are short, narrow and bent ventrally (
|
||
<figureCitation id="4BC24485FF8F393248D4F90BFEBDF923" captionStart="FIGURES 38 50" captionStartId="22.[264,383,1403,1427]" captionTargetBox="[306,1285,460,1385]" captionTargetId="figure@22.[294,1294,449,1393]" captionTargetPageId="22" captionText="FIGURES 38 50. Genitalia of Geostiba appalachigena Gusarov, sp. n. (paratypes from Black Mountain (38, 40 45, 47 50) and Back Allegheny Mountain (39, 46), West Virginia). 38 – details of internal sac retracted into median lobe, ventral view; 39 – copulatory piece of internal sac, dorsal view; 40 – medial lamellae of internal sac, ventral view; 41 distal diverticula of everted internal sac, ventral view; 42 – everted internal sac, ventral view; 43 – spermatheca; 44 45 – details of internal sac retracted into median lobe, lateral view; 46 – copulatory piece of internal sac, lateral view; 47 – medial lamella of internal sac, lateral view; 48 – everted internal sac, lateral view; 49 – right distal diverticulum of everted internal sac, lateral view; 50 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (38, 42, 44 45, 48), 0.1 mm (39 41, 43, 46 47, 49 50)." httpUri="https://zenodo.org/record/155709/files/figure.png" pageId="17" pageNumber="18">Figs. 40, 47</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FF8F39324D47F963FE79F923" box="[354,401,1759,1785]" captionStart="FIGURES 197 203" captionStartId="49.[264,383,1650,1674]" captionTargetBox="[311,1273,594,1629]" captionTargetId="figure@49.[305,1283,588,1638]" captionTargetPageId="49" captionText="FIGURES 197 203. Genitalia of Geostiba balsamensis Pace (Cataloochee Balsam Mountain, North Carolina). 197 – details of internal sac retracted into median lobe, ventral view; 198 – everted internal sac, ventral view; 199 – completely everted internal sac, ventral view; 200 – copulatory piece of internal sac, dorsal view; 201 – medial lamellae of internal sac, ventral view; 202 – distal diverticula of everted internal sac, ventral view; 203 – spermatheca. Scale bar 0.2 mm (197 199), 0.1 mm (200 203). CP – copulatory piece, DD – distal diverticula of internal sac, LW – lateral wall of internal sac, ML – medial lamellae, PD – proximal diverticula of internal sac, S – suspensoria, UP – Ushaped plate." httpUri="https://zenodo.org/record/155730/files/figure.png" pageId="17" pageNumber="18">201</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FF8F39324DBBF963FE27F923" box="[414,463,1759,1785]" captionStart="FIGURES 204 212" captionStartId="50.[264,383,1644,1668]" captionTargetBox="[311,1279,700,1618]" captionTargetId="figure@50.[300,1287,689,1625]" captionTargetPageId="50" captionText="FIGURES 204 212. Aedeagus of Geostiba balsamensis Pace (Cataloochee Balsam Mountain, North Carolina). 204 206 – details of internal sac retracted into median lobe, lateral view; 207 – copulatory piece of internal sac, lateral view; 208 – medial lamella of internal sac, lateral view; 209 – everted internal sac, lateral view; 210 – completely everted internal sac, lateral view; 211 – right distal diverticulum of everted internal sac, lateral view; 212 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (204 206, 209 210), 0.1 mm (207 208, 211 212)." httpUri="https://zenodo.org/record/155731/files/figure.png" pageId="17" pageNumber="18">208</figureCitation>
|
||
). This form of the internal sac is very different from the forms found in Palaearctic
|
||
<emphasis id="E18D8412FF8F39324DB5F8BBFE45F8FA" box="[400,429,1799,1824]" italics="true" pageId="17" pageNumber="18">G.</emphasis>
|
||
(
|
||
<emphasis id="E18D8412FF8F39324D9AF8BBFE1EF8FA" box="[447,502,1799,1824]" italics="true" pageId="17" pageNumber="18">s. str</emphasis>
|
||
.)
