1181 lines
215 KiB
XML
1181 lines
215 KiB
XML
<document id="6398C7049846EAA7A65BA2F6920D1E4A" ID-CLB-Dataset="22296" ID-DOI="10.5852/ejt.2018.456" ID-GBIF-Dataset="0566a11e-d6a3-4283-89c7-c7feea96fdbf" ID-Zenodo-Dep="3817823" ID-ZooBank="DE1B1DEE-9871-4803-9F67-025F2B439560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588965964057" checkinUser="valdenar" docAuthor="Sørensen, Martin V., Rohal, Melissa & Thistle, David" docDate="2018" docId="F64287A2505DFF8D1464FED5FD4E0901" docLanguage="en" docName="ejt-456_sorensen_rohal_thistle.pdf.imf" docOrigin="European Journal of Taxonomy 456" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes dubiosus Sørensen & Rohal & Thistle 2018, sp. nov." docType="treatment" docUuid="312B7A46-B12C-49DE-B11B-D36EE2FCA488" docUuidSource="ZooBank" docVersion="8" lastPageNumber="20" masterDocId="0A7BFFDA5056FF9E1600FFDEFFDE0C64" masterDocTitle="Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific" masterLastPageNumber="75" masterPageNumber="1" pageNumber="12" updateTime="1698839057356" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="11C68CD75E773D18F43CF3108EAEA25B">
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<mods:title id="6D6A3460AE94C66673012932B95C8CC2">Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific</mods:title>
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<mods:name id="D16132D286A9916EA5E00746559C0A17" type="personal">
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<mods:roleTerm id="F4380C1FA6FBA9740ACE273930ADFD90">Author</mods:roleTerm>
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<mods:namePart id="8878DB968D86F71543F959D8A20012E2">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="FFC5A01DE482185DA7296AF65AF70E8F">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<mods:nameIdentifier id="09530C72A4756DCD5A8334DA6FF7B6EC" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="4EE752EF5B4624D8A3EBA24B2E4CAA79">Rohal, Melissa</mods:namePart>
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<mods:nameIdentifier id="1214CD30D9C7C53FD143F8B765C796D5" type="email">mrohal@islander.tamucc.edu</mods:nameIdentifier>
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<mods:namePart id="3BF653D31FD6D28B7C3CCBFDB94CAACB">Thistle, David</mods:namePart>
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<mods:nameIdentifier id="6754892AE42DA758052820759C37ED96" type="email">dthistle@fsu.edu</mods:nameIdentifier>
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<mods:title id="5AC09A9EE8C108DC9463DAE04A47CE08">European Journal of Taxonomy</mods:title>
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<mods:part id="53420E0B4EE057F3EBAA0B534DBD5F0B">
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<mods:date id="5E17AA86CD725A035EDC71542FE56482">2018</mods:date>
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<mods:number id="B0F9A0D001FBE2F55C90C1E7FBEBBC71">2018-08-17</mods:number>
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<mods:number id="05AA908C1B1B8E927BAB9ABC7979B949">456</mods:number>
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<treatment id="F64287A2505DFF8D1464FED5FD4E0901" ID-DOI="http://doi.org/10.5281/zenodo.3818846" ID-GBIF-Taxon="163993849" ID-Zenodo-Dep="3818846" LSID="urn:lsid:zoobank.org:act:312B7A46-B12C-49DE-B11B-D36EE2FCA488" httpUri="http://treatment.plazi.org/id/F64287A2505DFF8D1464FED5FD4E0901" lastPageId="19" lastPageNumber="20" pageId="11" pageNumber="12">
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<subSubSection id="36F1653F505DFF951464FED5FB120D2D" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="7E5436B4505DFF951464FED5FC0E0D42" blockId="11.[360,1228,267,366]" box="[612,976,267,294]" pageId="11" pageNumber="12">
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<heading id="251C81D8505DFF951464FED5FC0E0D42" box="[612,976,267,294]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
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<taxonomicName id="B9EB4D37505DFF951464FED5FCB10D42" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[612,879,267,294]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="12" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
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<emphasis id="4C9FEAA6505DFF951464FED5FCB10D42" bold="true" box="[612,879,267,294]" italics="true" pageId="11" pageNumber="12">Echinoderes dubiosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="57AC57DD505DFF951576FED2FC0E0D42" box="[886,976,268,294]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="7E5436B4505DFF951768FEF1FB120D2D" blockId="11.[360,1228,267,366]" box="[360,1228,303,329]" pageId="11" pageNumber="12">
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urn:lsid:zoobank.org:act:
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<uuid id="0A4D0C61505DFF951486FEF1FB120D2D" box="[646,1228,303,329]" pageId="11" pageNumber="12">312B7A46-B12C-49DE-B11B-D36EE2FCA488</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F505DFF9514A0FE8AFC4A0D0A" box="[672,916,340,366]" pageId="11" pageNumber="12" type="description">
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<paragraph id="7E5436B4505DFF9514A0FE8AFC4A0D0A" blockId="11.[360,1228,267,366]" box="[672,916,340,366]" pageId="11" pageNumber="12">
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<figureCitation id="E6D02A31505DFF9514A0FE8AFCD70D0A" box="[672,777,340,366]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="12.[189,232,1791,1817]" captionStartId-1="13.[189,232,1683,1709]" captionStartId-2="15.[189,232,1576,1602]" captionTargetBox-0="[261,1323,265,1769]" captionTargetBox-1="[236,1351,265,1655]" captionTargetId-0="figure@12.[260,1327,265,1769]" captionTargetId-1="figure@13.[236,1351,265,1656]" captionTargetId-2="figure@16.[195,1389,265,2019]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="16" captionText-0="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." captionText-1="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." captionText-2="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi-0="http://doi.org/10.5281/zenodo.3817831" figureDoi-1="http://doi.org/10.5281/zenodo.3817833" figureDoi-2="http://doi.org/10.5281/zenodo.3817835" httpUri-0="https://zenodo.org/record/3817831/files/figure.png" httpUri-1="https://zenodo.org/record/3817833/files/figure.png" httpUri-2="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">
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||
<figureCitation id="E6D02A31505DFF9514A0FE8AFD370D0A" box="[672,745,340,366]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4</figureCitation>
|
||
–6
|
||
</figureCitation>
|
||
,
|
||
<tableCitation id="3369030F505DFF951514FE8AFC4A0D0A" box="[788,916,340,366]" captionStart-0="Table 4" captionStart-1="Table 5" captionStart-2="Table 6" captionStartId-0="15.[189,258,268,294]" captionStartId-1="17.[189,258,268,294]" captionStartId-2="23.[189,258,268,294]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionTargetPageId-2="23" captionText-0="Table 4. Measurements from light microscopy of Echinoderes dubiosus sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-7 = maximum sternal width, measured on segment 7 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." captionText-1="Table 5. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes dubiosus sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco 1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; pf = papillary flap; po = pore; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters." captionText-2="Table 6. Measurements from light microscopy of Echinoderes hamiltonorum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." httpUri-0="http://table.plazi.org/id/2A94663C5059FF9116BDFED2FB6F0DD6" pageId="11" pageNumber="12" tableUuid-0="2A94663C5059FF9116BDFED2FB6F0DD6">Tables 4–6</tableCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F505DFF9516BDFE48FC590EDC" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="7E5436B4505DFF9516BDFE48FEE80DD4" blockId="11.[189,310,406,432]" box="[189,310,406,432]" pageId="11" pageNumber="12">
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<heading id="251C81D8505DFF9516BDFE48FEE80DD4" bold="true" box="[189,310,406,432]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
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<emphasis id="4C9FEAA6505DFF9516BDFE48FEE80DD4" bold="true" box="[189,310,406,432]" pageId="11" pageNumber="12">Diagnosis</emphasis>
|
||
</heading>
|
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</paragraph>
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<paragraph id="7E5436B4505DFF9516BDFE18FC590EDC" blockId="11.[189,1399,453,697]" pageId="11" pageNumber="12">
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||
<taxonomicName id="B9EB4D37505DFF9516BDFE18FE910D84" authorityName="Claparede" authorityYear="1863" box="[189,335,454,480]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="12" phylum="Cephalorhyncha" rank="genus">
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||
<emphasis id="4C9FEAA6505DFF9516BDFE18FE910D84" box="[189,335,454,480]" italics="true" pageId="11" pageNumber="12">Echinoderes</emphasis>
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||
</taxonomicName>
|
||
with middorsal spines on segments 4 to 8 and spines in lateroventral positions on segments 6 to 9, with spines on segment 9 always extending beyond the terminal segment. Tubes present in lateroventral positions on segment 5, sublateral positions on segment 8 and laterodorsal positions on segment 10; tubes on segment 10 do not show sexual dimorphism. Minute glandular cell outlets
|
||
<typeStatus id="A1508816505DFF951344FDEDFAA90E29" box="[1348,1399,563,589]" pageId="11" pageNumber="12">type</typeStatus>
|
||
2 present in midlateral positions on segment 2. Segment 9 with middorsal cuticular structures, forming an anterior pore, and a posterior papillary flap, flanked by two sensory spots. Males with three pairs of penile spines; females with lateral terminal accessory spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F505DFF9516BDFD3FFB770F2A" pageId="11" pageNumber="12" type="etymology">
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||
<paragraph id="7E5436B4505DFF9516BDFD3FFE9A0E9F" blockId="11.[189,324,737,763]" box="[189,324,737,763]" pageId="11" pageNumber="12">
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<heading id="251C81D8505DFF9516BDFD3FFE9A0E9F" bold="true" box="[189,324,737,763]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
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||
<emphasis id="4C9FEAA6505DFF9516BDFD3FFE9A0E9F" bold="true" box="[189,324,737,763]" pageId="11" pageNumber="12">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDFCCEFB770F2A" blockId="11.[189,1399,784,847]" pageId="11" pageNumber="12">
|
||
This species name
|
||
<taxonomicName id="B9EB4D37505DFF95179EFCCEFDD80F4E" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[414,518,784,810]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="12" phylum="Cephalorhyncha" rank="species" species="dubiosus">
|
||
<emphasis id="4C9FEAA6505DFF95179EFCCEFDD80F4E" box="[414,518,784,810]" italics="true" pageId="11" pageNumber="12">dubiosus</emphasis>
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||
</taxonomicName>
|
||
is derived from the Latin ‘dubium’, because of its very close resemblance with
|
||
<taxonomicName id="B9EB4D37505DFF9516F8FCEAFE230F2A" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[248,509,820,846]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="11" pageNumber="12" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA6505DFF9516F8FCEAFE230F2A" box="[248,509,820,846]" italics="true" pageId="11" pageNumber="12">Echinoderes bathyalis</emphasis>
|
||
</taxonomicName>
|
||
, and thus slightly questionable status as a distinct species.
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="36F1653F505DFF9516BDFCA9FD6A0902" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="7E5436B4505DFF9516BDFCA9FE740FF5" blockId="11.[189,426,887,913]" box="[189,426,887,913]" pageId="11" pageNumber="12">
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<heading id="251C81D8505DFF9516BDFCA9FE740FF5" bold="true" box="[189,426,887,913]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="4C9FEAA6505DFF9516BDFCA9FE740FF5" bold="true" box="[189,426,887,913]" pageId="11" pageNumber="12">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="CE833CE9505DFF9516EAFC79FAC50849" ID-GBIF-Occurrence="2611490312" collectingDate="2008-09-20" collectionCode="US" collectorName="St. & Fluoromount G & See" country="United States of America" latitude="39.999443" location="West Coast" longLatPrecision="19" longitude="-125.87417" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
|
||
<paragraph id="7E5436B4505DFF9516EAFC79FE820FA5" blockId="11.[189,1399,935,1070]" box="[234,348,935,961]" pageId="11" pageNumber="12">
|
||
<emphasis id="4C9FEAA6505DFF9516EAFC79FE820FA5" bold="true" box="[234,348,935,961]" pageId="11" pageNumber="12">
|
||
<typeStatus id="A1508816505DFF9516EAFC79FE820FA5" box="[234,348,935,961]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDFC15FAC50849" blockId="11.[189,1399,935,1070]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="06FC7624505DFF9516BDFC15FDBA0F81" box="[189,612,971,997]" name="United States of America" pageId="11" pageNumber="12">UNITED STATES OF AMERICA</collectingCountry>
|
||
: adult
|
||
<specimenCount id="68EDFD3D505DFF9514BAFC15FD0E0F81" box="[698,720,971,997]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
|
||
,
|
||
<collectionCode id="18FAAE71505DFF9514E3FC15FCD60F81" box="[739,776,971,997]" country="South Africa" name="University of Stellenbosch" pageId="11" pageNumber="12">US</collectionCode>
|
||
<location id="7B34606F505DFF951514FC15FC7E0F81" LSID="urn:lsid:plazi:treatment:F64287A2505DFF8D1464FED5FD4E0901:7B34606F505DFF951514FC15FC7E0F81" box="[788,928,971,997]" country="United States of America" latitude="39.999443" longLatPrecision="19" longitude="-125.87417" name="West Coast" pageId="11" pageNumber="12" stateProvince="California">West Coast</location>
|
||
, off northern
|
||
<collectingRegion id="BC2FF856505DFF95124DFC14FB190F80" box="[1101,1223,970,996]" country="United States of America" name="California" pageId="11" pageNumber="12">California</collectingRegion>
|
||
,
|
||
<geoCoordinate id="1BDF5073505DFF9512D8FC17FAAC0F81" box="[1240,1394,969,997]" degrees="39" direction="north" minutes="59" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="58" value="39.999443">39°59′58″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1BDF5073505DFF9516BDFC33FEB4086D" box="[189,362,1005,1033]" degrees="125" direction="west" minutes="52" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="27" value="-125.87417">125°52′27″ W</geoCoordinate>
|
||
,
|
||
<collectorName id="D31E5362505DFF95177AFC31FE44086D" box="[378,410,1007,1033]" pageId="11" pageNumber="12">St.</collectorName>
|
||
3, 3675 m deep, collected from mud,
|
||
<date id="0A551074505DFF951581FC31FBF6086D" box="[897,1064,1007,1033]" pageId="11" pageNumber="12" value="2008-09-20">
|
||
<collectingDate id="1A11E99C505DFF951581FC31FBF6086D" box="[897,1064,1007,1033]" pageId="11" pageNumber="12" value="2008-09-20">20 Sep. 2008</collectingDate>
|
||
</date>
|
||
, mounted on glass slide in
|
||
<collectorName id="D31E5362505DFF9516BDFBCDFEAD0849" box="[189,371,1043,1069]" pageId="11" pageNumber="12">Fluoromount G</collectorName>
|
||
(NHMD-213583).
|
||
<collectorName id="D31E5362505DFF951458FBCDFD5C0849" box="[600,642,1043,1069]" pageId="11" pageNumber="12">See</collectorName>
|
||
<figureCitation id="E6D02A31505DFF951489FBCDFD100849" box="[649,718,1043,1069]" captionStart="Fig" captionStartId="2.[189,232,2000,2026]" captionTargetBox="[241,1347,1019,1957]" captionTargetId="figure@2.[241,1347,1019,1958]" captionTargetPageId="2" captionText="Fig. 1. Map showing the eight sampling stations off the US West Coast. See Table 1 for further details." figureDoi="http://doi.org/10.5281/zenodo.3817825" httpUri="https://zenodo.org/record/3817825/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
|
||
for localities and
|
||
<tableCitation id="3369030F505DFF9515A1FBCDFC260849" box="[929,1016,1043,1069]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="11" pageNumber="12" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
|
||
for detailed station data.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="7E5436B4505DFF9516EAFB8BFEB9080B" blockId="11.[189,1399,1109,1244]" box="[234,359,1109,1135]" pageId="11" pageNumber="12">
|
||
<materialsCitation id="CE833CE9505DFF9516EAFB8BFEB9080B" ID-GBIF-Occurrence="2611490308" box="[234,359,1109,1135]" pageId="11" pageNumber="12" typeStatus="paratype">
|
||
<emphasis id="4C9FEAA6505DFF9516EAFB8BFEB9080B" bold="true" box="[234,359,1109,1135]" pageId="11" pageNumber="12">
|
||
<typeStatus id="A1508816505DFF9516EAFB8BFEB9080B" box="[234,359,1109,1135]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDFBA4FBCC08B8" blockId="11.[189,1399,1109,1244]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="06FC7624505DFF9516BDFBA4FD8508F7" box="[189,603,1145,1172]" name="United States of America" pageId="11" pageNumber="12">UNITED STATES OF AMERICA</collectingCountry>
|
||
:
|
||
<specimenCount id="68EDFD3D505DFF951468FBA7FD4B08F0" box="[616,661,1145,1172]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||
, 1 ♂, same collecting data as for
|
||
<typeStatus id="A1508816505DFF95122DFBA7FB4908F7" box="[1069,1175,1145,1171]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="68EDFD3D505DFF9512A4FBA7FB3708F0" box="[1188,1257,1145,1172]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
|
||
, St. 1;
|
||
<specimenCount id="68EDFD3D505DFF951341FBA7FAB108F0" box="[1345,1391,1145,1172]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||
, St. 2;
|
||
<specimenCount id="68EDFD3D505DFF951703FB40FEEE08DC" box="[259,304,1182,1208]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||
, St. 4;
|
||
<specimenCount id="68EDFD3D505DFF951785FB40FE1608DC" box="[389,456,1182,1208]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
|
||
, St. 5; 1 ♂, St. 6;
|
||
<specimenCount id="68EDFD3D505DFF95149FFB40FD3C08DC" box="[671,738,1182,1208]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
|
||
, 1 ♂, St. 8 (NHMD-213584–213595). All
|
||
<typeStatus id="A1508816505DFF9512DAFB40FA9408DC" box="[1242,1354,1182,1208]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
|
||
are mounted in Fluoromount G, nine on glass slides and three on H-S slides.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516EAFADAFD89097A" blockId="11.[189,1399,1284,1382]" box="[234,599,1284,1310]" pageId="11" pageNumber="12">
|
||
<emphasis id="4C9FEAA6505DFF9516EAFADAFD89097A" bold="true" box="[234,599,1284,1310]" pageId="11" pageNumber="12">Additional non-type material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDFAF7FD6A0902" blockId="11.[189,1399,1284,1382]" pageId="11" pageNumber="12">
|
||
<collectingCountry id="06FC7624505DFF9516BDFAF7FD870926" box="[189,601,1320,1347]" name="United States of America" pageId="11" pageNumber="12">UNITED STATES OF AMERICA</collectingCountry>
|
||
:
|
||
<specimenCount id="68EDFD3D505DFF951464FAF6FD4F0927" box="[612,657,1320,1347]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||
, 1 ♂, St. 6;
|
||
<specimenCount id="68EDFD3D505DFF951524FAF6FC8F0927" box="[804,849,1320,1347]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
|
||
, St. 7. All mounted for SEM and stored in the first author’s personal reference collection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F505DFF8F16BDFA51FCBB0AC2" lastPageId="17" lastPageNumber="18" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="7E5436B4505DFF9516BDFA51FE9009CD" blockId="11.[189,334,1423,1449]" box="[189,334,1423,1449]" pageId="11" pageNumber="12">
|
||
<heading id="251C81D8505DFF9516BDFA51FE9009CD" bold="true" box="[189,334,1423,1449]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="4C9FEAA6505DFF9516BDFA51FE9009CD" bold="true" box="[189,334,1423,1449]" pageId="11" pageNumber="12">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDFA60FB430A21" blockId="11.[189,1399,1470,1605]" pageId="11" pageNumber="12">
|
||
Adults with head, neck and eleven trunk segments (
|
||
<figureCitation id="E6D02A31505DFF951525FA60FCAF09BC" box="[805,881,1470,1496]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4</figureCitation>
|
||
A–B, 5A, 6A). The trunk appears relatively slender, with very long lateral terminal spines, often exceeding the trunk length (
|
||
<figureCitation id="E6D02A31505DFF95126DFA3CFB660999" box="[1133,1208,1506,1533]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4</figureCitation>
|
||
A–C, 5A). For a complete overview of measurements and dimensions, see
|
||
<tableCitation id="3369030F505DFF95156EF9D8FC140A45" box="[878,970,1542,1569]" captionStart="Table 4" captionStartId="15.[189,258,268,294]" captionTargetPageId="15" captionText="Table 4. Measurements from light microscopy of Echinoderes dubiosus sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-7 = maximum sternal width, measured on segment 7 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." httpUri="http://table.plazi.org/id/2A94663C5059FF9116BDFED2FB6F0DD6" pageId="11" pageNumber="12" tableUuid="2A94663C5059FF9116BDFED2FB6F0DD6">Table 4</tableCitation>
|
||
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, is summarized in
|
||
<tableCitation id="3369030F505DFF95123EF9F5FB490A21" box="[1086,1175,1578,1605]" captionStart="Table 5" captionStartId="17.[189,258,268,294]" captionTargetPageId="17" captionText="Table 5. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes dubiosus sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco 1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; pf = papillary flap; po = pore; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters." pageId="11" pageNumber="12">Table 5</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDF9B0FAB40A97" blockId="11.[189,1399,1645,1780]" pageId="11" pageNumber="12">
|
||
None of the specimens mounted for SEM allowed a detailed examination of the head morphology. The neck has 16 placids, measuring
|
||
<quantity id="B9139B51505DFF95142FF94FFDA50AC8" box="[559,635,1681,1708]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="13.0">13 µm</quantity>
|
||
in length. The midventral placid is broadest, measuring
|
||
<quantity id="B9139B51505DFF95130CF94FFA860AC8" box="[1292,1368,1681,1708]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="10.0">10 µm</quantity>
|
||
in width at its base, whereas all others are narrower, measuring
|
||
<quantity id="B9139B51505DFF951577F96BFC6D0AB4" box="[887,947,1717,1744]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="12" unit="mm" value="8.0">8 µm</quantity>
|
||
in width at their bases. Four dorsal and two ventral trichoscalid plates are present, with the ventral pair being broader than the two dorsal pairs.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505DFF9516BDF8C2FE0F0B88" blockId="11.[189,1399,1820,2028]" pageId="11" pageNumber="12">
|
||
Segment 1 consists of a complete cuticular ring (
|
||
<figureCitation id="E6D02A31505DFF9514F5F8C2FCE10B52" box="[757,831,1820,1846]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4</figureCitation>
|
||
A–B, 5B–C). Sensory spots are located medially on the segment in subdorsal and laterodorsal positions (
|
||
<figureCitation id="E6D02A31505DFF951545F89EFC790B3E" box="[837,935,1856,1882]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505DFF9515B2F89EFC080B3F" box="[946,982,1856,1883]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="11" pageNumber="12">5B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505DFF9515E3F89EFBD60B3F" box="[995,1032,1856,1883]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">6B</figureCitation>
|
||
), and two glandular cell outlets
|
||
<typeStatus id="A1508816505DFF9516BDF8BBFF2E0B1B" box="[189,240,1893,1919]" pageId="11" pageNumber="12">type</typeStatus>
|
||
<quantity id="B9139B51505DFF9516F7F8BAFEFD0B1A" box="[247,291,1892,1918]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="12" unit="in" value="1.0">1 in</quantity>
|
||
middorsal positions (
|
||
<figureCitation id="E6D02A31505DFF951420F8BAFDA10B1A" box="[544,639,1892,1918]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
|
||
), and one pair in sublateral positions (
|
||
<figureCitation id="E6D02A31505DFF95123AF8BAFB440B1A" box="[1082,1178,1892,1918]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="11" pageNumber="12">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505DFF9512A6F8BAFB120B1B" box="[1190,1228,1892,1919]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">6C</figureCitation>
|
||
). The segment is completely devoid of cuticular hairs, except for long hairs around the sensory spots (
|
||
<figureCitation id="E6D02A31505DFF9512B6F856FB220BC7" box="[1206,1276,1928,1955]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||
B–C). The posterior segment margin is nearly straight, terminating in a pectinate fringe with thin, but quite long fringe tips (
|
||
<figureCitation id="E6D02A31505DFF951745F80FFE550B88" box="[325,395,2001,2028]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||
B–C).
|
||
</paragraph>
|
||
<caption id="2A94663C505AFF9216BDF921FDDA0B88" ID-DOI="http://doi.org/10.5281/zenodo.3817831" ID-Zenodo-Dep="3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1791,1817]" targetBox="[261,1323,265,1769]" targetPageId="12">
|
||
<paragraph id="7E5436B4505AFF9216BDF921FDDA0B88" blockId="12.[189,1399,1791,2029]" pageId="12" pageNumber="13">
|
||
<emphasis id="4C9FEAA6505AFF9216BDF921FED30B7D" bold="true" box="[189,269,1791,1817]" pageId="12" pageNumber="13">Fig. 4.</emphasis>
|
||
Line art illustrations of
|
||
<taxonomicName id="B9EB4D37505AFF921436F921FCE70B7D" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[566,825,1791,1817]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="12" pageNumber="13" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505AFF921436F921FCE70B7D" box="[566,825,1791,1817]" italics="true" pageId="12" pageNumber="13">Echinoderes dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505AFF921543F8DEFC7E0B7E" box="[835,928,1792,1818]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="4C9FEAA6505AFF9215AAF921FC610B7D" bold="true" box="[938,959,1791,1817]" pageId="12" pageNumber="13">A</emphasis>
|
||
. ♀, dorsal view.
|
||
<emphasis id="4C9FEAA6505AFF921291F921FB7B0B7D" bold="true" box="[1169,1189,1791,1817]" pageId="12" pageNumber="13">B</emphasis>
|
||
. ♀, ventral view.
|
||
<emphasis id="4C9FEAA6505AFF9216BDF8FCFF0C0B58" bold="true" box="[189,210,1826,1852]" pageId="12" pageNumber="13">C</emphasis>
|
||
. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view.
|
||
<emphasis id="4C9FEAA6505AFF9212C0F8FCFB0B0B58" bold="true" box="[1216,1237,1826,1852]" pageId="12" pageNumber="13">D</emphasis>
|
||
. ♂, segments 10 to 11, dorsal view.
|
||
<emphasis id="4C9FEAA6505AFF9217C0F898FE0A0B04" bold="true" box="[448,468,1862,1888]" pageId="12" pageNumber="13">E</emphasis>
|
||
. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C505BFF9316BDF94DFCC00B8E" ID-DOI="http://doi.org/10.5281/zenodo.3817833" ID-Zenodo-Dep="3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1683,1709]" targetBox="[236,1351,265,1655]" targetPageId="13">
|
||
<paragraph id="7E5436B4505BFF9316BDF94DFCC00B8E" blockId="13.[189,1399,1683,2026]" pageId="13" pageNumber="14">
|
||
<emphasis id="4C9FEAA6505BFF9316BDF94DFED40AC9" bold="true" box="[189,266,1683,1709]" pageId="13" pageNumber="14">Fig. 5.</emphasis>
|
||
Light micrographs showing overviews and details of
|
||
<taxonomicName id="B9EB4D37505BFF93157DF94DFBA30AC9" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[893,1149,1683,1709]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="13" pageNumber="14" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505BFF93157DF94DFBA30AC9" box="[893,1149,1683,1709]" italics="true" pageId="13" pageNumber="14">Echinoderes dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505BFF931284F94DFB3F0AC9" box="[1156,1249,1683,1709]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592).
|
||
<emphasis id="4C9FEAA6505BFF9315EDF968FBDC0AB4" bold="true" box="[1005,1026,1718,1744]" pageId="13" pageNumber="14">A</emphasis>
|
||
. Ventral overview.
|
||
<emphasis id="4C9FEAA6505BFF9312E3F968FB290AB4" bold="true" box="[1251,1271,1718,1744]" pageId="13" pageNumber="14">B</emphasis>
|
||
. Segments 1 to 6, dorsal view.
|
||
<emphasis id="4C9FEAA6505BFF9317ABF907FE1E0A97" bold="true" box="[427,448,1753,1779]" pageId="13" pageNumber="14">C</emphasis>
|
||
. Segments 1 to 6, ventral view.
|
||
<emphasis id="4C9FEAA6505BFF931543F907FC860A97" bold="true" box="[835,856,1753,1779]" pageId="13" pageNumber="14">D</emphasis>
|
||
. Segments 7 to 10, dorsal view.
|
||
<emphasis id="4C9FEAA6505BFF9312E1F907FB2B0A97" bold="true" box="[1249,1269,1753,1779]" pageId="13" pageNumber="14">E</emphasis>
|
||
. Segments 8 to 11, ventral view.
|
||
<emphasis id="4C9FEAA6505BFF9317D0F923FE3C0B73" bold="true" box="[464,482,1789,1815]" pageId="13" pageNumber="14">F</emphasis>
|
||
. Segments 8 to 11, ventral view.
|
||
<emphasis id="4C9FEAA6505BFF931585F922FC420B72" bold="true" box="[901,924,1788,1814]" pageId="13" pageNumber="14">G</emphasis>
|
||
. Segments 10 to 11 in female, ventral view.
|
||
<emphasis id="4C9FEAA6505BFF931703F8FEFEC40B5E" bold="true" box="[259,282,1824,1850]" pageId="13" pageNumber="14">H</emphasis>
|
||
. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B45058FF9016BDFED5FAA90E4E" blockId="14.[189,1399,267,554]" pageId="14" pageNumber="15">
|
||
Segment 2 consists of a complete cuticular ring, with minute glandular cell outlets
|
||
<typeStatus id="A15088165058FF9012ACFED2FB010D42" box="[1196,1247,268,294]" pageId="14" pageNumber="15">type</typeStatus>
|
||
2 located in midlateral positions (
|
||
<figureCitation id="E6D02A315058FF9017BBFEEEFDFE0D2F" box="[443,544,304,331]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF901430FEEEFD8A0D2F" box="[560,596,304,331]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">5B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF901464FEEEFDAC0D2F" box="[612,626,304,331]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">6</figureCitation>
|
||
D–E). Pachycyclus of the anterior segment margin is of regular thickness and uninterrupted. Sensory spots are located in middorsal, laterodorsal (
|
||
<figureCitation id="E6D02A315058FF9012D0FE8BFAED0D14" box="[1232,1331,341,368]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6D</figureCitation>
|
||
) and ventromedial (
|
||
<figureCitation id="E6D02A315058FF901767FEA5FE1E0DF2" box="[359,448,379,406]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6C</figureCitation>
|
||
) positions. Secondary pectinate fringe not detected on this or any of the following segments. On this and the eight following segments, the cuticular hairs are bracteate. The fairly long hairs are arranged in only two to three rows across the tergal plate (
|
||
<figureCitation id="E6D02A315058FF9015CEFE1BFB950DBB" box="[974,1099,453,479]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6B, D</figureCitation>
|
||
) and on the lateral halves of the sternal plates. The posterior segment margin is nearly straight and consists of a pectinate fringe, with fringe tips being longer and thicker than those on preceding segment, and very flexible (
|
||
<figureCitation id="E6D02A315058FF9012EAFDD1FAF00E4E" box="[1258,1326,527,554]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
|
||
B–D).
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDFD8DFC340EB8" blockId="14.[189,1399,594,732]" pageId="14" pageNumber="15">
|
||
Segment 3, and remaining segments, consisting of one tergal and two sternal plates (
|
||
<figureCitation id="E6D02A315058FF901299FD8CFB3A0E09" box="[1177,1252,594,621]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
A–B, 5A, C, E–G). Pachycyclus of the anterior segment margin is well-developed and interrupted at the tergosternal and midsternal junctions, but also middorsally. No conspicuous cuticular structures present. Cuticular hairs and posterior segment margin straight as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDFCDBFB230F0E" blockId="14.[189,1399,773,874]" pageId="14" pageNumber="15">
|
||
Segment 4 with acicular spine in middorsal position (
|
||
<figureCitation id="E6D02A315058FF901542FCDBFC790F7B" box="[834,935,773,799]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF9015B5FCDBFC070F44" box="[949,985,773,800]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">5B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF9015E9FCDBFBCE0F44" box="[1001,1040,773,800]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">6G</figureCitation>
|
||
). Sensory spots not observed. Anterior part of segment with glandular cell outlets
|
||
<typeStatus id="A15088165058FF901533FCF5FCB80F21" box="[819,870,811,837]" pageId="14" pageNumber="15">type</typeStatus>
|
||
<quantity id="B9139B515058FF90156FFCF4FC410F20" box="[879,927,810,836]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="15" unit="in" value="1.0">1 in</quantity>
|
||
paradorsal and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDFC4DFD180F9C" blockId="14.[189,1399,915,1016]" pageId="14" pageNumber="15">
|
||
Segment 5 with acicular spine in middorsal position and minute tubes in lateroventral positions (
|
||
<figureCitation id="E6D02A315058FF9016C5FC66FECE0FB6" box="[197,272,952,978]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
A–B, 5B–C, 6G). Glandular cell outlets
|
||
<typeStatus id="A15088165058FF9014EDFC66FCFE0FB6" box="[749,800,952,978]" pageId="14" pageNumber="15">type</typeStatus>
|
||
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDFBFEFBB008E1" blockId="14.[189,1399,1056,1157]" pageId="14" pageNumber="15">
|
||
Segment 6 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A315058FF901202FBFEFB95085E" box="[1026,1099,1056,1082]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
A–B, 5B, 6F–G). Sensory spots present in paradorsal (
|
||
<figureCitation id="E6D02A315058FF901407FB9BFD800804" box="[519,606,1093,1120]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6F</figureCitation>
|
||
) and sublateral positions. Glandular cell outlets
|
||
<typeStatus id="A15088165058FF901298FB98FB150804" box="[1176,1227,1094,1120]" pageId="14" pageNumber="15">type</typeStatus>
|
||
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDFB70FB220977" blockId="14.[189,1399,1198,1299]" pageId="14" pageNumber="15">
|
||
Segment 7 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A315058FF90124EFB70FB4008AC" box="[1102,1182,1198,1224]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
A–B, 5D, 6F–G). Sensory spots present in paradorsal (
|
||
<figureCitation id="E6D02A315058FF90146DFB0DFD1A088A" box="[621,708,1235,1262]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6F</figureCitation>
|
||
) and ventromedial positions. Glandular cell outlets
|
||
<typeStatus id="A15088165058FF901326FB0AFA87088A" box="[1318,1369,1236,1262]" pageId="14" pageNumber="15">type</typeStatus>
|
||
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDFAE2FC4E0A1B" blockId="14.[189,1399,1339,1663]" pageId="14" pageNumber="15">
|
||
Segment 8 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A315058FF901211FAE5FB820932" box="[1041,1116,1339,1366]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
A–B, 5D–E). Middorsal spine long, sometimes reaching or even exceeding the posterior margin of segment 11 (
|
||
<figureCitation id="E6D02A315058FF9012B6FABFFACB091F" box="[1206,1301,1377,1403]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Fig. 4A</figureCitation>
|
||
). Rather thick and rigid tubes, without differentiated bases or lateral wings, present in sublateral positions (
|
||
<figureCitation id="E6D02A315058FF9016C5FA75FED109A2" box="[197,271,1451,1478]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">Figs 5</figureCitation>
|
||
E–F, 6F). The exact position of the tubes varied between the examined specimens; they always stay within the sublateral area, but in some specimens, including the
|
||
<typeStatus id="A15088165058FF9015ECFA0EFB8D098E" box="[1004,1107,1488,1514]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
and some
|
||
<typeStatus id="A15088165058FF9012D3FA0EFA98098E" box="[1235,1350,1488,1514]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
|
||
, the tubes are very close to the midlateral line, whereas in others the tubes are almost in lateral accessory positions (compare
|
||
<figureCitation id="E6D02A315058FF9017A5F9C4FD860A51" box="[421,600,1562,1589]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">Fig. 5E and 5F</figureCitation>
|
||
). The variation is not related to gender or population. Sensory spots are present in paradorsal positions only. Glandular cell outlets type 1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45058FF9016BDF976FC230B8F" blockId="14.[189,1399,1704,2027]" pageId="14" pageNumber="15">
|
||
Segment 9 with acicular spines in lateroventral positions; the spines are thicker than other spines in the lateroventral series, and also long, always extending beyond the terminal segment (
|
||
<figureCitation id="E6D02A315058FF9012BEF913FAC00A83" box="[1214,1310,1741,1767]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF90132BF913FA910A8C" box="[1323,1359,1741,1768]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">5E</figureCitation>
|
||
). A couple of uncommon cuticular structures are present in middorsal position (
|
||
<figureCitation id="E6D02A315058FF90122BF92CFB520B68" box="[1067,1164,1778,1804]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF901297F92CFB610B69" box="[1175,1215,1778,1805]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">6K</figureCitation>
|
||
). Most anterior, it has an unpaired pore, flanked by cuticular ridges forming an inverted wedge. Posterior to this pore, a papillary flap or tuft of papillae, flanked by a pair of paradorsal sensory spots, is present. The structures are very conspicuous in SEM (
|
||
<figureCitation id="E6D02A315058FF90142FF8BFFD500B18" box="[559,654,1889,1916]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="14" pageNumber="15">Fig. 6K</figureCitation>
|
||
), but hardly visible in LM (
|
||
<figureCitation id="E6D02A315058FF9015DCF8BFFBE30B18" box="[988,1085,1889,1916]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">Fig. 5D</figureCitation>
|
||
). Additional sensory spots present in subdorsal, laterodorsal and ventrolateral positions (
|
||
<figureCitation id="E6D02A315058FF9015A8F858FC280BC5" box="[936,1014,1926,1953]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
|
||
A–B, 5D). Small rounded sieve plates are present in sublateral positions (
|
||
<figureCitation id="E6D02A315058FF901494F875FD2F0BA2" box="[660,753,1963,1990]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="14" pageNumber="15">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315058FF9014FCF875FCC10BA2" box="[764,799,1963,1990]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="14" pageNumber="15">5F</figureCitation>
|
||
). Glandular cell outlets
|
||
<typeStatus id="A15088165058FF90122EF872FBBF0BA2" box="[1070,1121,1964,1990]" pageId="14" pageNumber="15">type</typeStatus>
|
||
1, pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<caption id="2A94663C5059FF9116BDFED2FB6F0DD6" ID-Table-UUID="2A94663C5059FF9116BDFED2FB6F0DD6" httpUri="http://table.plazi.org/id/2A94663C5059FF9116BDFED2FB6F0DD6" pageId="15" pageNumber="16" targetBox="[355,1204,464,1477]" targetIsTable="true" targetPageId="15">
|
||
<paragraph id="7E5436B45059FF9116BDFED2FB6F0DD6" blockId="15.[189,1399,267,435]" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF9116BDFED2FEFC0D41" bold="true" box="[189,290,267,294]" pageId="15" pageNumber="16">Table 4.</emphasis>
|
||
Measurements from light microscopy of
|
||
<taxonomicName id="B9EB4D375059FF911525FED5FBF60D41" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[805,1064,267,293]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="15" pageNumber="16" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65059FF911525FED5FBF60D41" box="[805,1064,267,293]" italics="true" pageId="15" pageNumber="16">Echinoderes dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5059FF911233FED2FB4E0D42" box="[1075,1168,268,294]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-7 = maximum sternal width, measured on segment 7 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B45059FF911766FE0EFB7F09A1" pageId="15" pageNumber="16">
|
||
<table id="0CEBC414505900611763FE0EFB6A09A1" box="[355,1204,464,1477]" colsContinueIn="23.[357,1199,470,1400]" gridcols="5" gridrows="29" pageId="15" pageNumber="16">
|
||
<tr id="C0DB34F6505900611763FE0EFB6A0D8F" box="[355,1204,464,491]" gridrow="0" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FE0EFE3B0D8F" box="[355,485,464,491]" gridcol="0" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF911766FE0EFE3F0D8F" bold="true" box="[358,481,464,491]" pageId="15" pageNumber="16">Character</emphasis>
|
||
</th>
|
||
<th id="830A5D8A50590061143AFE0EFD940D8F" box="[570,586,464,491]" gridcol="1" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF91143AFE0FFD940D8F" bold="true" box="[570,586,465,491]" pageId="15" pageNumber="16">n</emphasis>
|
||
</th>
|
||
<th id="830A5D8A505900611497FE0EFCF50D8F" box="[663,811,464,491]" gridcol="2" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF9114BBFE0FFCD90D8F" bold="true" box="[699,775,465,491]" pageId="15" pageNumber="16">Range</emphasis>
|
||
</th>
|
||
<th id="830A5D8A505900611589FE0EFC3E0D8F" box="[905,992,464,491]" gridcol="3" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF911592FE0FFC080D8F" bold="true" box="[914,982,465,491]" pageId="15" pageNumber="16">Mean</emphasis>
|
||
</th>
|
||
<th id="830A5D8A50590061125DFE0EFB6A0D8F" box="[1117,1204,464,491]" gridcol="4" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF911276FE0EFB440D8F" bold="true" box="[1142,1178,464,491]" pageId="15" pageNumber="16">SD</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FDDCFB6A0E79" box="[355,1204,514,541]" gridrow="1" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FDDCFE3B0E79" box="[355,485,514,541]" gridcol="0" gridrow="1" pageId="15" pageNumber="16">TL</th>
|
||
<td id="830A5D8A50590061143AFDDCFD940E79" box="[570,586,514,541]" gridcol="1" gridrow="1" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FDDCFCF50E79" box="[663,811,514,541]" gridcol="2" gridrow="1" pageId="15" pageNumber="16">238–281</td>
|
||
<td id="830A5D8A505900611589FDDCFC3E0E79" box="[905,992,514,541]" gridcol="3" gridrow="1" pageId="15" pageNumber="16">259</td>
|
||
<td id="830A5D8A50590061125DFDDCFB6A0E79" box="[1117,1204,514,541]" gridcol="4" gridrow="1" pageId="15" pageNumber="16">13.22</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FDFAFB6A0E5B" box="[355,1204,548,575]" gridrow="2" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FDFAFE3B0E5B" box="[355,485,548,575]" gridcol="0" gridrow="2" pageId="15" pageNumber="16">MSW-7</th>
|
||
<td id="830A5D8A50590061143AFDFAFD940E5B" box="[570,586,548,575]" gridcol="1" gridrow="2" pageId="15" pageNumber="16">5</td>
|
||
<td id="830A5D8A505900611497FDFAFCF50E5B" box="[663,811,548,575]" gridcol="2" gridrow="2" pageId="15" pageNumber="16">60–70</td>
|
||
<td id="830A5D8A505900611589FDFAFC3E0E5B" box="[905,992,548,575]" gridcol="3" gridrow="2" pageId="15" pageNumber="16">64</td>
|
||
<td id="830A5D8A50590061125DFDFAFB6A0E5B" box="[1117,1204,548,575]" gridcol="4" gridrow="2" pageId="15" pageNumber="16">3.91</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FD9BFB6A0E04" box="[355,1204,581,608]" gridrow="3" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FD9BFE3B0E04" box="[355,485,581,608]" gridcol="0" gridrow="3" pageId="15" pageNumber="16">MSW-7/TL</th>
|
||
<td id="830A5D8A50590061143AFD9BFD940E04" box="[570,586,581,608]" gridcol="1" gridrow="3" pageId="15" pageNumber="16">5</td>
|
||
<td id="830A5D8A505900611497FD9BFCF50E04" box="[663,811,581,608]" gridcol="2" gridrow="3" pageId="15" pageNumber="16">22.8–25.4%</td>
|
||
<td id="830A5D8A505900611589FD9BFC3E0E04" box="[905,992,581,608]" gridcol="3" gridrow="3" pageId="15" pageNumber="16">24.2%</td>
|
||
<td id="830A5D8A50590061125DFD9BFB6A0E04" box="[1117,1204,581,608]" gridcol="4" gridrow="3" pageId="15" pageNumber="16">1.11%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FDB9FB6A0EE6" box="[355,1204,615,642]" gridrow="4" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FDB9FE3B0EE6" box="[355,485,615,642]" gridcol="0" gridrow="4" pageId="15" pageNumber="16">SW-10</th>
|
||
<td id="830A5D8A50590061143AFDB9FD940EE6" box="[570,586,615,642]" gridcol="1" gridrow="4" pageId="15" pageNumber="16">6</td>
|
||
<td id="830A5D8A505900611497FDB9FCF50EE6" box="[663,811,615,642]" gridcol="2" gridrow="4" pageId="15" pageNumber="16">51–57</td>
|
||
<td id="830A5D8A505900611589FDB9FC3E0EE6" box="[905,992,615,642]" gridcol="3" gridrow="4" pageId="15" pageNumber="16">53</td>
|
||
<td id="830A5D8A50590061125DFDB9FB6A0EE6" box="[1117,1204,615,642]" gridcol="4" gridrow="4" pageId="15" pageNumber="16">2.25%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FD56FB6A0EC7" box="[355,1204,648,675]" gridrow="5" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FD56FE3B0EC7" box="[355,485,648,675]" gridcol="0" gridrow="5" pageId="15" pageNumber="16">SW-10/TL</th>
|
||
<td id="830A5D8A50590061143AFD56FD940EC7" box="[570,586,648,675]" gridcol="1" gridrow="5" pageId="15" pageNumber="16">6</td>
|
||
<td id="830A5D8A505900611497FD56FCF50EC7" box="[663,811,648,675]" gridcol="2" gridrow="5" pageId="15" pageNumber="16">19.5–22.2%</td>
|
||
<td id="830A5D8A505900611589FD56FC3E0EC7" box="[905,992,648,675]" gridcol="3" gridrow="5" pageId="15" pageNumber="16">20.4%</td>
|
||
<td id="830A5D8A50590061125DFD56FB6A0EC7" box="[1117,1204,648,675]" gridcol="4" gridrow="5" pageId="15" pageNumber="16">0.92%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FD75FB6A0EA1" box="[355,1204,683,709]" gridrow="6" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FD75FE3B0EA1" box="[355,485,683,709]" gridcol="0" gridrow="6" pageId="15" pageNumber="16">S1</th>
|
||
<td id="830A5D8A50590061143AFD75FD940EA1" box="[570,586,683,709]" gridcol="1" gridrow="6" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FD75FCF50EA1" box="[663,811,683,709]" gridcol="2" gridrow="6" pageId="15" pageNumber="16">30–37</td>
|
||
<td id="830A5D8A505900611589FD75FC3E0EA1" box="[905,992,683,709]" gridcol="3" gridrow="6" pageId="15" pageNumber="16">32</td>
|
||
<td id="830A5D8A50590061125DFD75FB6A0EA1" box="[1117,1204,683,709]" gridcol="4" gridrow="6" pageId="15" pageNumber="16">1.94</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FD12FB6A0E83" box="[355,1204,716,743]" gridrow="7" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FD12FE3B0E83" box="[355,485,716,743]" gridcol="0" gridrow="7" pageId="15" pageNumber="16">S2</th>
|
||
<td id="830A5D8A50590061143AFD12FD940E83" box="[570,586,716,743]" gridcol="1" gridrow="7" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FD12FCF50E83" box="[663,811,716,743]" gridcol="2" gridrow="7" pageId="15" pageNumber="16">25–31</td>
|
||
<td id="830A5D8A505900611589FD12FC3E0E83" box="[905,992,716,743]" gridcol="3" gridrow="7" pageId="15" pageNumber="16">27</td>
|
||
<td id="830A5D8A50590061125DFD12FB6A0E83" box="[1117,1204,716,743]" gridcol="4" gridrow="7" pageId="15" pageNumber="16">2.06</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FD30FB6A0F6C" box="[355,1204,750,776]" gridrow="8" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FD30FE3B0F6C" box="[355,485,750,776]" gridcol="0" gridrow="8" pageId="15" pageNumber="16">S3</th>
|
||
<td id="830A5D8A50590061143AFD30FD940F6C" box="[570,586,750,776]" gridcol="1" gridrow="8" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FD30FCF50F6C" box="[663,811,750,776]" gridcol="2" gridrow="8" pageId="15" pageNumber="16">27–31</td>
|
||
<td id="830A5D8A505900611589FD30FC3E0F6C" box="[905,992,750,776]" gridcol="3" gridrow="8" pageId="15" pageNumber="16">29</td>
|
||
<td id="830A5D8A50590061125DFD30FB6A0F6C" box="[1117,1204,750,776]" gridcol="4" gridrow="8" pageId="15" pageNumber="16">1.22</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FCCEFB6A0F4E" box="[355,1204,784,810]" gridrow="9" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FCCEFE3B0F4E" box="[355,485,784,810]" gridcol="0" gridrow="9" pageId="15" pageNumber="16">S4</th>
|
||
<td id="830A5D8A50590061143AFCCEFD940F4E" box="[570,586,784,810]" gridcol="1" gridrow="9" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FCCEFCF50F4E" box="[663,811,784,810]" gridcol="2" gridrow="9" pageId="15" pageNumber="16">29–41</td>
|
||
<td id="830A5D8A505900611589FCCEFC3E0F4E" box="[905,992,784,810]" gridcol="3" gridrow="9" pageId="15" pageNumber="16">33</td>
|
||
<td id="830A5D8A50590061125DFCCEFB6A0F4E" box="[1117,1204,784,810]" gridcol="4" gridrow="9" pageId="15" pageNumber="16">3.46</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FCEEFB6A0F2F" box="[355,1204,816,843]" gridrow="10" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FCEEFE3B0F2F" box="[355,485,816,843]" gridcol="0" gridrow="10" pageId="15" pageNumber="16">S5</th>
|
||
<td id="830A5D8A50590061143AFCEEFD940F2F" box="[570,586,816,843]" gridcol="1" gridrow="10" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FCEEFCF50F2F" box="[663,811,816,843]" gridcol="2" gridrow="10" pageId="15" pageNumber="16">32–46</td>
|
||
<td id="830A5D8A505900611589FCEEFC3E0F2F" box="[905,992,816,843]" gridcol="3" gridrow="10" pageId="15" pageNumber="16">37</td>
|
||
<td id="830A5D8A50590061125DFCEEFB6A0F2F" box="[1117,1204,816,843]" gridcol="4" gridrow="10" pageId="15" pageNumber="16">4.21</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FC8CFB6A0F09" box="[355,1204,850,877]" gridrow="11" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FC8CFE3B0F09" box="[355,485,850,877]" gridcol="0" gridrow="11" pageId="15" pageNumber="16">S6</th>
|
||
<td id="830A5D8A50590061143AFC8CFD940F09" box="[570,586,850,877]" gridcol="1" gridrow="11" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FC8CFCF50F09" box="[663,811,850,877]" gridcol="2" gridrow="11" pageId="15" pageNumber="16">34–47</td>
|
||
<td id="830A5D8A505900611589FC8CFC3E0F09" box="[905,992,850,877]" gridcol="3" gridrow="11" pageId="15" pageNumber="16">40</td>
|
||
<td id="830A5D8A50590061125DFC8CFB6A0F09" box="[1117,1204,850,877]" gridcol="4" gridrow="11" pageId="15" pageNumber="16">3.50</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FCAAFB6A0FEB" box="[355,1204,884,911]" gridrow="12" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FCAAFE3B0FEB" box="[355,485,884,911]" gridcol="0" gridrow="12" pageId="15" pageNumber="16">S7</th>
|
||
<td id="830A5D8A50590061143AFCAAFD940FEB" box="[570,586,884,911]" gridcol="1" gridrow="12" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FCAAFCF50FEB" box="[663,811,884,911]" gridcol="2" gridrow="12" pageId="15" pageNumber="16">38–49</td>
|
||
<td id="830A5D8A505900611589FCAAFC3E0FEB" box="[905,992,884,911]" gridcol="3" gridrow="12" pageId="15" pageNumber="16">43</td>
|
||
<td id="830A5D8A50590061125DFCAAFB6A0FEB" box="[1117,1204,884,911]" gridcol="4" gridrow="12" pageId="15" pageNumber="16">3.16</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FC4BFB6A0FD4" box="[355,1204,917,944]" gridrow="13" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FC4BFE3B0FD4" box="[355,485,917,944]" gridcol="0" gridrow="13" pageId="15" pageNumber="16">S8</th>
|
||
<td id="830A5D8A50590061143AFC4BFD940FD4" box="[570,586,917,944]" gridcol="1" gridrow="13" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FC4BFCF50FD4" box="[663,811,917,944]" gridcol="2" gridrow="13" pageId="15" pageNumber="16">39–48</td>
|
||
<td id="830A5D8A505900611589FC4BFC3E0FD4" box="[905,992,917,944]" gridcol="3" gridrow="13" pageId="15" pageNumber="16">45</td>
|
||
<td id="830A5D8A50590061125DFC4BFB6A0FD4" box="[1117,1204,917,944]" gridcol="4" gridrow="13" pageId="15" pageNumber="16">2.79</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FC66FB6A0FB6" box="[355,1204,952,978]" gridrow="14" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FC66FE3B0FB6" box="[355,485,952,978]" gridcol="0" gridrow="14" pageId="15" pageNumber="16">S9</th>
|
||
<td id="830A5D8A50590061143AFC66FD940FB6" box="[570,586,952,978]" gridcol="1" gridrow="14" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FC66FCF50FB6" box="[663,811,952,978]" gridcol="2" gridrow="14" pageId="15" pageNumber="16">40–46</td>
|
||
<td id="830A5D8A505900611589FC66FC3E0FB6" box="[905,992,952,978]" gridcol="3" gridrow="14" pageId="15" pageNumber="16">43</td>
|
||
<td id="830A5D8A50590061125DFC66FB6A0FB6" box="[1117,1204,952,978]" gridcol="4" gridrow="14" pageId="15" pageNumber="16">1.96</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FC07FB6A0F97" box="[355,1204,985,1011]" gridrow="15" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FC07FE3B0F97" box="[355,485,985,1011]" gridcol="0" gridrow="15" pageId="15" pageNumber="16">S10</th>
|
||
<td id="830A5D8A50590061143AFC07FD940F97" box="[570,586,985,1011]" gridcol="1" gridrow="15" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FC07FCF50F97" box="[663,811,985,1011]" gridcol="2" gridrow="15" pageId="15" pageNumber="16">34–43</td>
|
||
<td id="830A5D8A505900611589FC07FC3E0F97" box="[905,992,985,1011]" gridcol="3" gridrow="15" pageId="15" pageNumber="16">37</td>
|
||
<td id="830A5D8A50590061125DFC07FB6A0F97" box="[1117,1204,985,1011]" gridcol="4" gridrow="15" pageId="15" pageNumber="16">3.24</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FC24FB6A0871" box="[355,1204,1018,1045]" gridrow="16" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FC24FE3B0871" box="[355,485,1018,1045]" gridcol="0" gridrow="16" pageId="15" pageNumber="16">S11</th>
|
||
<td id="830A5D8A50590061143AFC24FD940871" box="[570,586,1018,1045]" gridcol="1" gridrow="16" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FC24FCF50871" box="[663,811,1018,1045]" gridcol="2" gridrow="16" pageId="15" pageNumber="16">29–35</td>
|
||
<td id="830A5D8A505900611589FC24FC3E0871" box="[905,992,1018,1045]" gridcol="3" gridrow="16" pageId="15" pageNumber="16">31</td>
|
||
<td id="830A5D8A50590061125DFC24FB6A0871" box="[1117,1204,1018,1045]" gridcol="4" gridrow="16" pageId="15" pageNumber="16">1.96</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FBFBFB6A0825" box="[355,1204,1061,1089]" gridrow="17" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FBFBFE3B0825" box="[355,485,1061,1089]" gridcol="0" gridrow="17" pageId="15" pageNumber="16">MD4 (ac)</th>
|
||
<td id="830A5D8A50590061143AFBFBFD940825" box="[570,586,1061,1089]" gridcol="1" gridrow="17" pageId="15" pageNumber="16">7</td>
|
||
<td id="830A5D8A505900611497FBFBFCF50825" box="[663,811,1061,1089]" gridcol="2" gridrow="17" pageId="15" pageNumber="16">30–52</td>
|
||
<td id="830A5D8A505900611589FBFBFC3E0825" box="[905,992,1061,1089]" gridcol="3" gridrow="17" pageId="15" pageNumber="16">38</td>
|
||
<td id="830A5D8A50590061125DFBFBFB6A0825" box="[1117,1204,1061,1089]" gridcol="4" gridrow="17" pageId="15" pageNumber="16">8.65</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FB99FB6A0806" box="[355,1204,1095,1122]" gridrow="18" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FB99FE3B0806" box="[355,485,1095,1122]" gridcol="0" gridrow="18" pageId="15" pageNumber="16">MD5 (ac)</th>
|
||
<td id="830A5D8A50590061143AFB99FD940806" box="[570,586,1095,1122]" gridcol="1" gridrow="18" pageId="15" pageNumber="16">6</td>
|
||
<td id="830A5D8A505900611497FB99FCF50806" box="[663,811,1095,1122]" gridcol="2" gridrow="18" pageId="15" pageNumber="16">41–78</td>
|
||
<td id="830A5D8A505900611589FB99FC3E0806" box="[905,992,1095,1122]" gridcol="3" gridrow="18" pageId="15" pageNumber="16">57</td>
|
||
<td id="830A5D8A50590061125DFB99FB6A0806" box="[1117,1204,1095,1122]" gridcol="4" gridrow="18" pageId="15" pageNumber="16">12.19</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FBB7FB6A08E0" box="[355,1204,1129,1156]" gridrow="19" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FBB7FE3B08E0" box="[355,485,1129,1156]" gridcol="0" gridrow="19" pageId="15" pageNumber="16">MD6 (ac)</th>
|
||
<td id="830A5D8A50590061143AFBB7FD9408E0" box="[570,586,1129,1156]" gridcol="1" gridrow="19" pageId="15" pageNumber="16">7</td>
|
||
<td id="830A5D8A505900611497FBB7FCF508E0" box="[663,811,1129,1156]" gridcol="2" gridrow="19" pageId="15" pageNumber="16">64–87</td>
|
||
<td id="830A5D8A505900611589FBB7FC3E08E0" box="[905,992,1129,1156]" gridcol="3" gridrow="19" pageId="15" pageNumber="16">70</td>
|
||
<td id="830A5D8A50590061125DFBB7FB6A08E0" box="[1117,1204,1129,1156]" gridcol="4" gridrow="19" pageId="15" pageNumber="16">8.12</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FB55FB6A08C1" box="[355,1204,1163,1189]" gridrow="20" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FB55FE3B08C1" box="[355,485,1163,1189]" gridcol="0" gridrow="20" pageId="15" pageNumber="16">MD7 (ac)</th>
|
||
<td id="830A5D8A50590061143AFB55FD9408C1" box="[570,586,1163,1189]" gridcol="1" gridrow="20" pageId="15" pageNumber="16">8</td>
|
||
<td id="830A5D8A505900611497FB55FCF508C1" box="[663,811,1163,1189]" gridcol="2" gridrow="20" pageId="15" pageNumber="16">68–93</td>
|
||
<td id="830A5D8A505900611589FB55FC3E08C1" box="[905,992,1163,1189]" gridcol="3" gridrow="20" pageId="15" pageNumber="16">80</td>
|
||
<td id="830A5D8A50590061125DFB55FB6A08C1" box="[1117,1204,1163,1189]" gridcol="4" gridrow="20" pageId="15" pageNumber="16">8.69</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FB72FB6A08A3" box="[355,1204,1196,1223]" gridrow="21" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FB72FE3B08A3" box="[355,485,1196,1223]" gridcol="0" gridrow="21" pageId="15" pageNumber="16">MD8 (ac)</th>
|
||
<td id="830A5D8A50590061143AFB72FD9408A3" box="[570,586,1196,1223]" gridcol="1" gridrow="21" pageId="15" pageNumber="16">8</td>
|
||
<td id="830A5D8A505900611497FB72FCF508A3" box="[663,811,1196,1223]" gridcol="2" gridrow="21" pageId="15" pageNumber="16">96–136</td>
|
||
<td id="830A5D8A505900611589FB72FC3E08A3" box="[905,992,1196,1223]" gridcol="3" gridrow="21" pageId="15" pageNumber="16">110</td>
|
||
<td id="830A5D8A50590061125DFB72FB6A08A3" box="[1117,1204,1196,1223]" gridcol="4" gridrow="21" pageId="15" pageNumber="16">12.08</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FB09FB6A0896" box="[355,1204,1239,1266]" gridrow="22" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FB09FE3B0896" box="[355,485,1239,1266]" gridcol="0" gridrow="22" pageId="15" pageNumber="16">LV6 (ac)</th>
|
||
<td id="830A5D8A50590061143AFB09FD940896" box="[570,586,1239,1266]" gridcol="1" gridrow="22" pageId="15" pageNumber="16">5</td>
|
||
<td id="830A5D8A505900611497FB09FCF50896" box="[663,811,1239,1266]" gridcol="2" gridrow="22" pageId="15" pageNumber="16">28–56</td>
|
||
<td id="830A5D8A505900611589FB09FC3E0896" box="[905,992,1239,1266]" gridcol="3" gridrow="22" pageId="15" pageNumber="16">39</td>
|
||
<td id="830A5D8A50590061125DFB09FB6A0896" box="[1117,1204,1239,1266]" gridcol="4" gridrow="22" pageId="15" pageNumber="16">10.44</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FB27FB6A0970" box="[355,1204,1273,1300]" gridrow="23" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FB27FE3B0970" box="[355,485,1273,1300]" gridcol="0" gridrow="23" pageId="15" pageNumber="16">LV7 (ac)</th>
|
||
<td id="830A5D8A50590061143AFB27FD940970" box="[570,586,1273,1300]" gridcol="1" gridrow="23" pageId="15" pageNumber="16">6</td>
|
||
<td id="830A5D8A505900611497FB27FCF50970" box="[663,811,1273,1300]" gridcol="2" gridrow="23" pageId="15" pageNumber="16">32–68</td>
|
||
<td id="830A5D8A505900611589FB27FC3E0970" box="[905,992,1273,1300]" gridcol="3" gridrow="23" pageId="15" pageNumber="16">48</td>
|
||
<td id="830A5D8A50590061125DFB27FB6A0970" box="[1117,1204,1273,1300]" gridcol="4" gridrow="23" pageId="15" pageNumber="16">11.63</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FAC4FB6A0951" box="[355,1204,1306,1333]" gridrow="24" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FAC4FE3B0951" box="[355,485,1306,1333]" gridcol="0" gridrow="24" pageId="15" pageNumber="16">LV8 (ac)</th>
|
||
<td id="830A5D8A50590061143AFAC4FD940951" box="[570,586,1306,1333]" gridcol="1" gridrow="24" pageId="15" pageNumber="16">8</td>
|
||
<td id="830A5D8A505900611497FAC4FCF50951" box="[663,811,1306,1333]" gridcol="2" gridrow="24" pageId="15" pageNumber="16">39–91</td>
|
||
<td id="830A5D8A505900611589FAC4FC3E0951" box="[905,992,1306,1333]" gridcol="3" gridrow="24" pageId="15" pageNumber="16">58</td>
|
||
<td id="830A5D8A50590061125DFAC4FB6A0951" box="[1117,1204,1306,1333]" gridcol="4" gridrow="24" pageId="15" pageNumber="16">17.61</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FAE2FB6A0933" box="[355,1204,1340,1367]" gridrow="25" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FAE2FE3B0933" box="[355,485,1340,1367]" gridcol="0" gridrow="25" pageId="15" pageNumber="16">LV9 (ac)</th>
|
||
<td id="830A5D8A50590061143AFAE2FD940933" box="[570,586,1340,1367]" gridcol="1" gridrow="25" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FAE2FCF50933" box="[663,811,1340,1367]" gridcol="2" gridrow="25" pageId="15" pageNumber="16">79–120</td>
|
||
<td id="830A5D8A505900611589FAE2FC3E0933" box="[905,992,1340,1367]" gridcol="3" gridrow="25" pageId="15" pageNumber="16">90</td>
|
||
<td id="830A5D8A50590061125DFAE2FB6A0933" box="[1117,1204,1340,1367]" gridcol="4" gridrow="25" pageId="15" pageNumber="16">12.45</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FAB9FB6A09E6" box="[355,1204,1383,1410]" gridrow="26" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FAB9FE3B09E6" box="[355,485,1383,1410]" gridcol="0" gridrow="26" pageId="15" pageNumber="16">LTS</th>
|
||
<td id="830A5D8A50590061143AFAB9FD9409E6" box="[570,586,1383,1410]" gridcol="1" gridrow="26" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FAB9FCF509E6" box="[663,811,1383,1410]" gridcol="2" gridrow="26" pageId="15" pageNumber="16">186–323</td>
|
||
<td id="830A5D8A505900611589FAB9FC3E09E6" box="[905,992,1383,1410]" gridcol="3" gridrow="26" pageId="15" pageNumber="16">263</td>
|
||
<td id="830A5D8A50590061125DFAB9FB6A09E6" box="[1117,1204,1383,1410]" gridcol="4" gridrow="26" pageId="15" pageNumber="16">42.57</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FA56FB6A09C0" box="[355,1204,1416,1444]" gridrow="27" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FA56FE3B09C0" box="[355,485,1416,1444]" gridcol="0" gridrow="27" pageId="15" pageNumber="16">LTS/TL</th>
|
||
<td id="830A5D8A50590061143AFA56FD9409C0" box="[570,586,1416,1444]" gridcol="1" gridrow="27" pageId="15" pageNumber="16">7</td>
|
||
<td id="830A5D8A505900611497FA56FCF509C0" box="[663,811,1416,1444]" gridcol="2" gridrow="27" pageId="15" pageNumber="16">75.0–118.9%</td>
|
||
<td id="830A5D8A505900611589FA56FC3E09C0" box="[905,992,1416,1444]" gridcol="3" gridrow="27" pageId="15" pageNumber="16">101.4%</td>
|
||
<td id="830A5D8A50590061125DFA56FB6A09C0" box="[1117,1204,1416,1444]" gridcol="4" gridrow="27" pageId="15" pageNumber="16">14.54%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505900611763FA75FB6A09A1" box="[355,1204,1451,1477]" gridrow="28" pageId="15" pageNumber="16">
|
||
<th id="830A5D8A505900611763FA75FE3B09A1" box="[355,485,1451,1477]" gridcol="0" gridrow="28" pageId="15" pageNumber="16">LTAS</th>
|
||
<td id="830A5D8A50590061143AFA75FD9409A1" box="[570,586,1451,1477]" gridcol="1" gridrow="28" pageId="15" pageNumber="16">9</td>
|
||
<td id="830A5D8A505900611497FA75FCF509A1" box="[663,811,1451,1477]" gridcol="2" gridrow="28" pageId="15" pageNumber="16">61–78</td>
|
||
<td id="830A5D8A505900611589FA75FC3E09A1" box="[905,992,1451,1477]" gridcol="3" gridrow="28" pageId="15" pageNumber="16">68</td>
|
||
<td id="830A5D8A50590061125DFA75FB6A09A1" box="[1117,1204,1451,1477]" gridcol="4" gridrow="28" pageId="15" pageNumber="16">7.76</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="2A94663C5059FF9116BDF9F6FE270B8D" ID-DOI="http://doi.org/10.5281/zenodo.3817835" ID-Zenodo-Dep="3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="15" pageNumber="16" subCaptionStartIDs="15.[452,623,1823,1849]" subCaptionStarts="Abbr" targetBox="[195,1389,266,2019]" targetPageId="16">
|
||
<paragraph id="7E5436B45059FF9116BDF9F6FE270B8D" blockId="15.[189,1399,1576,2025]" pageId="15" pageNumber="16">
|
||
<emphasis id="4C9FEAA65059FF9116BDF9F6FEDB0A26" bold="true" box="[189,261,1576,1602]" pageId="15" pageNumber="16">Fig. 6</emphasis>
|
||
(opposite page). Scanning electron micrographs showing overviews and details of
|
||
<taxonomicName id="B9EB4D375059FF9112E5F9F6FEFB0A02" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="15" pageNumber="16" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65059FF9112E5F9F6FEFB0A02" italics="true" pageId="15" pageNumber="16">Echinoderes dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5059FF91172BF992FE5B0A02" box="[299,389,1612,1638]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="4C9FEAA65059FF91178BF995FE7E0A01" bold="true" box="[395,416,1611,1637]" pageId="15" pageNumber="16">A</emphasis>
|
||
. Lateral overview.
|
||
<emphasis id="4C9FEAA65059FF91147FF992FD4D0A02" bold="true" box="[639,659,1612,1638]" pageId="15" pageNumber="16">B</emphasis>
|
||
. Segments 1 to 3, laterodorsal view.
|
||
<emphasis id="4C9FEAA65059FF91123CF995FB8F0A01" bold="true" box="[1084,1105,1611,1637]" pageId="15" pageNumber="16">C</emphasis>
|
||
. Segments 1 to 3, ventral view.
|
||
<emphasis id="4C9FEAA65059FF911704F9B1FEC70AED" bold="true" box="[260,281,1647,1673]" pageId="15" pageNumber="16">D</emphasis>
|
||
. Segments 2 to 3, lateral view.
|
||
<emphasis id="4C9FEAA65059FF911493F9B1FD790AED" bold="true" box="[659,679,1647,1673]" pageId="15" pageNumber="16">E</emphasis>
|
||
. Detail showing minute midlateral glandular cell outlet
|
||
<typeStatus id="A15088165059FF911344F9B1FAA90AED" box="[1348,1399,1647,1673]" pageId="15" pageNumber="16">type</typeStatus>
|
||
2 on segment 2.
|
||
<emphasis id="4C9FEAA65059FF91177FF94CFE4F0AC8" bold="true" box="[383,401,1682,1708]" pageId="15" pageNumber="16">F</emphasis>
|
||
. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7.
|
||
<emphasis id="4C9FEAA65059FF9116BDF96BFF0A0AAB" bold="true" box="[189,212,1717,1743]" pageId="15" pageNumber="16">G</emphasis>
|
||
. Segments 4 to 7, left side middorsal to laterodorsal areas.
|
||
<emphasis id="4C9FEAA65059FF911593F96BFC740AAB" bold="true" box="[915,938,1717,1743]" pageId="15" pageNumber="16">H</emphasis>
|
||
. Segments 8, lateral view.
|
||
<emphasis id="4C9FEAA65059FF9112E9F96BFB2A0AAB" bold="true" box="[1257,1268,1717,1743]" pageId="15" pageNumber="16">I</emphasis>
|
||
. Segments
|
||
<quantity id="B9139B515059FF9116BDF906FE9D0A96" box="[189,323,1752,1779]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="15" pageNumber="16" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10 to 11 in</quantity>
|
||
male, laterodorsal view.
|
||
<emphasis id="4C9FEAA65059FF911474F907FD5D0A97" bold="true" box="[628,643,1753,1779]" pageId="15" pageNumber="16">J</emphasis>
|
||
. Segments
|
||
<quantity id="B9139B515059FF911511F906FC490A96" box="[785,919,1752,1779]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="15" pageNumber="16" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10 to 11 in</quantity>
|
||
female, laterodorsal view.
|
||
<emphasis id="4C9FEAA65059FF9112DFF907FB280A97" bold="true" box="[1247,1270,1753,1779]" pageId="15" pageNumber="16">K</emphasis>
|
||
. Mid- and paradorsal regions of segment 9.
|
||
<emphasis id="4C9FEAA65059FF91144AF922FD800B72" bold="true" box="[586,606,1788,1814]" pageId="15" pageNumber="16">L</emphasis>
|
||
. Segments
|
||
<quantity id="B9139B515059FF9114E8F922FC850B71" box="[744,859,1787,1814]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="15" pageNumber="16" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9 to 11 in</quantity>
|
||
male, ventral view.
|
||
<emphasis id="4C9FEAA65059FF91124EF922FBB40B72" bold="true" box="[1102,1130,1788,1814]" pageId="15" pageNumber="16">M</emphasis>
|
||
. Segments
|
||
<quantity id="B9139B515059FF9112F5F922FAA90B71" box="[1269,1399,1787,1814]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="15" pageNumber="16" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10 to 11 in</quantity>
|
||
female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet
|
||
<typeStatus id="A15088165059FF911508F8B8FCE50BE4" box="[776,827,1894,1920]" pageId="15" pageNumber="16">type</typeStatus>
|
||
2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet
|
||
<typeStatus id="A15088165059FF91170BF872FEE00BA2" box="[267,318,1964,1990]" pageId="15" pageNumber="16">type</typeStatus>
|
||
1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C5047FF8F16BDFED2FD180D9C" pageId="17" pageNumber="18" targetBox="[204,1386,547,993]" targetIsTable="true" targetPageId="17">
|
||
<paragraph id="7E5436B45047FF8F16BDFED2FD180D9C" blockId="17.[189,1399,267,504]" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F16BDFED2FEFD0D42" bold="true" box="[189,291,268,294]" pageId="17" pageNumber="18">Table 5.</emphasis>
|
||
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
|
||
<taxonomicName id="B9EB4D375047FF8F17CDFEF1FD0D0D2D" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[461,723,303,329]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="17" pageNumber="18" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65047FF8F17CDFEF1FD0D0D2D" box="[461,723,303,329]" italics="true" pageId="17" pageNumber="18">Echinoderes dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5047FF8F14E0FEF1FC9E0D2D" box="[736,832,303,329]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco 1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; pf = papillary flap; po = pore; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B45047FF8F14EBFDFDFBA20F84" pageId="17" pageNumber="18">
|
||
<table id="0CEBC4145047006116CCFDFDFAB40F85" box="[204,1386,547,993]" colsContinueFrom="9.[216,1389,503,948]" colsContinueIn="57.[210,1369,536,1014]" gridcols="11" gridrows="13" pageId="17" pageNumber="18">
|
||
<tr id="C0DB34F65047006116CCFDFDFAB40E5F" box="[204,1386,547,571]" gridrow="0" pageId="17" pageNumber="18">
|
||
<th id="830A5D8A5047006116CCFDFDFAB40E5F" box="[204,1386,547,571]" colspan="11" colspanRight="10" gridcol="0" gridrow="0" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F14EBFDFDFC990E5F" bold="true" box="[747,839,547,571]" pageId="17" pageNumber="18">Position</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFD8EFAB40E0D" box="[204,1386,592,617]" gridrow="1" pageId="17" pageNumber="18">
|
||
<th id="830A5D8A5047006116CCFD8EFEF00E0D" box="[204,302,592,617]" gridcol="0" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F16CCFD8EFEF00E0C" bold="true" box="[204,302,592,616]" pageId="17" pageNumber="18">Segment</emphasis>
|
||
</th>
|
||
<td id="830A5D8A50470061174CFD8EFE1F0E0D" box="[332,449,592,617]" gridcol="1" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F1770FD8FFE420E0D" bold="true" box="[368,412,593,617]" pageId="17" pageNumber="18">MD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A5047006117D8FD8EFDF00E0D" box="[472,558,592,617]" gridcol="2" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F17F2FD8FFDC80E0D" bold="true" box="[498,534,593,617]" pageId="17" pageNumber="18">PD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504700611456FD8EFDA60E0D" box="[598,632,592,617]" gridcol="3" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F1456FD8EFDA60E0C" bold="true" box="[598,632,592,616]" pageId="17" pageNumber="18">SD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A5047006114B9FD8EFD000E0D" box="[697,734,592,617]" gridcol="4" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F14B9FD8FFD000E0D" bold="true" box="[697,734,593,617]" pageId="17" pageNumber="18">LD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504700611506FD8EFCB30E0D" box="[774,877,592,617]" gridcol="5" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F1524FD8FFC910E0D" bold="true" box="[804,847,593,617]" pageId="17" pageNumber="18">ML</emphasis>
|
||
</td>
|
||
<td id="830A5D8A50470061158DFD8EFC1F0E0D" box="[909,961,592,617]" gridcol="6" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F1597FD8EFC660E0C" bold="true" box="[919,952,592,616]" pageId="17" pageNumber="18">SL</emphasis>
|
||
</td>
|
||
<td id="830A5D8A5047006115E7FD8EFBED0E0D" box="[999,1075,592,617]" gridcol="7" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F15FBFD8EFBFE0E0C" bold="true" box="[1019,1056,592,616]" pageId="17" pageNumber="18">LA</emphasis>
|
||
</td>
|
||
<td id="830A5D8A50470061125EFD8EFBA00E0D" box="[1118,1150,592,617]" gridcol="8" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F125EFD8FFBA00E0D" bold="true" box="[1118,1150,593,617]" pageId="17" pageNumber="18">LV</emphasis>
|
||
</td>
|
||
<td id="830A5D8A5047006112C3FD8EFB360E0D" box="[1219,1256,592,617]" gridcol="9" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F12C3FD8FFB360E0D" bold="true" box="[1219,1256,593,617]" pageId="17" pageNumber="18">VL</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504700611314FD8EFAB40E0D" box="[1300,1386,592,617]" gridcol="10" gridrow="1" pageId="17" pageNumber="18">
|
||
<emphasis id="4C9FEAA65047FF8F1329FD8FFA8B0E0D" bold="true" box="[1321,1365,593,617]" pageId="17" pageNumber="18">VM</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFDA6FAB40EF4" box="[204,1386,632,656]" gridrow="2" pageId="17" pageNumber="18" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFDA6FEF00EF4" box="[204,302,632,656]" gridcol="0" gridrow="2" pageId="17" pageNumber="18">1</th>
|
||
<td id="830A5D8A50470061174CFDA6FE1F0EF4" box="[332,449,632,656]" gridcol="1" gridrow="2" pageId="17" pageNumber="18">gco1, gco1</td>
|
||
<td id="830A5D8A504700611456FDA6FDA60EF4" box="[598,632,632,656]" gridcol="3" gridrow="2" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A5047006114B9FDA6FD000EF4" box="[697,734,632,656]" gridcol="4" gridrow="2" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A50470061158DFDA6FC1F0EF4" box="[909,961,632,656]" gridcol="6" gridrow="2" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFD44FAB40ED6" box="[204,1386,666,690]" gridrow="3" pageId="17" pageNumber="18" rowspan-2="1" rowspan-3="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFD44FEF00ED6" box="[204,302,666,690]" gridcol="0" gridrow="3" pageId="17" pageNumber="18">2</th>
|
||
<td id="830A5D8A50470061174CFD44FE1F0ED6" box="[332,449,666,690]" gridcol="1" gridrow="3" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A5047006114B9FD44FD000ED6" box="[697,734,666,690]" gridcol="4" gridrow="3" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A504700611506FD44FCB30ED6" box="[774,877,666,690]" gridcol="5" gridrow="3" pageId="17" pageNumber="18">gco2</td>
|
||
<td id="830A5D8A504700611314FD44FAB40ED6" box="[1300,1386,666,690]" gridcol="10" gridrow="3" pageId="17" pageNumber="18">ss</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFD65FAB40EB7" box="[204,1386,699,723]" gridrow="4" pageId="17" pageNumber="18" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFD65FEF00EB7" box="[204,302,699,723]" gridcol="0" gridrow="4" pageId="17" pageNumber="18">3</th>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFD03FAB40E91" box="[204,1386,733,757]" gridrow="5" pageId="17" pageNumber="18" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFD03FEF00E91" box="[204,302,733,757]" gridcol="0" gridrow="5" pageId="17" pageNumber="18">4</th>
|
||
<td id="830A5D8A50470061174CFD03FE1F0E91" box="[332,449,733,757]" gridcol="1" gridrow="5" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A5047006117D8FD03FDF00E91" box="[472,558,733,757]" gridcol="2" gridrow="5" pageId="17" pageNumber="18">gco1</td>
|
||
<td id="830A5D8A504700611314FD03FAB40E91" box="[1300,1386,733,757]" gridcol="10" gridrow="5" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFD20FAB40F73" box="[204,1386,766,791]" gridrow="6" pageId="17" pageNumber="18" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFD20FEF00F73" box="[204,302,766,791]" gridcol="0" gridrow="6" pageId="17" pageNumber="18">5</th>
|
||
<td id="830A5D8A50470061174CFD20FE1F0F73" box="[332,449,766,791]" gridcol="1" gridrow="6" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A5047006117D8FD20FDF00F73" box="[472,558,766,791]" gridcol="2" gridrow="6" pageId="17" pageNumber="18">gco1</td>
|
||
<td id="830A5D8A50470061125EFD20FBA00F73" box="[1118,1150,766,791]" gridcol="8" gridrow="6" pageId="17" pageNumber="18">tu</td>
|
||
<td id="830A5D8A504700611314FD20FAB40F73" box="[1300,1386,766,791]" gridcol="10" gridrow="6" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFCFEFAB40F5C" box="[204,1386,800,824]" gridrow="7" pageId="17" pageNumber="18" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFCFEFEF00F5C" box="[204,302,800,824]" gridcol="0" gridrow="7" pageId="17" pageNumber="18">6</th>
|
||
<td id="830A5D8A50470061174CFCFEFE1F0F5C" box="[332,449,800,824]" gridcol="1" gridrow="7" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A5047006117D8FCFEFDF00F5C" box="[472,558,800,824]" gridcol="2" gridrow="7" pageId="17" pageNumber="18">ss, gco1</td>
|
||
<td id="830A5D8A50470061158DFCFEFC1F0F5C" box="[909,961,800,824]" gridcol="6" gridrow="7" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A50470061125EFCFEFBA00F5C" box="[1118,1150,800,824]" gridcol="8" gridrow="7" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A504700611314FCFEFAB40F5C" box="[1300,1386,800,824]" gridcol="10" gridrow="7" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFC9CFAB40F3E" box="[204,1386,834,858]" gridrow="8" pageId="17" pageNumber="18" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFC9CFEF00F3E" box="[204,302,834,858]" gridcol="0" gridrow="8" pageId="17" pageNumber="18">7</th>
|
||
<td id="830A5D8A50470061174CFC9CFE1F0F3E" box="[332,449,834,858]" gridcol="1" gridrow="8" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A5047006117D8FC9CFDF00F3E" box="[472,558,834,858]" gridcol="2" gridrow="8" pageId="17" pageNumber="18">ss, gco1</td>
|
||
<td id="830A5D8A50470061125EFC9CFBA00F3E" box="[1118,1150,834,858]" gridcol="8" gridrow="8" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A504700611314FC9CFAB40F3E" box="[1300,1386,834,858]" gridcol="10" gridrow="8" pageId="17" pageNumber="18">ss, gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFCBAFAB40F18" box="[204,1386,868,892]" gridrow="9" pageId="17" pageNumber="18" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFCBAFEF00F18" box="[204,302,868,892]" gridcol="0" gridrow="9" pageId="17" pageNumber="18">8</th>
|
||
<td id="830A5D8A50470061174CFCBAFE1F0F18" box="[332,449,868,892]" gridcol="1" gridrow="9" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A5047006117D8FCBAFDF00F18" box="[472,558,868,892]" gridcol="2" gridrow="9" pageId="17" pageNumber="18">ss, gco1</td>
|
||
<td id="830A5D8A50470061158DFCBAFC1F0F18" box="[909,961,868,892]" gridcol="6" gridrow="9" pageId="17" pageNumber="18">tu</td>
|
||
<td id="830A5D8A50470061125EFCBAFBA00F18" box="[1118,1150,868,892]" gridcol="8" gridrow="9" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A504700611314FCBAFAB40F18" box="[1300,1386,868,892]" gridcol="10" gridrow="9" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFC5BFAB40FF9" box="[204,1386,901,925]" gridrow="10" pageId="17" pageNumber="18" rowspan-5="1" rowspan-7="1">
|
||
<th id="830A5D8A5047006116CCFC5BFEF00FF9" box="[204,302,901,925]" gridcol="0" gridrow="10" pageId="17" pageNumber="18">9</th>
|
||
<td id="830A5D8A50470061174CFC5BFE1F0FF9" box="[332,449,901,925]" gridcol="1" gridrow="10" pageId="17" pageNumber="18">po,pf</td>
|
||
<td id="830A5D8A5047006117D8FC5BFDF00FF9" box="[472,558,901,925]" gridcol="2" gridrow="10" pageId="17" pageNumber="18">ss, gco1</td>
|
||
<td id="830A5D8A504700611456FC5BFDA60FF9" box="[598,632,901,925]" gridcol="3" gridrow="10" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A5047006114B9FC5BFD000FF9" box="[697,734,901,925]" gridcol="4" gridrow="10" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A50470061158DFC5BFC1F0FF9" box="[909,961,901,925]" gridcol="6" gridrow="10" pageId="17" pageNumber="18">si</td>
|
||
<td id="830A5D8A50470061125EFC5BFBA00FF9" box="[1118,1150,901,925]" gridcol="8" gridrow="10" pageId="17" pageNumber="18">ac</td>
|
||
<td id="830A5D8A5047006112C3FC5BFB360FF9" box="[1219,1256,901,925]" gridcol="9" gridrow="10" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A504700611314FC5BFAB40FF9" box="[1300,1386,901,925]" gridcol="10" gridrow="10" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFC79FAB40FDB" box="[204,1386,935,959]" gridrow="11" pageId="17" pageNumber="18" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="830A5D8A5047006116CCFC79FEF00FDB" box="[204,302,935,959]" gridcol="0" gridrow="11" pageId="17" pageNumber="18">10</th>
|
||
<td id="830A5D8A50470061174CFC79FE1F0FDB" box="[332,449,935,959]" gridcol="1" gridrow="11" pageId="17" pageNumber="18">gco1, gco1</td>
|
||
<td id="830A5D8A5047006117D8FC79FDF00FDB" box="[472,558,935,959]" gridcol="2" gridrow="11" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A5047006114B9FC79FD000FDB" box="[697,734,935,959]" gridcol="4" gridrow="11" pageId="17" pageNumber="18">tu</td>
|
||
<td id="830A5D8A5047006112C3FC79FB360FDB" box="[1219,1256,935,959]" gridcol="9" gridrow="11" pageId="17" pageNumber="18">ss</td>
|
||
<td id="830A5D8A504700611314FC79FAB40FDB" box="[1300,1386,935,959]" gridcol="10" gridrow="11" pageId="17" pageNumber="18">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F65047006116CCFC19FAB40F85" box="[204,1386,967,993]" gridrow="12" pageId="17" pageNumber="18" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="830A5D8A5047006116CCFC19FEF00F85" box="[204,302,967,993]" gridcol="0" gridrow="12" pageId="17" pageNumber="18">11</th>
|
||
<td id="830A5D8A50470061174CFC19FE1F0F85" box="[332,449,967,993]" gridcol="1" gridrow="12" pageId="17" pageNumber="18">gco1, gco1</td>
|
||
<td id="830A5D8A504700611506FC19FCB30F85" box="[774,877,967,993]" gridcol="5" gridrow="12" pageId="17" pageNumber="18">3 x pe(♂)</td>
|
||
<td id="830A5D8A5047006115E7FC19FBED0F85" box="[999,1075,967,993]" gridcol="7" gridrow="12" pageId="17" pageNumber="18">ltas(♀)</td>
|
||
<td id="830A5D8A50470061125EFC19FBA00F85" box="[1118,1150,967,993]" gridcol="8" gridrow="12" pageId="17" pageNumber="18">lts</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45047FF8F16BDFBE5FC2F092F" blockId="17.[189,1399,1082,1355]" pageId="17" pageNumber="18">
|
||
Segment 10 with laterodorsal tubes near posterior segment margin; tubes are long and thin, with bases and lateral wings in both sexes (
|
||
<figureCitation id="E6D02A315047FF8F1401FB80FD58081C" box="[513,646,1118,1144]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="17" pageNumber="18">Figs 4A, D</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315047FF8F1492FB83FD66081C" box="[658,696,1117,1144]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="17" pageNumber="18">5H</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315047FF8F14C4FB80FD0D081D" box="[708,723,1118,1145]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="17" pageNumber="18">6</figureCitation>
|
||
I–J). Sensory spots present in paradorsal and ventrolateral positions (
|
||
<figureCitation id="E6D02A315047FF8F1738FB5FFE5B08FF" box="[312,389,1153,1179]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="17" pageNumber="18">Figs 4</figureCitation>
|
||
A–B, 6I, L–M). Glandular cell outlets
|
||
<typeStatus id="A15088165047FF8F1553FB5FFC5808FF" box="[851,902,1153,1179]" pageId="17" pageNumber="18">type</typeStatus>
|
||
1 present as two middorsal ones and one pair in ventromedial positions (
|
||
<figureCitation id="E6D02A315047FF8F142FFB7AFDAD08DB" box="[559,627,1188,1215]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||
E–F, H). Cuticular hairs scarcer than on preceding segments. The posterior segment margin of the tergal plate is straight; in females it terminates in a pectinate fringe with shorter fringe tips, whereas the fringe is hardly seen in males (
|
||
<figureCitation id="E6D02A315047FF8F159CFB34FC310961" box="[924,1007,1258,1285]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="17" pageNumber="18">Fig. 6I</figureCitation>
|
||
). The margins of the sternal ones are concave, extending posteriorly near the midventral junction (
|
||
<figureCitation id="E6D02A315047FF8F15C5FAD0FBF5094D" box="[965,1067,1294,1321]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="17" pageNumber="18">Fig. 6M</figureCitation>
|
||
); pectinate fringe with long fringe tips present in both sexes. Pachycycli as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45047FF8F16BDFAACFCBB0AC2" blockId="17.[189,1399,1394,1702]" pageId="17" pageNumber="18">
|
||
Segment 11 with lateral terminal spines, nearly equal in length to or longer than trunk length (
|
||
<figureCitation id="E6D02A315047FF8F1311FAACFAAC09E8" box="[1297,1394,1394,1420]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="17" pageNumber="18">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315047FF8F16BDFA4BFF3B09D4" box="[189,229,1429,1456]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="17" pageNumber="18">5A</figureCitation>
|
||
). Males with three pairs of penile spines; dorsal and ventral penile spines are thin, flexible tubes, whereas the median ones are thicker and conical, yet flexible (
|
||
<figureCitation id="E6D02A315047FF8F15B8FA67FBD809B7" box="[952,1030,1465,1491]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="17" pageNumber="18">Figs 4</figureCitation>
|
||
D–E, 5H, 6I, L); females with lateral terminal accessory spines (
|
||
<figureCitation id="E6D02A315047FF8F1445FA02FD510992" box="[581,655,1500,1526]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="17" pageNumber="18">Figs 4</figureCitation>
|
||
A–B, 5G, 6J). No sensory spots observed. Glandular cell outlets
|
||
<typeStatus id="A15088165047FF8F16BDF9DEFF2E0A7E" box="[189,240,1536,1562]" pageId="17" pageNumber="18">type</typeStatus>
|
||
1 present as two middorsal ones (
|
||
<figureCitation id="E6D02A315047FF8F1478FA21FD080A7E" box="[632,726,1535,1562]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="17" pageNumber="18">Fig. 5H</figureCitation>
|
||
). The segment is completely devoid of cuticular hairs, but densely covered with short hair-like extensions, in middorsal to laterodorsal areas, and along the ventral margin (
|
||
<figureCitation id="E6D02A315047FF8F171FF99BFEB10A3B" box="[287,367,1605,1631]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="17" pageNumber="18">Fig. 6I</figureCitation>
|
||
, L–M). Tergal extensions are short and pointed, with an extra tooth at the inferior margin (
|
||
<figureCitation id="E6D02A315047FF8F16C5F9B7FECD0AE7" box="[197,275,1641,1667]" captionStart="Fig" captionStartId="12.[189,232,1791,1817]" captionTargetBox="[261,1323,265,1769]" captionTargetId="figure@12.[260,1327,265,1769]" captionTargetPageId="12" captionText="Fig. 4. Line art illustrations of Echinoderes dubiosus sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♀, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♂, segments 10 to 11, dorsal view. E. ♂, segments 10 to 11, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pe = penile spines; pf = papillary flap; po = pore; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817831" httpUri="https://zenodo.org/record/3817831/files/figure.png" pageId="17" pageNumber="18">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315047FF8F1721F9B6FE990AE7" box="[289,327,1640,1667]" captionStart="Fig" captionStartId="13.[189,232,1683,1709]" captionTargetBox="[236,1351,265,1655]" captionTargetId="figure@13.[236,1351,265,1656]" captionTargetPageId="13" captionText="Fig. 5. Light micrographs showing overviews and details of Echinoderes dubiosus sp. nov., holotype, ♀ (A–E, G) (NHMD-213583) and paratype, ♂ (F, H) (NHMD-213592). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 7 to 10, dorsal view. E. Segments 8 to 11, ventral view. F. Segments 8 to 11, ventral view. G. Segments 10 to 11 in female, ventral view. H. Segments 10 to 11 in male, dorsal view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdss = subdorsal sensory spot; si = sieve plate; slt = sublateral tube; te = tergal extensions; vmgco1 = ventromedial glandular cell outlet type 1." figureDoi="http://doi.org/10.5281/zenodo.3817833" httpUri="https://zenodo.org/record/3817833/files/figure.png" pageId="17" pageNumber="18">5G</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315047FF8F1755F9B7FEAE0AE0" box="[341,368,1641,1668]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="17" pageNumber="18">6I</figureCitation>
|
||
). Sternal extensions not present. Posterior margin of sternal plates with slightly longer fringe tips near the insertion of the lateral terminal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F5047FF8D16BDF910FD4E0901" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="7E5436B45047FF8F16BDF910FEF10A8C" blockId="17.[189,303,1742,1768]" box="[189,303,1742,1768]" pageId="17" pageNumber="18">
|
||
<heading id="251C81D85047FF8F16BDF910FEF10A8C" bold="true" box="[189,303,1742,1768]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="4C9FEAA65047FF8F16BDF910FEF10A8C" bold="true" box="[189,303,1742,1768]" pageId="17" pageNumber="18">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45047FF8C16BDF922FE0D0D2E" blockId="17.[189,1399,1788,2026]" lastBlockId="18.[189,1399,267,330]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
With middorsal spines on segments 4 to 8 and lateroventral tubes/spines on segments 5 to 9,
|
||
<taxonomicName id="B9EB4D375047FF8F12EEF923FAA80B72" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1262,1398,1788,1814]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="17" pageNumber="18" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65047FF8F12EEF923FAA80B72" box="[1262,1398,1788,1814]" italics="true" pageId="17" pageNumber="18">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5047FF8F16BDF8FEFEC20B5E" box="[189,284,1824,1850]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has the most common spine formula among species of
|
||
<emphasis id="4C9FEAA65047FF8F15D7F8C1FBAE0B5D" box="[983,1136,1823,1849]" italics="true" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B9EB4D375047FF8F15D7F8C1FBB20B5D" authorityName="Claparede" authorityYear="1863" box="[983,1132,1823,1849]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="17" pageNumber="18" phylum="Cephalorhyncha" rank="genus">Echinoderes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
But still, the species is easily distinguished from most of its congeners by its thick and very long lateroventral spines on segment 9, combined with the presence of only midlateral glandular cell outlets
|
||
<typeStatus id="A15088165047FF8F1282F8B8FB6B0BE4" box="[1154,1205,1894,1920]" pageId="17" pageNumber="18">type</typeStatus>
|
||
2 on segment 2 and sublateral tubes on segment 8. Even though the spines on segment 9 usually are the longest in the lateroventral series of
|
||
<taxonomicName id="B9EB4D375047FF8F17C2F872FD8A0BA2" authorityName="Claparede" authorityYear="1863" box="[450,596,1964,1990]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="17" pageNumber="18" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA65047FF8F17C2F872FD8A0BA2" box="[450,596,1964,1990]" italics="true" pageId="17" pageNumber="18">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
, they rarely reach the posterior margin of segment 10. Lateroventral spines of segment 9 extending beyond the posterior margin of the terminal segment is only known from
|
||
<taxonomicName id="B9EB4D375044FF8C16BDFED2FE980D41" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[189,326,267,293]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C16BDFED2FE980D41" box="[189,326,267,293]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C174DFED2FE750D42" box="[333,427,268,294]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="B9EB4D375044FF8C17B6FED2FD9D0D41" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[438,579,267,293]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C17B6FED2FD9D0D41" box="[438,579,267,293]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
and females of
|
||
<taxonomicName id="B9EB4D375044FF8C14FEFED5FB6F0D42" authority="Karling, 1954" authorityName="Karling" authorityYear="1954" box="[766,1201,267,294]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="levanderi">
|
||
<emphasis id="4C9FEAA65044FF8C14FEFED5FBD80D41" box="[766,1030,267,293]" italics="true" pageId="18" pageNumber="19">Echinoderes levanderi</emphasis>
|
||
Karling, 1954
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1A7A4B455044FF8C12C0FED5FED60D2E" author="Yamasaki H. & Neuhaus B. & George K. H." pageId="18" pageNumber="19" pagination="79 - 106" refId="ref38720" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 a. Echinoderidae (Cyclorhagida: Kinorhyncha) from two seamounts and the adjacent deep-sea floor in the Northeast Atlantic Ocean, including descriptions of three new species. Cahiers de Biologie Marine 59: 79 - 106." type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA65044FF8C133CFED2FAA90D41" box="[1340,1399,267,293]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
2018a
|
||
</bibRefCitation>
|
||
; Sørensen 2018).
|
||
</paragraph>
|
||
<paragraph id="7E5436B45044FF8C16BDFEADFDB30EE8" blockId="18.[189,1399,370,653]" pageId="18" pageNumber="19">
|
||
Moreover, the presence of glandular cell outlets
|
||
<typeStatus id="A15088165044FF8C1519FEADFC930DE9" box="[793,845,371,397]" pageId="18" pageNumber="19">type</typeStatus>
|
||
<quantity id="B9139B515044FF8C1558FEADFC570DE8" box="[856,905,370,397]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">2 in</quantity>
|
||
midlateral positions only on segment 2 is nearly unique among its congeners, and shared only with
|
||
<taxonomicName id="B9EB4D375044FF8C15D2FE46FBB60DD5" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[978,1128,407,433]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C15D2FE46FBB60DD5" box="[978,1128,407,433]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
. In general, whereas laterodorsal or sublateral glandular cell outlets
|
||
<typeStatus id="A15088165044FF8C14EFFE62FCFD0DB2" box="[751,803,444,470]" pageId="18" pageNumber="19">type</typeStatus>
|
||
2 on segment 2 are fairly common among species of
|
||
<taxonomicName id="B9EB4D375044FF8C16E0FE3EFEAB0D9E" authorityName="Claparede" authorityYear="1863" box="[224,373,480,506]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA65044FF8C16E0FE3EFEAB0D9E" box="[224,373,480,506]" italics="true" pageId="18" pageNumber="19">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
, only two other species,
|
||
<taxonomicName id="B9EB4D375044FF8C14AEFE3FFB6A0D9E" authority="Higgins & Kristensen, 1988" authorityName="Higgins & Kristensen" authorityYear="1988" box="[686,1204,480,506]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="aquilonius">
|
||
<emphasis id="4C9FEAA65044FF8C14AEFE3FFC8F0D9E" box="[686,849,480,506]" italics="true" pageId="18" pageNumber="19">E. aquilonius</emphasis>
|
||
<bibRefCitation id="1A7A4B455044FF8C155BFE3EFB6A0D9E" author="Higgins R. P. & Kristensen R. M." box="[859,1204,480,506]" pageId="18" pageNumber="19" pagination="1 - 56" refId="ref36745" refString="Higgins R. P. & Kristensen R. M. 1988. Kinorhyncha from Disko Island, West Greenland. Smithsonian Contributions to Zoology 458: 1 - 56." type="journal article" year="1988">Higgins & Kristensen, 1988</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9EB4D375044FF8C12F4FE3FFEB20E7A" authority="Higgins, 1960" authorityName="Higgins" authorityYear="1960" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="pennaki">
|
||
<emphasis id="4C9FEAA65044FF8C12F4FE3FFAA90D9E" box="[1268,1399,480,506]" italics="true" pageId="18" pageNumber="19">E. pennaki</emphasis>
|
||
<bibRefCitation id="1A7A4B455044FF8C16BDFDDBFEB20E7A" author="Higgins R. P." box="[189,364,516,543]" pageId="18" pageNumber="19" pagination="85 - 91" refId="ref36525" refString="Higgins R. P. 1960. A new species of Echinoderes (Kinorhyncha) from Puget Sound. Transactions of the American Microscopical Society 79: 85 - 91." type="journal article" year="1960">Higgins, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
, have glandular cell outlets
|
||
<typeStatus id="A15088165044FF8C14C6FDDBFD240E7B" box="[710,762,517,543]" pageId="18" pageNumber="19">type</typeStatus>
|
||
<quantity id="B9139B515044FF8C1503FDDBFCED0E7B" box="[771,819,517,543]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">2 in</quantity>
|
||
midlateral positions of this segment. However, these two species have 2 and 3 pairs of glandular cell outlets
|
||
<typeStatus id="A15088165044FF8C15CBFDF4FC210E20" box="[971,1023,554,580]" pageId="18" pageNumber="19">type</typeStatus>
|
||
2 on segment 2, respectively, and additional pairs on segments 4, 5 and 8 (for
|
||
<taxonomicName id="B9EB4D375044FF8C1515FD90FC660E0C" authorityName="Higgins & Kristensen" authorityYear="1988" box="[789,952,590,616]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="aquilonius">
|
||
<emphasis id="4C9FEAA65044FF8C1515FD90FC660E0C" box="[789,952,590,616]" italics="true" pageId="18" pageNumber="19">E. aquilonius</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="1A7A4B455044FF8C15F1FD90FA980E0C" author="Grzelak K. & Sorensen M. V." box="[1009,1350,590,616]" pageId="18" pageNumber="19" pagination="113 - 147" refId="ref36159" refString="Grzelak K. & Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096 [published online in 2017]." type="journal article" year="2018">Grzelak and Sørensen 2018</bibRefCitation>
|
||
; for
|
||
<taxonomicName id="B9EB4D375044FF8C16BDFDADFEE30EE8" authorityName="Higgins" authorityYear="1960" box="[189,317,626,652]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="pennaki">
|
||
<emphasis id="4C9FEAA65044FF8C16BDFDADFEE30EE8" box="[189,317,626,652]" italics="true" pageId="18" pageNumber="19">E. pennaki</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="1A7A4B455044FF8C1773FDADFDBE0EE8" author="Herranz M. & Yangel E. & Leander B." box="[371,608,626,653]" pageId="18" pageNumber="19" pagination="303 - 325" refId="ref36456" refString="Herranz M., Yangel E. & Leander B. 2018. Echinoderes hakaiensis sp. nov.: a new mud dragon (Kinorhyncha, Echinoderidae) from the northeastern Pacific Ocean with the redescription of Echinoderes pennaki Higgins, 1960. Marine Biodiversity 48: 303 - 325. https: // doi. org / 10.1007 / s 12526 - 017 - 0726 - z" type="journal article" year="2018">
|
||
Herranz
|
||
<emphasis id="4C9FEAA65044FF8C17DBFDADFDC60EE8" box="[475,536,626,652]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E5436B45044FF8C16BDFD6BFD760B5E" blockId="18.[189,1399,693,1851]" pageId="18" pageNumber="19">
|
||
Hence,
|
||
<taxonomicName id="B9EB4D375044FF8C1718FD68FE7C0EAB" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[280,418,693,719]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C1718FD68FE7C0EAB" box="[280,418,693,719]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C17ACFD6BFDD60EAB" box="[428,520,693,719]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is very easily distinguished from most congeners, except for
|
||
<taxonomicName id="B9EB4D375044FF8C12E9FD68FAA90EAB" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[1257,1399,693,719]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C12E9FD68FAA90EAB" box="[1257,1399,693,719]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
with which it shows a very close resemblance.
|
||
<taxonomicName id="B9EB4D375044FF8C14DDFD07FC3F0E97" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[733,993,729,755]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C14DDFD07FC3F0E97" box="[733,993,729,755]" italics="true" pageId="18" pageNumber="19">Echinoderes bathyalis</emphasis>
|
||
</taxonomicName>
|
||
is also described from deep waters (
|
||
<quantity id="B9139B515044FF8C16C5FD20FEB50F7C" box="[197,363,766,793]" metricMagnitude="3" metricUnit="m" metricValue="2.798" metricValueMax="2.875" metricValueMin="2.721" pageId="18" pageNumber="19" unit="m" value="2798.0" valueMax="2875.0" valueMin="2721.0">2721–2875 m</quantity>
|
||
deep), but in the East Atlantic, near Sedlo Seamount, off the
|
||
<collectingRegion id="BC2FF8565044FF8C1252FD20FB770F7C" box="[1106,1193,766,792]" country="Portugal" name="Acores" pageId="18" pageNumber="19">Azores</collectingRegion>
|
||
. The two species have identical spine patterns, and
|
||
<taxonomicName id="B9EB4D375044FF8C144DFCFDFD040F58" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[589,730,802,828]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C144DFCFDFD040F58" box="[589,730,802,828]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
is also characterized by having long lateroventral spines on segment 9, extending well beyond the terminal segment. However, the middorsal spine of segment
|
||
<quantity id="B9139B515044FF8C1368FC99FF0A0FE1" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="18" pageNumber="19" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="B9EB4D375044FF8C16DEFCB2FEB30FE1" authorityName="Yamasaki, Neuhaus & George" authorityYear="2018" box="[222,365,875,901]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C16DEFCB2FEB30FE1" box="[222,365,875,901]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
appears to be longer than the one in
|
||
<taxonomicName id="B9EB4D375044FF8C1534FCB2FC610FE1" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[820,959,875,901]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C1534FCB2FC610FE1" box="[820,959,875,901]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C15C9FCB2FBF70FE2" box="[969,1065,876,902]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and always extending well beyond the terminal segment. In
|
||
<taxonomicName id="B9EB4D375044FF8C1445FC4FFD110FCE" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[581,719,912,938]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C1445FC4FFD110FCE" box="[581,719,912,938]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C14D8FC4EFCEA0FCE" box="[728,820,912,938]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
it extends to the posterior margin of segment 11 only. Both species have tubes on segment 5, but, whereas they are present in lateroventral positions in
|
||
<taxonomicName id="B9EB4D375044FF8C16DEFC04FEB70F97" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[222,361,985,1011]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C16DEFC04FEB70F97" box="[222,361,985,1011]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C1774FC07FE0A0F97" box="[372,468,985,1011]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the tubes are slightly displaced to lateral accessory positions in
|
||
<taxonomicName id="B9EB4D375044FF8C12E7FC04FE080873" authority="(Yamasaki et al. 2018 a)" baseAuthorityName="Yamasaki" baseAuthorityYear="2018" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C12E7FC04FAA90F97" box="[1255,1399,985,1011]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
(
|
||
<bibRefCitation id="1A7A4B455044FF8C16C5FC23FE130873" author="Yamasaki H. & Neuhaus B. & George K. H." box="[197,461,1021,1047]" pageId="18" pageNumber="19" pagination="79 - 106" refId="ref38720" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 a. Echinoderidae (Cyclorhagida: Kinorhyncha) from two seamounts and the adjacent deep-sea floor in the Northeast Atlantic Ocean, including descriptions of three new species. Cahiers de Biologie Marine 59: 79 - 106." type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA65044FF8C1740FC20FEA40873" box="[320,378,1021,1047]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
2018a
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Furthermore, unlike
|
||
<taxonomicName id="B9EB4D375044FF8C14D2FC20FC800873" box="[722,862,1021,1047]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C14D2FC20FC800873" box="[722,862,1021,1047]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9EB4D375044FF8C156CFC20FC2A0873" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[876,1012,1021,1047]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C156CFC20FC2A0873" box="[876,1012,1021,1047]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C15FBFC20FB8B087C" box="[1019,1109,1022,1048]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also has sublateral tubes on segment 8. We would treat this latter difference with some caution though, because even though
|
||
<bibRefCitation id="1A7A4B455044FF8C16BDFB98FE050804" author="Yamasaki H. & Neuhaus B. & George K. H." box="[189,475,1094,1120]" pageId="18" pageNumber="19" pagination="79 - 106" refId="ref38720" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 a. Echinoderidae (Cyclorhagida: Kinorhyncha) from two seamounts and the adjacent deep-sea floor in the Northeast Atlantic Ocean, including descriptions of three new species. Cahiers de Biologie Marine 59: 79 - 106." type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA65044FF8C1739FB99FEA80804" box="[313,374,1094,1120]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
(2018a)
|
||
</bibRefCitation>
|
||
confirm the lack of such a tube in
|
||
<taxonomicName id="B9EB4D375044FF8C1586FB99FBCB0804" authorityName="Sanchez, Pardos & Sorensen" authorityYear="2014" box="[902,1045,1094,1120]" class="Kinorhyncha" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Homalorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="abyssalis">
|
||
<emphasis id="4C9FEAA65044FF8C1586FB99FBCB0804" box="[902,1045,1094,1120]" italics="true" pageId="18" pageNumber="19">E. abyssalis</emphasis>
|
||
</taxonomicName>
|
||
, they note that one specimen has indications of a glandular cell outlet
|
||
<typeStatus id="A15088165044FF8C14A5FBB5FD0608E1" box="[677,728,1131,1157]" pageId="18" pageNumber="19">type</typeStatus>
|
||
<quantity id="B9139B515044FF8C14E1FBB5FCD108E1" box="[737,783,1131,1157]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="19" unit="in" value="2.0">2 in</quantity>
|
||
lateral accessory positions on segment 8. Since the attachment sites of tubes sometimes resemble glandular cell outlets
|
||
<typeStatus id="A15088165044FF8C15EFFB4EFBFC08CE" box="[1007,1058,1168,1194]" pageId="18" pageNumber="19">type</typeStatus>
|
||
2, and the actual tubes may be difficult to visualize in LM, we cannot reject the possibility that some specimens of
|
||
<taxonomicName id="B9EB4D375044FF8C12E7FB6BFAA908AA" box="[1255,1399,1204,1230]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C12E7FB6BFAA908AA" box="[1255,1399,1204,1230]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
have tubes on segment
|
||
<quantity id="B9139B515044FF8C17D0FB07FE230896" box="[464,509,1240,1267]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="18" pageNumber="19" unit="in" value="8.0">8 in</quantity>
|
||
nearly the same positions as
|
||
<taxonomicName id="B9EB4D375044FF8C1557FB07FC3E0896" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[855,992,1240,1266]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C1557FB07FC3E0896" box="[855,992,1240,1266]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C15E8FB07FB9D0897" box="[1000,1091,1241,1267]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
During the preparation of the present description we contacted Dr Yamasaki regarding the nature of this structure in
|
||
<taxonomicName id="B9EB4D375044FF8C12EAFB20FAA90973" box="[1258,1399,1277,1303]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C12EAFB20FAA90973" box="[1258,1399,1277,1303]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
and he confirmed that it was only observed in one specimen (out of three), that it seemed to be a glandular cell outlet
|
||
<typeStatus id="A15088165044FF8C17B0FA98FE3D0904" box="[432,483,1350,1376]" pageId="18" pageNumber="19">type</typeStatus>
|
||
2, rather than a tube, and that the position was clearly lateral accessory rather than sublateral (H. Yamasaki, pers. comm.). Hence, even though the differences are subtle, we would still consider them as diagnostic.
|
||
<taxonomicName id="B9EB4D375044FF8C145BFA51FCBE09CD" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[603,864,1423,1449]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C145BFA51FCBE09CD" box="[603,864,1423,1449]" italics="true" pageId="18" pageNumber="19">Echinoderes dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C156DFA51FC1209CD" box="[877,972,1423,1449]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
furthermore has laterodorsal tubes on segment 10. Such tubes are missing in
|
||
<taxonomicName id="B9EB4D375044FF8C14BFFA6AFC9009A9" box="[703,846,1459,1485]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C14BFFA6AFC9009A9" box="[703,846,1459,1485]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
. However, Yamasaki (pers. comm.) could not confirm the absence of such tubes in
|
||
<taxonomicName id="B9EB4D375044FF8C1475FA07FCDC0996" box="[629,770,1496,1522]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C1475FA07FCDC0996" box="[629,770,1496,1522]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
with complete certainty, because they sometimes are very difficult to see with LM. The two species also seem to differ from each other in their distribution of sensory spots though.
|
||
<taxonomicName id="B9EB4D375044FF8C17EFF9FFFD290A5F" box="[495,759,1569,1595]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C17EFF9FFFD290A5F" box="[495,759,1569,1595]" italics="true" pageId="18" pageNumber="19">Echinoderes bathyalis</emphasis>
|
||
</taxonomicName>
|
||
has no laterodorsal sensory spots on segment 2, but quite distinct subdorsal sensory spots on segment 3 (
|
||
<bibRefCitation id="1A7A4B455044FF8C151EF99BFBFA0A04" author="Yamasaki H. & Neuhaus B. & George K. H." box="[798,1060,1605,1632]" pageId="18" pageNumber="19" pagination="79 - 106" refId="ref38720" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 a. Echinoderidae (Cyclorhagida: Kinorhyncha) from two seamounts and the adjacent deep-sea floor in the Northeast Atlantic Ocean, including descriptions of three new species. Cahiers de Biologie Marine 59: 79 - 106." type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA65044FF8C1599F998FC0D0A3B" box="[921,979,1605,1631]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
2018a
|
||
</bibRefCitation>
|
||
, and subsequently confirmed by Yamasaki, pers. comm.). SEM images of
|
||
<taxonomicName id="B9EB4D375044FF8C14DCF9B5FCB90AE0" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[732,871,1642,1668]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C14DCF9B5FCB90AE0" box="[732,871,1642,1668]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C1571F9B4FC110AE0" box="[881,975,1642,1668]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
quite clearly show the presence of laterodorsal sensory spots on segment 2 (
|
||
<figureCitation id="E6D02A315044FF8C14A6F950FCDD0ACD" box="[678,771,1678,1705]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="18" pageNumber="19">Fig. 6D</figureCitation>
|
||
) and confirm their absence in subdorsal positions on segment 3 (
|
||
<figureCitation id="E6D02A315044FF8C1744F96DFE7E0AAA" box="[324,416,1715,1742]" captionStart="Fig" captionStartId="15.[189,232,1576,1602]" captionTargetId="figure@16.[195,1389,265,2019]" captionTargetPageId="16" captionText="Fig. 6 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes dubiosus sp. nov. A. Lateral overview. B. Segments 1 to 3, laterodorsal view. C. Segments 1 to 3, ventral view. D. Segments 2 to 3, lateral view. E. Detail showing minute midlateral glandular cell outlet type 2 on segment 2. F. Detail showing middorsal spines and paradorsal sensory spots on segments 6 and 7. G. Segments 4 to 7, left side middorsal to laterodorsal areas. H. Segments 8, lateral view. I. Segments 10 to 11 in male, laterodorsal view. J. Segments 10 to 11 in female, laterodorsal view. K. Mid- and paradorsal regions of segment 9. L. Segments 9 to 11 in male, ventral view. M. Segments 10 to 11 in female, ventral view. Abbreviations: ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; pf = papillary flap; po = pore; sdss = subdorsal sensory spot; slgco1 = sublateral glandular cell outlet type 1; slt = sublateral tube; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817835" httpUri="https://zenodo.org/record/3817835/files/figure.png" pageId="18" pageNumber="19">Fig. 6B</figureCitation>
|
||
). Finally,
|
||
<taxonomicName id="B9EB4D375044FF8C1414F96AFD420AA9" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[532,668,1715,1741]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C1414F96AFD420AA9" box="[532,668,1715,1741]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C14A4F96DFD200AA9" box="[676,766,1715,1741]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has an unusual papillary flap in middorsal position on segment 9. This structure seems to be absent in
|
||
<taxonomicName id="B9EB4D375044FF8C14EDF906FCA40A95" box="[749,890,1751,1777]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C14EDF906FCA40A95" box="[749,890,1751,1777]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
, but as stressed above, the structure is very difficult to visualize with LM. According to Yamasaki (pers. comm.) a similar middorsal structure could “possibly be present” in
|
||
<taxonomicName id="B9EB4D375044FF8C17DDF8FFFDB40B5E" box="[477,618,1824,1850]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C17DDF8FFFDB40B5E" box="[477,618,1824,1850]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
also.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45044FF8C16BDF8BAFAA80B8F" blockId="18.[189,1399,1891,2027]" pageId="18" pageNumber="19">
|
||
Hence, in summary,
|
||
<taxonomicName id="B9EB4D375044FF8C17B6F8BAFD9E0B19" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[438,576,1891,1917]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65044FF8C17B6F8BAFD9E0B19" box="[438,576,1891,1917]" italics="true" pageId="18" pageNumber="19">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5044FF8C144AF8BAFD790B1A" box="[586,679,1892,1918]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="B9EB4D375044FF8C14E4F8BAFCAD0B19" box="[740,883,1891,1917]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C14E4F8BAFCAD0B19" box="[740,883,1891,1917]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the proportionally longer middorsal spine of segment
|
||
<quantity id="B9139B515044FF8C145BF856FD560BC6" box="[603,648,1928,1954]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="18" pageNumber="19" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="B9EB4D375044FF8C1490F857FCC20BC6" box="[656,796,1928,1954]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="18" pageNumber="19" phylum="Cephalorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="4C9FEAA65044FF8C1490F857FCC20BC6" box="[656,796,1928,1954]" italics="true" pageId="18" pageNumber="19">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
, and in differences in sensory spot positions on the tergal plates of segments 2 and 3. Additional potential differences are the presence/absence of tubes on segments 8 and 10, and a middorsal papillary flap on segment 9. Despite the subtle differences, we
|
||
</paragraph>
|
||
<paragraph id="7E5436B45045FF8D16BDFED5FCD60D28" blockId="19.[189,1398,267,332]" pageId="19" pageNumber="20">would consider the two species to be distinct, but the resemblance clearly indicates that they are very closely related in spite of the geographic distance.</paragraph>
|
||
<paragraph id="7E5436B45045FF8D16BDFEA6FD4E0901" blockId="19.[189,1399,376,1382]" pageId="19" pageNumber="20">
|
||
One of the noteworthy traits in
|
||
<taxonomicName id="B9EB4D375045FF8D1432FEA7FD650DF6" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[562,699,376,402]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65045FF8D1432FEA7FD650DF6" box="[562,699,376,402]" italics="true" pageId="19" pageNumber="20">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5045FF8D14C4FEA6FCC10DF6" box="[708,799,376,402]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the unusual presence of middorsal structures on segment 9. Middorsal structures in
|
||
<taxonomicName id="B9EB4D375045FF8D1464FE41FD280DDD" authorityName="Claparede" authorityYear="1863" box="[612,758,415,441]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA65045FF8D1464FE41FD280DDD" box="[612,758,415,441]" italics="true" pageId="19" pageNumber="20">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
usually include acicular spines, glandular cell outlets
|
||
<typeStatus id="A15088165045FF8D16BDFE19FF2E0D85" box="[189,240,455,481]" pageId="19" pageNumber="20">type</typeStatus>
|
||
1 and sensory spots. Some species also have an extending protuberance, but this structure would always be protruding from the intersegmental joint between segments 10 and 11 (see, e.g.,
|
||
<taxonomicName id="B9EB4D375045FF8D12ECFE30FDD20E4B" authority="Sorensen & Landers, 2014" authorityName="Sorensen & Landers" authorityYear="2014" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="augustae">
|
||
<emphasis id="4C9FEAA65045FF8D12ECFE30FAA90E63" box="[1260,1399,494,519]" italics="true" pageId="19" pageNumber="20">E. augustae</emphasis>
|
||
<bibRefCitation id="1A7A4B455045FF8D16BDFDCBFDD20E4B" author="Sorensen M. V. & Landers S. C." box="[189,524,533,559]" pageId="19" pageNumber="20" pagination="1 - 18" refId="ref38030" refString="Sorensen M. V. & Landers S. C. 2014. Two new species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Gulf of Mexico. Frontiers in Marine Science 1 (8): 1 - 18. https: // doi. org / 10.3389 / fmars. 2014.00008" type="journal article" year="2014">Sørensen & Landers, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9EB4D375045FF8D141DFDCBFCB10E4A" authority="Higgins, 1964" authorityName="Higgins" authorityYear="1964" box="[541,879,532,559]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="bookhouti">
|
||
<emphasis id="4C9FEAA65045FF8D141DFDCBFD6C0E4B" box="[541,690,533,559]" italics="true" pageId="19" pageNumber="20">E bookhouti</emphasis>
|
||
<bibRefCitation id="1A7A4B455045FF8D14C0FDCBFCB10E4A" author="Higgins R. P." box="[704,879,532,559]" pageId="19" pageNumber="20" pagination="479 - 493" refId="ref36588" refString="Higgins R. P. 1964. Three new kinorhynchs from the North Carolina coast. Bulletin of Marine Science of the Gulf and Caribbean 14: 479 - 493." type="journal article" year="1964">Higgins, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B9EB4D375045FF8D15B9FDCBFEB80E32" authority="Sorensen, Herranz & Landers, 2016" authorityName="Sorensen, Herranz & Landers" authorityYear="2016" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="charlotteae">
|
||
<emphasis id="4C9FEAA65045FF8D15B9FDCBFBB80E4B" box="[953,1126,533,559]" italics="true" pageId="19" pageNumber="20">E. charlotteae</emphasis>
|
||
<bibRefCitation id="1A7A4B455045FF8D1273FDCBFEB80E32" author="Sorensen M. V. & Herranz M. & Landers S. C." pageId="19" pageNumber="20" pagination="48 - 68" refId="ref38383" refString="Sorensen M. V., Herranz M. & Landers S. C. 2016 a. A new species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Gulf of Mexico, with a redescription of Echinoderes bookhouti Higgins, 1964. Zoologischer Anzeiger 265: 48 - 68. https: // doi. org / 10.1016 / j. jcz. 2016.04.004" type="journal article" year="2016">Sørensen, Herranz & Landers, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1A7A4B455045FF8D1775FDE2FD700E32" author="Sorensen M. V. & Landers S. C." box="[373,686,572,598]" pageId="19" pageNumber="20" pagination="1 - 18" refId="ref38030" refString="Sorensen M. V. & Landers S. C. 2014. Two new species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Gulf of Mexico. Frontiers in Marine Science 1 (8): 1 - 18. https: // doi. org / 10.3389 / fmars. 2014.00008" type="journal article" year="2014">Sørensen & Landers 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A7A4B455045FF8D14BBFDE2FC600E33" author="Sorensen M. V. & Herranz M. & Landers S. C." box="[699,958,572,599]" pageId="19" pageNumber="20" pagination="48 - 68" refId="ref38383" refString="Sorensen M. V., Herranz M. & Landers S. C. 2016 a. A new species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Gulf of Mexico, with a redescription of Echinoderes bookhouti Higgins, 1964. Zoologischer Anzeiger 265: 48 - 68. https: // doi. org / 10.1016 / j. jcz. 2016.04.004" type="journal article" year="2016">
|
||
Sørensen
|
||
<emphasis id="4C9FEAA65045FF8D152FFDE3FCB40E32" box="[815,874,572,598]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
|
||
2016a
|
||
</bibRefCitation>
|
||
)). Segment 9 of
|
||
<taxonomicName id="B9EB4D375045FF8D1283FDE3FAD20E32" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1155,1292,572,598]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65045FF8D1283FDE3FAD20E32" box="[1155,1292,572,598]" italics="true" pageId="19" pageNumber="20">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5045FF8D1314FDE2FAAC0E32" box="[1300,1394,572,598]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, however, has an unpaired, middorsal pore, flanked by cuticular ridges anterior to a papillary flap or tuft of papillae, which again is flanked by a pair of paradorsal sensory spots. Such an arrangement has not been reported from any other kinorhynch. We do know, however, about three other cases of uncommon middorsal structures among species of
|
||
<taxonomicName id="B9EB4D375045FF8D1498FD06FCF40E96" authorityName="Claparede" authorityYear="1863" box="[664,810,728,754]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA65045FF8D1498FD06FCF40E96" box="[664,810,728,754]" italics="true" pageId="19" pageNumber="20">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
, and, interestingly, they also appear on segment 9. The description of
|
||
<taxonomicName id="B9EB4D375045FF8D17C0FCDFFC400F7E" authority="Thormar & Sorensen, 2010" authorityName="Thormar & Sorensen" authorityYear="2010" box="[448,926,768,794]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="gizoensis">
|
||
<emphasis id="4C9FEAA65045FF8D17C0FCDFFD900F7E" box="[448,590,769,794]" italics="true" pageId="19" pageNumber="20">E. gizoensis</emphasis>
|
||
<bibRefCitation id="1A7A4B455045FF8D1457FCDEFC400F7E" author="Thormar J. & Sorensen M. V." box="[599,926,768,794]" pageId="19" pageNumber="20" pagination="67 - 96" refId="ref38502" refString="Thormar J. & Sorensen M. V. 2010. Two new species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Solomon Islands. Meiofauna Marina 18: 67 - 96." type="journal article" year="2010">Thormar & Sørensen, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
reports on segment 9 the presence of a middorsal: “…elongate fringed area of filiform cuticular extensions, with a small pore”, situated between a pair of paradorsal sensory spots (
|
||
<bibRefCitation id="1A7A4B455045FF8D146EFC90FC620F0C" author="Thormar J. & Sorensen M. V." box="[622,956,846,872]" pageId="19" pageNumber="20" pagination="67 - 96" refId="ref38502" refString="Thormar J. & Sorensen M. V. 2010. Two new species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Solomon Islands. Meiofauna Marina 18: 67 - 96." type="journal article" year="2010">Thormar & Sørensen 2010</bibRefCitation>
|
||
). This structure somehow resembles the anterior middorsal pore in
|
||
<taxonomicName id="B9EB4D375045FF8D1434FCA8FD1E0FEB" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[564,704,885,911]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65045FF8D1434FCA8FD1E0FEB" box="[564,704,885,911]" italics="true" pageId="19" pageNumber="20">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5045FF8D14CCFCA8FCF30FF4" box="[716,813,886,912]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, even though the new species lacks the filiform cuticular extensions. But then again, the structure in
|
||
<taxonomicName id="B9EB4D375045FF8D1534FC43FC1C0FD2" authorityName="Thormar & Sorensen" authorityYear="2010" box="[820,962,925,950]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="gizoensis">
|
||
<emphasis id="4C9FEAA65045FF8D1534FC43FC1C0FD2" box="[820,962,925,950]" italics="true" pageId="19" pageNumber="20">E. gizoensis</emphasis>
|
||
</taxonomicName>
|
||
might also simply be a single-pored sensory spot with very long papillae. The other unusual middorsal structure on segment 9 was reported from
|
||
<taxonomicName id="B9EB4D375045FF8D1700FC32FE4C0861" authorityName="Karling" authorityYear="1954" box="[256,402,1003,1029]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="levanderi">
|
||
<emphasis id="4C9FEAA65045FF8D1700FC32FE4C0861" box="[256,402,1003,1029]" italics="true" pageId="19" pageNumber="20">E. levanderi</emphasis>
|
||
</taxonomicName>
|
||
. In the species’ recent redescription, Sørensen (2018) mentioned that a “..cuticular depression or scar, lined with short hairs is present in middorsal position, in between the paradorsal sensory spots”, and a nearly similar structure is found in
|
||
<taxonomicName id="B9EB4D375045FF8D155BFBE4FC2D0837" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[859,1011,1081,1107]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="yamasakii" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65045FF8D155BFBE4FC2D0837" box="[859,1011,1081,1107]" italics="true" pageId="19" pageNumber="20">E. yamasakii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5045FF8D15FBFBE4FB880830" box="[1019,1110,1082,1108]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see description below). Again, it is suggestive that this puzzling middorsal structure also appears on segment 9, and, as in the case of
|
||
<taxonomicName id="B9EB4D375045FF8D171AFB56FE7608C5" authorityName="Thormar & Sorensen" authorityYear="2010" box="[282,424,1160,1185]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="gizoensis">
|
||
<emphasis id="4C9FEAA65045FF8D171AFB56FE7608C5" box="[282,424,1160,1185]" italics="true" pageId="19" pageNumber="20">E. gizoensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9EB4D375045FF8D17E4FB56FDB008C5" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[484,622,1159,1185]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="dubiosus" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65045FF8D17E4FB56FDB008C5" box="[484,622,1159,1185]" italics="true" pageId="19" pageNumber="20">E. dubiosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5045FF8D1477FB56FD0808C6" box="[631,726,1160,1186]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, is flanked by paradorsal sensory spots, but besides this, it is hard to argue for or against a homology between any of these structures. Sørensen (2018) was furthermore slightly reluctant about the nature of this structure because it was visible in SEM only, and only one SEM specimen was orientated in a way that allowed examination of the middorsal position of segment 9. The observations from the four species suggest though, that extra attention should be paid to similar structures in future descriptions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |