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<document id="1987D6D235F634114F798BAE6C2F6D77" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="1175­5326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FF8039014D2DF97FFDFEFD4C" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Geostiba (Sibiota) nimbicola Lohse &amp; Smetana 1988" docType="treatment" docVersion="14" lastPageNumber="35" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="31" updateTime="1698188143631" updateUser="plazi">
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<mods:title id="D8903778F5326307DCDA9B0D7255AFBB">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<paragraph id="D3465800FF80393D4D2DF97FFBB5F907" blockId="30.[264,1117,1731,1757]" box="[264,1117,1731,1757]" pageId="30" pageNumber="31">
<heading id="880EEF6CFF80393D4D2DF97FFBB5F907" bold="true" box="[264,1117,1731,1757]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<keyStep id="680D40A5FF80393D4D2DF97FFBB5F907" box="[264,1117,1731,1757]" pageId="30" pageNumber="31">
<key id="A7680F31FF80393D4D2DF97FFBB5F907" box="[264,1117,1731,1757]" pageId="30" pageNumber="31">
<keyLead id="6808FB35FF80393D4D2DF97FFBB5F907" box="[264,1117,1731,1757]" pageId="30" pageNumber="31">
<emphasis id="E18D8412FF80393D4D2DF97FFBB5F907" bold="true" box="[264,1117,1731,1757]" pageId="30" pageNumber="31">
4.
<taxonomicName id="14F92383FF80393D4D03F97FFC46F907" ID-CoL="6KD75" authority="Lohse &amp; Smetana, 1988" authorityName="Lohse &amp; Smetana" authorityYear="1988" box="[294,942,1731,1757]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nimbicola" subGenus="Sibiota">
<emphasis id="E18D8412FF80393D4D03F97FFE78F907" bold="true" box="[294,400,1731,1757]" italics="true" pageId="30" pageNumber="31">Geostiba</emphasis>
(
<emphasis id="E18D8412FF80393D4D85F97FFE1DF907" bold="true" box="[416,501,1731,1757]" italics="true" pageId="30" pageNumber="31">Sibiota</emphasis>
)
<emphasis id="E18D8412FF80393D4E23F97FFD97F907" bold="true" box="[518,639,1731,1757]" italics="true" pageId="30" pageNumber="31">nimbicola</emphasis>
<bibRefCitation id="B76825F1FF80393D4EA3F97FFC46F907" author="Lohse" box="[646,942,1731,1757]" pageId="30" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse &amp; Smetana, 1988</bibRefCitation>
</taxonomicName>
(
<figureCitation id="4BC24485FF80393D4F9BF97FFBBDF907" box="[958,1109,1731,1757]" captionStart-0="FIGURES 94 ­ 99" captionStart-1="FIGURES 100 ­ 107" captionStart-2="FIGURES 108 ­ 119" captionStartId-0="31.[264,382,1619,1643]" captionStartId-1="33.[264,383,1742,1766]" captionStartId-2="34.[264,383,1639,1663]" captionTargetBox-0="[315,1275,519,1594]" captionTargetBox-1="[319,1271,659,1705]" captionTargetBox-2="[344,1245,762,1606]" captionTargetId-0="figure@31.[301,1285,505,1602]" captionTargetId-1="figure@33.[308,1278,649,1715]" captionTargetId-2="figure@34.[335,1252,750,1614]" captionTargetPageId-0="31" captionTargetPageId-1="33" captionTargetPageId-2="34" captionText-0="FIGURES 94 ­ 99. Abdominal segments 7 ­ 8 of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94 ­ 97, 99), 0.1 mm (98)." captionText-1="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." captionText-2="FIGURES 108 ­ 119. Genitalia of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 108 details of internal sac retracted into median lobe, ventral view; 109 copulatory piece of internal sac, dorsal view; 110 medial lamellae of internal sac, ventral view; 111 distal diverticula of everted internal sac, ventral view; 112 everted internal sac, ventral view; 113 spermatheca; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 copulatory piece of internal sac, lateral view; 117 medial lamella of internal sac, lateral view; 118 everted internal sac, lateral view; 119 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (108, 112, 114 ­ 115, 118), 0.1 mm (109 ­ 111, 113, 116 ­ 117, 119)." httpUri-0="https://zenodo.org/record/155716/files/figure.png" httpUri-1="https://zenodo.org/record/155717/files/figure.png" httpUri-2="https://zenodo.org/record/155718/files/figure.png" pageId="30" pageNumber="31">Figs. 94­119</figureCitation>
)
</emphasis>
</keyLead>
</key>
</keyStep>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFF80393D4D2DF8B2FD4FF8BF" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="D3465800FF80393D4D2DF8B2FCCEF8FF" blockId="30.[264,1130,1806,1893]" box="[264,806,1806,1829]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="F3E97F2EFF80393D4D2DF8B2FCCEF8FF" box="[264,806,1806,1829]" pageId="30" pageNumber="31">
<taxonomicName id="14F92383FF80393D4D2DF8B2FCCAF8FF" ID-CoL="6KD75" authority="Lohse &amp; Smetana, 1988: 275" authorityName="Lohse &amp; Smetana" authorityPageNumber="275" authorityYear="1988" box="[264,802,1806,1829]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FF80393D4D2DF8B2FE31F8FF" box="[264,473,1806,1829]" italics="true" pageId="30" pageNumber="31">Geostiba nimbicola</emphasis>
<treatmentCitation id="52587E11FF80393D4DC5F8B2FCCAF8FF" author="Lohse" box="[480,802,1806,1829]" page="275" pageId="30" pageNumber="31" year="1988">
<bibRefCitation id="B76825F1FF80393D4DC5F8B2FD0EF8FF" author="Lohse" box="[480,742,1806,1829]" pageId="30" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse &amp; Smetana, 1988</bibRefCitation>
: 275
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D3465800FF80393D4D2DF892FD4FF8BF" blockId="30.[264,1130,1806,1893]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="F3E97F2EFF80393D4D2DF892FD4FF8BF" pageId="30" pageNumber="31">
<taxonomicName id="14F92383FF80393D4D2DF892FCDCF89F" ID-CoL="3FSKZ" authority="Lohse &amp; Smetana, 1988: 273" authorityName="Lohse &amp; Smetana" authorityPageNumber="273" authorityYear="1988" box="[264,820,1838,1861]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="E18D8412FF80393D4D2DF892FE36F89F" box="[264,478,1838,1861]" italics="true" pageId="30" pageNumber="31">Geostiba bicarinata</emphasis>
:
<treatmentCitation id="52587E11FF80393D4DD6F892FCDCF89F" author="Lohse" box="[499,820,1838,1861]" page="273" pageId="30" pageNumber="31" year="1988">
<bibRefCitation id="B76825F1FF80393D4DD6F892FD11F89F" author="Lohse" box="[499,761,1838,1861]" pageId="30" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse &amp; Smetana, 1988</bibRefCitation>
: 273
</treatmentCitation>
</taxonomicName>
,
<emphasis id="E18D8412FF80393D4F1AF892FC7DF89F" box="[831,917,1838,1861]" italics="true" pageId="30" pageNumber="31">ex parte</emphasis>
(misidentification).
<taxonomicName id="14F92383FF80393D4D2DF8F2FD4BF8BF" ID-CoL="6KD75" authority="Pace, 1997: 106" authorityName="Pace" authorityPageNumber="106" authorityYear="1997" box="[264,675,1870,1893]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FF80393D4D2DF8F2FE31F8BF" box="[264,473,1870,1893]" italics="true" pageId="30" pageNumber="31">Geostiba nimbicola</emphasis>
:
<treatmentCitation id="52587E11FF80393D4DCAF8F2FD4BF8BF" author="Pace" box="[495,675,1870,1893]" page="106" pageId="30" pageNumber="31" year="1997">
<bibRefCitation id="B76825F1FF80393D4DCAF8F2FD80F8BF" author="Pace" box="[495,616,1870,1893]" pageId="30" pageNumber="88" refString="Pace, R. (1997) Aleocharinae attere del North Carolina, Tennessee e Oregon (Coleoptera, Staphylinidae). Bollettino del Museo Regionale di Scienze Naturali, Torino, 15 (1), 101 - 110." type="journal article" year="1997">Pace, 1997</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFF8139034D2DFEA7FBBEFD1C" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="D3465800FF81393C4D2DFEA7FCADFE74" blockId="31.[264,1324,283,430]" pageId="31" pageNumber="32">
<emphasis id="E18D8412FF81393C4D2DFEA7FE56FEEF" bold="true" box="[264,446,283,309]" pageId="31" pageNumber="32">
<typeStatus id="0C42E6A2FF81393C4D2DFEA7FEADFEEF" box="[264,325,283,309]" pageId="31" pageNumber="32">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0C42E6A2FF81393C4DEAFEA0FDD7FEEC" box="[463,575,284,310]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
,,
<emphasis id="E18D8412FF81393C4E4AFEA7FB38FEEF" bold="true" box="[623,1232,283,309]" pageId="31" pageNumber="32">
<collectingCountry id="ABEE1890FF81393C4E4AFEA7FCB2FEEF" box="[623,858,283,309]" name="United States of America" pageId="31" pageNumber="32">UNITED STATES</collectingCountry>
: Tennessee / North Carolina:
</emphasis>
Sevier Co. / Swain Co.: Great Smoky Mountains National Park, Clingmans Dome [
<geoCoordinate id="B6CD3EC7FF81393C48BAFEF8FAC4FE84" box="[1183,1324,324,350]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="35.56278">35°33'46&quot;N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FF81393C4D2DFED0FE75FE5C" box="[264,413,364,390]" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="15" value="-83.498604">83°29'55&quot;W</geoCoordinate>
],
<quantity id="1401F5E5FF81393C4D94FED0FDB9FE5C" box="[433,593,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.985" metricValueMax="2.02" metricValueMin="1.95" pageId="31" pageNumber="32" unit="m" value="1985.0" valueMax="2020.0" valueMin="1950.0">1950­2020 m</quantity>
, (A.Smetana)
<date id="A7477EC0FF81393C4EDEFED0FC82FE5C" box="[763,874,364,390]" pageId="31" pageNumber="32" value="1986-06-02">2.vi.1986</date>
(
<collectionCode id="B5E8C0C5FF81393C4F5FFED0FC2FFE5C" box="[890,967,364,390]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="31" pageNumber="32">CNCI</collectionCode>
); allotype,, same data as the
<typeStatus id="0C42E6A2FF81393C4D2DFE28FE9AFE74" box="[264,370,404,430]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
;
<typeStatus id="0C42E6A2FF81393C4D5BFE28FE1BFE74" box="[382,499,404,430]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
: 3,, ditto (all
<collectionCode id="B5E8C0C5FF81393C4EC8FE28FCD1FE74" box="[749,825,404,430]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="31" pageNumber="32">CNCI</collectionCode>
).
</paragraph>
<caption id="87860888FF81393C4D2DF9EFFCC1F910" httpUri="https://zenodo.org/record/155716/files/figure.png" pageId="31" pageNumber="32" targetBox="[315,1275,519,1594]" targetPageId="31">
<paragraph id="D3465800FF81393C4D2DF9EFFCC1F910" blockId="31.[264,1323,1619,1738]" pageId="31" pageNumber="32">
<emphasis id="E18D8412FF81393C4D2DF9EFFE2BF9B1" bold="true" box="[264,451,1619,1643]" pageId="31" pageNumber="32">FIGURES 94­99</emphasis>
. Abdominal segments 7­8 of
<taxonomicName id="14F92383FF81393C4F20F9EFFB72F9B0" authority="Lohse &amp; Smetana" authorityName="Lohse &amp; Smetana" box="[773,1178,1619,1642]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FF81393C4F20F9EFFC3EF9B0" box="[773,982,1619,1642]" italics="true" pageId="31" pageNumber="32">Geostiba nimbicola</emphasis>
Lohse &amp; Smetana
</taxonomicName>
(Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94­97, 99), 0.1 mm (98).
</paragraph>
</caption>
<paragraph id="D3465800FF8139034D1DF8A9FB1EFE24" blockId="31.[264,1323,1813,1880]" lastBlockId="32.[264,1324,283,1830]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="E18D8412FF81393C4D1DF8A9FC56F8F5" bold="true" box="[312,958,1813,1839]" pageId="31" pageNumber="32">
Additional material.
<collectingCountry id="ABEE1890FF81393C4E60F8A9FCC7F8F5" box="[581,815,1813,1839]" name="United States of America" pageId="31" pageNumber="32">UNITED STATES</collectingCountry>
: Tennessee
</emphasis>
/
<emphasis id="E18D8412FF81393C4FF3F8A9FB48F8F5" bold="true" box="[982,1184,1813,1839]" pageId="31" pageNumber="32">North Carolina:</emphasis>
Sevier Co. / Swain Co.: (with missing abdomen), same data as the
<typeStatus id="0C42E6A2FF81393C4FE8F882FBDBF882" box="[973,1075,1854,1880]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
(
<collectionCode id="B5E8C0C5FF81393C4860F882FB79F882" box="[1093,1169,1854,1880]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="31" pageNumber="32">CNCI</collectionCode>
); 2, 3, ditto but (W.S.Suter)
<date id="A7477EC0FFBE39034E23FEA0FD92FEEC" box="[518,634,284,310]" pageId="32" pageNumber="33" value="1982-05-29">29.v.1982</date>
(
<collectionCode id="B5E8C0C5FFBE39034EAEFEA0FD0BFEEC" box="[651,739,284,310]" pageId="32" pageNumber="33">KSEM</collectionCode>
);
<emphasis id="E18D8412FFBE39034EDDFEA7FC96FEEF" bold="true" box="[760,894,283,309]" pageId="32" pageNumber="33">Tennessee:</emphasis>
Sevier Co.:, 4, Great Smoky Mts. National Park, Ramsey Cascade, [
<geoCoordinate id="B6CD3EC7FFBE39034EF3FEF8FC8AFE84" box="[726,866,324,350]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="35.708336">35°42'30&quot;N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFBE39034F4FFEF8FBE8FE84" box="[874,1024,324,350]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.30056">83°18'02&quot;W</geoCoordinate>
], moss on rock (Lackey),
<date id="A7477EC0FFBE39034D2DFED0FE66FE5C" box="[264,398,364,390]" pageId="32" pageNumber="33" value="1956-07-21">21.vii.1956</date>
(
<collectionCode id="B5E8C0C5FFBE39034DB9FED0FE1DFE5C" LSID="urn:lsid:biocol.org:col:34795" box="[412,501,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="32" pageNumber="33">FMNH</collectionCode>
,
<collectionCode id="B5E8C0C5FFBE39034E24FED0FDA4FE5C" box="[513,588,364,390]" pageId="32" pageNumber="33">SPSU</collectionCode>
);
<emphasis id="E18D8412FFBE39034E45FED7FCC2FE5F" bold="true" box="[608,810,363,389]" pageId="32" pageNumber="33">North Carolina:</emphasis>
Haywood Co.:
<specimenCount id="C5FF9389FFBE39034FD5FED0FB7AFE5C" box="[1008,1170,364,390]" pageId="32" pageNumber="33" type="generic">15 specimens</specimenCount>
, Mount Sterling Ridge,
<quantity id="1401F5E5FFBE39034DB8FE28FE06FE74" box="[413,494,404,430]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="32" pageNumber="33" unit="km" value="26.0">26 km</quantity>
NNW Waynesville,
<geoCoordinate id="B6CD3EC7FFBE39034ED6FE28FC82FE74" box="[755,874,404,430]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="92" value="35.698334">35°41.9'N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFBE39034F52FE28FC1DFE74" box="[887,1013,404,430]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="92" value="-83.125">83°07.5'W</geoCoordinate>
,
<quantity id="1401F5E5FFBE39034822FE28FB81FE74" box="[1031,1129,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="33" unit="m" value="1700.0">1700 m</quantity>
, in forest litter,
<taxonomicName id="14F92383FFBE39034D2DFE00FEA2FE0F" box="[264,330,444,469]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034D2DFE00FEA2FE0F" box="[264,330,444,469]" italics="true" pageId="32" pageNumber="33">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBE39034D40FE00FE4EFE0F" box="[357,422,444,469]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034D40FE00FE4EFE0F" box="[357,422,444,469]" italics="true" pageId="32" pageNumber="33">Abies</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBE39034DE4FE00FDCBFE0F" box="[449,547,444,469]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034DE4FE00FDCBFE0F" box="[449,547,444,469]" italics="true" pageId="32" pageNumber="33">Quercus</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBE39034E18FE00FCA8FE0C" authority="V.I.Gusarov" authorityName="V.I.Gusarov" box="[573,832,444,470]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034E18FE00FD62FE0F" box="[573,650,444,469]" italics="true" pageId="32" pageNumber="33">Betula</emphasis>
(V.I.Gusarov)
</taxonomicName>
,
<date id="A7477EC0FFBE39034F7DFE00FC31FE0C" box="[856,985,444,470]" pageId="32" pageNumber="33" value="2001-09-22">22.ix.2001</date>
;
<specimenCount id="C5FF9389FFBE39034FD4FE00FB76FE0C" box="[1009,1182,444,470]" pageId="32" pageNumber="33" type="generic">14 specimens</specimenCount>
, ditto but,
<geoCoordinate id="B6CD3EC7FFBE39034D2DFE58FE97FE24" box="[264,383,484,510]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="92" value="35.701668">35°42.1'N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFBE39034DA3FE58FDECFE24" box="[390,516,484,510]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="92" value="-83.113335">83°06.8'W</geoCoordinate>
,
<quantity id="1401F5E5FFBE39034E35FE58FD84FE24" box="[528,620,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="32" pageNumber="33" unit="m" value="1550.0">1550 m</quantity>
,
<taxonomicName id="14F92383FFBE39034E52FE58FD52FE27" box="[631,698,484,509]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034E52FE58FD52FE27" box="[631,698,484,509]" italics="true" pageId="32" pageNumber="33">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBE39034EEDFE58FC9DFE27" box="[712,885,484,509]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034EEDFE58FC9DFE27" box="[712,885,484,509]" italics="true" pageId="32" pageNumber="33">Rhododendron</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBE39034FA1FE58FC38FE27" box="[900,976,484,509]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBE39034FA1FE58FC38FE27" box="[900,976,484,509]" italics="true" pageId="32" pageNumber="33">Betula</emphasis>
</taxonomicName>
(all
<collectionCode id="B5E8C0C5FFBE39034839FE58FB85FE24" box="[1052,1133,484,510]" pageId="32" pageNumber="33">KSEM</collectionCode>
and
<collectionCode id="B5E8C0C5FFBE39034883FE58FB06FE24" box="[1190,1262,484,510]" pageId="32" pageNumber="33">SPSU</collectionCode>
)
</paragraph>
<paragraph id="D3465800FFBE39034D1DFDB0FBBEFD1C" blockId="32.[264,1324,283,1830]" pageId="32" pageNumber="33">
One of the
<typeStatus id="0C42E6A2FFBE39034DE6FDB0FDDBFDFC" box="[451,563,524,550]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="14F92383FFBE39034E7AFDB0FCE8FDFF" box="[607,768,524,549]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="E18D8412FFBE39034E7AFDB0FCE8FDFF" box="[607,768,524,549]" italics="true" pageId="32" pageNumber="33">G. bicarinata</emphasis>
</taxonomicName>
listed by
<bibRefCitation id="B76825F1FFBE39034F5FFDB0FB19FDFC" author="Lohse" box="[890,1265,524,550]" pageId="32" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse &amp; Smetana (1988: 275)</bibRefCitation>
(not found in
<collectionCode id="B5E8C0C5FFBE39034D53FD88FE56FD94" box="[374,446,564,590]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="32" pageNumber="33">CNCI</collectionCode>
) must also belong to
<taxonomicName id="14F92383FFBE39034EE7FD88FCB5FD97" box="[706,861,564,589]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBE39034EE7FD88FCB5FD97" box="[706,861,564,589]" italics="true" pageId="32" pageNumber="33">G. nimbicola</emphasis>
</taxonomicName>
(
<specimenCount id="C5FF9389FFBE39034F57FD88FC12FD94" box="[882,1018,564,590]" pageId="32" pageNumber="33" type="generic">1 specimen</specimenCount>
, Great Smoky Mountains National Park, Clingmans Dome,
<quantity id="1401F5E5FFBE39034EB3FDE0FCDEFDAC" box="[662,822,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.985" metricValueMax="2.02" metricValueMin="1.95" pageId="32" pageNumber="33" unit="m" value="1985.0" valueMax="2020.0" valueMin="1950.0">1950­2020 m</quantity>
, (A.Smetana)
<date id="A7477EC0FFBE39034FFBFDE0FBB9FDAC" box="[990,1105,604,630]" pageId="32" pageNumber="33" value="1986-06-02">2.vi.1986</date>
). This assertion is based on the results of my extensive collecting efforts which demonstrated that
<taxonomicName id="14F92383FFBE39034893FD38FED4FD1F" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="E18D8412FFBE39034893FD38FED4FD1F" italics="true" pageId="32" pageNumber="33">G. bicarinata</emphasis>
</taxonomicName>
is restricted to the Great Balsam Mountains Pisgah Ridge massif.
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFBE39034D1DFD6FFBBEFB14" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="D3465800FFBE39034D1DFD6FFDF6FC6C" blockId="32.[264,1324,283,1830]" pageId="32" pageNumber="33">
<emphasis id="E18D8412FFBE39034D1DFD6FFE50FD37" bold="true" box="[312,440,723,749]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
<taxonomicName id="14F92383FFBE39034DE9FD68FD5CFD37" box="[460,692,724,749]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBE39034DE9FD68FD5CFD37" box="[460,692,724,749]" italics="true" pageId="32" pageNumber="33">Geostiba nimbicola</emphasis>
</taxonomicName>
can be distinguished from other Nearctic species of
<taxonomicName id="14F92383FFBE39034D2DFD40FE9AFCCF" box="[264,370,764,789]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FFBE39034D2DFD40FE9AFCCF" box="[264,370,764,789]" italics="true" pageId="32" pageNumber="33">Geostiba</emphasis>
</taxonomicName>
by having small eyes (temple length to eye length ratio 3.0­4.0), pronotal pubescence of
<typeStatus id="0C42E6A2FFBE39034D56FC98FE4DFCE4" box="[371,421,804,830]" pageId="32" pageNumber="33">type</typeStatus>
V, reduced wings, short elytra (pronotum length to elytron length ratio 1.3), the presence of two short parallel carinae in the middle of abdominal tergum
<quantity id="1401F5E5FFBE390348B9FCF0FB22FCBC" box="[1180,1226,844,870]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="32" pageNumber="33" unit="in" value="7.0">7 in</quantity>
front of posterior margin, the shape of the aedeagus (
<figureCitation id="4BC24485FFBE39034F3CFCC8FBC5FC54" box="[793,1069,884,910]" captionStart-0="FIGURES 100 ­ 107" captionStart-1="FIGURES 108 ­ 119" captionStartId-0="33.[264,383,1742,1766]" captionStartId-1="34.[264,383,1639,1663]" captionTargetBox-0="[319,1271,659,1705]" captionTargetBox-1="[344,1245,762,1606]" captionTargetId-0="figure@33.[308,1278,649,1715]" captionTargetId-1="figure@34.[335,1252,750,1614]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." captionText-1="FIGURES 108 ­ 119. Genitalia of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 108 details of internal sac retracted into median lobe, ventral view; 109 copulatory piece of internal sac, dorsal view; 110 medial lamellae of internal sac, ventral view; 111 distal diverticula of everted internal sac, ventral view; 112 everted internal sac, ventral view; 113 spermatheca; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 copulatory piece of internal sac, lateral view; 117 medial lamella of internal sac, lateral view; 118 everted internal sac, lateral view; 119 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (108, 112, 114 ­ 115, 118), 0.1 mm (109 ­ 111, 113, 116 ­ 117, 119)." httpUri-0="https://zenodo.org/record/155717/files/figure.png" httpUri-1="https://zenodo.org/record/155718/files/figure.png" pageId="32" pageNumber="33">Figs. 100­112, 114­119</figureCitation>
) and the shape of the spermatheca (
<figureCitation id="4BC24485FFBE39034D8EFC20FDF8FC6C" box="[427,528,924,950]" captionStart="FIGURES 108 ­ 119" captionStartId="34.[264,383,1639,1663]" captionTargetBox="[344,1245,762,1606]" captionTargetId="figure@34.[335,1252,750,1614]" captionTargetPageId="34" captionText="FIGURES 108 ­ 119. Genitalia of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 108 details of internal sac retracted into median lobe, ventral view; 109 copulatory piece of internal sac, dorsal view; 110 medial lamellae of internal sac, ventral view; 111 distal diverticula of everted internal sac, ventral view; 112 everted internal sac, ventral view; 113 spermatheca; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 copulatory piece of internal sac, lateral view; 117 medial lamella of internal sac, lateral view; 118 everted internal sac, lateral view; 119 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (108, 112, 114 ­ 115, 118), 0.1 mm (109 ­ 111, 113, 116 ­ 117, 119)." httpUri="https://zenodo.org/record/155718/files/figure.png" pageId="32" pageNumber="33">Fig. 113</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBE39034D1DFC78FBBEFB14" blockId="32.[264,1324,283,1830]" pageId="32" pageNumber="33">
<taxonomicName id="14F92383FFBE39034D1DFC78FDF5FC07" box="[312,541,964,989]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBE39034D1DFC78FDF5FC07" box="[312,541,964,989]" italics="true" pageId="32" pageNumber="33">Geostiba nimbicola</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="14F92383FFBE39034F29FC78FC4AFC07" box="[780,930,964,989]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pluvigena">
<emphasis id="E18D8412FFBE39034F29FC78FC4AFC07" box="[780,930,964,989]" italics="true" pageId="32" pageNumber="33">G. pluvigena</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBE39034F95FC78FBBDFC07" box="[944,1109,964,989]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nebuligena">
<emphasis id="E18D8412FFBE39034F95FC78FBBDFC07" box="[944,1109,964,989]" italics="true" pageId="32" pageNumber="33">G. nebuligena</emphasis>
</taxonomicName>
, and
<taxonomicName id="14F92383FFBE390348B0FC78FEB9FBDF" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="crepusculigena">
<emphasis id="E18D8412FFBE390348B0FC78FEB9FBDF" italics="true" pageId="32" pageNumber="33">G. crepusculigena</emphasis>
</taxonomicName>
.
<taxonomicName id="14F92383FFBE39034D4FFC50FDBAFBDF" box="[362,594,1004,1029]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBE39034D4FFC50FDBAFBDF" box="[362,594,1004,1029]" italics="true" pageId="32" pageNumber="33">Geostiba nimbicola</emphasis>
</taxonomicName>
differs from
<taxonomicName id="14F92383FFBE39034ED7FC50FC62FBDF" box="[754,906,1004,1029]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pluvigena">
<emphasis id="E18D8412FFBE39034ED7FC50FC62FBDF" box="[754,906,1004,1029]" italics="true" pageId="32" pageNumber="33">G. pluvigena</emphasis>
</taxonomicName>
in having longer and more narrow apex of median lobe without apical denticle (in lateral view) (
<figureCitation id="4BC24485FFBE39034FD9FBA8FB4DFBF4" box="[1020,1189,1044,1070]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="32" pageNumber="33">Figs. 100­107</figureCitation>
,
<figureCitation id="4BC24485FFBE39034890FBA8FAF3FBF4" box="[1205,1307,1044,1070]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="32" pageNumber="33">126­133</figureCitation>
); from
<taxonomicName id="14F92383FFBE39034D6FFB80FE1AFB8F" box="[330,498,1084,1109]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nebuligena">
<emphasis id="E18D8412FFBE39034D6FFB80FE1AFB8F" box="[330,498,1084,1109]" italics="true" pageId="32" pageNumber="33">G. nebuligena</emphasis>
</taxonomicName>
in having deeper apical emargination in female sternum 8 (
<figureCitation id="4BC24485FFBE390348E5FB80FEC0FBA4" captionStart="FIGURES 94 ­ 99" captionStartId="31.[264,382,1619,1643]" captionTargetBox="[315,1275,519,1594]" captionTargetId="figure@31.[301,1285,505,1602]" captionTargetPageId="31" captionText="FIGURES 94 ­ 99. Abdominal segments 7 ­ 8 of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94 ­ 97, 99), 0.1 mm (98)." httpUri="https://zenodo.org/record/155716/files/figure.png" pageId="32" pageNumber="33">Figs. 97­ 98</figureCitation>
,
<figureCitation id="4BC24485FFBE39034D10FBD8FE73FBA4" box="[309,411,1124,1150]" captionStart="FIGURES 146 ­ 151" captionStartId="39.[264,382,1567,1591]" captionTargetBox="[332,1258,573,1533]" captionTargetId="figure@39.[323,1263,564,1541]" captionTargetPageId="39" captionText="FIGURES 146 ­ 151. Abdominal segments 7 ­ 8 of Geostiba nebuligena Gusarov, sp. n. (paratypes from the Appalachian National Scenic Trail W of Clingmans Dome, Tennessee / North Carolina). 146 male tergum 8; 147 male sternum 8; 148 female tergum 8; 149 female sternum 8, 150 apex of female sternum 8; 151 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (146 ­ 149, 151), 0.1 mm (150)." httpUri="https://zenodo.org/record/155722/files/figure.png" pageId="32" pageNumber="33">149­150</figureCitation>
), longer and more narrow apex of median lobe without apical denticle (in lateral view) (
<figureCitation id="4BC24485FFBE39034DB2FB30FDD6FB7C" box="[407,574,1164,1190]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="32" pageNumber="33">Figs. 100­107</figureCitation>
,
<figureCitation id="4BC24485FFBE39034E6FFB30FD58FB7C" box="[586,688,1164,1190]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 median lobe, ventral view; 153 ­ 154 apex of median lobe, ventral view; 155 median lobe, lateral view; 156 apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="32" pageNumber="33">152­156</figureCitation>
); from
<taxonomicName id="14F92383FFBE39034F23FB30FC34FB7F" box="[774,988,1164,1189]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="crepusculigena">
<emphasis id="E18D8412FFBE39034F23FB30FC34FB7F" box="[774,988,1164,1189]" italics="true" pageId="32" pageNumber="33">G. crepusculigena</emphasis>
</taxonomicName>
in lacking obtuse projection on the ventral side of the apex of median lobe (
<figureCitation id="4BC24485FFBE39034F15FB08FC3FFB14" box="[816,983,1204,1230]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="32" pageNumber="33">Figs. 102­103</figureCitation>
,
<figureCitation id="4BC24485FFBE39034FC6FB08FBA1FB14" box="[995,1097,1204,1230]" captionStart="FIGURES 174 ­ 177" captionStartId="45.[264,383,1766,1790]" captionTargetBox="[317,1274,1164,1735]" captionTargetId="figure@45.[305,1283,1154,1742]" captionTargetPageId="45" captionText="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." httpUri="https://zenodo.org/record/155726/files/figure.png" pageId="32" pageNumber="33">176­177</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFBE39014D1DFB67FD95FE0C" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
<paragraph id="D3465800FFBE39034D1DFB67FE1EFA9C" blockId="32.[264,1324,283,1830]" pageId="32" pageNumber="33">
<emphasis id="E18D8412FFBE39034D1DFB67FE38FB2F" bold="true" box="[312,464,1243,1269]" pageId="32" pageNumber="33">Description.</emphasis>
Length 1.6­2.0 mm. Light brown or brown, apex of abdomen often lighter, antennae brownish yellow or light brown, legs and mouthparts brownish yellow. Body parallel­sided.
</paragraph>
<paragraph id="D3465800FFBE39034D1DFAE8FD6DF9D4" blockId="32.[264,1324,283,1830]" pageId="32" pageNumber="33">
Head as wide as long, surface on disk with fine isodiametric microsculpture, puncturation very fine, distance between punctures equal to 3­4 times their diameter. Temple length to eye length ratio 3.0­4.0. Antennal article 2 longer than article 3, article 4 transverse (width to length ratio 1.6), articles 5­10 strongly transverse, last article as long as 9 and 10 combined (as in
<figureCitation id="4BC24485FFBE39034E04FA48FD9FF9D4" box="[545,631,1524,1550]" captionStart="FIGURES 8 ­ 17" captionStartId="6.[264,383,1709,1733]" captionTargetBox="[307,1277,321,1687]" captionTargetId="figure@6.[297,1289,310,1697]" captionTargetPageId="6" captionText="FIGURES 8 ­ 17. Mouthparts of Geostiba circellaris (Gravenhorst) (8, Moscow Region, Russia), G. infirma (Weise) (9, Ivano­Frankovsk Region, Ukraine), G. carteriensis Pace (10 ­ 15, Tennessee), G. appalachigena Gusarov, sp. n. (16, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (17, paratype from North Carolina). 8 ­ 10, prementum; 11 hypopharynx; 12 left maxilla, ventral view; 13 left galea, dorsal view; 14 left lacinia, dorsal view; 15 mentum; 16 ­ 17 right antenna. Scale bar 0.1 mm (8 ­ 15), 0.2 mm (16 ­ 17). L Length of antennal article." httpUri="https://zenodo.org/record/155703/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBE39034D1DF9A0FDC3F8FC" blockId="32.[264,1324,283,1830]" pageId="32" pageNumber="33">
Pronotum as wide as long, width
<quantity id="1401F5E5FFBE39034EEBF9A0FC91F9EC" box="[718,889,1564,1590]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="32" pageNumber="33" unit="mm" value="0.385" valueMax="0.4" valueMin="0.37">0.37­0.40 mm</quantity>
, wider than head (pronotal width to head width ratio 1.1); microsculpture and puncturation as on head. Pronotal pubescence of
<typeStatus id="0C42E6A2FFBE39034D2DF9D0FED2F95C" box="[264,314,1644,1670]" pageId="32" pageNumber="33">type</typeStatus>
V. Elytra measured from humeral angle shorter than pronotum (pronotal length to elytral length ratio 1.3), wider than long (1.5), with fine isodiametric microsculpture and fine asperate puncturation, distance between punctures equals 2­3 times their diameter. Elytral suture behind scutellum slightly raised in both sexes. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="D3465800FFBF39024D1DFEA0FBFBFE24" blockId="33.[264,1324,284,590]" pageId="33" pageNumber="34">
Abdominal terga with fine microsculpture of transverse meshes, with fine and sparse puncturation, puncturation becoming finer towards abdomen apex, on terga 3­5 distance between punctures equals 3­7 times their diameter. Tergum 7 without white edge. Male tergum 7 with two medial carinae in front of posterior margin. Male tergum 8 with two weak and short carinae in front of posterior margin, posterior margin convex (
<figureCitation id="4BC24485FFBF39024D34FE58FE80FE24" box="[273,360,484,510]" captionStart="FIGURES 94 ­ 99" captionStartId="31.[264,382,1619,1643]" captionTargetBox="[315,1275,519,1594]" captionTargetId="figure@31.[301,1285,505,1602]" captionTargetPageId="31" captionText="FIGURES 94 ­ 99. Abdominal segments 7 ­ 8 of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94 ­ 97, 99), 0.1 mm (98)." httpUri="https://zenodo.org/record/155716/files/figure.png" pageId="33" pageNumber="34">Fig. 94</figureCitation>
). Male sternum 8 with convex posterior margin (
<figureCitation id="4BC24485FFBF39024F88FE58FBEDFE24" box="[941,1029,484,510]" captionStart="FIGURES 94 ­ 99" captionStartId="31.[264,382,1619,1643]" captionTargetBox="[315,1275,519,1594]" captionTargetId="figure@31.[301,1285,505,1602]" captionTargetPageId="31" captionText="FIGURES 94 ­ 99. Abdominal segments 7 ­ 8 of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94 ­ 97, 99), 0.1 mm (98)." httpUri="https://zenodo.org/record/155716/files/figure.png" pageId="33" pageNumber="34">Fig. 95</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBF39024D1DFDB0FD9AFD94" blockId="33.[264,1324,284,590]" pageId="33" pageNumber="34">
Female tergum 8 with convex posterior margin (
<figureCitation id="4BC24485FFBF39024F59FDB0FC3EFDFC" box="[892,982,524,550]" captionStart="FIGURES 94 ­ 99" captionStartId="31.[264,382,1619,1643]" captionTargetBox="[315,1275,519,1594]" captionTargetId="figure@31.[301,1285,505,1602]" captionTargetPageId="31" captionText="FIGURES 94 ­ 99. Abdominal segments 7 ­ 8 of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94 ­ 97, 99), 0.1 mm (98)." httpUri="https://zenodo.org/record/155716/files/figure.png" pageId="33" pageNumber="34">Fig. 96</figureCitation>
), sternum 8 with emarginate posterior margin (
<figureCitation id="4BC24485FFBF39024DFFFD88FD8DFD94" box="[474,613,564,590]" captionStart="FIGURES 94 ­ 99" captionStartId="31.[264,382,1619,1643]" captionTargetBox="[315,1275,519,1594]" captionTargetId="figure@31.[301,1285,505,1602]" captionTargetPageId="31" captionText="FIGURES 94 ­ 99. Abdominal segments 7 ­ 8 of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 94 male tergum 8; 95 male sternum 8; 96 female tergum 8; 97 female sternum 8, 98 apex of female sternum 8; 99 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (94 ­ 97, 99), 0.1 mm (98)." httpUri="https://zenodo.org/record/155716/files/figure.png" pageId="33" pageNumber="34">Figs. 97­98</figureCitation>
).
</paragraph>
<caption id="87860888FFBF39024D2DF972FAE0F89F" httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="33" pageNumber="34" targetBox="[319,1271,659,1705]" targetPageId="33">
<paragraph id="D3465800FFBF39024D2DF972FAE0F89F" blockId="33.[264,1323,1742,1861]" pageId="33" pageNumber="34">
<emphasis id="E18D8412FFBF39024D2DF972FE0AF93C" bold="true" box="[264,482,1742,1766]" pageId="33" pageNumber="34">FIGURES 100­107</emphasis>
. Aedeagus of
<taxonomicName id="14F92383FFBF39024EB4F972FBD0F93F" authority="Lohse &amp; Smetana" authorityName="Lohse &amp; Smetana" box="[657,1080,1742,1765]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBF39024EB4F972FC8FF93F" box="[657,871,1742,1765]" italics="true" pageId="33" pageNumber="34">Geostiba nimbicola</emphasis>
Lohse &amp; Smetana
</taxonomicName>
(100­103 holotype; 104­107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107).
</paragraph>
</caption>
<paragraph id="D3465800FFBC39014D1DFEA0FCF4FE74" blockId="34.[264,1324,284,662]" pageId="34" pageNumber="35">
Aedeagus as in
<figureCitation id="4BC24485FFBC39014DD7FEA0FCEFFEEC" box="[498,775,284,310]" captionStart-0="FIGURES 100 ­ 107" captionStart-1="FIGURES 108 ­ 119" captionStartId-0="33.[264,383,1742,1766]" captionStartId-1="34.[264,383,1639,1663]" captionTargetBox-0="[319,1271,659,1705]" captionTargetBox-1="[344,1245,762,1606]" captionTargetId-0="figure@33.[308,1278,649,1715]" captionTargetId-1="figure@34.[335,1252,750,1614]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." captionText-1="FIGURES 108 ­ 119. Genitalia of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 108 details of internal sac retracted into median lobe, ventral view; 109 copulatory piece of internal sac, dorsal view; 110 medial lamellae of internal sac, ventral view; 111 distal diverticula of everted internal sac, ventral view; 112 everted internal sac, ventral view; 113 spermatheca; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 copulatory piece of internal sac, lateral view; 117 medial lamella of internal sac, lateral view; 118 everted internal sac, lateral view; 119 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (108, 112, 114 ­ 115, 118), 0.1 mm (109 ­ 111, 113, 116 ­ 117, 119)." httpUri-0="https://zenodo.org/record/155717/files/figure.png" httpUri-1="https://zenodo.org/record/155718/files/figure.png" pageId="34" pageNumber="35">Figs. 100­112, 114­119</figureCitation>
. Apex of median lobe in ventral view narrow, its outline convex apically (
<figureCitation id="4BC24485FFBC39014E77FEF8FC86FE84" box="[594,878,324,350]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="34" pageNumber="35">Figs. 100­101, 104­105</figureCitation>
), in lateral view strait, without apical denticle or obtuse projection ventrally (
<figureCitation id="4BC24485FFBC39014EF0FED0FC04FE5C" box="[725,1004,364,390]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="34" pageNumber="35">Figs. 102­103, 106­107</figureCitation>
), distal diverticula of internal sac in ventral view broad (
<figureCitation id="4BC24485FFBC39014E4FFE28FCE7FE74" box="[618,783,404,430]" captionStart="FIGURES 108 ­ 119" captionStartId="34.[264,383,1639,1663]" captionTargetBox="[344,1245,762,1606]" captionTargetId="figure@34.[335,1252,750,1614]" captionTargetPageId="34" captionText="FIGURES 108 ­ 119. Genitalia of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 108 details of internal sac retracted into median lobe, ventral view; 109 copulatory piece of internal sac, dorsal view; 110 medial lamellae of internal sac, ventral view; 111 distal diverticula of everted internal sac, ventral view; 112 everted internal sac, ventral view; 113 spermatheca; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 copulatory piece of internal sac, lateral view; 117 medial lamella of internal sac, lateral view; 118 everted internal sac, lateral view; 119 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (108, 112, 114 ­ 115, 118), 0.1 mm (109 ­ 111, 113, 116 ­ 117, 119)." httpUri="https://zenodo.org/record/155718/files/figure.png" pageId="34" pageNumber="35">Figs. 111­112</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBC39014D1DFE00FD95FE0C" blockId="34.[264,1324,284,662]" box="[312,637,444,470]" pageId="34" pageNumber="35">
Spermatheca as in
<figureCitation id="4BC24485FFBC39014E30FE00FD92FE0C" box="[533,634,444,470]" captionStart="FIGURES 108 ­ 119" captionStartId="34.[264,383,1639,1663]" captionTargetBox="[344,1245,762,1606]" captionTargetId="figure@34.[335,1252,750,1614]" captionTargetPageId="34" captionText="FIGURES 108 ­ 119. Genitalia of Geostiba nimbicola Lohse &amp; Smetana (Mount Sterling Ridge Trail, Tennessee). 108 details of internal sac retracted into median lobe, ventral view; 109 copulatory piece of internal sac, dorsal view; 110 medial lamellae of internal sac, ventral view; 111 distal diverticula of everted internal sac, ventral view; 112 everted internal sac, ventral view; 113 spermatheca; 114 ­ 115 details of internal sac retracted into median lobe, lateral view; 116 copulatory piece of internal sac, lateral view; 117 medial lamella of internal sac, lateral view; 118 everted internal sac, lateral view; 119 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (108, 112, 114 ­ 115, 118), 0.1 mm (109 ­ 111, 113, 116 ­ 117, 119)." httpUri="https://zenodo.org/record/155718/files/figure.png" pageId="34" pageNumber="35">Fig. 113</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFBC39014D1DFE5DFDFEFD4C" pageId="34" pageNumber="35" type="distribution">
<paragraph id="D3465800FFBC39014D1DFE5DFDA9FDF9" blockId="34.[264,1324,284,662]" pageId="34" pageNumber="35">
<emphasis id="E18D8412FFBC39014D1DFE5DFE30FE21" bold="true" box="[312,472,481,507]" pageId="34" pageNumber="35">Distribution.</emphasis>
Known from the Great Smoky Mountains massif (Tennessee and North Carolina) (
<figureCitation id="4BC24485FFBC39014DA2FDB5FE10FDF9" box="[391,504,521,547]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="34" pageNumber="35">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FFBC39014E21FDB5FDDDFDF9" box="[516,565,521,547]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="34" pageNumber="35">340</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBC39014D1DFD8DFDFEFD4C" blockId="34.[264,1324,284,662]" pageId="34" pageNumber="35">
<emphasis id="E18D8412FFBC39014D1DFD8DFDE2FD91" bold="true" box="[312,522,561,587]" pageId="34" pageNumber="35">Natural History.</emphasis>
<taxonomicName id="14F92383FFBC39014E04FD8EFCE5FD91" box="[545,781,562,587]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBC39014E04FD8EFCE5FD91" box="[545,781,562,587]" italics="true" pageId="34" pageNumber="35">Geostiba nimbicola</emphasis>
</taxonomicName>
was collected at altitudes above
<quantity id="1401F5E5FFBC3901488FFD8DFAE0FD91" box="[1194,1288,561,587]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">1500 m</quantity>
in moss or in leaf litter in mixed forest with red spruce (
<taxonomicName id="14F92383FFBC39014FBFFDEBFBDFFDAA" box="[922,1079,599,624]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="rubens">
<emphasis id="E18D8412FFBC39014FBFFDEBFBDFFDAA" box="[922,1079,599,624]" italics="true" pageId="34" pageNumber="35">Picea rubens</emphasis>
</taxonomicName>
), Frasers fir (
<taxonomicName id="14F92383FFBC390148CCFDEBFEBEFD4F" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="fraseri">
<emphasis id="E18D8412FFBC390148CCFDEBFEBEFD4F" italics="true" pageId="34" pageNumber="35">Abies fraseri</emphasis>
</taxonomicName>
), birch and oak.
</paragraph>
</subSubSection>
</treatment>
</document>