|
||
<taxonomicName id="14F92383FF8F39324E2AF8BBFD6EF8FA" box="[527,646,1799,1824]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="circellaris">
|
||
<emphasis id="E18D8412FF8F39324E2AF8BBFD6EF8FA" box="[527,646,1799,1824]" italics="true" pageId="17" pageNumber="18">circellaris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4BC24485FF8F39324EB2F8BBFCC9F8FB" box="[663,801,1799,1825]" captionStart="FIGURES 21 25" captionStartId="10.[264,383,1697,1721]" captionTargetBox="[310,1273,423,1660]" captionTargetId="figure@10.[302,1285,409,1671]" captionTargetPageId="10" captionText="FIGURES 21 25. Partially everted internal sac of aedeagus of Geostiba circellaris (Gravenhorst) (21 23, St. Petersburg Region, Russia) and G. infirma (Weise) (24 25, IvanoFrankovsk Region, Ukraine). 21, 24 – ventral view; 22, 25 – lateral view; 23 – dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155705/files/figure.png" pageId="17" pageNumber="18">Figs. 2122</figureCitation>
|
||
),
|
||
<taxonomicName id="14F92383FF8F39324F12F8BBFBB5F8FA" box="[823,1117,1799,1825]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="infirma" subGenus="Sipalotricha">
|
||
<emphasis id="E18D8412FF8F39324F12F8BBFCBBF8FA" box="[823,851,1799,1824]" italics="true" pageId="17" pageNumber="18">G.</emphasis>
|
||
(
|
||
<emphasis id="E18D8412FF8F39324F40F8BBFC1EF8FA" box="[869,1014,1799,1824]" italics="true" pageId="17" pageNumber="18">Sipalotricha</emphasis>
|
||
)
|
||
<emphasis id="E18D8412FF8F39324822F8BBFBB5F8FA" box="[1031,1117,1799,1824]" italics="true" pageId="17" pageNumber="18">infirma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4BC24485FF8F3932484BF8BBFB11F8FB" box="[1134,1273,1799,1825]" captionStart="FIGURES 21 25" captionStartId="10.[264,383,1697,1721]" captionTargetBox="[310,1273,423,1660]" captionTargetId="figure@10.[302,1285,409,1671]" captionTargetPageId="10" captionText="FIGURES 21 25. Partially everted internal sac of aedeagus of Geostiba circellaris (Gravenhorst) (21 23, St. Petersburg Region, Russia) and G. infirma (Weise) (24 25, IvanoFrankovsk Region, Ukraine). 21, 24 – ventral view; 22, 25 – lateral view; 23 – dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155705/files/figure.png" pageId="17" pageNumber="18">Figs. 2425</figureCitation>
|
||
),
|
||
<taxonomicName id="14F92383FF8F3932492BF8BBFE3BF892" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="padana" subGenus="Sibiota">
|
||
<emphasis id="E18D8412FF8F3932492BF8BBFAC3F8FA" box="[1294,1323,1799,1824]" italics="true" pageId="17" pageNumber="18">G.</emphasis>
|
||
(
|
||
<emphasis id="E18D8412FF8F39324D37F893FE8DF892" box="[274,357,1839,1864]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
)
|
||
<emphasis id="E18D8412FF8F39324D5EF893FE3BF892" box="[379,467,1839,1864]" italics="true" pageId="17" pageNumber="18">padana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4BC24485FF8F39324DCDF893FD9CF893" box="[488,628,1839,1865]" captionStart="FIGURES 26 28" captionStartId="12.[264,383,1733,1757]" captionTargetBox="[306,1283,630,1710]" captionTargetId="figure@12.[298,1290,623,1720]" captionTargetPageId="12" captionText="FIGURES 26 28. Partially everted internal sac of aedeagus of Geostiba padana (Weise) (26 27, Crissolo, Italy) and G. oertzeni (Eppelsheim) (28, Parnassos, Greece). 26 – lateral view; 27 28 – ventral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155706/files/figure.png" pageId="17" pageNumber="18">Figs. 2627</figureCitation>
|
||
) and
|
||
<taxonomicName id="14F92383FF8F39324E9AF893FC50F892" box="[703,952,1839,1865]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="oertzeni" subGenus="Sibiota">
|
||
<emphasis id="E18D8412FF8F39324E9AF893FD33F892" box="[703,731,1839,1864]" italics="true" pageId="17" pageNumber="18">G.</emphasis>
|
||
(
|
||
<emphasis id="E18D8412FF8F39324ED4F893FCACF892" box="[753,836,1839,1864]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
|
||
)
|
||
<emphasis id="E18D8412FF8F39324F7FF893FC50F892" box="[858,952,1839,1864]" italics="true" pageId="17" pageNumber="18">oertzeni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4BC24485FF8F39324FE8F893FBC2F893" box="[973,1066,1839,1865]" captionStart="FIGURES 26 28" captionStartId="12.[264,383,1733,1757]" captionTargetBox="[306,1283,630,1710]" captionTargetId="figure@12.[298,1290,623,1720]" captionTargetPageId="12" captionText="FIGURES 26 28. Partially everted internal sac of aedeagus of Geostiba padana (Weise) (26 27, Crissolo, Italy) and G. oertzeni (Eppelsheim) (28, Parnassos, Greece). 26 – lateral view; 27 28 – ventral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155706/files/figure.png" pageId="17" pageNumber="18">Fig. 28</figureCitation>
|
||
). The hypothesis of monophyly of the Nearctic group of species can be tested only if representatives from other lineages are included in the analysis, which is outside the scope of the present paper. However, if this hypothesis is accepted and both
|
||
<taxonomicName id="14F92383FF8C39314F7CFEF8FC44FE87" box="[857,940,324,349]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314F7CFEF8FC44FE87" box="[857,940,324,349]" italics="true" pageId="18" pageNumber="19">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="14F92383FF8C39314FCFFEF8FB93FE87" box="[1002,1147,324,349]" class="Insecta" family="Staphylinidae" genus="Sipalotricha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314FCFFEF8FB93FE87" box="[1002,1147,324,349]" italics="true" pageId="18" pageNumber="19">Sipalotricha</emphasis>
|
||
</taxonomicName>
|
||
are monophyletic groups, then all native Nearctic species must be placed in the same subgenus of
|
||
<taxonomicName id="14F92383FF8C393148D5FED0FEA9FE77" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C393148D5FED0FEA9FE77" italics="true" pageId="18" pageNumber="19">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8C39314D1DFE00FE73FBA4" blockId="18.[264,1324,284,1270]" pageId="18" pageNumber="19">
|
||
Among the twelve Appalachian species,
|
||
<taxonomicName id="14F92383FF8C39314F31FE00FC0DFE0F" box="[788,997,444,469]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="appalachigena">
|
||
<emphasis id="E18D8412FF8C39314F31FE00FC0DFE0F" box="[788,997,444,469]" italics="true" pageId="18" pageNumber="19">G. appalachigena</emphasis>
|
||
</taxonomicName>
|
||
is the least adapted to cryptic habitats such as soil and leaf litter and has the following plesiomorphies: well developed wings, long elytra (longer than pronotum), large eyes (temple length / eye length ratio 2.32.7) and large body (pronotal width
|
||
<quantity id="1401F5E5FF8C39314F3FFD88FC2CFD94" box="[794,964,564,590]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="18" pageNumber="19" unit="mm" value="0.435" valueMax="0.47" valueMin="0.4">0.400.47 mm</quantity>
|
||
).
|
||
<taxonomicName id="14F92383FF8C39314FFBFD88FB15FD97" box="[990,1277,564,589]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="appalachigena">
|
||
<emphasis id="E18D8412FF8C39314FFBFD88FB15FD97" box="[990,1277,564,589]" italics="true" pageId="18" pageNumber="19">Geostiba appalachigena</emphasis>
|
||
</taxonomicName>
|
||
has well developed carinae on the male tergum 7 and could be considered a typical representative of the subgenus
|
||
<taxonomicName id="14F92383FF8C39314E25FD38FDBBFD47" box="[512,595,644,669]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314E25FD38FDBBFD47" box="[512,595,644,669]" italics="true" pageId="18" pageNumber="19">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
. The four species of
|
||
<taxonomicName id="14F92383FF8C39314F7EFD38FC2DFD47" box="[859,965,644,669]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314F7EFD38FC2DFD47" box="[859,965,644,669]" italics="true" pageId="18" pageNumber="19">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
without carinae can be considered as more adapted to cryptic habitats. In comparison to
|
||
<taxonomicName id="14F92383FF8C39314FF9FD10FB47FD1F" box="[988,1199,684,709]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="appalachigena">
|
||
<emphasis id="E18D8412FF8C39314FF9FD10FB47FD1F" box="[988,1199,684,709]" italics="true" pageId="18" pageNumber="19">G. appalachigena</emphasis>
|
||
</taxonomicName>
|
||
they have the following apomorphies: reduced wings, short elytra (shorter than pronotum), small eyes (temple length / eye length ratio 3.86.0) and smaller body (pronotal width
|
||
<quantity id="1401F5E5FF8C3931489FFD40FED3FCE4" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.4" metricValueMin="3.4" pageId="18" pageNumber="19" unit="mm" value="0.39" valueMax="0.44" valueMin="0.34">0.340.44 mm</quantity>
|
||
). One can hypothesize that the four species without carinae on male tergum 7 originated from an ancestor or ancestors with carinae. Two arguments confirm that this scenario is possible.
|
||
<taxonomicName id="14F92383FF8C39314DC4FCC8FD0BFC57" box="[481,739,884,909]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="balsamensis">
|
||
<emphasis id="E18D8412FF8C39314DC4FCC8FD0BFC57" box="[481,739,884,909]" italics="true" pageId="18" pageNumber="19">Geostiba balsamensis</emphasis>
|
||
</taxonomicName>
|
||
, one of the smaller species (pronotal width 0.36 0.39), has weak enough carinae on tergum 7 that
|
||
<bibRefCitation id="B76825F1FF8C39314F6FFC20FC30FC6C" author="Pace" box="[842,984,924,950]" pageId="18" pageNumber="88" refString="Pace, R. (1997) Aleocharinae attere del North Carolina, Tennessee e Oregon (Coleoptera, Staphylinidae). Bollettino del Museo Regionale di Scienze Naturali, Torino, 15 (1), 101 - 110." type="journal article" year="1997">Pace (1997)</bibRefCitation>
|
||
overlooked them and placed this species in
|
||
<taxonomicName id="14F92383FF8C39314D9EFC78FD86FC07" box="[443,622,964,989]" class="Insecta" family="Staphylinidae" genus="Lioglutosipalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314D9EFC78FD86FC07" box="[443,622,964,989]" italics="true" pageId="18" pageNumber="19">Lioglutosipalia</emphasis>
|
||
</taxonomicName>
|
||
. The state of the carinae in
|
||
<taxonomicName id="14F92383FF8C39314FE8FC78FB6BFC07" box="[973,1155,964,989]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="balsamensis">
|
||
<emphasis id="E18D8412FF8C39314FE8FC78FB6BFC07" box="[973,1155,964,989]" italics="true" pageId="18" pageNumber="19">G. balsamensis</emphasis>
|
||
</taxonomicName>
|
||
can be considered intermediate between the well developed and the absent states. In many species of
|
||
<taxonomicName id="14F92383FF8C39314D2DFBA8FE9AFBF7" box="[264,370,1044,1069]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314D2DFBA8FE9AFBF7" box="[264,370,1044,1069]" italics="true" pageId="18" pageNumber="19">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
the small males are known to lack secondary sexual characters present in the large males (
|
||
<bibRefCitation id="B76825F1FF8C39314D83FB80FDADFB8C" author="Assing" box="[422,581,1084,1110]" pageId="18" pageNumber="86" refString="Assing, V. (2000) On some species of Geostiba Thomson, 1858 from the eastern Alps (Coleoptera: Staphylinidae, Aleocharinae). Koleopterologische Rundschau, 70, 79 - 85." type="journal article" year="2000">Assing 2000</bibRefCitation>
|
||
). The same trend may take place when species evolve to a smaller size.
|
||
</paragraph>
|
||
<paragraph id="D3465800FF8C39314D1DFB30FB97FB2C" blockId="18.[264,1324,284,1270]" pageId="18" pageNumber="19">
|
||
Taking into account the above arguments, I place all sixteen native Nearctic species of
|
||
<taxonomicName id="14F92383FF8C39314D2DFB08FE9AFB17" box="[264,370,1204,1229]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314D2DFB08FE9AFB17" box="[264,370,1204,1229]" italics="true" pageId="18" pageNumber="19">Geostiba</emphasis>
|
||
</taxonomicName>
|
||
in the subgenus
|
||
<taxonomicName id="14F92383FF8C39314E63FB08FD71FB17" box="[582,665,1204,1229]" class="Insecta" family="Staphylinidae" genus="Sibiota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E18D8412FF8C39314E63FB08FD71FB17" box="[582,665,1204,1229]" italics="true" pageId="18" pageNumber="19">Sibiota</emphasis>
|
||
</taxonomicName>
|
||
. This subgenus is also represented in the Palaearctic region from Europe to the Caucasus, Central Asia, Siberia and the Far East.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |