781 lines
133 KiB
XML
781 lines
133 KiB
XML
<document id="A94378F7242C02DB206F99AADE649742" ID-CLB-Dataset="22296" ID-DOI="10.5852/ejt.2018.456" ID-GBIF-Dataset="0566a11e-d6a3-4283-89c7-c7feea96fdbf" ID-Zenodo-Dep="3817823" ID-ZooBank="DE1B1DEE-9871-4803-9F67-025F2B439560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588965964057" checkinUser="valdenar" docAuthor="Sørensen, Martin V., Rohal, Melissa & Thistle, David" docDate="2018" docId="F64287A25053FF941470FD9FFC8A0B8F" docLanguage="en" docName="ejt-456_sorensen_rohal_thistle.pdf.imf" docOrigin="European Journal of Taxonomy 456" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes anniae Sørensen & Rohal & Thistle 2018, sp. nov." docType="treatment" docUuid="45B173E7-B6E9-443D-8C8F-3C424183DAFD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="11" masterDocId="0A7BFFDA5056FF9E1600FFDEFFDE0C64" masterDocTitle="Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific" masterLastPageNumber="75" masterPageNumber="1" pageNumber="6" updateTime="1698839057356" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="A40BAAE6C96122ABD1DFC0C81B8A98EF">
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<mods:title id="B40412E92ADC437725B3F5AB3D4D5BFE">Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific</mods:title>
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<mods:name id="AB1E568DAB79F1951FBC6FD154247397" type="personal">
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<mods:roleTerm id="8D882AC19AF5428C8F55CE448281215E">Author</mods:roleTerm>
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<mods:namePart id="23F6538C52E59F70CA8E1D71646C3ABD">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="7D80FA0AEBB8B6D1F433863E4C4AD743">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<mods:nameIdentifier id="55D828DACA78EFF68304A0760318C0BF" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="2A6F9F06E4718B3E4AE82BD32D3291E7">Rohal, Melissa</mods:namePart>
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<mods:nameIdentifier id="6B81424329BFB7215DB7DD26B0724632" type="email">mrohal@islander.tamucc.edu</mods:nameIdentifier>
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<mods:namePart id="C50133E029D42D84F790A526332278C3">Thistle, David</mods:namePart>
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<mods:nameIdentifier id="1DE852E15719E4C3CCBFCC75193CA5F4" type="email">dthistle@fsu.edu</mods:nameIdentifier>
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<mods:title id="C0F091B0BD57251CC4960E1773CEE955">European Journal of Taxonomy</mods:title>
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<mods:date id="6C7A1F8332F92FFF180153E7CABBD0A7">2018</mods:date>
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<mods:number id="85396B7366FB22CBB05D687F70391AFD">2018-08-17</mods:number>
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<mods:number id="6DF9D9640F3C4C97C647BA6CBFF4BF35">456</mods:number>
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<treatment id="F64287A25053FF941470FD9FFC8A0B8F" ID-DOI="http://doi.org/10.5281/zenodo.3818848" ID-GBIF-Taxon="163993850" ID-Zenodo-Dep="3818848" LSID="urn:lsid:zoobank.org:act:45B173E7-B6E9-443D-8C8F-3C424183DAFD" httpUri="http://treatment.plazi.org/id/F64287A25053FF941470FD9FFC8A0B8F" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6">
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<subSubSection id="36F1653F5053FF9B1470FD9FFC1A0E38" box="[624,964,577,604]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="7E5436B45053FF9B1470FD9FFC1A0E38" blockId="5.[365,1222,577,676]" box="[624,964,577,604]" pageId="5" pageNumber="6">
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<heading id="251C81D85053FF9B1470FD9FFC1A0E38" box="[624,964,577,604]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
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<taxonomicName id="B9EB4D375053FF9B1470FD9FFCBC0E38" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[624,866,577,604]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="5" pageNumber="6" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
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<emphasis id="4C9FEAA65053FF9B1470FD9FFCBC0E38" bold="true" box="[624,866,577,604]" italics="true" pageId="5" pageNumber="6">Echinoderes anniae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="57AC57DD5053FF9B1569FD9CFC1A0E38" box="[873,964,578,604]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F5053FF9B176DFDB8FB180EE5" box="[365,1222,614,641]" pageId="5" pageNumber="6" type="description">
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<paragraph id="7E5436B45053FF9B176DFDB8FB180EE5" blockId="5.[365,1222,577,676]" box="[365,1222,614,641]" pageId="5" pageNumber="6">
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<uri id="0A7A3AB65053FF9B176DFDB8FB180EE5" box="[365,1222,614,641]" pageId="5" pageNumber="6">
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urn:lsid:zoobank.org:act:
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<uuid id="0A4D0C615053FF9B148FFDB8FB180EE5" box="[655,1222,614,641]" pageId="5" pageNumber="6">45B173E7-B6E9-443D-8C8F-3C424183DAFD</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F5053FF9B14A0FD54FC4A0EC0" box="[672,916,650,676]" pageId="5" pageNumber="6" type="description">
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<paragraph id="7E5436B45053FF9B14A0FD54FC4A0EC0" blockId="5.[365,1222,577,676]" box="[672,916,650,676]" pageId="5" pageNumber="6">
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<figureCitation id="E6D02A315053FF9B14A0FD54FCD70EC0" box="[672,777,650,676]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1754,1780]" captionStartId-1="7.[190,233,1649,1675]" captionTargetBox-0="[213,1376,269,1731]" captionTargetBox-1="[194,1396,291,1609]" captionTargetId-0="figure@6.[211,1377,264,1731]" captionTargetId-1="figure@7.[194,1396,291,1609]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." captionText-1="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi-0="http://doi.org/10.5281/zenodo.3817827" figureDoi-1="http://doi.org/10.5281/zenodo.3817829" httpUri-0="https://zenodo.org/record/3817827/files/figure.png" httpUri-1="https://zenodo.org/record/3817829/files/figure.png" pageId="5" pageNumber="6">Figs 2–3</figureCitation>
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,
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<tableCitation id="3369030F5053FF9B1514FD54FC4A0EC0" box="[788,916,650,676]" captionStart-0="Table 2" captionStart-1="Table 3" captionStartId-0="8.[189,258,268,294]" captionStartId-1="9.[189,258,268,294]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Table 2. Measurements from light microscopy for male holotype of Echinoderes anniae sp. nov. (in µm). Abbreviations: (ac) = acicular spine; LTS = lateral terminal spine; LV = lateroventral; MD, middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW- 10 = standard width, always measured on segment 10; TL = trunk length." captionText-1="Table 3. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes anniae sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♂) = putative male condition of sexually dimorphic character." httpUri-0="http://table.plazi.org/id/2A94663C505EFF9616BDFED2FBC70DEB" httpUri-1="http://table.plazi.org/id/2A94663C505FFF9716BDFED2FD300DB1" pageId="5" pageNumber="6" tableUuid-0="2A94663C505EFF9616BDFED2FBC70DEB" tableUuid-1="2A94663C505FFF9716BDFED2FD300DB1">Tables 2–3</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F5053FF9B16BDFD12FEEC0F94" pageId="5" pageNumber="6" type="diagnosis">
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<paragraph id="7E5436B45053FF9B16BDFD12FEE80E82" blockId="5.[189,310,716,742]" box="[189,310,716,742]" pageId="5" pageNumber="6">
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<heading id="251C81D85053FF9B16BDFD12FEE80E82" bold="true" box="[189,310,716,742]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis id="4C9FEAA65053FF9B16BDFD12FEE80E82" bold="true" box="[189,310,716,742]" pageId="5" pageNumber="6">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7E5436B45053FF9B16BDFD22FEEC0F94" blockId="5.[189,1399,764,1008]" pageId="5" pageNumber="6">
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<taxonomicName id="B9EB4D375053FF9B16BDFD22FE910F72" authorityName="Claparede" authorityYear="1863" box="[189,335,764,790]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="5" pageNumber="6" phylum="Cephalorhyncha" rank="genus">
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<emphasis id="4C9FEAA65053FF9B16BDFD22FE910F72" box="[189,335,764,790]" italics="true" pageId="5" pageNumber="6">Echinoderes</emphasis>
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</taxonomicName>
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with middorsal spines on segments 4, 6 and 8, and spines in lateroventral positions on segments 6 to 9. Tubes present in laterodorsal positions on segment 10, and glandular cell outlets
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<typeStatus id="A15088165053FF9B1344FCFFFAA90F5F" box="[1348,1399,801,827]" pageId="5" pageNumber="6">type</typeStatus>
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<quantity id="B9139B515053FF9B16BDFC9BFF340F3B" box="[189,234,837,863]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="6" unit="in" value="2.0">2 in</quantity>
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midlateral positions on segment 1, in subdorsal, laterodorsal, sublateral and ventrolateral positions on segment 2, in lateral accessory positions on segment 5, and in sublateral positions on segment 8. Tergal extensions of segment 11 extend into thin, flexible tips; sternal extensions short, rounded, with ventrolateral seta-like tuft of extended fringe tips. Males with three pairs of penile spines; females unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F5053FF9B16BDFBC6FBC80806" box="[189,1046,1048,1122]" pageId="5" pageNumber="6" type="etymology">
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<paragraph id="7E5436B45053FF9B16BDFBC6FE9A0856" blockId="5.[189,324,1048,1074]" box="[189,324,1048,1074]" pageId="5" pageNumber="6">
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<heading id="251C81D85053FF9B16BDFBC6FE9A0856" bold="true" box="[189,324,1048,1074]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis id="4C9FEAA65053FF9B16BDFBC6FE9A0856" bold="true" box="[189,324,1048,1074]" pageId="5" pageNumber="6">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7E5436B45053FF9B16BDFB96FBC80806" blockId="5.[189,1046,1096,1122]" box="[189,1046,1096,1122]" pageId="5" pageNumber="6">The first author (MVS) dedicates this species to his mother, Anni Larsen.</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F5053FF9B16BDFB54FABD0924" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="7E5436B45053FF9B16BDFB54FE7408C0" blockId="5.[189,426,1162,1188]" box="[189,426,1162,1188]" pageId="5" pageNumber="6">
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<heading id="251C81D85053FF9B16BDFB54FE7408C0" bold="true" box="[189,426,1162,1188]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis id="4C9FEAA65053FF9B16BDFB54FE7408C0" bold="true" box="[189,426,1162,1188]" pageId="5" pageNumber="6">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="CE833CE95053FF9B16EAFB64FABD0924" ID-GBIF-Occurrence="2611490305" collectingDate="2008-09-16" collectionCode="US" collectorName="St. & Fluoromount G & No & See" country="United States of America" latitude="43.997223" location="West Coast" longLatPrecision="19" longitude="-130.39333" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
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<paragraph id="7E5436B45053FF9B16EAFB64FE8208B0" blockId="5.[189,1398,1210,1345]" box="[234,348,1210,1236]" pageId="5" pageNumber="6">
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<emphasis id="4C9FEAA65053FF9B16EAFB64FE8208B0" bold="true" box="[234,348,1210,1236]" pageId="5" pageNumber="6">
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<typeStatus id="A15088165053FF9B16EAFB64FE8208B0" box="[234,348,1210,1236]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="7E5436B45053FF9B16BDFB01FABD0924" blockId="5.[189,1398,1210,1345]" pageId="5" pageNumber="6">
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<collectingCountry id="06FC76245053FF9B16BDFB01FD8C089C" box="[189,594,1246,1273]" name="United States of America" pageId="5" pageNumber="6">UNITED STATES OF AMERICA</collectingCountry>
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: adult ♂, off
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<collectingRegion id="BC2FF8565053FF9B14E7FB00FC9C089C" box="[743,834,1246,1272]" country="United States of America" name="Oregon" pageId="5" pageNumber="6">Oregon</collectingRegion>
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,
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<collectionCode id="18FAAE715053FF9B154CFB00FCAF089C" box="[844,881,1246,1272]" country="South Africa" name="University of Stellenbosch" pageId="5" pageNumber="6">US</collectionCode>
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<location id="7B34606F5053FF9B1577FB01FC23089C" LSID="urn:lsid:plazi:treatment:F64287A25053FF941470FD9FFC8A0B8F:7B34606F5053FF9B1577FB01FC23089C" box="[887,1021,1246,1273]" country="United States of America" latitude="43.997223" longLatPrecision="19" longitude="-130.39333" name="West Coast" pageId="5" pageNumber="6" stateProvince="Oregon">West Coast</location>
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,
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<geoCoordinate id="1BDF50735053FF9B1207FB02FB45089D" box="[1031,1179,1244,1273]" degrees="43" direction="north" minutes="59" orientation="latitude" pageId="5" pageNumber="6" precision="15" seconds="50" value="43.997223">43°59′50″ N</geoCoordinate>
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,
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<geoCoordinate id="1BDF50735053FF9B12A6FB02FA92089D" box="[1190,1356,1244,1273]" degrees="130" direction="west" minutes="23" orientation="longitude" pageId="5" pageNumber="6" precision="15" seconds="36" value="-130.39333">130°23′36″ W</geoCoordinate>
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,
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<collectorName id="D31E53625053FF9B1357FB00FAA8089C" box="[1367,1398,1246,1272]" pageId="5" pageNumber="6">St.</collectorName>
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1, 3250 m deep,collected from mud,
|
||
<date id="0A5510745053FF9B1467FADCFCDC0978" box="[615,770,1282,1308]" pageId="5" pageNumber="6" value="2008-09-16">
|
||
<collectingDate id="1A11E99C5053FF9B1467FADCFCDC0978" box="[615,770,1282,1308]" pageId="5" pageNumber="6" value="2008-09-16">16 Sep. 2008</collectingDate>
|
||
</date>
|
||
, mounted on an H-S slide in
|
||
<collectorName id="D31E53625053FF9B124EFADCFADD0978" box="[1102,1283,1282,1308]" pageId="5" pageNumber="6">Fluoromount G</collectorName>
|
||
(NHMD- 226483).
|
||
<collectorName id="D31E53625053FF9B172DFAF9FE8F0925" box="[301,337,1319,1345]" pageId="5" pageNumber="6">No</collectorName>
|
||
other specimens were found.
|
||
<collectorName id="D31E53625053FF9B14B1FAF9FD050925" box="[689,731,1319,1345]" pageId="5" pageNumber="6">See</collectorName>
|
||
<figureCitation id="E6D02A315053FF9B14E3FAF8FCF60925" box="[739,808,1318,1345]" captionStart="Fig" captionStartId="2.[189,232,2000,2026]" captionTargetBox="[241,1347,1019,1957]" captionTargetId="figure@2.[241,1347,1019,1958]" captionTargetPageId="2" captionText="Fig. 1. Map showing the eight sampling stations off the US West Coast. See Table 1 for further details." figureDoi="http://doi.org/10.5281/zenodo.3817825" httpUri="https://zenodo.org/record/3817825/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
|
||
for locality and
|
||
<tableCitation id="3369030F5053FF9B15E9FAF8FB9E0925" box="[1001,1088,1318,1345]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="5" pageNumber="6" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
|
||
for detailed station data.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F5053FF9416BDFAB7FDAC0F5B" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="description">
|
||
<paragraph id="7E5436B45053FF9B16BDFAB7FE9009E7" blockId="5.[189,334,1385,1411]" box="[189,334,1385,1411]" pageId="5" pageNumber="6">
|
||
<heading id="251C81D85053FF9B16BDFAB7FE9009E7" bold="true" box="[189,334,1385,1411]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="4C9FEAA65053FF9B16BDFAB7FE9009E7" bold="true" box="[189,334,1385,1411]" pageId="5" pageNumber="6">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45053FF9B16BDFA47FAC30A44" blockId="5.[189,1399,1433,1568]" pageId="5" pageNumber="6">
|
||
Adults with head, neck and eleven trunk segments (
|
||
<figureCitation id="E6D02A315053FF9B1519FA47FCBC09D7" box="[793,866,1433,1459]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
|
||
A–B, 3A). The trunk appears stout, with very well-developed pachycycli, almost parallel-sided, and round in cross-section. Only male morphology is known. For complete overview of measurements and dimensions, see
|
||
<tableCitation id="3369030F5053FF9B15F5FA3FFB91099F" box="[1013,1103,1505,1531]" captionStart="Table 2" captionStartId="8.[189,258,268,294]" captionTargetPageId="8" captionText="Table 2. Measurements from light microscopy for male holotype of Echinoderes anniae sp. nov. (in µm). Abbreviations: (ac) = acicular spine; LTS = lateral terminal spine; LV = lateroventral; MD, middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW- 10 = standard width, always measured on segment 10; TL = trunk length." httpUri="http://table.plazi.org/id/2A94663C505EFF9616BDFED2FBC70DEB" pageId="5" pageNumber="6" tableUuid="2A94663C505EFF9616BDFED2FBC70DEB">Table 2</tableCitation>
|
||
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, is summarized in
|
||
<tableCitation id="3369030F5053FF9B12BFF9DBFAC60A44" box="[1215,1304,1541,1568]" captionStart="Table 3" captionStartId="9.[189,258,268,294]" captionTargetPageId="9" captionText="Table 3. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes anniae sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♂) = putative male condition of sexually dimorphic character." httpUri="http://table.plazi.org/id/2A94663C505FFF9716BDFED2FD300DB1" pageId="5" pageNumber="6" tableUuid="2A94663C505FFF9716BDFED2FD300DB1">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45053FF9B16BDF996FE390A97" blockId="5.[189,1399,1608,1779]" pageId="5" pageNumber="6">
|
||
The head was fully retracted in the holotypic specimen; hence, no information on head morphology is available. The neck has 16 placids, measuring
|
||
<quantity id="B9139B515053FF9B150CF9B2FC840AE3" box="[780,858,1644,1671]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="mm" value="12.0">12 µm</quantity>
|
||
in length. The midventral placid is broadest, measuring
|
||
<quantity id="B9139B515053FF9B173EF94FFE550ACF" box="[318,395,1681,1707]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="6" unit="mm" value="13.0">13 µm</quantity>
|
||
in width at its base, whereas all others are narrower, measuring
|
||
<quantity id="B9139B515053FF9B1278F94FFB680ACF" box="[1144,1206,1681,1707]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="6" unit="mm" value="9.0">9 µm</quantity>
|
||
in width at their bases. Four dorsal and two ventral trichoscalid plates are present, with the ventral ones being broader than the four dorsal ones.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45053FF9B16BDF8C2FBF80B8F" blockId="5.[189,1399,1819,2027]" pageId="5" pageNumber="6">
|
||
Segment 1 consists of a complete cuticular ring. Sensory spots are located on anterior segment half, but not immediately at the anterior margin, in subdorsal and laterodorsal positions (
|
||
<figureCitation id="E6D02A315053FF9B129FF89EFADC0B3E" box="[1183,1282,1856,1882]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="5" pageNumber="6">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315053FF9B1310F89EFAEB0B3E" box="[1296,1333,1856,1882]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="5" pageNumber="6">3B</figureCitation>
|
||
), and more posteriorly, in ventrolateral positions (
|
||
<figureCitation id="E6D02A315053FF9B14CAF8BAFCF20B1A" box="[714,812,1892,1918]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="5" pageNumber="6">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315053FF9B153AF8BAFCBE0B1A" box="[826,864,1892,1918]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="5" pageNumber="6">3C</figureCitation>
|
||
). Glandular cell outlets
|
||
<typeStatus id="A15088165053FF9B1280F8BAFB6D0B1A" box="[1152,1203,1892,1918]" pageId="5" pageNumber="6">type</typeStatus>
|
||
2 are present in midlateral positions (
|
||
<figureCitation id="E6D02A315053FF9B17B9F856FDDA0BC6" box="[441,516,1928,1954]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
|
||
A–B, 3B–C), and glandular cell outlets
|
||
<typeStatus id="A15088165053FF9B15E3F857FBC80BC7" box="[995,1046,1929,1955]" pageId="5" pageNumber="6">type</typeStatus>
|
||
<quantity id="B9139B515053FF9B121EF856FB930BC6" box="[1054,1101,1928,1954]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="6" unit="in" value="1.0">1 in</quantity>
|
||
middorsal (
|
||
<figureCitation id="E6D02A315053FF9B12DEF856FAE40BC6" box="[1246,1338,1928,1954]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="5" pageNumber="6">Fig. 3B</figureCitation>
|
||
) and lateroventral positions. The segment appears to be devoid of cuticular hairs. The posterior segment margin is nearly straight, terminating in a well-developed pectinate fringe.
|
||
</paragraph>
|
||
<caption id="2A94663C5050FF9816BDF904FD440B8F" ID-DOI="http://doi.org/10.5281/zenodo.3817827" ID-Zenodo-Dep="3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1754,1780]" targetBox="[213,1376,269,1731]" targetPageId="6">
|
||
<paragraph id="7E5436B45050FF9816BDF904FD440B8F" blockId="6.[189,1399,1753,2028]" pageId="6" pageNumber="7">
|
||
<emphasis id="4C9FEAA65050FF9816BDF904FED00A90" bold="true" box="[189,270,1754,1780]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
|
||
Line art illustrations of
|
||
<taxonomicName id="B9EB4D375050FF98143EF905FCF50A90" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[574,811,1755,1781]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="6" pageNumber="7" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65050FF98143EF905FCF50A90" box="[574,811,1755,1781]" italics="true" pageId="6" pageNumber="7">Echinoderes anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5050FF981536F905FC4B0A91" box="[822,917,1755,1781]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="4C9FEAA65050FF9815A0F904FC6B0A90" bold="true" box="[928,949,1754,1780]" pageId="6" pageNumber="7">A</emphasis>
|
||
. ♂, dorsal view.
|
||
<emphasis id="4C9FEAA65050FF98128DF905FB7F0A91" bold="true" box="[1165,1185,1755,1781]" pageId="6" pageNumber="7">B</emphasis>
|
||
. ♂, ventral view.
|
||
<emphasis id="4C9FEAA65050FF9816BDF920FF0C0B7C" bold="true" box="[189,210,1790,1816]" pageId="6" pageNumber="7">C</emphasis>
|
||
. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C5051FF9916BEF9AFFAF70B88" ID-DOI="http://doi.org/10.5281/zenodo.3817829" ID-Zenodo-Dep="3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="7" pageNumber="8" startId="7.[190,233,1649,1675]" targetBox="[194,1396,291,1609]" targetPageId="7">
|
||
<paragraph id="7E5436B45051FF9916BEF9AFFAF70B88" blockId="7.[190,1400,1649,2028]" pageId="7" pageNumber="8">
|
||
<emphasis id="4C9FEAA65051FF9916BEF9AFFECE0AEF" bold="true" box="[190,272,1649,1675]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
|
||
Light micrographs showing overviews and details of male holotype of
|
||
<taxonomicName id="B9EB4D375051FF99128AF9AFFAA90AEF" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1162,1399,1649,1675]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="7" pageNumber="8" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65051FF99128AF9AFFAA90AEF" box="[1162,1399,1649,1675]" italics="true" pageId="7" pageNumber="8">Echinoderes anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5051FF9916BEF94BFEC20ACB" box="[190,284,1685,1711]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NHMD-226483).
|
||
<emphasis id="4C9FEAA65051FF991407F94AFDC20ACA" bold="true" box="[519,540,1684,1710]" pageId="7" pageNumber="8">A</emphasis>
|
||
. Ventral overview.
|
||
<emphasis id="4C9FEAA65051FF99150AF94BFCC00ACB" bold="true" box="[778,798,1685,1711]" pageId="7" pageNumber="8">B</emphasis>
|
||
. Segments 1 to 4, dorsal view.
|
||
<emphasis id="4C9FEAA65051FF9912A3F94AFB660ACA" bold="true" box="[1187,1208,1684,1710]" pageId="7" pageNumber="8">C</emphasis>
|
||
. Segments 1 to 4, ventral view.
|
||
<emphasis id="4C9FEAA65051FF99177FF966FE4A0AB6" bold="true" box="[383,404,1720,1746]" pageId="7" pageNumber="8">D</emphasis>
|
||
. Segments 8 to 9, dorsal view.
|
||
<emphasis id="4C9FEAA65051FF991511F966FCFB0AB6" bold="true" box="[785,805,1720,1746]" pageId="7" pageNumber="8">E</emphasis>
|
||
. Segments 5 to 7, ventral view.
|
||
<emphasis id="4C9FEAA65051FF9912AAF966FB620AB6" bold="true" box="[1194,1212,1720,1746]" pageId="7" pageNumber="8">F</emphasis>
|
||
. Segments 9 to 11, dorsal view.
|
||
<emphasis id="4C9FEAA65051FF991781F904FE460A90" bold="true" box="[385,408,1754,1780]" pageId="7" pageNumber="8">G</emphasis>
|
||
. Segments 10 to 11, ventral view.
|
||
<emphasis id="4C9FEAA65051FF991534F905FC950A91" bold="true" box="[820,843,1755,1781]" pageId="7" pageNumber="8">H</emphasis>
|
||
. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C505EFF9616BDFED2FBC70DEB" ID-Table-UUID="2A94663C505EFF9616BDFED2FBC70DEB" httpUri="http://table.plazi.org/id/2A94663C505EFF9616BDFED2FBC70DEB" pageId="8" pageNumber="9" targetBox="[594,971,441,1392]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="7E5436B4505EFF9616BDFED2FBC70DEB" blockId="8.[189,1398,267,399]" pageId="8" pageNumber="9">
|
||
<emphasis id="4C9FEAA6505EFF9616BDFED2FEC00D41" bold="true" box="[189,286,267,294]" pageId="8" pageNumber="9">Table 2.</emphasis>
|
||
Measurements from light microscopy for male holotype of
|
||
<taxonomicName id="B9EB4D37505EFF9615CDFED5FB6A0D41" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[973,1204,267,293]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="8" pageNumber="9" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505EFF9615CDFED5FB6A0D41" box="[973,1204,267,293]" italics="true" pageId="8" pageNumber="9">Echinoderes anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505EFF9612B9FED2FACC0D42" box="[1209,1298,268,294]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(in µm). Abbreviations: (ac) = acicular spine; LTS = lateral terminal spine; LV = lateroventral; MD, middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW- 10 = standard width, always measured on segment 10; TL = trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B4505EFF961455FE67FC1D0914" pageId="8" pageNumber="9">
|
||
<table id="0CEBC414505E00611452FE67FC150914" box="[594,971,441,1392]" gridcols="2" gridrows="26" pageId="8" pageNumber="9">
|
||
<tr id="C0DB34F6505E00611452FE67FC150DB0" box="[594,971,441,468]" gridrow="0" pageId="8" pageNumber="9">
|
||
<th id="830A5D8A505E00611452FE67FD040DB0" box="[594,730,441,468]" gridcol="0" gridrow="0" pageId="8" pageNumber="9">
|
||
<emphasis id="4C9FEAA6505EFF961455FE67FD080DB7" bold="true" box="[597,726,441,467]" pageId="8" pageNumber="9">Character</emphasis>
|
||
</th>
|
||
<th id="830A5D8A505E00611570FE67FC150DB0" box="[880,971,441,468]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
|
||
<emphasis id="4C9FEAA6505EFF961571FE64FC150DB0" bold="true" box="[881,971,442,468]" pageId="8" pageNumber="9">Length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FE32FC150E63" box="[594,971,492,519]" gridrow="1" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FE32FD040E63" box="[594,730,492,519]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">TL</td>
|
||
<td id="830A5D8A505E00611570FE32FC150E63" box="[880,971,492,519]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">353</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FDD1FC150E4D" box="[594,971,527,553]" gridrow="2" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FDD1FD040E4D" box="[594,730,527,553]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">MSW-8</td>
|
||
<td id="830A5D8A505E00611570FDD1FC150E4D" box="[880,971,527,553]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">61</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FDECFC150E28" box="[594,971,562,588]" gridrow="3" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FDECFD040E28" box="[594,730,562,588]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">MSW-8/TL</td>
|
||
<td id="830A5D8A505E00611570FDECFC150E28" box="[880,971,562,588]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">17.3%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FD88FC150E14" box="[594,971,598,624]" gridrow="4" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FD88FD040E14" box="[594,730,598,624]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">SW-10</td>
|
||
<td id="830A5D8A505E00611570FD88FC150E14" box="[880,971,598,624]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">49</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FDA7FC150EF7" box="[594,971,633,659]" gridrow="5" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FDA7FD040EF7" box="[594,730,633,659]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">SW-10/TL</td>
|
||
<td id="830A5D8A505E00611570FDA7FC150EF7" box="[880,971,633,659]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">13.9%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FD42FC150ED2" box="[594,971,668,694]" gridrow="6" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FD42FD040ED2" box="[594,730,668,694]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">S1</td>
|
||
<td id="830A5D8A505E00611570FD42FC150ED2" box="[880,971,668,694]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">38</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FD61FC150EBD" box="[594,971,703,729]" gridrow="7" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FD61FD040EBD" box="[594,730,703,729]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">S2</td>
|
||
<td id="830A5D8A505E00611570FD61FC150EBD" box="[880,971,703,729]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">32</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FD3CFC150E98" box="[594,971,738,764]" gridrow="8" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FD3CFD040E98" box="[594,730,738,764]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">S3</td>
|
||
<td id="830A5D8A505E00611570FD3CFC150E98" box="[880,971,738,764]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">36</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FCD8FC150F44" box="[594,971,774,800]" gridrow="9" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FCD8FD040F44" box="[594,730,774,800]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">S4</td>
|
||
<td id="830A5D8A505E00611570FCD8FC150F44" box="[880,971,774,800]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">37</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FCF7FC150F20" box="[594,971,809,836]" gridrow="10" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FCF7FD040F20" box="[594,730,809,836]" gridcol="0" gridrow="10" pageId="8" pageNumber="9">S5</td>
|
||
<td id="830A5D8A505E00611570FCF7FC150F20" box="[880,971,809,836]" gridcol="1" gridrow="10" pageId="8" pageNumber="9">40</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FC92FC150F02" box="[594,971,844,870]" gridrow="11" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FC92FD040F02" box="[594,730,844,870]" gridcol="0" gridrow="11" pageId="8" pageNumber="9">S6</td>
|
||
<td id="830A5D8A505E00611570FC92FC150F02" box="[880,971,844,870]" gridcol="1" gridrow="11" pageId="8" pageNumber="9">42</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FCB1FC150FEE" box="[594,971,879,906]" gridrow="12" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FCB1FD040FEE" box="[594,730,879,906]" gridcol="0" gridrow="12" pageId="8" pageNumber="9">S7</td>
|
||
<td id="830A5D8A505E00611570FCB1FC150FEE" box="[880,971,879,906]" gridcol="1" gridrow="12" pageId="8" pageNumber="9">51</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FC4CFC150FC9" box="[594,971,914,941]" gridrow="13" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FC4CFD040FC9" box="[594,730,914,941]" gridcol="0" gridrow="13" pageId="8" pageNumber="9">S8</td>
|
||
<td id="830A5D8A505E00611570FC4CFC150FC9" box="[880,971,914,941]" gridcol="1" gridrow="13" pageId="8" pageNumber="9">52</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FC6BFC150FB4" box="[594,971,949,976]" gridrow="14" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FC6BFD040FB4" box="[594,730,949,976]" gridcol="0" gridrow="14" pageId="8" pageNumber="9">S9</td>
|
||
<td id="830A5D8A505E00611570FC6BFC150FB4" box="[880,971,949,976]" gridcol="1" gridrow="14" pageId="8" pageNumber="9">55</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FC07FC150F97" box="[594,971,985,1011]" gridrow="15" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FC07FD040F97" box="[594,730,985,1011]" gridcol="0" gridrow="15" pageId="8" pageNumber="9">S10</td>
|
||
<td id="830A5D8A505E00611570FC07FC150F97" box="[880,971,985,1011]" gridcol="1" gridrow="15" pageId="8" pageNumber="9">44</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FC22FC150872" box="[594,971,1020,1046]" gridrow="16" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FC22FD040872" box="[594,730,1020,1046]" gridcol="0" gridrow="16" pageId="8" pageNumber="9">S11</td>
|
||
<td id="830A5D8A505E00611570FC22FC150872" box="[880,971,1020,1046]" gridcol="1" gridrow="16" pageId="8" pageNumber="9">38</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FBF7FC150827" box="[594,971,1065,1091]" gridrow="17" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FBF7FD040827" box="[594,730,1065,1091]" gridcol="0" gridrow="17" pageId="8" pageNumber="9">MD4 (ac)</td>
|
||
<td id="830A5D8A505E00611570FBF7FC150827" box="[880,971,1065,1091]" gridcol="1" gridrow="17" pageId="8" pageNumber="9">36</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FB92FC150802" box="[594,971,1100,1126]" gridrow="18" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FB92FD040802" box="[594,730,1100,1126]" gridcol="0" gridrow="18" pageId="8" pageNumber="9">MD6 (ac)</td>
|
||
<td id="830A5D8A505E00611570FB92FC150802" box="[880,971,1100,1126]" gridcol="1" gridrow="18" pageId="8" pageNumber="9">missing</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FBB1FC1508ED" box="[594,971,1135,1161]" gridrow="19" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FBB1FD0408ED" box="[594,730,1135,1161]" gridcol="0" gridrow="19" pageId="8" pageNumber="9">MD8 (ac)</td>
|
||
<td id="830A5D8A505E00611570FBB1FC1508ED" box="[880,971,1135,1161]" gridcol="1" gridrow="19" pageId="8" pageNumber="9">107</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FB42FC1508D2" box="[594,971,1180,1206]" gridrow="20" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FB42FD0408D2" box="[594,730,1180,1206]" gridcol="0" gridrow="20" pageId="8" pageNumber="9">LV6 (ac)</td>
|
||
<td id="830A5D8A505E00611570FB42FC1508D2" box="[880,971,1180,1206]" gridcol="1" gridrow="20" pageId="8" pageNumber="9">32</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FB61FC1508BE" box="[594,971,1215,1242]" gridrow="21" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FB61FD0408BE" box="[594,730,1215,1242]" gridcol="0" gridrow="21" pageId="8" pageNumber="9">LV7 (ac)</td>
|
||
<td id="830A5D8A505E00611570FB61FC1508BE" box="[880,971,1215,1242]" gridcol="1" gridrow="21" pageId="8" pageNumber="9">33</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FB3CFC150898" box="[594,971,1250,1276]" gridrow="22" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FB3CFD040898" box="[594,730,1250,1276]" gridcol="0" gridrow="22" pageId="8" pageNumber="9">LV8 (ac)</td>
|
||
<td id="830A5D8A505E00611570FB3CFC150898" box="[880,971,1250,1276]" gridcol="1" gridrow="22" pageId="8" pageNumber="9">36</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FAD8FC150944" box="[594,971,1286,1312]" gridrow="23" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FAD8FD040944" box="[594,730,1286,1312]" gridcol="0" gridrow="23" pageId="8" pageNumber="9">LV9 (ac)</td>
|
||
<td id="830A5D8A505E00611570FAD8FC150944" box="[880,971,1286,1312]" gridcol="1" gridrow="23" pageId="8" pageNumber="9">41</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FAECFC150929" box="[594,971,1330,1357]" gridrow="24" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FAECFD040929" box="[594,730,1330,1357]" gridcol="0" gridrow="24" pageId="8" pageNumber="9">LTS</td>
|
||
<td id="830A5D8A505E00611570FAECFC150929" box="[880,971,1330,1357]" gridcol="1" gridrow="24" pageId="8" pageNumber="9">351</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505E00611452FA88FC150914" box="[594,971,1366,1392]" gridrow="25" pageId="8" pageNumber="9">
|
||
<td id="830A5D8A505E00611452FA88FD040914" box="[594,730,1366,1392]" gridcol="0" gridrow="25" pageId="8" pageNumber="9">LTS/TL</td>
|
||
<td id="830A5D8A505E00611570FA88FC150914" box="[880,971,1366,1392]" gridcol="1" gridrow="25" pageId="8" pageNumber="9">99.4%</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505EFF9616BDFA39FB510AB0" blockId="8.[189,1399,1511,1749]" pageId="8" pageNumber="9">
|
||
Segment 2 consists of a complete cuticular ring, with glandular cell outlets
|
||
<typeStatus id="A1508816505EFF961234FA39FBB90A65" box="[1076,1127,1511,1537]" pageId="8" pageNumber="9">type</typeStatus>
|
||
2 located in subdorsal, laterodorsal, sublateral and ventrolateral positions (
|
||
<figureCitation id="E6D02A31505EFF96152DF9D4FCA50A40" box="[813,891,1546,1572]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="8" pageNumber="9">Figs 2</figureCitation>
|
||
A–B, 3B–C). Pachycyclus of the anterior segment margin is well-developed, with weak indications of interruptions in middorsal and lateroventral positions. Sensory spots are located in laterodorsal and ventromedial positions. A single glandular cell outlet
|
||
<typeStatus id="A1508816505EFF961708F9AAFEE50AEA" box="[264,315,1652,1678]" pageId="8" pageNumber="9">type</typeStatus>
|
||
1 is located in middorsal position. Secondary pectinate fringe not detected on this or any of the following segments. They are evenly distributed around the segment. The posterior segment margin is nearly straight, and consists of a pectinate fringe with well-developed fringe tips.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505EFF9616BDF922FD190B8D" blockId="8.[189,1399,1787,2025]" pageId="8" pageNumber="9">
|
||
Segment 3, and remaining segments, consist of one tergal and two sternal plates (
|
||
<figureCitation id="E6D02A31505EFF961293F925FB280B72" box="[1171,1270,1787,1814]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="8" pageNumber="9">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505EFF961304F922FAAF0B72" box="[1284,1393,1788,1814]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="8" pageNumber="9">3A, C, E</figureCitation>
|
||
, G–H). Pachycyclus of the anterior segment margin is well-developed, and interrupted at the tergosternal junctions, but also middorsally. Sensory spots are located in subdorsal and midlateral positions (
|
||
<figureCitation id="E6D02A31505EFF96130FF89CFA8C0B38" box="[1295,1362,1858,1884]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
|
||
A– B). Glandular cell outlets
|
||
<typeStatus id="A1508816505EFF9617EEF8B8FDFF0BE4" box="[494,545,1894,1920]" pageId="8" pageNumber="9">type</typeStatus>
|
||
1 are located in ventromedial positions (
|
||
<figureCitation id="E6D02A31505EFF961201F8BBFB830B1B" box="[1025,1117,1893,1919]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="8" pageNumber="9">Fig. 3B</figureCitation>
|
||
). Cuticular hairs evenly distributed over tergal plate, between secondary fringe and intersegmental joint line, whereas the sternal plates only have hairs on their most lateral halves. Posterior segment margin straight, with pectinate fringe formed by well-developed fringe tips.
|
||
</paragraph>
|
||
<caption id="2A94663C505FFF9716BDFED2FD300DB1" ID-Table-UUID="2A94663C505FFF9716BDFED2FD300DB1" httpUri="http://table.plazi.org/id/2A94663C505FFF9716BDFED2FD300DB1" pageId="9" pageNumber="10" targetBox="[216,1389,503,948]" targetIsTable="true" targetPageId="9">
|
||
<paragraph id="7E5436B4505FFF9716BDFED2FD300DB1" blockId="9.[189,1399,267,469]" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9716BDFED2FEC20D41" bold="true" box="[189,284,267,294]" pageId="9" pageNumber="10">Table 3.</emphasis>
|
||
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
|
||
<taxonomicName id="B9EB4D37505FFF97174CFEF1FDE80D2C" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[332,566,303,329]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="9" pageNumber="10" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505FFF97174CFEF1FDE80D2C" box="[332,566,303,329]" italics="true" pageId="9" pageNumber="10">Echinoderes anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505FFF97143EFEF1FD470D2D" box="[574,665,303,329]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♂) = putative male condition of sexually dimorphic character.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B4505FFF9714ECFE29FBA40FD0" pageId="9" pageNumber="10">
|
||
<table id="0CEBC414505F006116D8FE29FAB30FD0" box="[216,1389,503,948]" colsContinueIn="17.[204,1386,547,993]" gridcols="11" gridrows="13" pageId="9" pageNumber="10">
|
||
<tr id="C0DB34F6505F006116D8FE29FAB30E6B" box="[216,1389,503,527]" gridrow="0" pageId="9" pageNumber="10">
|
||
<th id="830A5D8A505F006116D8FE29FAB30E6B" box="[216,1389,503,527]" colspan="11" colspanRight="10" gridcol="0" gridrow="0" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9714ECFE29FC960E6B" bold="true" box="[748,840,503,527]" pageId="9" pageNumber="10">Position</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FDFAFAB30E59" box="[216,1389,548,573]" gridrow="1" pageId="9" pageNumber="10">
|
||
<th id="830A5D8A505F006116D8FDFAFEE40E59" box="[216,314,548,573]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9716D8FDFAFEE40E58" bold="true" box="[216,314,548,572]" pageId="9" pageNumber="10">Segment</emphasis>
|
||
</th>
|
||
<td id="830A5D8A505F0061175CFDFAFE680E59" box="[348,438,548,573]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF971773FDFBFE410E59" bold="true" box="[371,415,549,573]" pageId="9" pageNumber="10">MD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F006117CBFDFAFDC50E59" box="[459,539,548,573]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9717E1FDFBFDDB0E59" bold="true" box="[481,517,549,573]" pageId="9" pageNumber="10">PD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F00611443FDFAFDA90E59" box="[579,631,548,573]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF97144CFDFAFDB00E58" bold="true" box="[588,622,548,572]" pageId="9" pageNumber="10">SD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F0061149CFDFAFD070E59" box="[668,729,548,573]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9714B4FDFBFD070E59" bold="true" box="[692,729,549,573]" pageId="9" pageNumber="10">LD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F00611500FDFAFCBE0E59" box="[768,864,548,573]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF97151CFDFBFC990E59" bold="true" box="[796,839,549,573]" pageId="9" pageNumber="10">ML</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F00611580FDFAFC6A0E59" box="[896,948,548,573]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF97158BFDFAFC720E58" bold="true" box="[907,940,548,572]" pageId="9" pageNumber="10">SL</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F006115EAFDFAFBC00E59" box="[1002,1054,548,573]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9715F2FDFAFBC90E58" bold="true" box="[1010,1047,548,572]" pageId="9" pageNumber="10">LA</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F00611254FDFAFB560E59" box="[1108,1160,548,573]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF97125DFDFBFBA30E59" bold="true" box="[1117,1149,549,573]" pageId="9" pageNumber="10">LV</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F006112BEFDFAFB2C0E59" box="[1214,1266,548,573]" gridcol="9" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF9712C6FDFBFB350E59" bold="true" box="[1222,1259,549,573]" pageId="9" pageNumber="10">VL</emphasis>
|
||
</td>
|
||
<td id="830A5D8A505F00611317FDFAFAB30E59" box="[1303,1389,548,573]" gridcol="10" gridrow="1" pageId="9" pageNumber="10">
|
||
<emphasis id="4C9FEAA6505FFF97132BFDFBFA890E59" bold="true" box="[1323,1367,549,573]" pageId="9" pageNumber="10">VM</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FD92FAB30E00" box="[216,1389,588,612]" gridrow="2" pageId="9" pageNumber="10" rowspan-10="1" rowspan-2="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="830A5D8A505F006116D8FD92FEE40E00" box="[216,314,588,612]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">1</th>
|
||
<td id="830A5D8A505F0061175CFD92FE680E00" box="[348,438,588,612]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F00611443FD92FDA90E00" box="[579,631,588,612]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F0061149CFD92FD070E00" box="[668,729,588,612]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611500FD92FCBE0E00" box="[768,864,588,612]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">gco2</td>
|
||
<td id="830A5D8A505F00611254FD92FB560E00" box="[1108,1160,588,612]" gridcol="8" gridrow="2" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F006112BEFD92FB2C0E00" box="[1214,1266,588,612]" gridcol="9" gridrow="2" pageId="9" pageNumber="10">ss</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FDB0FAB30EE2" box="[216,1389,622,646]" gridrow="3" pageId="9" pageNumber="10" rowspan-2="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="830A5D8A505F006116D8FDB0FEE40EE2" box="[216,314,622,646]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">2</th>
|
||
<td id="830A5D8A505F0061175CFDB0FE680EE2" box="[348,438,622,646]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F00611443FDB0FDA90EE2" box="[579,631,622,646]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">gco2</td>
|
||
<td id="830A5D8A505F0061149CFDB0FCBE0EE2" box="[668,864,622,646]" colspan="2" colspanRight="1" gridcol="4" gridrow="3" pageId="9" pageNumber="10">gco2, ss</td>
|
||
<td id="830A5D8A505F00611580FDB0FC6A0EE2" box="[896,948,622,646]" gridcol="6" gridrow="3" pageId="9" pageNumber="10">gco2</td>
|
||
<td id="830A5D8A505F006112BEFDB0FB2C0EE2" box="[1214,1266,622,646]" gridcol="9" gridrow="3" pageId="9" pageNumber="10">gco2</td>
|
||
<td id="830A5D8A505F00611317FDB0FAB30EE2" box="[1303,1389,622,646]" gridcol="10" gridrow="3" pageId="9" pageNumber="10">ss</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FD51FAB30EC3" box="[216,1389,655,679]" gridrow="4" pageId="9" pageNumber="10" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FD51FEE40EC3" box="[216,314,655,679]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">3</th>
|
||
<td id="830A5D8A505F00611443FD51FDA90EC3" box="[579,631,655,679]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611500FD51FCBE0EC3" box="[768,864,655,679]" gridcol="5" gridrow="4" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611317FD51FAB30EC3" box="[1303,1389,655,679]" gridcol="10" gridrow="4" pageId="9" pageNumber="10">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FD6FFAB30EAD" box="[216,1389,689,713]" gridrow="5" pageId="9" pageNumber="10" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FD6FFEE40EAD" box="[216,314,689,713]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">4</th>
|
||
<td id="830A5D8A505F0061175CFD6FFE680EAD" box="[348,438,689,713]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F006117CBFD6FFDC50EAD" box="[459,539,689,713]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F00611317FD6FFAB30EAD" box="[1303,1389,689,713]" gridcol="10" gridrow="5" pageId="9" pageNumber="10">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FD0CFAB30E8F" box="[216,1389,722,747]" gridrow="6" pageId="9" pageNumber="10" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FD0CFEE40E8F" box="[216,314,722,747]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">5</th>
|
||
<td id="830A5D8A505F0061175CFD0CFE680E8F" box="[348,438,722,747]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F00611443FD0CFDA90E8F" box="[579,631,722,747]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611500FD0CFCBE0E8F" box="[768,864,722,747]" gridcol="5" gridrow="6" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F006115EAFD0CFBC00E8F" box="[1002,1054,722,747]" gridcol="7" gridrow="6" pageId="9" pageNumber="10">gco2</td>
|
||
<td id="830A5D8A505F00611317FD0CFAB30E8F" box="[1303,1389,722,747]" gridcol="10" gridrow="6" pageId="9" pageNumber="10">ss, gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FD2AFAB30F68" box="[216,1389,756,780]" gridrow="7" pageId="9" pageNumber="10" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FD2AFEE40F68" box="[216,314,756,780]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">6</th>
|
||
<td id="830A5D8A505F0061175CFD2AFE680F68" box="[348,438,756,780]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F006117CBFD2AFDC50F68" box="[459,539,756,780]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F00611500FD2AFCBE0F68" box="[768,864,756,780]" gridcol="5" gridrow="7" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611254FD2AFB560F68" box="[1108,1160,756,780]" gridcol="8" gridrow="7" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F00611317FD2AFAB30F68" box="[1303,1389,756,780]" gridcol="10" gridrow="7" pageId="9" pageNumber="10">ss, gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FCC8FAB30F4A" box="[216,1389,790,814]" gridrow="8" pageId="9" pageNumber="10" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FCC8FEE40F4A" box="[216,314,790,814]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">7</th>
|
||
<td id="830A5D8A505F0061175CFCC8FE680F4A" box="[348,438,790,814]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F00611443FCC8FDA90F4A" box="[579,631,790,814]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611500FCC8FCBE0F4A" box="[768,864,790,814]" gridcol="5" gridrow="8" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611254FCC8FB560F4A" box="[1108,1160,790,814]" gridcol="8" gridrow="8" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F00611317FCC8FAB30F4A" box="[1303,1389,790,814]" gridcol="10" gridrow="8" pageId="9" pageNumber="10">ss, gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FCE9FAB30F2B" box="[216,1389,823,847]" gridrow="9" pageId="9" pageNumber="10" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FCE9FEE40F2B" box="[216,314,823,847]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">8</th>
|
||
<td id="830A5D8A505F0061175CFCE9FE680F2B" box="[348,438,823,847]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F006117CBFCE9FDC50F2B" box="[459,539,823,847]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">gco1,ss</td>
|
||
<td id="830A5D8A505F00611580FCE9FC6A0F2B" box="[896,948,823,847]" gridcol="6" gridrow="9" pageId="9" pageNumber="10">gco2</td>
|
||
<td id="830A5D8A505F00611254FCE9FB560F2B" box="[1108,1160,823,847]" gridcol="8" gridrow="9" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F00611317FCE9FAB30F2B" box="[1303,1389,823,847]" gridcol="10" gridrow="9" pageId="9" pageNumber="10">ss, gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FC87FAB30F15" box="[216,1389,857,881]" gridrow="10" pageId="9" pageNumber="10" rowspan-1="1" rowspan-5="1" rowspan-7="1">
|
||
<th id="830A5D8A505F006116D8FC87FEE40F15" box="[216,314,857,881]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">9</th>
|
||
<td id="830A5D8A505F006117CBFC87FDC50F15" box="[459,539,857,881]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611443FC87FDA90F15" box="[579,631,857,881]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">gco1</td>
|
||
<td id="830A5D8A505F0061149CFC87FD070F15" box="[668,729,857,881]" gridcol="4" gridrow="10" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611580FC87FC6A0F15" box="[896,948,857,881]" gridcol="6" gridrow="10" pageId="9" pageNumber="10">si</td>
|
||
<td id="830A5D8A505F00611254FC87FB560F15" box="[1108,1160,857,881]" gridcol="8" gridrow="10" pageId="9" pageNumber="10">ac</td>
|
||
<td id="830A5D8A505F006112BEFC87FB2C0F15" box="[1214,1266,857,881]" gridcol="9" gridrow="10" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611317FC87FAB30F15" box="[1303,1389,857,881]" gridcol="10" gridrow="10" pageId="9" pageNumber="10">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FCA5FAB30FF7" box="[216,1389,891,915]" gridrow="11" pageId="9" pageNumber="10" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="830A5D8A505F006116D8FCA5FEE40FF7" box="[216,314,891,915]" gridcol="0" gridrow="11" pageId="9" pageNumber="10">10</th>
|
||
<td id="830A5D8A505F0061175CFCA5FE680FF7" box="[348,438,891,915]" gridcol="1" gridrow="11" pageId="9" pageNumber="10">2 x gco1</td>
|
||
<td id="830A5D8A505F00611443FCA5FDA90FF7" box="[579,631,891,915]" gridcol="3" gridrow="11" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F0061149CFCA5FD070FF7" box="[668,729,891,915]" gridcol="4" gridrow="11" pageId="9" pageNumber="10">tu</td>
|
||
<td id="830A5D8A505F006112BEFCA5FB2C0FF7" box="[1214,1266,891,915]" gridcol="9" gridrow="11" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611317FCA5FAB30FF7" box="[1303,1389,891,915]" gridcol="10" gridrow="11" pageId="9" pageNumber="10">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6505F006116D8FC44FAB30FD0" box="[216,1389,922,948]" gridrow="12" pageId="9" pageNumber="10" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A505F006116D8FC44FEE40FD0" box="[216,314,922,948]" gridcol="0" gridrow="12" pageId="9" pageNumber="10">11</th>
|
||
<td id="830A5D8A505F00611443FC44FDA90FD0" box="[579,631,922,948]" gridcol="3" gridrow="12" pageId="9" pageNumber="10">ss</td>
|
||
<td id="830A5D8A505F00611500FC44FCBE0FD0" box="[768,864,922,948]" gridcol="5" gridrow="12" pageId="9" pageNumber="10">3 x pe(♂)</td>
|
||
<td id="830A5D8A505F00611254FC44FB560FD0" box="[1108,1160,922,948]" gridcol="8" gridrow="12" pageId="9" pageNumber="10">lts</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505FFF9716BDFBD0FBB0080B" blockId="9.[189,1398,1038,1135]" pageId="9" pageNumber="10">
|
||
Segment 4 with acicular spine in middorsal position (
|
||
<figureCitation id="E6D02A31505FFF971525FBD0FC58084C" box="[805,902,1038,1064]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505FFF971591FBD0FC68084C" box="[913,950,1038,1064]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">3B</figureCitation>
|
||
). Sensory spots not observed. Anterior part of segment with glandular cell outlets
|
||
<typeStatus id="A1508816505FFF9714B3FBECFD380828" box="[691,742,1074,1100]" pageId="9" pageNumber="10">type</typeStatus>
|
||
<quantity id="B9139B51505FFF9714EDFBECFCC4082F" box="[749,794,1073,1100]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="10" unit="in" value="1.0">1 in</quantity>
|
||
paradorsal and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505FFF9716BDFB48FCFE097E" blockId="9.[189,1399,1174,1306]" pageId="9" pageNumber="10">
|
||
Segment 5 without spines or tubes. Glandular cell outlets
|
||
<typeStatus id="A1508816505FFF971570FB49FC7D08D5" box="[880,931,1175,1201]" pageId="9" pageNumber="10">type</typeStatus>
|
||
2 present in lateral accessory positions (
|
||
<figureCitation id="E6D02A31505FFF9716C5FB67FEFB08B7" box="[197,293,1209,1235]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505FFF971731FB67FE8808B7" box="[305,342,1209,1235]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">3E</figureCitation>
|
||
). Sensory spots present in subdorsal, midlateral and ventromedial positions, and glandular cell outlets
|
||
<typeStatus id="A1508816505FFF97174AFB03FEA30893" box="[330,381,1245,1271]" pageId="9" pageNumber="10">type</typeStatus>
|
||
<quantity id="B9139B51505FFF971787FB03FE680892" box="[391,438,1244,1271]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="10" unit="in" value="1.0">1 in</quantity>
|
||
middorsal and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505FFF9716BDFA9FFE7C09A1" blockId="9.[189,1399,1345,1477]" pageId="9" pageNumber="10">
|
||
Segment 6 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A31505FFF971241FA9FFB51093F" box="[1089,1167,1345,1371]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Figs 2</figureCitation>
|
||
A–B, 3E). Sensory spots present in midlateral and ventromedial positions, and glandular cell outlets
|
||
<typeStatus id="A1508816505FFF971289FABBFB62091B" box="[1161,1212,1381,1407]" pageId="9" pageNumber="10">type</typeStatus>
|
||
<quantity id="B9139B51505FFF9712C4FABBFB2D091A" box="[1220,1267,1380,1407]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="10" unit="in" value="1.0">1 in</quantity>
|
||
paradorsal and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505FFF9716BDFA32FE790A14" blockId="9.[189,1399,1516,1648]" pageId="9" pageNumber="10">
|
||
Segment 7 with acicular spines in lateroventral positions (
|
||
<figureCitation id="E6D02A31505FFF971594FA32FC240A62" box="[916,1018,1516,1542]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505FFF97120AFA32FBF10A62" box="[1034,1071,1516,1542]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">3E</figureCitation>
|
||
). Sensory spots present in subdorsal, midlateral and ventromedial positions, and glandular cell outlets
|
||
<typeStatus id="A1508816505FFF971256F9CEFB540A4E" box="[1110,1162,1552,1578]" pageId="9" pageNumber="10">type</typeStatus>
|
||
<quantity id="B9139B51505FFF971293F9CEFB1C0A4D" box="[1171,1218,1551,1578]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="10" unit="in" value="1.0">1 in</quantity>
|
||
middorsal and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505FFF9716BDF949FC230B5B" blockId="9.[189,1399,1687,1855]" pageId="9" pageNumber="10">
|
||
Segment 8 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A31505FFF971217F949FB880AD5" box="[1047,1110,1687,1713]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Fig 2</figureCitation>
|
||
A–B, 3D, H). Middorsal spine is long, reaching the posterior margin of segment 10, and slightly thicker than the middorsal spine of segment 4. Glandular cell outlets
|
||
<typeStatus id="A1508816505FFF971461F900FD4A0A9C" box="[609,660,1758,1784]" pageId="9" pageNumber="10">type</typeStatus>
|
||
2 present in sublateral positions (
|
||
<figureCitation id="E6D02A31505FFF971218F900FBB20A9C" box="[1048,1132,1758,1784]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Fig 2B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505FFF971277F900FB1C0A9C" box="[1143,1218,1758,1784]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">3D, H</figureCitation>
|
||
). Sensory spots and glandular cell outlets
|
||
<typeStatus id="A1508816505FFF9717EAF8DFFDC30B7F" box="[490,541,1793,1819]" pageId="9" pageNumber="10">type</typeStatus>
|
||
1 present in paradorsal and ventromedial positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505FFF9416BDF8B8FB960E40" blockId="9.[189,1399,1893,2026]" lastBlockId="10.[189,1399,267,548]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
Segment 9 with acicular spines in lateroventral positions (
|
||
<figureCitation id="E6D02A31505FFF971590F8BBFC280BE4" box="[912,1014,1893,1920]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="9" pageNumber="10">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505FFF971208F8B8FBF10BE4" box="[1032,1071,1894,1920]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">3H</figureCitation>
|
||
). Sensory spots present in paradorsal, laterodorsal (
|
||
<figureCitation id="E6D02A31505FFF9717E7F857FDB70BC7" box="[487,617,1929,1955]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">Fig. 3D, F</figureCitation>
|
||
) and ventrolateral positions, and glandular cell outlets
|
||
<typeStatus id="A1508816505FFF971309F857FAE20BC7" box="[1289,1340,1929,1955]" pageId="9" pageNumber="10">type</typeStatus>
|
||
<quantity id="B9139B51505FFF971346F857FAA90BC7" box="[1350,1399,1929,1955]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="10" unit="in" value="1.0">1 in</quantity>
|
||
subdorsal and ventromedial positions. Small rounded sieve plates are present in sublateral positions (
|
||
<figureCitation id="E6D02A31505FFF9716C6F811FEE30B8E" box="[198,317,1999,2026]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="9" pageNumber="10">Fig. 3F, H</figureCitation>
|
||
). Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment. Segment 10 with laterodorsal tubes at posterior segment margin (
|
||
<figureCitation id="E6D02A31505CFF9415BEFED5FBFE0D41" box="[958,1056,267,293]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="10" pageNumber="11">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505CFF94122DFED5FB8E0D41" box="[1069,1104,267,293]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="10" pageNumber="11">3F</figureCitation>
|
||
); tubes in males are long and well-developed (ca
|
||
<quantity id="B9139B51505CFF9417D2FEEEFDCD0D2E" box="[466,531,304,330]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="9.0">9 µm</quantity>
|
||
); female condition of tubes is unknown. Sensory spots present in subdorsal and ventrolateral positions. Glandular cell outlets
|
||
<typeStatus id="A1508816505CFF94150AFE8AFCE30D0A" box="[778,829,340,366]" pageId="10" pageNumber="11">type</typeStatus>
|
||
1 present as two middorsal ones and one pair in ventromedial positions. Tergal plate only with a few cuticular hairs on the most posterior part; sternal plates with cuticular hairs on the lateral halves, but fewer than on preceding segment. The posterior segment margin of the tergal plate is straight, and only interrupted where the laterodorsal tubes attach, whereas the margins of the sternal ones are concave, extending posteriorly near the midventral junction. Pachycycli and pectinate fringe of posterior margin as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505CFF9416BDFD92FDAC0F5B" blockId="10.[189,1399,587,831]" pageId="10" pageNumber="11">
|
||
Segment 11 with lateral terminal spines, almost equal to trunk in length (
|
||
<figureCitation id="E6D02A31505CFF941235FD95FB4A0E02" box="[1077,1172,587,614]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
|
||
). Males with three pairs of thin and flexible penile spines (
|
||
<figureCitation id="E6D02A31505CFF941488FDAEFD370EEE" box="[648,745,624,650]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="10" pageNumber="11">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31505CFF9414F5FDAEFCC20EEE" box="[757,796,624,650]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="10" pageNumber="11">3G</figureCitation>
|
||
); female condition unknown. Sensory spots present in subdorsal positions near the posterior margin of the segment. The segment is completely devoid of cuticular hairs. Tergal extensions narrow gradually into long, slightly flexible tips (
|
||
<figureCitation id="E6D02A31505CFF9412AAFD66FAD50EB6" box="[1194,1291,696,722]" captionStart="Fig" captionStartId="7.[190,233,1649,1675]" captionTargetBox="[194,1396,291,1609]" captionTargetId="figure@7.[194,1396,291,1609]" captionTargetPageId="7" captionText="Fig. 3. Light micrographs showing overviews and details of male holotype of Echinoderes anniae sp. nov. (NHMD-226483). A. Ventral overview. B. Segments 1 to 4, dorsal view. C. Segments 1 to 4, ventral view. D. Segments 8 to 9, dorsal view. E. Segments 5 to 7, ventral view. F. Segments 9 to 11, dorsal view. G. Segments 10 to 11, ventral view. H. Segments 8 to 9, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; lts = lateral terminal spine; lvgco1 = lateroventral glandular cell outlet type 1; lvs = lateroventral spine; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; mlgco2 = midlateral glandular cell outlet type 2; pdss = paradorsal sensory spot; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817829" httpUri="https://zenodo.org/record/3817829/files/figure.png" pageId="10" pageNumber="11">Fig. 3G</figureCitation>
|
||
). Sternal extensions are short and rounded. Posterior margin of sternal plates with paired tufts of extended fringe tips (
|
||
<figureCitation id="E6D02A31505CFF9416F7FCDFFE8D0F7F" box="[247,339,769,795]" captionStart="Fig" captionStartId="6.[189,232,1754,1780]" captionTargetBox="[213,1376,269,1731]" captionTargetId="figure@6.[211,1377,264,1731]" captionTargetPageId="6" captionText="Fig. 2. Line art illustrations of Echinoderes anniae sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. Segments 10 to 11 in male, with lateral terminal spines drawn in full length, dorsal view.Abbreviations: eft = extended fringe tips; lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldt = laterodorsal tube; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine (*broken); mlgco2 = midlateral glandular cell outlet type 2; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlgco2 = ventrolateral glandular cell outlet type 2." figureDoi="http://doi.org/10.5281/zenodo.3817827" httpUri="https://zenodo.org/record/3817827/files/figure.png" pageId="10" pageNumber="11">Fig. 2B</figureCitation>
|
||
); they resemble ventrolateral setae, but the structures appear to be formed by long, merged fringe tips from the posterior margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F505CFF9416BDFCB6FC8A0B8F" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="7E5436B4505CFF9416BDFCB6FEF10FE6" blockId="10.[189,303,872,898]" box="[189,303,872,898]" pageId="10" pageNumber="11">
|
||
<heading id="251C81D8505CFF9416BDFCB6FEF10FE6" bold="true" box="[189,303,872,898]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="4C9FEAA6505CFF9416BDFCB6FEF10FE6" bold="true" box="[189,303,872,898]" pageId="10" pageNumber="11">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505CFF9416BDFC49FBDA0802" blockId="10.[189,1399,919,1127]" pageId="10" pageNumber="11">
|
||
The new species is very easily recognized by the nature of its cuticular structures in the lateral series of segment 5. The presence of a glandular cell outlet
|
||
<typeStatus id="A1508816505CFF94152DFC62FCBE0FB2" box="[813,864,956,982]" pageId="10" pageNumber="11">type</typeStatus>
|
||
<quantity id="B9139B51505CFF941567FC62FC4A0FB1" box="[871,916,955,982]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="11" unit="in" value="2.0">2 in</quantity>
|
||
lateral accessory positions is rare among congeners and shared with only
|
||
<taxonomicName id="B9EB4D37505CFF941435FC3FFBDF0F9F" authority="(Yamasaki et al. 2018 b)" baseAuthorityName="Yamasaki" baseAuthorityYear="2018" box="[565,1025,992,1019]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="4C9FEAA6505CFF941435FC3FFD3E0F9E" box="[565,736,993,1018]" italics="true" pageId="10" pageNumber="11">E. unispinosus</emphasis>
|
||
(
|
||
<bibRefCitation id="1A7A4B45505CFF9414EFFC3EFC270F9F" author="Yamasaki H. & Neuhaus B. & George K. H." box="[751,1017,992,1019]" pageId="10" pageNumber="11" pagination="541 - 566" refId="ref38773" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 b. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Mediterranean Seamounts and from the deep-sea floor in the Northeast Atlantic Ocean, including notes on two undescribed species. Zootaxa 4378: 541 - 566. https: // doi. org / 10.11646 / zootaxa. 4387.3.8" type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA6505CFF94156AFC3FFC7A0F9E" box="[874,932,992,1018]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2018b
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. A second trait, also exclusively shared with
|
||
<taxonomicName id="B9EB4D37505CFF941752FBDBFDDE087A" box="[338,512,1029,1054]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="4C9FEAA6505CFF941752FBDBFDDE087A" box="[338,512,1029,1054]" italics="true" pageId="10" pageNumber="11">E. unispinosus</emphasis>
|
||
</taxonomicName>
|
||
, is the presence of midlateral glandular cell outlets
|
||
<typeStatus id="A1508816505CFF941278FBDAFB75087A" box="[1144,1195,1028,1054]" pageId="10" pageNumber="11">type</typeStatus>
|
||
2 on segment 1. However,
|
||
<taxonomicName id="B9EB4D37505CFF941737FBF7FE3A0826" box="[311,484,1065,1091]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="4C9FEAA6505CFF941737FBF7FE970826" box="[311,329,1065,1090]" italics="true" pageId="10" pageNumber="11">E</emphasis>
|
||
.
|
||
<emphasis id="4C9FEAA6505CFF941759FBF7FE3A0826" box="[345,484,1065,1090]" italics="true" pageId="10" pageNumber="11">unispinosus</emphasis>
|
||
</taxonomicName>
|
||
has only a single middorsal spine (see below in the present contribution and
|
||
<bibRefCitation id="1A7A4B45505CFF9416BDFB92FE190803" author="Yamasaki H. & Neuhaus B. & George K. H." box="[189,455,1100,1127]" pageId="10" pageNumber="11" pagination="541 - 566" refId="ref38773" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 b. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Mediterranean Seamounts and from the deep-sea floor in the Northeast Atlantic Ocean, including notes on two undescribed species. Zootaxa 4378: 541 - 566. https: // doi. org / 10.11646 / zootaxa. 4387.3.8" type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA6505CFF941737FB93FEAF0802" box="[311,369,1100,1126]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2018b
|
||
</bibRefCitation>
|
||
); hence, the two species are easily distinguished.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4505CFF9416BDFB51FC8A0B8F" blockId="10.[189,1399,1167,2027]" pageId="10" pageNumber="11">
|
||
The lack of lateroventral tubes on segment 5, combined with the presence of lateroventral spines on segments 6 to 9, is rather rare among species of
|
||
<taxonomicName id="B9EB4D37505CFF941508FB6DFC4408A9" authorityName="Claparede" authorityYear="1863" box="[776,922,1203,1229]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA6505CFF941508FB6DFC4408A9" box="[776,922,1203,1229]" italics="true" pageId="10" pageNumber="11">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
, and only shared with
|
||
<taxonomicName id="B9EB4D37505CFF9412B0FB6AFF240895" authority="Higgins, 1966" authorityName="Higgins" authorityYear="1966" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF9412B0FB6AFAD708A9" box="[1200,1289,1203,1229]" italics="true" pageId="10" pageNumber="11">E. arlis</emphasis>
|
||
<bibRefCitation id="1A7A4B45505CFF941312FB6DFF240895" author="Higgins R. P." pageId="10" pageNumber="11" pagination="518 - 520" refId="ref36619" refString="Higgins R. P. 1966. Echinoderes arlis, a new kinorhynch from the Arctic Ocean. Pacific Science 20: 518 - 520." type="journal article" year="1966">Higgins, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9EB4D37505CFF941705FB06FE5E0895" authorityName="Grzelak & Sorensen" authorityYear="2017" box="[261,384,1239,1265]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="drogoni">
|
||
<emphasis id="4C9FEAA6505CFF941705FB06FE5E0895" box="[261,384,1239,1265]" italics="true" pageId="10" pageNumber="11">E. drogoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9EB4D37505CFF94178EFB06FC210896" authority="Adrianov in Adrianov & Malakhov, 1999" authorityName="Adrianov in Adrianov & Malakhov" authorityYear="1999" box="[398,1023,1239,1266]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="koreanus">
|
||
<emphasis id="4C9FEAA6505CFF94178EFB06FDC90895" box="[398,535,1239,1265]" italics="true" pageId="10" pageNumber="11">E. koreanus</emphasis>
|
||
Adrianov in
|
||
<bibRefCitation id="1A7A4B45505CFF9414AEFB09FC210896" author="Adrianov A. V. & Malakhov V. V." box="[686,1023,1239,1266]" pageId="10" pageNumber="11" refId="ref35928" refString="Adrianov A. V. & Malakhov V. V. 1999. Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow." type="book" year="1999">Adrianov & Malakhov, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9EB4D37505CFF94120AFB06FEBD0972" authority="Pardos, Sanchez & Herranz, 2016" authorityName="Pardos, Sanchez & Herranz" authorityYear="2016" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="orestauri">
|
||
<emphasis id="4C9FEAA6505CFF94120AFB06FB4A0895" box="[1034,1172,1240,1265]" italics="true" pageId="10" pageNumber="11">E. orestauri</emphasis>
|
||
Pardos,
|
||
<bibRefCitation id="1A7A4B45505CFF9412F8FB09FEBD0972" author="Sanchez N. & Yamasaki H." pageId="10" pageNumber="11" pagination="80 - 89" refId="ref37607" refString="Sanchez N. & Yamasaki H. 2016. Two new Pycnophyidae species (Kinorhyncha: Allomalorhagida) from Japan lacking ventral tubes in males. Zoologischer Anzeiger 265: 80 - 89. https: // doi. org / 10.1016 / j. jcz. 2016.04.001" type="journal article" year="2016">Sánchez & Herranz, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B9EB4D37505CFF941798FB23FBD60972" authority="Adrianov in Adrianov & Malakhov, 1999" authorityName="Adrianov in Adrianov & Malakhov" authorityYear="1999" box="[408,1032,1276,1302]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="svetlanae">
|
||
<emphasis id="4C9FEAA6505CFF941798FB23FDF80972" box="[408,550,1276,1302]" italics="true" pageId="10" pageNumber="11">E. svetlanae</emphasis>
|
||
Adrianov in
|
||
<bibRefCitation id="1A7A4B45505CFF9414BBFB22FBD60972" author="Adrianov A. V. & Malakhov V. V." box="[699,1032,1276,1302]" pageId="10" pageNumber="11" refId="ref35928" refString="Adrianov A. V. & Malakhov V. V. 1999. Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow." type="book" year="1999">Adrianov & Malakhov, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="1A7A4B45505CFF941242FB22FB3D0972" author="Higgins R. P." box="[1090,1251,1276,1302]" pageId="10" pageNumber="11" pagination="518 - 520" refId="ref36619" refString="Higgins R. P. 1966. Echinoderes arlis, a new kinorhynch from the Arctic Ocean. Pacific Science 20: 518 - 520." type="journal article" year="1966">Higgins 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A7A4B45505CFF9412EBFB22FEA4095E" author="Adrianov A. V. & Malakhov V. V." pageId="10" pageNumber="11" refId="ref35928" refString="Adrianov A. V. & Malakhov V. V. 1999. Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow." type="book" year="1999">Adrianov & Malakhov 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A7A4B45505CFF941786FAFEFDB5095E" author="Pardos F. & Herranz M. & Sanchez N." box="[390,619,1312,1338]" pageId="10" pageNumber="11" pagination="3 - 25" refId="ref37448" refString="Pardos F., Herranz M. & Sanchez N. 2016 a. Two sides of a coin: the phylum Kinorhyncha in Panama. I) Caribbean Panama. Zoologischer Anzeiger 265: 3 - 25. https: // doi. org / 10.1016 / j. jcz. 2016.06.005" type="journal article" year="2016">
|
||
Pardos
|
||
<emphasis id="4C9FEAA6505CFF9417DDFAFFFDC6095E" box="[477,536,1312,1338]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2016a
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1A7A4B45505CFF941477FAFEFC75095E" author="Grzelak K. & Sorensen M. V." box="[631,939,1312,1338]" pageId="10" pageNumber="11" pagination="113 - 147" refId="ref36159" refString="Grzelak K. & Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096 [published online in 2017]." type="journal article" year="2018">Grzelak & Sørensen 2018</bibRefCitation>
|
||
, in press). However,
|
||
<taxonomicName id="B9EB4D37505CFF9412A4FAFFFACB095E" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1188,1301,1313,1338]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF9412A4FAFFFACB095E" box="[1188,1301,1313,1338]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF94131CFAFEFAA9095E" box="[1308,1399,1312,1338]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is easily distinguished from the latter four, since they all have five middorsal spines, opposed to three in
|
||
<taxonomicName id="B9EB4D37505CFF9416DCFAB7FE9009E6" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[220,334,1385,1410]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF9416DCFAB7FE9009E6" box="[220,334,1385,1410]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF941757FAB7FE6D09E7" box="[343,435,1385,1411]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The species shows most resemblance with
|
||
<taxonomicName id="B9EB4D37505CFF9415BAFAB7FBCC09E6" authorityName="Higgins" authorityYear="1966" box="[954,1042,1384,1410]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF9415BAFAB7FBCC09E6" box="[954,1042,1384,1410]" italics="true" pageId="10" pageNumber="11">E. arlis</emphasis>
|
||
</taxonomicName>
|
||
. Both species have middorsal spines on segments 4, 6 and 8, and lateroventral spines on segments 6 to 9.
|
||
<taxonomicName id="B9EB4D37505CFF94123CFA53FAD509C3" authorityName="Higgins" authorityYear="1966" box="[1084,1291,1421,1447]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF94123CFA53FAD509C3" box="[1084,1291,1421,1447]" italics="true" pageId="10" pageNumber="11">Echinoderes arlis</emphasis>
|
||
</taxonomicName>
|
||
does not have lateral accessory glandular cell outlets
|
||
<typeStatus id="A1508816505CFF9414D2FA6FFCDB09AF" box="[722,773,1457,1483]" pageId="10" pageNumber="11">type</typeStatus>
|
||
2 on segment 5, but instead has sublateral tubes on this segment, but the attachment points of these tubes could be confused with gland outlets (
|
||
<bibRefCitation id="1A7A4B45505CFF9412F9FA0BFEB30A70" author="Grzelak K. & Sorensen M. V." pageId="10" pageNumber="11" pagination="113 - 147" refId="ref36159" refString="Grzelak K. & Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096 [published online in 2017]." type="journal article" year="2018">Grzelak & Sørensen 2018</bibRefCitation>
|
||
, in press). It furthermore shows the occasional presence of sublateral tubes on segment 8 (Grzelak & Sørensen in press), and the attachment points of these tubes could also be confused with gland outlets in
|
||
<taxonomicName id="B9EB4D37505CFF941781F99DFE2A0A38" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[385,500,1603,1628]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF941781F99DFE2A0A38" box="[385,500,1603,1628]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF9417FEF99CFD800A38" box="[510,606,1602,1628]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, although the gland outlets in
|
||
<taxonomicName id="B9EB4D37505CFF9415CFF99DFB9C0A38" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[975,1090,1603,1628]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF9415CFF99DFB9C0A38" box="[975,1090,1603,1628]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF94124CF99CFB770A38" box="[1100,1193,1602,1628]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are considerably larger than the tube attachments in
|
||
<taxonomicName id="B9EB4D37505CFF941456F9B9FD720AE4" authorityName="Higgins" authorityYear="1966" box="[598,684,1638,1664]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF941456F9B9FD720AE4" box="[598,684,1638,1664]" italics="true" pageId="10" pageNumber="11">E. arlis</emphasis>
|
||
</taxonomicName>
|
||
. Also the nearly parallel-sided trunks and the relatively welldeveloped pachycycli give the two species a fairly similar appearance. The main differences between the two species appear to be linked to the possession of either tubes or glandular cell outlets
|
||
<typeStatus id="A1508816505CFF941305F971FAE60AAD" box="[1285,1336,1711,1737]" pageId="10" pageNumber="11">type</typeStatus>
|
||
2. In addition to the differences on segments 5 and 8, the two species can be distinguished in a similar way on segment 2. Both species have glandular cell outlets
|
||
<typeStatus id="A1508816505CFF941515F926FC960B76" box="[789,840,1784,1810]" pageId="10" pageNumber="11">type</typeStatus>
|
||
<quantity id="B9139B51505CFF94154EF929FCA40B75" box="[846,890,1783,1809]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="11" unit="in" value="2.0">2 in</quantity>
|
||
subdorsal and sublateral positions, but they differ in the laterodorsal positions where only
|
||
<taxonomicName id="B9EB4D37505CFF9414E4F8C2FC880B51" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[740,854,1820,1845]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF9414E4F8C2FC880B51" box="[740,854,1820,1845]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF94155EF8C2FC640B52" box="[862,954,1820,1846]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has glandular cell outlets
|
||
<typeStatus id="A1508816505CFF9412F3F8C2FAF80B52" box="[1267,1318,1820,1846]" pageId="10" pageNumber="11">type</typeStatus>
|
||
2, and in the ventrolateral positions where
|
||
<taxonomicName id="B9EB4D37505CFF94145FF89FFD6B0B3E" authorityName="Higgins" authorityYear="1966" box="[607,693,1856,1882]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF94145FF89FFD6B0B3E" box="[607,693,1856,1882]" italics="true" pageId="10" pageNumber="11">E. arlis</emphasis>
|
||
</taxonomicName>
|
||
has tubes instead of glandular cell outlets
|
||
<typeStatus id="A1508816505CFF9412A3F89EFB080B3E" box="[1187,1238,1856,1882]" pageId="10" pageNumber="11">type</typeStatus>
|
||
2 (
|
||
<bibRefCitation id="1A7A4B45505CFF9412FAF89EFEB00B1A" author="Grzelak K. & Sorensen M. V." pageId="10" pageNumber="11" pagination="113 - 147" refId="ref36159" refString="Grzelak K. & Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096 [published online in 2017]." type="journal article" year="2018">Grzelak & Sørensen 2018</bibRefCitation>
|
||
, in press). Furthermore,
|
||
<taxonomicName id="B9EB4D37505CFF94148DF8BBFD200B1A" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[653,766,1893,1918]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF94148DF8BBFD200B1A" box="[653,766,1893,1918]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF941505F8BAFC810B1A" box="[773,863,1892,1918]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has glandular cell outlets
|
||
<typeStatus id="A1508816505CFF941292F8BAFB1B0B1A" box="[1170,1221,1892,1918]" pageId="10" pageNumber="11">type</typeStatus>
|
||
<quantity id="B9139B51505CFF9412CCF8BAFB260B1A" box="[1228,1272,1892,1918]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="11" unit="in" value="2.0">2 in</quantity>
|
||
midlateral positions on segment 1, which are certainly not present in
|
||
<taxonomicName id="B9EB4D37505CFF94155DF857FC6C0BC6" authorityName="Higgins" authorityYear="1966" box="[861,946,1928,1954]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF94155DF857FC6C0BC6" box="[861,946,1928,1954]" italics="true" pageId="10" pageNumber="11">E. arlis</emphasis>
|
||
</taxonomicName>
|
||
. Hence, it is possible to distinguish the two species in different ways, but the several similarities suggest a close relationship between
|
||
<taxonomicName id="B9EB4D37505CFF941307F873FAA90BA2" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1287,1399,1965,1990]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6505CFF941307F873FAA90BA2" box="[1287,1399,1965,1990]" italics="true" pageId="10" pageNumber="11">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD505CFF9416BDF80FFEC60B8F" box="[189,280,2001,2027]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="B9EB4D37505CFF941750F80CFE790B8F" authorityName="Higgins" authorityYear="1966" box="[336,423,2001,2027]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="arlis">
|
||
<emphasis id="4C9FEAA6505CFF941750F80CFE790B8F" box="[336,423,2001,2027]" italics="true" pageId="10" pageNumber="11">E. arlis</emphasis>
|
||
</taxonomicName>
|
||
, and potentially also
|
||
<taxonomicName id="B9EB4D37505CFF9414A2F80CFC900B8F" box="[674,846,2002,2027]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="10" pageNumber="11" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="4C9FEAA6505CFF9414A2F80CFC900B8F" box="[674,846,2002,2027]" italics="true" pageId="10" pageNumber="11">E. unispinosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |