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<document id="79AF2C0524450BF78413049DDA7D020A" ID-CLB-Dataset="118433" ID-DOI="10.35929/RSZ.0021" ID-GBIF-Dataset="0e9cc441-e337-4aca-bf79-d6335e798e0e" ID-ISSN="0035-418" ID-Zenodo-Dep="5743710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638276185617" checkinUser="carolina" docAuthor="Logunov, Dmitri V. &amp; Ponomarev, Alexander V." docDate="2020" docId="ED7D8797FF963F40FC7423936C9B8330" docLanguage="en" docName="RevueSuisseZool.127.2.275-313.pdf" docOrigin="Revue suisse de Zoologie 127 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Karakumosa Logunov &amp; Ponomarev 2020, gen. nov." docType="treatment" docVersion="8" lastPageNumber="280" masterDocId="1144FFEFFF943F46FFBE250E69008129" masterDocTitle="Karakumosa gen. nov., a new Central Asian genus of fossorial wolf spiders (Araneae: Lycosidae: Lycosinae)" masterLastPageNumber="313" masterPageNumber="275" pageNumber="276" updateTime="1699471511260" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0088EDE0D91941979D27FBAD83DA8E68">Karakumosa gen. nov., a new Central Asian genus of fossorial wolf spiders (Araneae: Lycosidae: Lycosinae)</mods:title>
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<mods:name id="9206D53EE5BF7F365D229451CDF6748B" type="personal">
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<mods:roleTerm id="990374833EE97C3487A8BDFB72B044D2">Author</mods:roleTerm>
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<mods:namePart id="ECCDDC939112E6E80AB8B66C016C4E15">Logunov, Dmitri V.</mods:namePart>
<mods:affiliation id="9069ACE0C47FAC4A06240A54C33FC579">The Manchester Museum, The University of Manchester, Oxford Road, Manchester M 13 9 PL, UK.</mods:affiliation>
</mods:name>
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<mods:namePart id="DAFD471CA736EF006A31FB90D80A3A67">Ponomarev, Alexander V.</mods:namePart>
<mods:affiliation id="EA2AC82EAEFBD680F9A2E05ED275DADF">Southern Scientific Centre of the Russian Academy of Sciences, Chekhov street 41, Rostov-on-Don, 344006, Russia. E-mail: ponomarev 1952 @ mail. ru</mods:affiliation>
</mods:name>
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<mods:title id="7588D17BE2FEDAF10C4BE365F4DEFE2A">Revue suisse de Zoologie</mods:title>
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<mods:date id="E6E58CC172F54C052F9095C6053C06FC">2020</mods:date>
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<mods:number id="93D405752CAAD24AE926C58C497A0E29">127</mods:number>
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<mods:classification id="F6C48A6D95EC8690588AD6E027AFC156">journal article</mods:classification>
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<mods:identifier id="A79583A461ECA3C13B067E190445326A" type="DOI">10.35929/RSZ.0021</mods:identifier>
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<treatment id="ED7D8797FF963F40FC7423936C9B8330" ID-DOI="http://doi.org/10.5281/zenodo.6983812" ID-GBIF-Taxon="198975222" ID-Zenodo-Dep="6983812" LSID="urn:lsid:plazi:treatment:ED7D8797FF963F40FC7423936C9B8330" httpUri="http://treatment.plazi.org/id/ED7D8797FF963F40FC7423936C9B8330" lastPageId="6" lastPageNumber="280" pageId="2" pageNumber="276">
<subSubSection id="2DCE650AFF963F44FC7423936C12879D" box="[970,1298,1692,1716]" pageId="2" pageNumber="276" type="nomenclature">
<paragraph id="656B3681FF963F44FC7423936C12879D" blockId="2.[970,1298,1692,1716]" box="[970,1298,1692,1716]" pageId="2" pageNumber="276">
<heading id="3E2381EDFF963F44FC7423936C12879D" bold="true" box="[970,1298,1692,1716]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="276" reason="2">
<emphasis id="57A0EA93FF963F44FC7423936C12879D" bold="true" box="[970,1298,1692,1716]" pageId="2" pageNumber="276">
Genus
<taxonomicName id="A2D44D02FF963F44FBA423926DA7879D" ID-CoL="98X5T" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1050,1191,1692,1716]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="276" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF963F44FBA423926DA7879D" bold="true" box="[1050,1191,1692,1716]" italics="true" pageId="2" pageNumber="276">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF963F44FB1323936C12879D" box="[1197,1298,1693,1716]" pageId="2" pageNumber="276" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF963F44FC8123D56C6D8618" pageId="2" pageNumber="276" type="type_taxon">
<paragraph id="656B3681FF963F44FC8123D56C6D8618" blockId="2.[831,1436,1755,1841]" pageId="2" pageNumber="276">
<emphasis id="57A0EA93FF963F44FC8123D56ADC87DB" bold="true" box="[831,988,1755,1778]" pageId="2" pageNumber="276">Type species:</emphasis>
<materialsCitation id="D5BC3CDCFF963F44FC5623D56C6A8618" ID-GBIF-Occurrence="3886766303" collectionCode="ISEA" country="Turkmenistan" county="Central Asia" location="Repetek Reserve" municipality="Karakumy Desert" pageId="2" pageNumber="276" specimenCount="1" typeStatus="holotype">
<taxonomicName id="A2D44D02FF963F44FC5623D56DCB87DB" ID-CoL="999W7" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1000,1227,1755,1778]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="276" phylum="Arthropoda" rank="species" species="repetek" status="sp. nov.">
<emphasis id="57A0EA93FF963F44FC5623D56DCB87DB" box="[1000,1227,1755,1778]" italics="true" pageId="2" pageNumber="276">
<originalDetermination id="46CA8E7DFF963F44FC5623D56DCB87DB" box="[1000,1227,1755,1778]" pageId="2" pageNumber="276">Karakumosa repetek</originalDetermination>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF963F44FB6923D56C3087DB" box="[1239,1328,1755,1778]" pageId="2" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
from the
<location id="600B605AFF963F44FC8123F46D058638" LSID="urn:lsid:plazi:treatment:ED7D8797FF963F40FC7423936C9B8330:600B605AFF963F44FC8123F46D058638" box="[831,1029,1786,1809]" country="Turkmenistan" county="Central Asia" municipality="Karakumy Desert" name="Repetek Reserve" pageId="2" pageNumber="276">Repetek Reserve</location>
,
<location id="600B605AFF963F44FBA523F46DEA8638" LSID="urn:lsid:plazi:treatment:ED7D8797FF963F40FC7423936C9B8330:600B605AFF963F44FBA523F46DEA8638" box="[1051,1258,1786,1809]" country="Turkmenistan" county="Central Asia" municipality="Karakumy Desert" name="Karakumy Desert" pageId="2" pageNumber="276">Karakumy Desert</location>
,
<collectingCountry id="1DC37611FF963F44FABE23F46C988638" box="[1280,1432,1786,1809]" name="Turkmenistan" pageId="2" pageNumber="276">Turkmenistan</collectingCountry>
,
<collectingCounty id="8C0A4E0DFF963F44FC8122146AC48618" box="[831,964,1818,1841]" pageId="2" pageNumber="276">Central Asia</collectingCounty>
(
<specimenCount id="73D2FD08FF963F44FC6C22146D078618" box="[978,1031,1818,1841]" pageId="2" pageNumber="276" type="male">male</specimenCount>
<typeStatus id="BA6F8823FF963F44FBB022146D6B8618" box="[1038,1131,1818,1841]" pageId="2" pageNumber="276" type="holotype">holotype</typeStatus>
deposited in the
<collectionCode id="03C5AE44FF963F44FA9C22146C628618" box="[1314,1378,1818,1841]" collectionName="Poland, Krakow, Polish Academy of Sciences, Institute of Systematic Zoology" pageId="2" pageNumber="276">ISEA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF963F45FC8122466B66837D" lastPageId="3" lastPageNumber="277" pageId="2" pageNumber="276" type="diagnosis">
<paragraph id="656B3681FF963F45FC8122466B66837D" blockId="2.[831,1437,1864,2077]" lastBlockId="3.[151,756,258,596]" lastPageId="3" lastPageNumber="277" pageId="2" pageNumber="276">
<emphasis id="57A0EA93FF963F44FC8122466AB78676" bold="true" box="[831,951,1864,1887]" pageId="2" pageNumber="276">Diagnosis:</emphasis>
The genus
<taxonomicName id="A2D44D02FF963F44FBFC22466DCB8676" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1090,1227,1864,1887]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="276" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF963F44FBFC22466DCB8676" box="[1090,1227,1864,1887]" italics="true" pageId="2" pageNumber="276">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF963F44FB6922466C3E8676" box="[1239,1342,1864,1887]" pageId="2" pageNumber="276" rank="genus">gen. nov.</taxonomicNameLabel>
belongs to the subfamily
<taxonomicName id="A2D44D02FF963F44FBA322666D8B8656" authorityName="Sundevall" authorityYear="1833" box="[1053,1163,1896,1919]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="276" phylum="Arthropoda" rank="subFamily" subFamily="Lycosinae">Lycosinae</taxonomicName>
(
<emphasis id="57A0EA93FF963F44FB1622666DE48656" box="[1192,1252,1896,1919]" italics="true" pageId="2" pageNumber="276">sensu</emphasis>
<bibRefCitation id="01454B70FF963F44FB4922666C988656" author="Zyuzin A. A." box="[1271,1432,1896,1919]" pageId="2" pageNumber="276" pagination="693 - 700" refId="ref24274" refString="Zyuzin A. A. 1993. Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and new species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum 33 (2): 693 - 700." type="journal article" year="1993">Zyuzin, 1993</bibRefCitation>
; see
<bibRefCitation id="01454B70FF963F44FCCE22896D1A86B7" author="Dondale C. D." box="[880,1050,1927,1950]" pageId="2" pageNumber="276" pagination="327 - 332" refId="ref21895" refString="Dondale C. D. 1986. The subfamilies of wolf spiders (Araneae: Lycosidae) [pp. 327 - 332]. In: Barrientos J. A. (ed.), Actas X. Congreso Internacional de Aracnologia, Jaca, Espana, Volume I. Juvenil, Barcelona, 73 pp." type="book chapter" year="1986">Dondale, 1986</bibRefCitation>
;
<bibRefCitation id="01454B70FF963F44FB9322896C1D86B7" author="Murphy N. P. &amp; Framenau V. W. &amp; Donnellan S. C. &amp; Harvey M. S. &amp; Park Y. C. &amp; Austin A. D." box="[1069,1309,1927,1950]" pageId="2" pageNumber="276" pagination="583 - 602" refId="ref22853" refString="Murphy N. P., Framenau V. W., Donnellan S. C., Harvey M. S., Park Y. C., Austin A. D. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution 28: 583 - 602." type="journal article" year="2006">
Murphy
<emphasis id="57A0EA93FF963F44FB2C22896DD086B7" box="[1170,1232,1927,1950]" italics="true" pageId="2" pageNumber="276">et al.</emphasis>
, 2006
</bibRefCitation>
;
<bibRefCitation id="01454B70FF963F44FA8E22896D098697" author="Piacentini L. N. &amp; Ramirez M. J." pageId="2" pageNumber="276" pagination="227 - 240" refId="ref23158" refString="Piacentini L. N., Ramirez M. J. 2019. Hunting the wolf: a molecular phylogeny of the wolf spiders (Araneae, Lycosidae). Molecular Phylogenetics and Evolution 136: 227 - 240." type="journal article" year="2019">Piacentini &amp; Ramírez, 2019</bibRefCitation>
) and is most similar to
<taxonomicName id="A2D44D02FF963F44FA8E22A96AF086F4" authority="Logunov, 2010" authorityName="Logunov" authorityYear="2010" class="Arachnida" family="Lycosidae" genus="Zyuzicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF963F44FA8E22A96C9C8697" box="[1328,1436,1959,1982]" italics="true" pageId="2" pageNumber="276">Zyuzicosa</emphasis>
<bibRefCitation id="01454B70FF963F44FC8122C86AF086F4" author="Logunov D. V." box="[831,1008,1990,2013]" pageId="2" pageNumber="276" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov, 2010</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="01454B70FF963F44FBFD22C86DF686F4" author="Logunov D. V." box="[1091,1270,1990,2013]" pageId="2" pageNumber="276" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov, 2010</bibRefCitation>
,
<bibRefCitation id="01454B70FF963F44FAB022C86C4986F4" author="Logunov D. V." box="[1294,1353,1990,2013]" pageId="2" pageNumber="276" pagination="349 - 362" refId="ref22421" refString="Logunov D. V. 2012. A synopsis of the genus Zyuzicosa Logunov, 2010 (Aranei: Lycosidae). Arthropoda Selecta 21 (4): 349 - 362." type="journal article" year="2012">2012</bibRefCitation>
). Both genera have a bipartite/biramous synembolus, the same SER/AER ratio (more than 1.3), tarsi of all legs with scopulae and spinules, and a prolatero-apical origin of the embolus. The new genus differs from all other Central Asian
<taxonomicName id="A2D44D02FF973F45FE3A244F68F18071" authorityName="Sundevall" authorityYear="1833" box="[388,497,321,344]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="276" phylum="Arthropoda" rank="subFamily" subFamily="Lycosinae">Lycosinae</taxonomicName>
genera (see
<bibRefCitation id="01454B70FF973F45FD30244F69D1805E" author="Logunov D. V." pageId="3" pageNumber="277" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov, 2010</bibRefCitation>
: table 2), including
<taxonomicName id="A2D44D02FF973F45FE04246E6B25805E" authorityName="Logunov" authorityYear="2010" box="[442,549,352,375]" class="Arachnida" family="Lycosidae" genus="Zyuzicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF973F45FE04246E6B25805E" box="[442,549,352,375]" italics="true" pageId="3" pageNumber="277">Zyuzicosa</emphasis>
</taxonomicName>
, by the following combination of characters: black ventral colour pattern of abdomen absent (
<figureCitation id="FDEF2A04FF973F45FE35249168E8809F" box="[395,488,415,438]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 25</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE4224916B19809F" box="[508,537,415,438]" captionStart="Figs 49-54" captionStartId="14.[151,192,1993,2014]" captionTargetBox="[155,1435,746,1948]" captionTargetId="figure-217@14.[151,1436,737,1955]" captionTargetPageId="14" captionText="Figs 49-54. Karakumosa gromovi sp. nov.; male paratype (49-50) and female paratype (51-54). (49, 52) Body, ventral view. (50-51) Ditto, dorsal view. (53) Female prosoma, frontal view. (54) Female body, lateral view. Scale bars 1 cm (49-52, 54), 0.5 cm (53)." figureDoi="http://doi.org/10.5281/zenodo.5743732" httpUri="https://zenodo.org/record/5743732/files/figure.png" pageId="3" pageNumber="277">52</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FD9324916B4A809F" box="[557,586,415,438]" captionStart="Figs 89-92" captionStartId="21.[151,192,970,991]" captionTargetBox="[153,1429,259,929]" captionTargetId="figure-469@21.[151,1436,257,930]" captionTargetPageId="21" captionText="Figs 89-92. Karakumosa repetek sp. nov.; male paratype (89-90) and female paratype (91-92). (89, 92). Body, dorsal view. (90, 91) Ditto, ventral view. Scale bars 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5743748" httpUri="https://zenodo.org/record/5743748/files/figure.png" pageId="3" pageNumber="277">91</figureCitation>
, etc.); median apophysis consisting of two flat plates (
<figureCitation id="FDEF2A04FF973F45FDE324B16BB580FF" box="[605,693,447,470]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">Figs 55</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FD7B24B16BF080FF" box="[709,752,447,470]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">118</figureCitation>
: OP, IP); synembolus with two acutely pointed lamellae (
<figureCitation id="FDEF2A04FF973F45FF2124F069F3833C" box="[159,243,510,533]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">Figs 57</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FF4024F0683F833C" box="[254,319,510,533]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">98-99</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FEF524F068A9833C" box="[331,425,510,533]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">116, 118</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE0B24F068DF833C" box="[437,479,510,533]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="3" pageNumber="277">129</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE5224F06B16833C" box="[492,534,510,533]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="3" pageNumber="277">149</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FD9C24F06B4C833C" box="[546,588,510,533]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="3" pageNumber="277">163</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FDE624F06B83833C" box="[600,643,510,533]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="3" pageNumber="277">175</figureCitation>
); epigynal atrium at least twice longer than wide; and septal pedicel absent (
<figureCitation id="FDEF2A04FF973F45FE8327336890837D" box="[317,400,573,596]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 27</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE25273368B8837D" box="[411,440,573,596]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="3" pageNumber="277">44</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE7D273368E0837D" box="[451,480,573,596]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">61</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE5527336B08837D" box="[491,520,573,596]" captionStart="Figs 70-75" captionStartId="17.[151,192,1222,1243]" captionTargetBox="[160,1436,257,1181]" captionTargetId="figure-337@17.[151,1439,257,1191]" captionTargetPageId="17" captionText="Figs 70-75. Karakumosa gromovi sp. nov.; male paratypes (74-75) and female paratype (70-73). (70) Body, dorsal view. (71) Ditto, ventral view. (72) Epigyne, ventral view. (73) Vulva, dorsal view. (74-75) Median tooth of MA, posterior view. Scale bars 1 cm (70-71), 0.5 mm (72), 0.25 mm (73), 0.1 mm (74-75)." figureDoi="http://doi.org/10.5281/zenodo.5743738" httpUri="https://zenodo.org/record/5743738/files/figure.png" pageId="3" pageNumber="277">72</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FDAD27336B30837D" box="[531,560,573,596]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="3" pageNumber="277">85</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FD8527336B59837D" box="[571,601,573,596]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">96</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF973F45FF2927626BAE8276" pageId="3" pageNumber="277" type="etymology">
<paragraph id="656B3681FF973F45FF2927626BAE8276" blockId="3.[151,756,620,863]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF292762681D83AA" bold="true" box="[151,285,620,643]" pageId="3" pageNumber="277">Etymology:</emphasis>
The new generic name is composed of two parts: ‘Karakum’, referring to the regional name for the Karakumy Desert where the
<typeStatus id="BA6F8823FF973F45FD8727A56B6783EB" box="[569,615,683,706]" pageId="3" pageNumber="277">type</typeStatus>
species and some other species were discovered, combined with the ending of the generic name
<emphasis id="57A0EA93FF973F45FE7D27E46B0E8228" box="[451,526,746,769]" italics="true" pageId="3" pageNumber="277">Lycosa</emphasis>
(meaning ‘tear like a wolf’; see
<bibRefCitation id="01454B70FF973F45FEB1260768F18209" author="Cameron H. D." box="[271,497,777,800]" pageId="3" pageNumber="277" pagination="274 - 330" refId="ref21792" refString="Cameron H. D. 2005. An etymological dictionary of North American spider genus names [pp. 274 - 330]. In: Ubick D., Paquin P., Cushing P. E., Roth V. (eds), Spiders of North America: an identification manual. American Arachnological Society, Keene, NH, 377 pp." type="book chapter" year="2005">Cameron, 2005: 303</bibRefCitation>
), to which the majority of large burrowing Holarctic wolf spiders are currently assigned. The generic name is feminine in gender.
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF973F45FF2926796C018701" pageId="3" pageNumber="277" type="description">
<paragraph id="656B3681FF973F45FF29267968C282E4" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF292679682482A7" bold="true" box="[151,292,887,910]" pageId="3" pageNumber="277">Description:</emphasis>
Large to very large fossorial wolf spiders, with body lengths 20.23±3.23 (n=8) in males, and 22.9±4.05 (n=7) in females.
</paragraph>
<paragraph id="656B3681FF973F45FF2926DB69DD8583" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF2926DB680082C5" box="[151,256,981,1004]" italics="true" pageId="3" pageNumber="277">Carapace</emphasis>
: In both sexes relatively low, with a clearly marked gradual descent from cephalic region towards abdomen (
<figureCitation id="FDEF2A04FF973F45FEB9211B68598505" box="[263,345,1045,1068]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 26</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FEDA211B68818505" box="[356,385,1045,1068]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="3" pageNumber="277">35</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE32211B68A98505" box="[396,425,1045,1068]" captionStart="Figs 49-54" captionStartId="14.[151,192,1993,2014]" captionTargetBox="[155,1435,746,1948]" captionTargetId="figure-217@14.[151,1436,737,1955]" captionTargetPageId="14" captionText="Figs 49-54. Karakumosa gromovi sp. nov.; male paratype (49-50) and female paratype (51-54). (49, 52) Body, ventral view. (50-51) Ditto, dorsal view. (53) Female prosoma, frontal view. (54) Female body, lateral view. Scale bars 1 cm (49-52, 54), 0.5 cm (53)." figureDoi="http://doi.org/10.5281/zenodo.5743732" httpUri="https://zenodo.org/record/5743732/files/figure.png" pageId="3" pageNumber="277">54</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE0D211B68D08505" box="[435,464,1045,1068]" captionStart="Figs 77-82" captionStartId="19.[151,192,1433,1454]" captionTargetBox="[151,1424,262,1385]" captionTargetId="figure-213@19.[151,1436,257,1409]" captionTargetPageId="19" captionText="Figs 77-82. Karakumosa medica (Pocock, 1889); female paralectotype (77-80) and male lectotype (81-82). (77, 81) Body, dorsal view. (78) Ditto, lateral view. (79) Prosoma, frontal view. (80, 82) Body, ventral view. Scale bars 1 cm (77-78, 80-82), 0.5 cm (79)." figureDoi="http://doi.org/10.5281/zenodo.5743744" httpUri="https://zenodo.org/record/5743744/files/figure.png" pageId="3" pageNumber="277">78</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE65211B6B058505" box="[475,517,1045,1068]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">138</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FDAE211B6B3B8505" box="[528,571,1045,1068]" captionStart="Figs 156-160" captionStartId="31.[151,192,1208,1229]" captionTargetBox="[159,1419,260,1158]" captionTargetId="figure-325@31.[151,1436,257,1163]" captionTargetPageId="31" captionText="Figs 156-160. Karakumosa turanica sp. nov., male holotype (156-158) and female paratype (159-160). (156, 160) Body, dorsal view. (157, 159) Ditto, ventral view. (158) Ditto, lateral view. Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5743766" httpUri="https://zenodo.org/record/5743766/files/figure.png" pageId="3" pageNumber="277">158</figureCitation>
), densely clothed with white or yellowish white setae, and with prominent dense white or yellowish marginal pubescence; all three characters are typical of lycosid burrowers (see
<bibRefCitation id="01454B70FF973F45FD1C217D69D18583" author="Zyuzin A. A." pageId="3" pageNumber="277" pagination="419 - 422" refId="ref24236" refString="Zyuzin A. A. 1990. Studies on burrowing spiders of the family Lycosidae (Araneae). I. Preliminary data on structural and functional features. Acta Zoologica Fennica 190: 419 - 422." type="journal article" year="1990">Zyuzin, 1990</bibRefCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FF2921BC6842846E" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF2921BC680C85E0" box="[151,268,1202,1225]" italics="true" pageId="3" pageNumber="277">Chelicerae</emphasis>
: Large, vertical, their proximal halves/twothirds of frontal side densely clothed with white or yellowish setae (
<figureCitation id="FDEF2A04FF973F45FEF421FF689E8421" box="[330,414,1265,1288]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="3" pageNumber="277">Figs 32</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE1421FF68C78421" box="[426,455,1265,1288]" captionStart="Figs 49-54" captionStartId="14.[151,192,1993,2014]" captionTargetBox="[155,1435,746,1948]" captionTargetId="figure-217@14.[151,1436,737,1955]" captionTargetPageId="14" captionText="Figs 49-54. Karakumosa gromovi sp. nov.; male paratype (49-50) and female paratype (51-54). (49, 52) Body, ventral view. (50-51) Ditto, dorsal view. (53) Female prosoma, frontal view. (54) Female body, lateral view. Scale bars 1 cm (49-52, 54), 0.5 cm (53)." figureDoi="http://doi.org/10.5281/zenodo.5743732" httpUri="https://zenodo.org/record/5743732/files/figure.png" pageId="3" pageNumber="277">53</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE6D21FF68F08421" box="[467,496,1265,1288]" captionStart="Figs 77-82" captionStartId="19.[151,192,1433,1454]" captionTargetBox="[151,1424,262,1385]" captionTargetId="figure-213@19.[151,1436,257,1409]" captionTargetPageId="19" captionText="Figs 77-82. Karakumosa medica (Pocock, 1889); female paralectotype (77-80) and male lectotype (81-82). (77, 81) Body, dorsal view. (78) Ditto, lateral view. (79) Prosoma, frontal view. (80, 82) Body, ventral view. Scale bars 1 cm (77-78, 80-82), 0.5 cm (79)." figureDoi="http://doi.org/10.5281/zenodo.5743744" httpUri="https://zenodo.org/record/5743744/files/figure.png" pageId="3" pageNumber="277">79</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE4521FF6B278421" box="[507,551,1265,1288]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">136</figureCitation>
); cheliceral groove with three promarginal and three retromarginal teeth (
<figureCitation id="FDEF2A04FF973F45FF21203E69FF846E" box="[159,255,1328,1351]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="3" pageNumber="277">Figs 144</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FEB4203E6836846E" box="[266,310,1328,1351]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="3" pageNumber="277">177</figureCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FF29205E69EA848F" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF29205E69C9844E" box="[151,201,1360,1383]" italics="true" pageId="3" pageNumber="277">Eyes</emphasis>
: AER procurved and distinctly (1.3-1.4 times) shorter than SER (
<figureCitation id="FDEF2A04FF973F45FED6206168C084AF" box="[360,448,1391,1414]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="3" pageNumber="277">Figs 32</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE71206168EC84AF" box="[463,492,1391,1414]" captionStart="Figs 49-54" captionStartId="14.[151,192,1993,2014]" captionTargetBox="[155,1435,746,1948]" captionTargetId="figure-217@14.[151,1436,737,1955]" captionTargetPageId="14" captionText="Figs 49-54. Karakumosa gromovi sp. nov.; male paratype (49-50) and female paratype (51-54). (49, 52) Body, ventral view. (50-51) Ditto, dorsal view. (53) Female prosoma, frontal view. (54) Female body, lateral view. Scale bars 1 cm (49-52, 54), 0.5 cm (53)." figureDoi="http://doi.org/10.5281/zenodo.5743732" httpUri="https://zenodo.org/record/5743732/files/figure.png" pageId="3" pageNumber="277">53</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE4520616B2784AF" box="[507,551,1391,1414]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">136</figureCitation>
); PME/AME ratio 1.7-2.5.
</paragraph>
<paragraph id="656B3681FF973F45FF2920A0687084CC" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF2920A069ED84EC" box="[151,237,1454,1477]" italics="true" pageId="3" pageNumber="277">Clypeus</emphasis>
: Narrow, its height equal or 1.3-1.7 times shorter than AME diameter.
</paragraph>
<paragraph id="656B3681FF973F45FF2920E369C9870D" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF2920E369E8872D" box="[151,232,1517,1540]" italics="true" pageId="3" pageNumber="277">Labium</emphasis>
: Visibly wider than long (length/width ratio 0.6- 0.8).
</paragraph>
<paragraph id="656B3681FF973F45FF2923226B28874A" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF29232269F0876A" box="[151,240,1580,1603]" italics="true" pageId="3" pageNumber="277">Sternum</emphasis>
: Ovoid, densely covered with white setae in both sexes (e.g.
<figureCitation id="FDEF2A04FF973F45FEFA23426895874A" box="[324,405,1612,1635]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 23</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE1E234268E5874A" box="[416,485,1612,1635]" captionStart="Figs 49-54" captionStartId="14.[151,192,1993,2014]" captionTargetBox="[155,1435,746,1948]" captionTargetId="figure-217@14.[151,1436,737,1955]" captionTargetPageId="14" captionText="Figs 49-54. Karakumosa gromovi sp. nov.; male paratype (49-50) and female paratype (51-54). (49, 52) Body, ventral view. (50-51) Ditto, dorsal view. (53) Female prosoma, frontal view. (54) Female body, lateral view. Scale bars 1 cm (49-52, 54), 0.5 cm (53)." figureDoi="http://doi.org/10.5281/zenodo.5743732" httpUri="https://zenodo.org/record/5743732/files/figure.png" pageId="3" pageNumber="277">49, 52</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE4E23426B1C874A" box="[496,540,1612,1635]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">137</figureCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FF2923656B5387C8" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF29236569FC87AB" box="[151,252,1643,1666]" italics="true" pageId="3" pageNumber="277">Abdomen</emphasis>
: Venter in both sexes without black pattern (e.g.
<figureCitation id="FDEF2A04FF973F45FF7123856822878B" box="[207,290,1675,1698]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 25</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FE912385684C878B" box="[303,332,1675,1698]" captionStart="Figs 49-54" captionStartId="14.[151,192,1993,2014]" captionTargetBox="[155,1435,746,1948]" captionTargetId="figure-217@14.[151,1436,737,1955]" captionTargetPageId="14" captionText="Figs 49-54. Karakumosa gromovi sp. nov.; male paratype (49-50) and female paratype (51-54). (49, 52) Body, ventral view. (50-51) Ditto, dorsal view. (53) Female prosoma, frontal view. (54) Female body, lateral view. Scale bars 1 cm (49-52, 54), 0.5 cm (53)." figureDoi="http://doi.org/10.5281/zenodo.5743732" httpUri="https://zenodo.org/record/5743732/files/figure.png" pageId="3" pageNumber="277">52</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FEE623856875878B" box="[344,373,1675,1698]" captionStart="Figs 89-92" captionStartId="21.[151,192,970,991]" captionTargetBox="[153,1429,259,929]" captionTargetId="figure-469@21.[151,1436,257,930]" captionTargetPageId="21" captionText="Figs 89-92. Karakumosa repetek sp. nov.; male paratype (89-90) and female paratype (91-92). (89, 92). Body, dorsal view. (90, 91) Ditto, ventral view. Scale bars 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5743748" httpUri="https://zenodo.org/record/5743748/files/figure.png" pageId="3" pageNumber="277">91</figureCitation>
), thus distinct from the majority of Palaearctic burrowing lycosine genera described to date (see
<bibRefCitation id="01454B70FF973F45FF7723C4685587C8" author="Simon E." box="[201,341,1738,1761]" pageId="3" pageNumber="277" pagination="57 - 91" refId="ref23665" refString="Simon E. 1876. Etudes arachnologiques. 4 e memoire. VII. Revision des especes europeennes du groupe de la Lycosa tarentula Rossi. Annales de la Societe Entomologique de France (5) 6: 57 - 91." type="journal article" year="1876">Simon, 1876</bibRefCitation>
;
<bibRefCitation id="01454B70FF973F45FEE123C46B0387C8" author="Logunov D. V." box="[351,515,1738,1761]" pageId="3" pageNumber="277" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov, 2010</bibRefCitation>
,
<bibRefCitation id="01454B70FF973F45FDB023C46B4887C8" author="Logunov D. V." box="[526,584,1738,1761]" pageId="3" pageNumber="277" pagination="349 - 362" refId="ref22421" refString="Logunov D. V. 2012. A synopsis of the genus Zyuzicosa Logunov, 2010 (Aranei: Lycosidae). Arthropoda Selecta 21 (4): 349 - 362." type="journal article" year="2012">2012</bibRefCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FF2923E76BDF86F4" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF2923E769C98629" box="[151,201,1769,1792]" italics="true" pageId="3" pageNumber="277">Legs</emphasis>
: leg formula IV,I,II,III in both sexes, rarely IV,II,I,III in some males; all segments densely covered with white setae; in both sexes, metatarsi and tarsi I-II ventrally with well-developed scopulae and longitudinal rows of spinules (sometimes poorly visible;
<figureCitation id="FDEF2A04FF973F45FD2922696BF08657" box="[663,752,1895,1918]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="3" pageNumber="277">Figs 36</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FF29228969B486B7" box="[151,180,1927,1950]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">63</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FF76228969F386B7" box="[200,243,1927,1950]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">140</figureCitation>
), tarsi III-IV only with ventral longitudinal rows of spinules (in some specimens a scopula could be developed on lateral sides of segments only;
<figureCitation id="FDEF2A04FF973F45FDC822C86BD286F4" box="[630,722,1990,2013]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">Fig. 141</figureCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FF2922E869F78934" blockId="3.[151,756,887,2077]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FF2922E8684B86D4" box="[151,331,2022,2045]" italics="true" pageId="3" pageNumber="277">Female pedipalp</emphasis>
: With a single tarsal claw (e.g.
<figureCitation id="FDEF2A04FF973F45FD2022E86BF086D4" box="[670,752,2022,2045]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 26</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FF292D0869B48934" box="[151,180,2054,2077]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="3" pageNumber="277">35</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FF012D0869EB8934" box="[191,235,2054,2077]" captionStart="Figs 136-141" captionStartId="26.[151,192,1422,1443]" captionTargetBox="[151,1435,257,1362]" captionTargetId="figure-218@26.[151,1436,257,1362]" captionTargetPageId="26" captionText="Figs 136-141. Karakumosa shmatkoi sp. nov., female from Azerbaijan, Absheron peninsula. (136) Prosoma, frontal view. (137) Ditto, ventral view. (138) Ditto, lateral view. (139) Epigyne, ventral view. (140) Tarsus I, ventral view. (141) Tarsus IV, ventral view. Scale bars 2 mm (137-138), 1 mm (136), 0.5 mm (140-141), 0.2 mm (139)." figureDoi="http://doi.org/10.5281/zenodo.5743758" httpUri="https://zenodo.org/record/5743758/files/figure.png" pageId="3" pageNumber="277">138</figureCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FC81240C6C298374" blockId="3.[831,1437,258,1576]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FC81240C6D798030" box="[831,1145,258,281]" italics="true" pageId="3" pageNumber="277">Female copulatory organs</emphasis>
: Epigyne with a well pronounced longitudinal atrium being at least two times longer than wide; edges at proximal end of atrium forming two round convergent lips, these usually markedly sclerotized and darker (brown or russet) than nearby cuticle (e.g.
<figureCitation id="FDEF2A04FF973F45FB9524AA6D858092" box="[1067,1157,420,443]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 27</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB2624AA6DB58092" box="[1176,1205,420,443]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">61</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB7624AA6DE68092" box="[1224,1254,420,443]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="3" pageNumber="277">85</figureCitation>
); septal pedicel absent; median septum essentially reduced to posterior transverse, slightly convex plate of variable shape (e.g.
<figureCitation id="FDEF2A04FF973F45FC81270B6A978335" box="[831,919,517,540]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 27</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC16270B6AC68335" box="[936,966,517,540]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">96</figureCitation>
); spermathecae tube-shaped, only slightly wider than or of same width as narrow and short insemination ducts (e.g.
<figureCitation id="FDEF2A04FF973F45FBFB27486D978374" box="[1093,1175,582,605]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="3" pageNumber="277">Figs 28</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB1C27486DBF8374" box="[1186,1215,582,605]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="3" pageNumber="277">43</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB7427486DE78374" box="[1226,1255,582,605]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">62</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB4C27486C1D8374" box="[1266,1309,582,605]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">121</figureCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FC8127686D298249" blockId="3.[831,1437,258,1576]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FC8127686AD88354" box="[831,984,614,637]" italics="true" pageId="3" pageNumber="277">Male pedipalp</emphasis>
: Femur length equal to that of patella+tibia (
<figureCitation id="FDEF2A04FF973F45FCF927886AC383B4" box="[839,963,646,669]" captionStart="Figs 12-21" captionStartId="8.[151,192,1900,1921]" captionTargetBox="[161,1426,261,1863]" captionTargetId="figure-9@8.[151,1436,257,1863]" captionTargetPageId="8" captionText="Figs 12-21. Karakumosa alticeps (Kroneberg, 1875); male specimens from Tartogay Vil. (12-15) and Zhankel’ Vil. (16), and the male lectotype (17-21). (12, 17) Left bulbus, ventral view. (13, 20) Ditto, retrolateral view. (14, 19) Ditto, posterior view. (15, 16) Left male palp, retrolateral view. (18) Left bulbus, apical view. (21) Original data labels of the lectotype. Scale bars 0.5 mm (15, 16), 0.2 mm (12-14, 17-20). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743720" httpUri="https://zenodo.org/record/5743720/files/figure.png" pageId="3" pageNumber="277">Figs 15-16</figureCitation>
); cymbium symmetrical, twice as long as wide, with almost round alveolus (
<figureCitation id="FDEF2A04FF973F45FB7727A96C1C8397" box="[1225,1308,679,702]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">Fig. 65</figureCitation>
), its length about equal to that of palpal tibia; distal part of cymbium 1.1-1.3 times shorter than alveolus length (
<figureCitation id="FDEF2A04FF973F45FA8627E66C8F83D6" box="[1336,1423,744,767]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">Fig. 65</figureCitation>
); cymbium with a cluster of blunt, rigid and straight bristles on its tips (
<figureCitation id="FDEF2A04FF973F45FC0226266D118216" box="[956,1041,808,831]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="3" pageNumber="277">Figs 45</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FBA026266D3A8216" box="[1054,1082,808,831]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="3" pageNumber="277">69</figureCitation>
) as typical of lycosid burrowers (
<bibRefCitation id="01454B70FF973F45FCF926476AD98249" author="Zyuzin A. A." box="[839,985,841,864]" pageId="3" pageNumber="277" pagination="419 - 422" refId="ref24236" refString="Zyuzin A. A. 1990. Studies on burrowing spiders of the family Lycosidae (Araneae). I. Preliminary data on structural and functional features. Acta Zoologica Fennica 190: 419 - 422." type="journal article" year="1990">Zyuzin, 1990</bibRefCitation>
,
<bibRefCitation id="01454B70FF973F45FC5A26476D1D8249" author="Zyuzin A. A." box="[996,1053,841,864]" pageId="3" pageNumber="277" pagination="693 - 700" refId="ref24274" refString="Zyuzin A. A. 1993. Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and new species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum 33 (2): 693 - 700." type="journal article" year="1993">1993</bibRefCitation>
).
</paragraph>
<paragraph id="656B3681FF973F45FC8126676C018701" blockId="3.[831,1437,258,1576]" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FC8126676D6582A9" box="[831,1125,873,896]" italics="true" pageId="3" pageNumber="277">Male copulatory organs</emphasis>
: Subtegulum round and comparatively small, situated in proximal-mesal position (
<figureCitation id="FDEF2A04FF973F45FCF926A46A9182E8" box="[839,913,938,961]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="3" pageNumber="277">Figs 9</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC1F26A46ABE82E8" box="[929,958,938,961]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="3" pageNumber="277">46</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC7026A46AEB82E8" box="[974,1003,938,961]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">56</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC4526A46D1982E8" box="[1019,1049,938,961]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">98</figureCitation>
: St); tegulum round and broad, in unexpanded palp clearly visible only on prolateralproximal side of bulbus (
<figureCitation id="FDEF2A04FF973F45FBE726E56DB0852B" box="[1113,1200,1003,1026]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">Figs 55</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB0126E56DDC852B" box="[1215,1244,1003,1026]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">98</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB5426E56C15852B" box="[1258,1301,1003,1026]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">118</figureCitation>
: T); median apophysis wide (usually wider than long) and broad (
<figureCitation id="FDEF2A04FF973F45FAD321056A4E856B" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="3" pageNumber="277">Figs 9</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FCE321256A7A856B" box="[861,890,1067,1090]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">55</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC3921256AB2856B" box="[903,946,1067,1090]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">118</figureCitation>
: OP, IP), folded along its proximal edge and consisting of two (outer and inner) flat plates, the outer plate bearing a proximal extension and a median tooth (
<figureCitation id="FDEF2A04FF973F45FCF921826A9B858A" box="[839,923,1164,1187]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">Figs 56</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC1921826AC5858A" box="[935,965,1164,1187]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">98</figureCitation>
: MT, PE); median tooth rarely singular and finger-shaped (
<figureCitation id="FDEF2A04FF973F45FC6321A36D2F85ED" box="[989,1071,1197,1220]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="3" pageNumber="277">Figs 30</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB8721A36D5785ED" box="[1081,1111,1197,1220]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="3" pageNumber="277">42</figureCitation>
), usually consisting of a lateral claw and a median edge bearing micro-teeth (indicated by arrow in
<figureCitation id="FDEF2A04FF973F45FC7721E36D2A842D" box="[969,1066,1261,1284]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">Figs 123</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB8621E36D61842D" box="[1080,1121,1261,1284]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="3" pageNumber="277">128</figureCitation>
) and in some species with a low serrate flange at its foot (indicated by arrow in
<figureCitation id="FDEF2A04FF973F45FAD020006A69846C" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">Figs 125</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FCC820206AA0846C" box="[886,928,1326,1349]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="3" pageNumber="277">134</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC1320206AD9846C" box="[941,985,1326,1349]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="3" pageNumber="277">152</figureCitation>
); synembolus biramous, with two acutely pointed lamellae (
<figureCitation id="FDEF2A04FF973F45FBB220416D66844F" box="[1036,1126,1359,1382]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">Figs 57</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FBC620416D95844F" box="[1144,1173,1359,1382]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="3" pageNumber="277">98</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB1920416DD2844F" box="[1191,1234,1359,1382]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">118</figureCitation>
: Se); a triangular hyaline conductor present, well-developed and pointed ventrad (
<figureCitation id="FDEF2A04FF973F45FC2320816AF0848F" box="[925,1008,1423,1446]" captionStart="Figs 12-21" captionStartId="8.[151,192,1900,1921]" captionTargetBox="[161,1426,261,1863]" captionTargetId="figure-9@8.[151,1436,257,1863]" captionTargetPageId="8" captionText="Figs 12-21. Karakumosa alticeps (Kroneberg, 1875); male specimens from Tartogay Vil. (12-15) and Zhankel’ Vil. (16), and the male lectotype (17-21). (12, 17) Left bulbus, ventral view. (13, 20) Ditto, retrolateral view. (14, 19) Ditto, posterior view. (15, 16) Left male palp, retrolateral view. (18) Left bulbus, apical view. (21) Original data labels of the lectotype. Scale bars 0.5 mm (15, 16), 0.2 mm (12-14, 17-20). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743720" httpUri="https://zenodo.org/record/5743720/files/figure.png" pageId="3" pageNumber="277">Figs 12</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FC4420816D17848F" box="[1018,1047,1423,1446]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">56</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB9C20816D4D848F" box="[1058,1101,1423,1446]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">118</figureCitation>
: C); embolus thin, with a rather wide and prominent pars pendula (
<figureCitation id="FDEF2A04FF973F45FB7020BE6C1984EE" box="[1230,1305,1456,1479]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="3" pageNumber="277">Figs 4</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FA9520BE6C5484EE" box="[1323,1364,1456,1479]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">116</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FADB20BE6C9084EE" box="[1381,1424,1456,1479]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="3" pageNumber="277">129</figureCitation>
), its origin in a prolatero-apical position, with only the embolic tip visible in between or beneath branches of the synembolus in unexpanded palp (
<figureCitation id="FDEF2A04FF973F45FB1B231F6DF58701" box="[1189,1269,1553,1576]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">Fig. 57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF973F42FC81234E6AC88330" lastPageId="4" lastPageNumber="278" pageId="3" pageNumber="277" type="discussion">
<paragraph id="656B3681FF973F42FC81234E6B6980B0" blockId="3.[831,1437,1600,2077]" lastBlockId="4.[151,757,258,1177]" lastPageId="4" lastPageNumber="278" pageId="3" pageNumber="277">
<emphasis id="57A0EA93FF973F45FC81234E6AC5877E" bold="true" box="[831,965,1600,1623]" pageId="3" pageNumber="277">Comments:</emphasis>
Within the Lycosoidea the transverse median apophysis is considered a typical feature of the
<taxonomicName id="A2D44D02FF973F45FC81238F6D6D87B1" authority="(Griswold, 1993)" baseAuthorityName="Griswold" baseAuthorityYear="1993" box="[831,1133,1665,1688]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="277" phylum="Arthropoda" rank="family">
Lycosidae (
<bibRefCitation id="01454B70FF973F45FC05238F6D6587B1" author="Griswold C. E." box="[955,1125,1665,1688]" pageId="3" pageNumber="277" pagination="1 - 39" refId="ref22040" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology, 539: 1 - 39." type="journal article" year="1993">Griswold, 1993</bibRefCitation>
)
</taxonomicName>
and a synapomorphy of the
<taxonomicName id="A2D44D02FF973F45FC8123AF6D6F8791" authority="(Dondale, 1986)" authorityName="Sundevall" authorityYear="1833" baseAuthorityName="Dondale" baseAuthorityYear="1986" box="[831,1135,1697,1720]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="276" phylum="Arthropoda" rank="subFamily" subFamily="Lycosinae">
Lycosinae (
<bibRefCitation id="01454B70FF973F45FC0123AF6D678791" author="Dondale C. D." box="[959,1127,1697,1720]" pageId="3" pageNumber="277" pagination="327 - 332" refId="ref21895" refString="Dondale C. D. 1986. The subfamilies of wolf spiders (Araneae: Lycosidae) [pp. 327 - 332]. In: Barrientos J. A. (ed.), Actas X. Congreso Internacional de Aracnologia, Jaca, Espana, Volume I. Juvenil, Barcelona, 73 pp." type="book chapter" year="1986">Dondale, 1986</bibRefCitation>
)
</taxonomicName>
, although its shape varies. In
<taxonomicName id="A2D44D02FF973F45FCE023CC6AE687F0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[862,998,1730,1753]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="277" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF973F45FCE023CC6AE687F0" box="[862,998,1730,1753]" italics="true" pageId="3" pageNumber="277">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF973F45FC5123CC6D5287F0" box="[1007,1106,1730,1753]" pageId="3" pageNumber="277" rank="genus">gen. nov.</taxonomicNameLabel>
the median apophysis is large and composite, consisting of two distinct plates that appear fused along their proximal edges: the outer and the inner plates (e.g.
<figureCitation id="FDEF2A04FF973F45FB8E222D6D788613" box="[1072,1144,1827,1850]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="3" pageNumber="277">Figs 9</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB36222D6DA58613" box="[1160,1189,1827,1850]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="3" pageNumber="277">56</figureCitation>
,
<figureCitation id="FDEF2A04FF973F45FB0A222D6DDF8613" box="[1204,1247,1827,1850]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="3" pageNumber="277">118</figureCitation>
: OP, IP). Such a complex structure of the median apophysis is currently regarded as unique within the
<taxonomicName id="A2D44D02FF973F45FB35226A6DF78652" authorityName="Sundevall" authorityYear="1833" box="[1163,1271,1892,1915]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="276" phylum="Arthropoda" rank="subFamily" subFamily="Lycosinae">Lycosinae</taxonomicName>
and within the entire family
<taxonomicName id="A2D44D02FF973F45FC66228A6D4886B2" baseAuthorityName="Griswold" baseAuthorityYear="1993" box="[984,1096,1924,1947]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="277" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
. Of the lycosid genera known to us, only two have a comparably complex structure of the median apophysis:
<taxonomicName id="A2D44D02FF973F45FB3F22CB6C9C86F5" authority="Zyuzin, 1993" authorityName="Zyuzin" authorityYear="1993" box="[1153,1436,1989,2012]" class="Arachnida" family="Lycosidae" genus="Oculicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF973F45FB3F22CB6DEF86F5" box="[1153,1263,1989,2012]" italics="true" pageId="3" pageNumber="277">Oculicosa</emphasis>
<bibRefCitation id="01454B70FF973F45FABF22CB6C9C86F5" author="Zyuzin A. A." box="[1281,1436,1989,2012]" pageId="3" pageNumber="277" pagination="693 - 700" refId="ref24274" refString="Zyuzin A. A. 1993. Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and new species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum 33 (2): 693 - 700." type="journal article" year="1993">Zyuzin, 1993</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="01454B70FF973F45FCF922EB6D7E86D5" author="Logunov D. V. &amp; Gromov A. V." box="[839,1150,2021,2044]" pageId="3" pageNumber="277" pagination="48 - 51" refId="ref22491" refString="Logunov D. V., Gromov A. V. 2011. Notes on the distribution of Oculicosa supermirabilis (Araneae, Lycosidae). Arachnologische Mitteilungen 42: 48 - 51." type="journal article" year="2011">Logunov &amp; Gromov, 2011</bibRefCitation>
: figs 1-2) and
<taxonomicName id="A2D44D02FF973F45FA8F22EB6C9D86D5" authorityName="Logunov" authorityYear="2010" box="[1329,1437,2021,2044]" class="Arachnida" family="Lycosidae" genus="Zyuzicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF973F45FA8F22EB6C9D86D5" box="[1329,1437,2021,2044]" italics="true" pageId="3" pageNumber="277">Zyuzicosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="01454B70FF973F45FCF92D086AF78934" author="Logunov D. V." box="[839,1015,2054,2077]" pageId="3" pageNumber="277" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov, 2010</bibRefCitation>
: figs 34-39). The outer plate of the median apophysis of
<taxonomicName id="A2D44D02FF903F42FE30240C6B178030" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[398,535,258,281]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FE30240C6B178030" box="[398,535,258,281]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FD9A240C6B8E8030" box="[548,654,258,281]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
, with its proximal extension and median tooth of MA, looks like an apomorphic modification of the prominent transverse chitinous ridge of the median apophysis in
<taxonomicName id="A2D44D02FF903F42FD3E246C6BED8050" authorityName="Zyuzin" authorityYear="1993" box="[640,749,354,377]" class="Arachnida" family="Lycosidae" genus="Oculicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF903F42FD3E246C6BED8050" box="[640,749,354,377]" italics="true" pageId="4" pageNumber="278">Oculicosa</emphasis>
</taxonomicName>
. Obviously, both structures are homologous.
</paragraph>
<paragraph id="656B3681FF903F42FF2924AC6BD38350" blockId="4.[151,757,258,1177]" pageId="4" pageNumber="278">
Another unique feature of
<taxonomicName id="A2D44D02FF903F42FE0124AC6B478090" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[447,583,418,441]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FE0124AC6B478090" box="[447,583,418,441]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FDEF24AC6BB68090" box="[593,694,418,441]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
is its biramous synembolus, consisting of two thin and very acutely pointed lamellae (e.g.
<figureCitation id="FDEF2A04FF903F42FE5C24EC6B2880D0" box="[482,552,482,505]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="4" pageNumber="278">Figs 4</figureCitation>
,
<figureCitation id="FDEF2A04FF903F42FD8824EC6B5380D0" box="[566,595,482,505]" captionStart="Figs 55-58" captionStartId="15.[151,192,2024,2045]" captionTargetBox="[151,1435,682,1983]" captionTargetId="figure-128@15.[151,1436,682,1983]" captionTargetPageId="15" captionText="Figs 55-58. Karakumosa gromovi sp. nov., bulbus of male paratype. (55) Ventral-retrolateral view. (56) Ventral view. (57)Apical view. (58) Retrolateral view. Scale bars 0.2 mm. Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743734" httpUri="https://zenodo.org/record/5743734/files/figure.png" pageId="4" pageNumber="278">57</figureCitation>
,
<figureCitation id="FDEF2A04FF903F42FDDF24EC6B7E80D0" box="[609,638,482,505]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="4" pageNumber="278">67</figureCitation>
,
<figureCitation id="FDEF2A04FF903F42FD3524EC6BB680D0" box="[651,694,482,505]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="4" pageNumber="278">116</figureCitation>
). The only other lycosid genus with a comparable conformation of the synembolus is
<taxonomicName id="A2D44D02FF903F42FE2C272C68FD8310" authorityName="Logunov" authorityYear="2010" box="[402,509,546,569]" class="Arachnida" family="Lycosidae" genus="Zyuzicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF903F42FE2C272C68FD8310" box="[402,509,546,569]" italics="true" pageId="4" pageNumber="278">Zyuzicosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="01454B70FF903F42FDAE272C6BBD8310" author="Logunov D. V." box="[528,701,546,569]" pageId="4" pageNumber="278" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov, 2010</bibRefCitation>
: figs 64, 66), but in the latter genus it consists of one acutely pointed lamella and a wide, strongly sclerotized base.
</paragraph>
<paragraph id="656B3681FF903F42FF29278C68888250" blockId="4.[151,757,258,1177]" pageId="4" pageNumber="278">
In the absence of a phylogenetic analysis of the Palaearctic genera of
<taxonomicName id="A2D44D02FF903F42FEAA27AC68808390" baseAuthorityName="Griswold" baseAuthorityYear="1993" box="[276,384,674,697]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
(but see
<bibRefCitation id="01454B70FF903F42FE4327AC6BF08390" author="Murphy N. P. &amp; Framenau V. W. &amp; Donnellan S. C. &amp; Harvey M. S. &amp; Park Y. C. &amp; Austin A. D." box="[509,752,674,697]" pageId="4" pageNumber="278" pagination="583 - 602" refId="ref22853" refString="Murphy N. P., Framenau V. W., Donnellan S. C., Harvey M. S., Park Y. C., Austin A. D. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution 28: 583 - 602." type="journal article" year="2006">
Murphy
<emphasis id="57A0EA93FF903F42FDDC27AC6BA18390" box="[610,673,674,697]" italics="true" pageId="4" pageNumber="278">et al.</emphasis>
, 2006
</bibRefCitation>
;
<bibRefCitation id="01454B70FF903F42FF2927CC68CF83F0" author="Piacentini L. N. &amp; Ramirez M. J." box="[151,463,706,729]" pageId="4" pageNumber="278" pagination="227 - 240" refId="ref23158" refString="Piacentini L. N., Ramirez M. J. 2019. Hunting the wolf: a molecular phylogeny of the wolf spiders (Araneae, Lycosidae). Molecular Phylogenetics and Evolution 136: 227 - 240." type="journal article" year="2019">Piacentini &amp; Ramírez, 2019</bibRefCitation>
), it is difficult to establish whether both unique features of
<taxonomicName id="A2D44D02FF903F42FE4127EC6B8783D0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[511,647,738,761]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FE4127EC6B8783D0" box="[511,647,738,761]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FD2E27EC6BF583D0" box="[656,757,738,761]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
are primitive or derived (they are likely to be derived). Yet, as a provisional hypothesis to be further tested, we consider both of them as the putative synapomorphies of
<taxonomicName id="A2D44D02FF903F42FF29266C681F8250" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[151,287,866,889]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FF29266C681F8250" box="[151,287,866,889]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FE98266C68888250" box="[294,392,866,889]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="656B3681FF903F42FF29268C6B5785B0" blockId="4.[151,757,258,1177]" pageId="4" pageNumber="278">
We speculate that if the genus
<taxonomicName id="A2D44D02FF903F42FE69268C6B5F82B0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[471,607,898,921]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FE69268C6B5F82B0" box="[471,607,898,921]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FDDB268C6BC582B0" box="[613,709,898,921]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
was included in one of the two latest phylogenetic analyses of the
<taxonomicName id="A2D44D02FF903F42FF0126CC682B82F0" baseAuthorityName="Griswold" baseAuthorityYear="1993" box="[191,299,962,985]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
based on DNA data (
<bibRefCitation id="01454B70FF903F42FDAA26CC6BF082F0" author="Murphy N. P. &amp; Framenau V. W. &amp; Donnellan S. C. &amp; Harvey M. S. &amp; Park Y. C. &amp; Austin A. D." box="[532,752,962,985]" pageId="4" pageNumber="278" pagination="583 - 602" refId="ref22853" refString="Murphy N. P., Framenau V. W., Donnellan S. C., Harvey M. S., Park Y. C., Austin A. D. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution 28: 583 - 602." type="journal article" year="2006">
Murphy
<emphasis id="57A0EA93FF903F42FDCD26CC6BAA82F0" box="[627,682,962,985]" italics="true" pageId="4" pageNumber="278">et al.</emphasis>
, 2006
</bibRefCitation>
;
<bibRefCitation id="01454B70FF903F42FF2926EC68CD82D0" author="Piacentini L. N. &amp; Ramirez M. J." box="[151,461,994,1017]" pageId="4" pageNumber="278" pagination="227 - 240" refId="ref23158" refString="Piacentini L. N., Ramirez M. J. 2019. Hunting the wolf: a molecular phylogeny of the wolf spiders (Araneae, Lycosidae). Molecular Phylogenetics and Evolution 136: 227 - 240." type="journal article" year="2019">Piacentini &amp; Ramírez, 2019</bibRefCitation>
), it would likely be placed either within the clade E (
<emphasis id="57A0EA93FF903F42FE14210C68E58530" box="[426,485,1026,1049]" italics="true" pageId="4" pageNumber="278">sensu</emphasis>
<bibRefCitation id="01454B70FF903F42FE52210C6BC38530" author="Murphy N. P. &amp; Framenau V. W. &amp; Donnellan S. C. &amp; Harvey M. S. &amp; Park Y. C. &amp; Austin A. D." box="[492,707,1026,1049]" pageId="4" pageNumber="278" pagination="583 - 602" refId="ref22853" refString="Murphy N. P., Framenau V. W., Donnellan S. C., Harvey M. S., Park Y. C., Austin A. D. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution 28: 583 - 602." type="journal article" year="2006">
Murphy
<emphasis id="57A0EA93FF903F42FDF6210C6B7E8530" box="[584,638,1026,1049]" italics="true" pageId="4" pageNumber="278">et al.</emphasis>
, 2006
</bibRefCitation>
: figs 2-3), somewhere close to
<emphasis id="57A0EA93FF903F42FE03212C6B778510" box="[445,631,1058,1081]" italics="true" pageId="4" pageNumber="278">Lycosa tarantula</emphasis>
(
<bibRefCitation id="01454B70FF903F42FD34212C69CE8570" author="Linnaeus C." pageId="4" pageNumber="278" refId="ref22333" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus differentiis, synonymis, locis. Editio decima, reformata. Impensis Direct. Laurentii Salvii, Holmiae, 821 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
) and the clade E1, or within the
<taxonomicName id="A2D44D02FF903F42FD85214C6BA78570" authorityName="Sundevall" authorityYear="1833" box="[571,679,1090,1113]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="subFamily" subFamily="Lycosinae">Lycosinae</taxonomicName>
(
<emphasis id="57A0EA93FF903F42FD07214C6BF48570" box="[697,756,1090,1113]" italics="true" pageId="4" pageNumber="278">sensu</emphasis>
<bibRefCitation id="01454B70FF903F42FF29216C68E18550" author="Piacentini L. N. &amp; Ramirez M. J." box="[151,481,1122,1145]" pageId="4" pageNumber="278" pagination="227 - 240" refId="ref23158" refString="Piacentini L. N., Ramirez M. J. 2019. Hunting the wolf: a molecular phylogeny of the wolf spiders (Araneae, Lycosidae). Molecular Phylogenetics and Evolution 136: 227 - 240." type="journal article" year="2019">Piacentini &amp; Ramírez, 2019</bibRefCitation>
: fig. 4), in the branch containing the Palaearctic
<emphasis id="57A0EA93FF903F42FE0F218C68FC85B0" box="[433,508,1154,1177]" italics="true" pageId="4" pageNumber="278">Lycosa</emphasis>
species.
</paragraph>
<paragraph id="656B3681FF903F42FF2921BF6AC88330" blockId="4.[151,756,1201,1256]" lastBlockId="4.[831,1437,258,537]" pageId="4" pageNumber="278">
<emphasis id="57A0EA93FF903F42FF2921BF683385E1" bold="true" box="[151,307,1201,1224]" pageId="4" pageNumber="278">Composition:</emphasis>
To date nine species are assigned to
<taxonomicName id="A2D44D02FF903F42FF2921DF682085C1" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[151,288,1233,1256]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FF2921DF682085C1" box="[151,288,1233,1256]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FE9521DF689485C1" box="[299,404,1233,1256]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
:
<taxonomicName id="A2D44D02FF903F42FE1C21DF6BF085C1" authority="(Kroneberg, 1875)" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[418,752,1233,1256]" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="alticeps">
<emphasis id="57A0EA93FF903F42FE1C21DF6B1785C1" box="[418,535,1233,1256]" italics="true" pageId="4" pageNumber="278">K. alticeps</emphasis>
(Kroneberg, 1875)
</taxonomicName>
, comb. nov. (
<specimenCount id="73D2FD08FF903F42FC79240C6ADF8030" box="[967,991,258,281]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FC61240C6AEF8030" box="[991,1007,258,281]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
,
<collectingCountry id="1DC37611FF903F42FC40240C6D768030" box="[1022,1142,258,281]" name="Uzbekistan" pageId="4" pageNumber="278">Uzbekistan</collectingCountry>
and southern
<collectingCountry id="1DC37611FF903F42FAAE240C6C918030" box="[1296,1425,258,281]" name="Kazakhstan" pageId="4" pageNumber="278">Kazakhstan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FC81242C6AD88010" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[831,984,290,313]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="badkhyzica" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FC81242C6AD88010" box="[831,984,290,313]" italics="true" pageId="4" pageNumber="278">K. badkhyzica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FC5E242C6D348010" box="[992,1076,290,313]" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FBFD242C6D5B8010" box="[1091,1115,290,313]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FBE5242C6D6B8010" box="[1115,1131,290,313]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
,
<collectingCountry id="1DC37611FF903F42FBC7242C6C118010" box="[1145,1297,290,313]" name="Turkmenistan" pageId="4" pageNumber="278">Turkmenistan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FA9A242C6C9C8010" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1316,1436,290,313]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="gromovi" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FA9A242C6C9C8010" box="[1316,1436,290,313]" italics="true" pageId="4" pageNumber="278">K. gromovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FC81244C6A938070" box="[831,915,322,345]" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FC1C244C6ABA8070" box="[930,954,322,345]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FC04244C6ACA8070" box="[954,970,322,345]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
, southern
<collectingCountry id="1DC37611FF903F42FB85244C6DB78070" box="[1083,1207,322,345]" name="Uzbekistan" pageId="4" pageNumber="278">Uzbekistan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FB74244C6A818050" authority="(Pocock, 1889)" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" baseAuthorityName="Pocock" baseAuthorityYear="1889" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="medica" status="comb. nov.">
<emphasis id="57A0EA93FF903F42FB74244C6C378070" box="[1226,1335,322,345]" italics="true" pageId="4" pageNumber="278">K. medica</emphasis>
(
<bibRefCitation id="01454B70FF903F42FAFB244C6A788050" author="Pocock R. I." pageId="4" pageNumber="278" pagination="110 - 121" refId="ref23240" refString="Pocock R. I. 1889. Arachnida, Chilopoda and Crustacea. In: Aitchison J. E. T. (ed.), The zoology of the Afghan Delimitation Commission. Transactions of the Linnean Society of London (Zoology, 2) 5 (3): 110 - 121, pl. 13." type="journal article" year="1889">Pocock, 1889</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="4C9357E8FF903F42FC2D246C6D128050" box="[915,1042,354,377]" pageId="4" pageNumber="278" rank="species">comb. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FB97246C6D418050" box="[1065,1089,354,377]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FBFE246C6D508050" box="[1088,1104,354,377]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
, north-western
<collectingCountry id="1DC37611FF903F42FAB7246C6C918050" box="[1289,1425,354,377]" name="Afghanistan" pageId="4" pageNumber="278">Afghanistan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FC81248C6AAC80B0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[831,940,386,409]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="repetek" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FC81248C6AAC80B0" box="[831,940,386,409]" italics="true" pageId="4" pageNumber="278">K. repetek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FC0A248C6D0780B0" box="[948,1031,386,409]" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FBA9248C6D2F80B0" box="[1047,1071,386,409]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FB91248C6D3F80B0" box="[1071,1087,386,409]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
,
<collectingCountry id="1DC37611FF903F42FBF2248C6DE480B0" box="[1100,1252,386,409]" name="Turkmenistan" pageId="4" pageNumber="278">Turkmenistan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FB48248C6C7680B0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1270,1398,386,409]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FB48248C6C7680B0" box="[1270,1398,386,409]" italics="true" pageId="4" pageNumber="278">K. shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FAC0248C6A6D8090" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FCC124AC6A978090" box="[895,919,418,441]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FC2924AC6AA78090" box="[919,935,418,441]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
,
<location id="600B605AFF903F42FC0624AC6D918090" LSID="urn:lsid:plazi:treatment:ED7D8797FF963F40FC7423936C9B8330:600B605AFF903F42FC0624AC6D918090" box="[952,1169,418,441]" country="Azerbaijan" name="northern Ciscaspian" pageId="4" pageNumber="278">northern Ciscaspian</location>
region and
<collectingCountry id="1DC37611FF903F42FAA624AC6C928090" box="[1304,1426,418,441]" name="Azerbaijan" pageId="4" pageNumber="278">Azerbaijan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FC8124CC6AD680F0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[831,982,450,473]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="tashkumyr" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FC8124CC6AD680F0" box="[831,982,450,473]" italics="true" pageId="4" pageNumber="278">K. tashkumyr</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FC5B24CC6D4080F0" box="[997,1088,450,473]" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FBE924CC6D6B80F0" box="[1111,1131,450,473]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
,
<collectingCountry id="1DC37611FF903F42FB3E24CC6C0080F0" box="[1152,1280,450,473]" name="Kyrgyzstan" pageId="4" pageNumber="278">Kyrgyzstan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FAA424CC6C9C80F0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1306,1436,450,473]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="turanica" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FAA424CC6C9C80F0" box="[1306,1436,450,473]" italics="true" pageId="4" pageNumber="278">K. turanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FC8124EC6A9680D0" box="[831,918,482,505]" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FC1624EC6AC080D0" box="[936,960,482,505]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FC7E24EC6AD080D0" box="[960,976,482,505]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
,
<collectingCountry id="1DC37611FF903F42FC5E24EC6D7880D0" box="[992,1144,482,505]" name="Turkmenistan" pageId="4" pageNumber="278">Turkmenistan</collectingCountry>
),
<taxonomicName id="A2D44D02FF903F42FB3024EC6DFA80D0" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1166,1274,482,505]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="57A0EA93FF903F42FB3024EC6DFA80D0" box="[1166,1274,482,505]" italics="true" pageId="4" pageNumber="278">K. zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FABB24EC6C5B80D0" box="[1285,1371,482,505]" pageId="4" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="73D2FD08FF903F42FAD324EC6C8580D0" box="[1389,1413,482,505]" pageId="4" pageNumber="278" type="male">♂</specimenCount>
<specimenCount id="73D2FD08FF903F42FA3B24EC6C9580D0" box="[1413,1429,482,505]" pageId="4" pageNumber="278" type="female">♀</specimenCount>
,
<collectingCountry id="1DC37611FF903F42FC81270C6ABC8330" box="[831,956,514,537]" name="Uzbekistan" pageId="4" pageNumber="278">Uzbekistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="2DCE650AFF903F42FC81273F6AA185E1" pageId="4" pageNumber="278" type="distribution">
<paragraph id="656B3681FF903F42FC81273F6D9E83C1" blockId="4.[831,1437,561,1224]" pageId="4" pageNumber="278">
<emphasis id="57A0EA93FF903F42FC81273F6AD78361" bold="true" box="[831,983,561,584]" pageId="4" pageNumber="278">Distribution:</emphasis>
Central Asia (
<figureCitation id="FDEF2A04FF903F42FB1E273F6DEF8361" box="[1184,1263,561,584]" captionStart="Fig" captionStartId="4.[151,186,2055,2076]" captionTargetBox="[155,1435,1347,2031]" captionTargetId="figure-536@4.[155,1435,1347,2031]" captionTargetPageId="4" captionText="Fig. 1. Geographical range of the genus Karakumosa gen. nov." figureDoi="http://doi.org/10.5281/zenodo.5743712" httpUri="https://zenodo.org/record/5743712/files/figure.png" pageId="4" pageNumber="278">Fig. 1</figureCitation>
): (semi)desert regions of the northern Ciscaspian Region, eastern
<collectingCountry id="1DC37611FF903F42FC81277F6ABB83A1" box="[831,955,625,648]" name="Azerbaijan" pageId="4" pageNumber="278">Azerbaijan</collectingCountry>
,
<collectingCountry id="1DC37611FF903F42FC67277F6D5C83A1" box="[985,1116,625,648]" name="Kazakhstan" pageId="4" pageNumber="278">Kazakhstan</collectingCountry>
,
<collectingCountry id="1DC37611FF903F42FBC5277F6DFC83A1" box="[1147,1276,625,648]" name="Kyrgyzstan" pageId="4" pageNumber="278">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="1DC37611FF903F42FAA4277F6C9883A1" box="[1306,1432,625,648]" name="Uzbekistan" pageId="4" pageNumber="278">Uzbekistan</collectingCountry>
,
<collectingCountry id="1DC37611FF903F42FC81279F6AD68381" box="[831,982,657,680]" name="Turkmenistan" pageId="4" pageNumber="278">Turkmenistan</collectingCountry>
and north-western
<collectingCountry id="1DC37611FF903F42FB69279F6C608381" box="[1239,1376,657,680]" name="Afghanistan" pageId="4" pageNumber="278">Afghanistan</collectingCountry>
; the occurrence of this genus in northern and north-eastern
<collectingCountry id="1DC37611FF903F42FC8127DF6A6A83C1" box="[831,874,721,744]" name="Iran" pageId="4" pageNumber="278">Iran</collectingCountry>
and
<collectingCountry id="1DC37611FF903F42FC2027DF6D0783C1" box="[926,1031,721,744]" name="Tajikistan" pageId="4" pageNumber="278">Tajikistan</collectingCountry>
is very likely.
</paragraph>
<paragraph id="656B3681FF903F42FC8127FF6AA185E1" blockId="4.[831,1437,561,1224]" pageId="4" pageNumber="278">
The genus is essentially restricted to the so-called Turan zoogeographic province (
<emphasis id="57A0EA93FF903F42FBDC261F6D9D8201" box="[1122,1181,785,808]" italics="true" pageId="4" pageNumber="278">sensu</emphasis>
<bibRefCitation id="01454B70FF903F42FB10261F6C958201" author="Kryzhanovsky O. L." box="[1198,1429,785,808]" pageId="4" pageNumber="278" refId="ref22258" refString="Kryzhanovsky O. L. 2002. Composition and distribution of entomofaunas of the globe. KMK Sci. Press, Moscow, 237 pp. [In Russian]" type="book" year="2002">Kryzhanovsky, 2002</bibRefCitation>
) (
<figureCitation id="FDEF2A04FF903F42FCF9263F6A8A8261" box="[839,906,817,840]" captionStart="Fig" captionStartId="4.[151,186,2055,2076]" captionTargetBox="[155,1435,1347,2031]" captionTargetId="figure-536@4.[155,1435,1347,2031]" captionTargetPageId="4" captionText="Fig. 1. Geographical range of the genus Karakumosa gen. nov." figureDoi="http://doi.org/10.5281/zenodo.5743712" httpUri="https://zenodo.org/record/5743712/files/figure.png" pageId="4" pageNumber="278">Fig. 1</figureCitation>
). It is the third genus of fossorial wolf spiders within the currently known
<taxonomicName id="A2D44D02FF903F42FBC7265F6DE58241" authorityName="Sundevall" authorityYear="1833" box="[1145,1253,849,872]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="subFamily" subFamily="Lycosinae">Lycosinae</taxonomicName>
that is confined to Central Asia (
<tableCitation id="2856033AFF903F42FC48267F6D4982A1" box="[1014,1097,881,904]" captionStart="Table 1" captionStartId="36.[151,204,256,277]" captionTargetBox="[162,1427,323,2056]" captionText="Table 1. The fossorial Lycosidae of Central Asia and their distribution." httpUri="http://table.plazi.org/id/31AB6609FFB03F62FF29240E6A3D803C" pageId="4" pageNumber="278" tableUuid="31AB6609FFB03F62FF29240E6A3D803C">Table 1</tableCitation>
). The other two are
<taxonomicName id="A2D44D02FF903F42FA8E267F6C9C82A1" authorityName="Zyuzin" authorityYear="1993" box="[1328,1436,881,904]" class="Arachnida" family="Lycosidae" genus="Oculicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF903F42FA8E267F6C9C82A1" box="[1328,1436,881,904]" italics="true" pageId="4" pageNumber="278">Oculicosa</emphasis>
</taxonomicName>
(see
<bibRefCitation id="01454B70FF903F42FCC8269F6D0B8281" author="Zyuzin A. A." box="[886,1035,913,936]" pageId="4" pageNumber="278" pagination="693 - 700" refId="ref24274" refString="Zyuzin A. A. 1993. Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and new species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum 33 (2): 693 - 700." type="journal article" year="1993">Zyuzin, 1993</bibRefCitation>
;
<bibRefCitation id="01454B70FF903F42FBA2269F6C498281" author="Logunov D. V. &amp; Gromov A. V." box="[1052,1353,913,936]" pageId="4" pageNumber="278" pagination="48 - 51" refId="ref22491" refString="Logunov D. V., Gromov A. V. 2011. Notes on the distribution of Oculicosa supermirabilis (Araneae, Lycosidae). Arachnologische Mitteilungen 42: 48 - 51." type="journal article" year="2011">Logunov &amp; Gromov, 2011</bibRefCitation>
: fig. 6) and
<taxonomicName id="A2D44D02FF903F42FCCB26BF6AE082E1" authorityName="Logunov" authorityYear="2010" box="[885,992,945,968]" class="Arachnida" family="Lycosidae" genus="Zyuzicosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="57A0EA93FF903F42FCCB26BF6AE082E1" box="[885,992,945,968]" italics="true" pageId="4" pageNumber="278">Zyuzicosa</emphasis>
</taxonomicName>
(see
<bibRefCitation id="01454B70FF903F42FB9626BF6DCF82E1" author="Logunov D. V." box="[1064,1231,945,968]" pageId="4" pageNumber="278" pagination="233 - 263" refId="ref22377" refString="Logunov D. V. 2010. On new central Asian genus and species of wolf spiders (Araneae: Lycosidae) exhibiting a pronounced sexual size dimorphism. Proceedings of the Zoological Institute RAS 314 (3): 233 - 263." type="journal article" year="2010">Logunov 2010</bibRefCitation>
,
<bibRefCitation id="01454B70FF903F42FB5C26BF6C1B82E1" author="Logunov D. V." box="[1250,1307,945,968]" pageId="4" pageNumber="278" pagination="349 - 362" refId="ref22421" refString="Logunov D. V. 2012. A synopsis of the genus Zyuzicosa Logunov, 2010 (Aranei: Lycosidae). Arthropoda Selecta 21 (4): 349 - 362." type="journal article" year="2012">2012</bibRefCitation>
: map), but their ranges are markedly smaller and of a different configuration.
<taxonomicName id="A2D44D02FF903F42FC5A26FF6D6C8521" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[996,1132,1009,1032]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FC5A26FF6D6C8521" box="[996,1132,1009,1032]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FBC526FF6DE58521" box="[1147,1253,1009,1032]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
seems to be a typical Turan genus (
<emphasis id="57A0EA93FF903F42FBA2211F6D578501" box="[1052,1111,1041,1064]" italics="true" pageId="4" pageNumber="278">sensu</emphasis>
<bibRefCitation id="01454B70FF903F42FBE3211F6C958501" author="Pravdin F. N. &amp; Mishchenko L. L." box="[1117,1429,1041,1064]" pageId="4" pageNumber="278" refId="ref23412" refString="Pravdin F. N., Mishchenko L. L. 1980. Formation and evolution of ecological insect faunas in Middle Asia. Nauka, Moscow, 156 pp. [In Russian]" type="book" year="1980">Pravdin &amp; Mishchenko, 1980</bibRefCitation>
) and an endemic to Central Asia; its geographical range lies within the Turan lowlands of the desert zone. We are unaware of any other Central Asian spider genus that has a distributional pattern similar to that of
<taxonomicName id="A2D44D02FF903F42FAAA219F6C9C8581" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1300,1436,1169,1192]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FAAA219F6C9C8581" box="[1300,1436,1169,1192]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FC8121BF6AA185E1" box="[831,929,1201,1224]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="31AB6609FF903F42FF292D096BFA8935" ID-DOI="http://doi.org/10.5281/zenodo.5743712" ID-Zenodo-Dep="5743712" box="[151,762,2055,2076]" httpUri="https://zenodo.org/record/5743712/files/figure.png" pageId="4" pageNumber="278" startId="4.[151,186,2055,2076]" targetBox="[155,1435,1347,2031]" targetPageId="4">
<paragraph id="656B3681FF903F42FF292D096BFA8935" blockId="4.[151,762,2055,2076]" box="[151,762,2055,2076]" pageId="4" pageNumber="278">
Fig. 1. Geographical range of the genus
<taxonomicName id="A2D44D02FF903F42FD9F2D096B9C8935" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[545,668,2055,2076]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF903F42FD9F2D096B9C8935" box="[545,668,2055,2076]" italics="true" pageId="4" pageNumber="278">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF903F42FD1F2D096BFA8935" box="[673,762,2055,2076]" pageId="4" pageNumber="278" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="2DCE650AFF913F40FF29240F6C9B8330" lastPageId="6" lastPageNumber="280" pageId="5" pageNumber="279" type="key">
<paragraph id="656B3681FF913F43FF29240F6B7D8031" blockId="5.[151,1055,256,312]" box="[151,637,256,280]" pageId="5" pageNumber="279">
<emphasis id="57A0EA93FF913F43FF29240F6B7D8031" bold="true" box="[151,637,256,280]" pageId="5" pageNumber="279">
Key to the species of
<taxonomicName id="A2D44D02FF913F43FE3B240E6B128031" ID-CoL="98X5T" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[389,530,256,280]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="57A0EA93FF913F43FE3B240E6B128031" bold="true" box="[389,530,256,280]" italics="true" pageId="5" pageNumber="279">Karakumosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FDA6240F6B7D8031" box="[536,637,257,280]" pageId="5" pageNumber="279" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="656B3681FF913F43FF29242F6D1F8011" blockId="5.[151,1055,256,312]" box="[151,1055,289,312]" pageId="5" pageNumber="279">
The female of
<taxonomicName id="A2D44D02FF913F43FE8D242F68C28011" ID-CoL="99B5D" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[307,450,289,312]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="tashkumyr" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FE8D242F68C28011" box="[307,450,289,312]" italics="true" pageId="5" pageNumber="279">K. tashkumyr</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FE76242F6B1B8011" box="[456,539,289,312]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
remains unknown and thus is not included here.
</paragraph>
<key id="114561B0FF913F40FF29246F6C9B8330" lastPageId="6" lastPageNumber="280" pageId="5" pageNumber="279">
<keyStep id="DE202E24FF913F43FF29246F6C9A80B1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29246F6C9A8051" blockId="5.[151,1434,353,408]" box="[151,1434,353,376]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29246F6C9A8051" box="[151,1434,353,376]" pageId="5" pageNumber="279">1A Males ........................................................................................................................................................................ 2</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29248F6C9A80B1" blockId="5.[151,1434,353,408]" box="[151,1434,385,408]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29248F6C9A80B1" box="[151,1434,385,408]" pageId="5" pageNumber="279">1B Females ..................................................................................................................................................................10</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF2924CF6C9A80D1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF2924CF6C9A80F1" blockId="5.[151,1434,449,504]" box="[151,1434,449,472]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2924CF6C9A80F1" box="[151,1434,449,472]" pageId="5" pageNumber="279">
2A Proximal extension of median apophysis clearly hook-shaped (
<figureCitation id="FDEF2A04FF913F43FC3F24CF6AD480F1" box="[897,980,449,472]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">Figs 60</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC6124CF6D0880F1" box="[991,1032,449,472]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="5" pageNumber="279">118</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FBAD24CF6D3C80F1" box="[1043,1084,449,472]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="5" pageNumber="279">150</figureCitation>
) ................................................ 3
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2924EF6C9A80D1" blockId="5.[151,1434,449,504]" box="[151,1434,481,504]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2924EF6C9A80D1" box="[151,1434,481,504]" pageId="5" pageNumber="279">
2B Proximal extension of median apophysis of a different shape (
<figureCitation id="FDEF2A04FF913F43FCC624EF6ABD80D1" box="[888,957,481,504]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="5" pageNumber="279">Figs 2</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC7624EF6AE580D1" box="[968,997,481,504]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">46</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC4E24EF6D0D80D1" box="[1008,1037,481,504]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">83</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FBA624EF6D3480D1" box="[1048,1076,481,504]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">98</figureCitation>
) ................................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29272F6C9B8351" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29272F6C9A8371" blockId="5.[151,1435,545,632]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29272F6C9A8371" pageId="5" pageNumber="279">
3A Median tooth of median apophysis with a low serrate flange at its foot (indicated by arrow in
<figureCitation id="FDEF2A04FF913F43FB79272F6C258311" box="[1223,1317,545,568]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="5" pageNumber="279">Figs 125</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FA8F272F6C5B8311" box="[1329,1371,545,568]" captionStart="Figs 126-135" captionStartId="25.[151,192,1838,1859]" captionTargetBox="[151,1435,257,1777]" captionTargetId="figure-9@25.[151,1436,257,1777]" captionTargetPageId="25" captionText="Figs 126-135. Karakumosa shmatkoi sp. nov.; male holotype (128, 132-133) and male paratype from Kalmykia, Utta Vil. (126-127, 129-131, 134-135). (126) Body, dorsal view. (127) Ditto, ventral view. (128, 134) Median tooth of MA, posterior view. (129) Embolic division. (130) Left male palp, ventral view. (131-132) Bulbus, ventral view. (133) Synembolus, apical view. (135) Bulbus, retrolateral view. Scale bars 1 cm (126-127), 0.5 mm (130-131, 135), 0.2 mm (132), 0.1 mm (128, 134)." figureDoi="http://doi.org/10.5281/zenodo.5743756" httpUri="https://zenodo.org/record/5743756/files/figure.png" pageId="5" pageNumber="279">134</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FAD8272F6C8F8311" box="[1382,1423,545,568]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="5" pageNumber="279">152</figureCitation>
) .................................................................................................................................................................................. 4
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29276F6C9B8351" blockId="5.[151,1435,545,632]" box="[151,1435,609,632]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29276F6C9B8351" box="[151,1435,609,632]" pageId="5" pageNumber="279">
3B Median tooth of median apophysis without such flange (
<figureCitation id="FDEF2A04FF913F43FC85276F6A8E8351" box="[827,910,609,632]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">Figs 66</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC27276F6AD98351" box="[921,985,609,632]" captionStart="Figs 70-75" captionStartId="17.[151,192,1222,1243]" captionTargetBox="[160,1436,257,1181]" captionTargetId="figure-337@17.[151,1439,257,1191]" captionTargetPageId="17" captionText="Figs 70-75. Karakumosa gromovi sp. nov.; male paratypes (74-75) and female paratype (70-73). (70) Body, dorsal view. (71) Ditto, ventral view. (72) Epigyne, ventral view. (73) Vulva, dorsal view. (74-75) Median tooth of MA, posterior view. Scale bars 1 cm (70-71), 0.5 mm (72), 0.25 mm (73), 0.1 mm (74-75)." figureDoi="http://doi.org/10.5281/zenodo.5743738" httpUri="https://zenodo.org/record/5743738/files/figure.png" pageId="5" pageNumber="279">74-75</figureCitation>
) ......................................
<taxonomicName id="A2D44D02FF913F43FB57276F6C418351" ID-CoL="99CFD" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1257,1345,609,632]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="gromovi" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB57276F6C418351" box="[1257,1345,609,632]" italics="true" pageId="5" pageNumber="279">gromovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF6276F6C9B8351" box="[1352,1435,609,632]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF2927AF6C9B8211" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF2927AF6C9B83F1" blockId="5.[151,1437,673,824]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2927AF6C9B83F1" pageId="5" pageNumber="279">
4A Inner plate of median apophysis as wide as outer plate, clearly visible in ventral view (indicated by arrow in
<figureCitation id="FDEF2A04FF913F43FACC27AF680E83F1" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="5" pageNumber="279">Fig. 148</figureCitation>
); tips of both synembolic lamellae markedly bent downward (
<figureCitation id="FDEF2A04FF913F43FC1A27CF6AFF83F1" box="[932,1023,705,728]" captionStart="Figs 142-154" captionStartId="28.[151,192,1972,1993]" captionTargetBox="[169,1419,257,1942]" captionTargetId="figure-9@28.[169,1419,257,1943]" captionTargetPageId="28" captionText="Figs 142-154. Karakumosa tashkumyr sp. nov., male paratypes, in ISEA. (142) Body, ventral view. (143) Ditto, dorsal view. (144) Left chelicera, ventral view. (145, 147, 152-154) Median tooth of MA, posterior view. (146) Left male palp, ventral view. (148, 150) Bulbus, ventral view. (151) Ditto, retrolateral view. (149) Embolic division. Scale bars 1 cm (142-143), 0.5 cm (144, 146, 150-151), 0.2 mm (148), 0.1 mm (145, 147, 149, 152-154)." figureDoi="http://doi.org/10.5281/zenodo.5743760" httpUri="https://zenodo.org/record/5743760/files/figure.png" pageId="5" pageNumber="279">Fig. 149</figureCitation>
) .............................
<taxonomicName id="A2D44D02FF913F43FB6C27CF6C4183F1" ID-CoL="99B5D" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1234,1345,705,728]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="tashkumyr" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB6C27CF6C4183F1" box="[1234,1345,705,728]" italics="true" pageId="5" pageNumber="279">tashkumyr</emphasis>
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<taxonomicNameLabel id="4C9357E8FF913F43FAF627CF6C9B83F1" box="[1352,1435,705,728]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2927EF6C9B8211" blockId="5.[151,1437,673,824]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2927EF6C9B8211" pageId="5" pageNumber="279">
4B Inner plate of median apophysis comparatively narrower and almost hidden beneath outer plate (indicated by arrow in
<figureCitation id="FDEF2A04FF913F43FEFD260F689F8231" box="[323,415,769,792]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="5" pageNumber="279">Fig. 118</figureCitation>
); synembolic lamellae straight or with tips only slightly bent downward (
<figureCitation id="FDEF2A04FF913F43FB12260F6C068231" box="[1196,1286,769,792]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="5" pageNumber="279">Fig. 116</figureCitation>
) ................... ........................................................................................................................................................
<taxonomicName id="A2D44D02FF913F43FB5F262F6C418211" ID-CoL="992FK" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1249,1345,801,824]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB5F262F6C418211" box="[1249,1345,801,824]" italics="true" pageId="5" pageNumber="279">shmatkoi</emphasis>
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<taxonomicNameLabel id="4C9357E8FF913F43FAF6262F6C9B8211" box="[1352,1435,801,824]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29266F6C9B82B1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29266F6C9A8251" blockId="5.[151,1435,865,920]" box="[151,1434,865,888]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29266F6C9A8251" box="[151,1434,865,888]" pageId="5" pageNumber="279">
5A Proximal extension of median apophysis prominent (
<figureCitation id="FDEF2A04FF913F43FCA9266F6A6A8251" box="[791,874,865,888]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">Figs 46</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FCCB266F6A928251" box="[885,914,865,888]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">83</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC23266F6AB98251" box="[925,953,865,888]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">98</figureCitation>
) ................................................................... 6
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29268F6C9B82B1" blockId="5.[151,1435,865,920]" box="[151,1435,897,920]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29268F6C9B82B1" box="[151,1435,897,920]" pageId="5" pageNumber="279">
5B Proximal extension of median apophysis not prominent (
<figureCitation id="FDEF2A04FF913F43FCFE268F6A9B82B1" box="[832,923,897,920]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="5" pageNumber="279">Fig. 174</figureCitation>
) ..................................................
<taxonomicName id="A2D44D02FF913F43FB49268F6C4182B1" ID-CoL="99C72" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1271,1345,897,920]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB49268F6C4182B1" box="[1271,1345,897,920]" italics="true" pageId="5" pageNumber="279">zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF6268F6C9B82B1" box="[1352,1435,897,920]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF2926CF6C9A82D1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF2926CF6C9A82F1" blockId="5.[151,1434,961,1016]" box="[151,1434,961,984]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2926CF6C9A82F1" box="[151,1434,961,984]" pageId="5" pageNumber="279">
6A Proximal extension of median apophysis spade/spatula-like, as wide as long (
<figureCitation id="FDEF2A04FF913F43FBA526CF6D6E82F1" box="[1051,1134,961,984]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">Figs 46</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FBC726CF6D9582F1" box="[1145,1173,961,984]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">98</figureCitation>
) .................................. 7
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2926EF6C9A82D1" blockId="5.[151,1434,961,1016]" box="[151,1434,993,1016]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2926EF6C9A82D1" box="[151,1434,993,1016]" pageId="5" pageNumber="279">
6B Proximal extension of median apophysis wider than long (
<figureCitation id="FDEF2A04FF913F43FCEF26EF6A9682D1" box="[849,918,993,1016]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="5" pageNumber="279">Figs 2</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC1C26EF6ABF82D1" box="[930,959,993,1016]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">83</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC7426EF6AF382D1" box="[970,1011,993,1016]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">161</figureCitation>
) .......................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29212F6C9B8571" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29212F6C9B8511" blockId="5.[151,1435,1057,1112]" box="[151,1435,1057,1080]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29212F6C9B8511" box="[151,1435,1057,1080]" pageId="5" pageNumber="279">
7A Median tooth of median apophysis finger-shaped (
<figureCitation id="FDEF2A04FF913F43FD45212F6A4E8511" box="[763,846,1057,1080]" captionStart="Figs 30-38" captionStartId="11.[151,192,1966,1987]" captionTargetBox="[156,1432,694,1916]" captionTargetId="figure-222@11.[151,1436,684,1916]" captionTargetPageId="11" captionText="Figs 30-38. Karakumosa badkhyzica sp. nov.; male paratype (30-31, 37-38) and female paratype (32-35), MMUE. (30-31) Median tooth of MA, posterior view. (32) Female prosoma, frontal view. (33, 37) Body, ventral view. (34, 38) Ditto, dorsal view. (35) Female prosoma, lateral view. (36) Female tarsus I, ventral view. Scale bars 0.5 cm (33-35, 37-38), 0.3 mm (36), 0.25 cm (32), 0.1 mm (30, 31)." figureDoi="http://doi.org/10.5281/zenodo.5743726" httpUri="https://zenodo.org/record/5743726/files/figure.png" pageId="5" pageNumber="279">Figs 31</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FCE7212F6A758511" box="[857,885,1057,1080]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">42</figureCitation>
) .................................................
<taxonomicName id="A2D44D02FF913F43FB77212F6C418511" ID-CoL="99BDZ" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1225,1345,1057,1080]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="badkhyzica" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB77212F6C418511" box="[1225,1345,1057,1080]" italics="true" pageId="5" pageNumber="279">badkhyzica</emphasis>
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<taxonomicNameLabel id="4C9357E8FF913F43FAF6212F6C9B8511" box="[1352,1435,1057,1080]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29214F6C9B8571" blockId="5.[151,1435,1057,1112]" box="[151,1435,1089,1112]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29214F6C9B8571" box="[151,1435,1089,1112]" pageId="5" pageNumber="279">
7B Median tooth of median apophysis bifurcated at its tip (
<figureCitation id="FDEF2A04FF913F43FC8D214F6A8E8571" box="[819,910,1089,1112]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">Fig. 100</figureCitation>
) ...................................................
<taxonomicName id="A2D44D02FF913F43FB4A214F6C418571" ID-CoL="999W7" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1268,1345,1089,1112]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="repetek" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB4A214F6C418571" box="[1268,1345,1089,1112]" italics="true" pageId="5" pageNumber="279">repetek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF6214F6C9B8571" box="[1352,1435,1089,1112]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="DE202E24FF913F43FF29218F6C9A8591" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29218F6C9A85B1" blockId="5.[151,1434,1153,1208]" box="[151,1434,1153,1176]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29218F6C9A85B1" box="[151,1434,1153,1176]" pageId="5" pageNumber="279">
8A Median tooth of median apophysis flat, without a ventral bulge (
<figureCitation id="FDEF2A04FF913F43FC2E218F6AD585B1" box="[912,981,1153,1176]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="5" pageNumber="279">Figs 5</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC5F218F6D0A85B1" box="[993,1034,1153,1176]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">164</figureCitation>
) ....................................................... 9
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2921AF6C9A8591" blockId="5.[151,1434,1153,1208]" box="[151,1434,1185,1208]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2921AF6C9A8591" box="[151,1434,1185,1208]" pageId="5" pageNumber="279">
8B Median tooth of median apophysis with a ventral bulge (
<figureCitation id="FDEF2A04FF913F43FC8121AF6A8D8591" box="[831,909,1185,1208]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">Fig. 87</figureCitation>
) ................................................................
<taxonomicName id="A2D44D02FF913F43FAF321AF6C9A8591" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" baseAuthorityName="Pocock" baseAuthorityYear="1889" box="[1357,1434,1185,1208]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="medica">
<emphasis id="57A0EA93FF913F43FAF321AF6C9A8591" box="[1357,1434,1185,1208]" italics="true" pageId="5" pageNumber="279">medica</emphasis>
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<keyStep id="DE202E24FF913F43FF2921EF6C9A8471" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF2921EF6C9B8431" blockId="5.[151,1437,1249,1368]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2921EF6C9B8431" pageId="5" pageNumber="279">
9A Proximal extension of median apophysis twice as wide as long, with a markedly pointed prolaterad-directed shoulder (
<figureCitation id="FDEF2A04FF913F43FEF3200F68AA8431" box="[333,426,1281,1304]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">Fig. 161</figureCitation>
); median tooth quadrangular (
<figureCitation id="FDEF2A04FF913F43FD52200F6A478431" box="[748,839,1281,1304]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">Fig. 164</figureCitation>
) ...........................................................
<taxonomicName id="A2D44D02FF913F43FB59200F6C418431" ID-CoL="99B65" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1255,1345,1281,1304]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="turanica" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB59200F6C418431" box="[1255,1345,1281,1304]" italics="true" pageId="5" pageNumber="279">turanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF6200F6C9B8431" box="[1352,1435,1281,1304]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29202F6C9A8471" blockId="5.[151,1437,1249,1368]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29202F6C9A8471" pageId="5" pageNumber="279">
9B Proximal extension of median apophysis triangular, with an obtuse prolaterad-directed shoulder (
<figureCitation id="FDEF2A04FF913F43FB55202F6C398411" box="[1259,1337,1313,1336]" captionStart="Figs 12-21" captionStartId="8.[151,192,1900,1921]" captionTargetBox="[161,1426,261,1863]" captionTargetId="figure-9@8.[151,1436,257,1863]" captionTargetPageId="8" captionText="Figs 12-21. Karakumosa alticeps (Kroneberg, 1875); male specimens from Tartogay Vil. (12-15) and Zhankel’ Vil. (16), and the male lectotype (17-21). (12, 17) Left bulbus, ventral view. (13, 20) Ditto, retrolateral view. (14, 19) Ditto, posterior view. (15, 16) Left male palp, retrolateral view. (18) Left bulbus, apical view. (21) Original data labels of the lectotype. Scale bars 0.5 mm (15, 16), 0.2 mm (12-14, 17-20). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743720" httpUri="https://zenodo.org/record/5743720/files/figure.png" pageId="5" pageNumber="279">Fig. 12</figureCitation>
); median tooth triangular, with a serrate prolateral edge (
<figureCitation id="FDEF2A04FF913F43FD66204F6A318471" box="[728,817,1345,1368]" captionStart="Figs 2-11" captionStartId="7.[151,192,1432,1453]" captionTargetBox="[151,1435,257,1390]" captionTargetId="figure-257@7.[151,1436,257,1390]" captionTargetPageId="7" captionText="Figs 2-11. Karakumosa alticeps (Kroneberg, 1875), left male palp; specimens from Chu river valley (2-4, 7-11) and Karamola Mt. (5- 6). (2, 9) Bulbus, ventral view. (3, 11) Ditto, retrolateral view. (4) Embolic division, ventral view. (5-8) Median tooth of MA, proximal view. (10) Bulbus, apical view. Scale bars 0.25 mm (2-3), 0.1 mm (4-11). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743716" httpUri="https://zenodo.org/record/5743716/files/figure.png" pageId="5" pageNumber="279">Figs 5-8</figureCitation>
) ............................................................................
<taxonomicName id="A2D44D02FF913F43FAF9204F6C9A8471" ID-CoL="3M6FY" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[1351,1434,1345,1368]" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="alticeps">
<emphasis id="57A0EA93FF913F43FAF9204F6C9A8471" box="[1351,1434,1345,1368]" italics="true" pageId="5" pageNumber="279">alticeps</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29208F6C9A84F1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29208F6C9A84B1" blockId="5.[151,1434,1409,1496]" box="[151,1434,1409,1432]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29208F6C9A84B1" box="[151,1434,1409,1432]" pageId="5" pageNumber="279">
10A Edges of epigynal atrium subparallel or slightly bent outward (
<figureCitation id="FDEF2A04FF913F43FC3E208F6AD384B1" box="[896,979,1409,1432]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">Figs 44</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC60208F6AFB84B1" box="[990,1019,1409,1432]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">61</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FBB8208F6D2384B1" box="[1030,1059,1409,1432]" captionStart="Figs 70-75" captionStartId="17.[151,192,1222,1243]" captionTargetBox="[160,1436,257,1181]" captionTargetId="figure-337@17.[151,1439,257,1191]" captionTargetPageId="17" captionText="Figs 70-75. Karakumosa gromovi sp. nov.; male paratypes (74-75) and female paratype (70-73). (70) Body, dorsal view. (71) Ditto, ventral view. (72) Epigyne, ventral view. (73) Vulva, dorsal view. (74-75) Median tooth of MA, posterior view. Scale bars 1 cm (70-71), 0.5 mm (72), 0.25 mm (73), 0.1 mm (74-75)." figureDoi="http://doi.org/10.5281/zenodo.5743738" httpUri="https://zenodo.org/record/5743738/files/figure.png" pageId="5" pageNumber="279">72</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FB90208F6D4A84B1" box="[1070,1098,1409,1432]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">85</figureCitation>
) ........................................... 11
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2920AF6C9A84F1" blockId="5.[151,1434,1409,1496]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2920AF6C9A84F1" pageId="5" pageNumber="279">
10B Edges of epigynal atrium slanted to each other; atrium anteriorly clearly narrower than posteriorly (
<figureCitation id="FDEF2A04FF913F43FAB920AF6C5A8491" box="[1287,1370,1441,1464]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="5" pageNumber="279">Figs 27</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FADB20AF6C818491" box="[1381,1409,1441,1464]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">96</figureCitation>
).. ................................................................................................................................................................................ 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29230F6C9B8771" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29230F6C9A8731" blockId="5.[151,1437,1537,1624]" box="[151,1434,1537,1560]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29230F6C9A8731" box="[151,1434,1537,1560]" pageId="5" pageNumber="279">
11A Edges of epigynal atrium subparallel (
<figureCitation id="FDEF2A04FF913F43FDC7230F6BCC8731" box="[633,716,1537,1560]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">Figs 61</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FD69230F6BF48731" box="[727,756,1537,1560]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">85</figureCitation>
), spermathecae as in
<figureCitation id="FDEF2A04FF913F43FC66230F6D2A8731" box="[984,1066,1537,1560]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">Figs 62</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FB8B230F6D508731" box="[1077,1104,1537,1560]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">86</figureCitation>
............................................ 12
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29232F6C9B8771" blockId="5.[151,1437,1537,1624]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29232F6C9B8771" pageId="5" pageNumber="279">
11B Edges of epigynal atrium slightly bent outward (biconvex), atrium barrel-shaped (
<figureCitation id="FDEF2A04FF913F43FBFA232F6D938711" box="[1092,1171,1569,1592]" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">Fig. 44</figureCitation>
), spermathecae as in
<figureCitation id="FDEF2A04FF913F43FACC232F69FE8771" captionStart="Figs 39-48" captionStartId="12.[151,192,1896,1917]" captionTargetBox="[151,1435,257,1846]" captionTargetId="figure-9@12.[151,1436,257,1846]" captionTargetPageId="12" captionText="Figs 39-48. Karakumosa badkhyzica sp. nov.; male paratype, MMUE (39-42), male holotype (45-48) and female paratype (43-44). (39-40, 46) Bulbus, ventral view. (41) Ditto, retrolateral view. (42, 48) Median tooth of MA, posterior view. (43) Vulva, dorsal view. (44) Epigyne, ventral view. (45) Cymbium tip, dorsal view. (47) Bulbus, apical view. Scale bars 0.5 mm (39, 44), 0.25 mm (40-41, 43), 0.2 mm (45-47), 0.1 mm (42), 0.05 mm (48). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743728" httpUri="https://zenodo.org/record/5743728/files/figure.png" pageId="5" pageNumber="279">Fig. 43</figureCitation>
.................................................................................................................................................
<taxonomicName id="A2D44D02FF913F43FB77234F6C418771" ID-CoL="99BDZ" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1225,1345,1601,1624]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="badkhyzica" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB77234F6C418771" box="[1225,1345,1601,1624]" italics="true" pageId="5" pageNumber="279">badkhyzica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF6234F6C9B8771" box="[1352,1435,1601,1624]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29238F6C9A87D1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29238F6C9B8791" blockId="5.[151,1437,1665,1784]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29238F6C9B8791" pageId="5" pageNumber="279">
12A Posterior transverse plate of epigyne in the shape of an inverted triangle (
<figureCitation id="FDEF2A04FF913F43FBB7238F6D5E87B1" box="[1033,1118,1665,1688]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">Figs 61</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FBD5238F6D8987B1" box="[1131,1161,1665,1688]" captionStart="Figs 70-75" captionStartId="17.[151,192,1222,1243]" captionTargetBox="[160,1436,257,1181]" captionTargetId="figure-337@17.[151,1439,257,1191]" captionTargetPageId="17" captionText="Figs 70-75. Karakumosa gromovi sp. nov.; male paratypes (74-75) and female paratype (70-73). (70) Body, dorsal view. (71) Ditto, ventral view. (72) Epigyne, ventral view. (73) Vulva, dorsal view. (74-75) Median tooth of MA, posterior view. Scale bars 1 cm (70-71), 0.5 mm (72), 0.25 mm (73), 0.1 mm (74-75)." figureDoi="http://doi.org/10.5281/zenodo.5743738" httpUri="https://zenodo.org/record/5743738/files/figure.png" pageId="5" pageNumber="279">72</figureCitation>
), spermathecae markedly widened anteriorly (
<figureCitation id="FDEF2A04FF913F43FE0423AF6B0D8791" box="[442,525,1697,1720]" captionStart="Figs 59-69" captionStartId="16.[153,194,1688,1709]" captionTargetBox="[151,1435,257,1621]" captionTargetId="figure-9@16.[151,1436,257,1621]" captionTargetPageId="16" captionText="Figs 59-69. Karakumosa gromovi sp. nov.; male paratypes (59-60, 64-69) and paratype female (61-63). (59-60) Bulbus, ventral view. (61) Epigyne, ventral view. (62) Vulva, dorsal view. (63) Female tarsus I, ventral view. (64, 66) Median tooth of MA, posterior view. (65) Cymbium, ventral view. (67) Embolic division, ventral view. (68) Bulbus, retrolateral view. (69) Cymbium tip, dorsal view.Scale bars 0.5 mm (59, 61), 0.3 mm (63, 65), 0.25 mm (62), 0.1 mm (60, 66-69), 0.05 mm (64). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743736" httpUri="https://zenodo.org/record/5743736/files/figure.png" pageId="5" pageNumber="279">Figs 62</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FDA623AF6B348791" box="[536,564,1697,1720]" captionStart="Figs 70-75" captionStartId="17.[151,192,1222,1243]" captionTargetBox="[160,1436,257,1181]" captionTargetId="figure-337@17.[151,1439,257,1191]" captionTargetPageId="17" captionText="Figs 70-75. Karakumosa gromovi sp. nov.; male paratypes (74-75) and female paratype (70-73). (70) Body, dorsal view. (71) Ditto, ventral view. (72) Epigyne, ventral view. (73) Vulva, dorsal view. (74-75) Median tooth of MA, posterior view. Scale bars 1 cm (70-71), 0.5 mm (72), 0.25 mm (73), 0.1 mm (74-75)." figureDoi="http://doi.org/10.5281/zenodo.5743738" httpUri="https://zenodo.org/record/5743738/files/figure.png" pageId="5" pageNumber="279">73</figureCitation>
) .....................................................................................................
<taxonomicName id="A2D44D02FF913F43FB5723AF6C418791" ID-CoL="99CFD" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1257,1345,1697,1720]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="gromovi" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB5723AF6C418791" box="[1257,1345,1697,1720]" italics="true" pageId="5" pageNumber="279">gromovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF623AF6C9B8791" box="[1352,1435,1697,1720]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2923CF6C9A87D1" blockId="5.[151,1437,1665,1784]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2923CF6C9A87D1" pageId="5" pageNumber="279">
12B Posterior transverse plate of epigyne anchor-shaped (
<figureCitation id="FDEF2A04FF913F43FCA823CF6A6587F1" box="[790,869,1729,1752]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">Fig. 85</figureCitation>
), spermathecae not widened anteriorly, worm-shaped (
<figureCitation id="FDEF2A04FF913F43FF5523EF683987D1" box="[235,313,1761,1784]" captionStart="Figs 83-88" captionStartId="20.[151,192,1303,1324]" captionTargetBox="[154,1435,257,1238]" captionTargetId="figure-325@20.[151,1436,257,1258]" captionTargetPageId="20" captionText="Figs 83-88. Karakumosa medica (Pocock, 1889); male lectotype (83-84, 87-88) and female paralectotype (85-86). (83) Bulbus, ventral view. (84) Ditto, retrolateral view. (85) Epigyne, ventral view. (86) Vulva, dorsal view. (87) Median tooth of MA, posterior view. (88) Synembolus, apical view. Scale bars 0.5 mm (83-86), 0.1 mm (87-88)." figureDoi="http://doi.org/10.5281/zenodo.5743746" httpUri="https://zenodo.org/record/5743746/files/figure.png" pageId="5" pageNumber="279">Fig. 86</figureCitation>
) .........................................................................................................................................................
<taxonomicName id="A2D44D02FF913F43FAF323EF6C9A87D1" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" baseAuthorityName="Pocock" baseAuthorityYear="1889" box="[1357,1434,1761,1784]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="medica">
<emphasis id="57A0EA93FF913F43FAF323EF6C9A87D1" box="[1357,1434,1761,1784]" italics="true" pageId="5" pageNumber="279">medica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF29222F6C9B8651" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF29222F6C9A8671" blockId="5.[151,1436,1825,1912]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29222F6C9A8671" pageId="5" pageNumber="279">
13A Posterior transverse plate more or less anchor-shaped (
<figureCitation id="FDEF2A04FF913F43FC8C222F6A868611" box="[818,902,1825,1848]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="5" pageNumber="279">Figs 27</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FC2C222F6AEE8611" box="[914,1006,1825,1848]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">167-168</figureCitation>
), spermathecae curved anticlockwise or subparallel (
<figureCitation id="FDEF2A04FF913F43FED8224F68B98671" box="[358,441,1857,1880]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="5" pageNumber="279">Figs 28</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FE7A224F68EE8671" box="[452,494,1857,1880]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="5" pageNumber="279">121</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FE47224F6B228671" box="[505,546,1857,1880]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">166</figureCitation>
) ..............................................................................................................................14
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF29226F6C9B8651" blockId="5.[151,1436,1825,1912]" box="[151,1435,1889,1912]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF29226F6C9B8651" box="[151,1435,1889,1912]" pageId="5" pageNumber="279">
13B Posterior transverse plate dumbbell-shaped (
<figureCitation id="FDEF2A04FF913F43FD03226F6A0D8651" box="[701,781,1889,1912]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">Fig. 96</figureCitation>
), spermathecae curved clockwise (
<figureCitation id="FDEF2A04FF913F43FB3E226F6DCE8651" box="[1152,1230,1889,1912]" captionStart="Figs 93-102" captionStartId="22.[151,192,1860,1881]" captionTargetBox="[151,1435,257,1809]" captionTargetId="figure-9@22.[151,1436,257,1809]" captionTargetPageId="22" captionText="Figs 93-102. Karakumosa repetek sp. nov.; male holotype (97-98), male paratype (93-94, 99-102) and female paratype (95-96). (93-94, 98). Bulbus, ventral view. (95) Vulva, dorsal view. (96) Epigyne, ventral view. (97) Median tooth of MA, ventral view. (99) Embolic division, ventral view. (100) Median tooth of MA, posterior view. (101) Embolic division, apical view. (102) Bulbus, retrolateral view. Scale bars 0.5 mm (93, 96), 0.25 mm (94, 102), 0.2 mm (98), 0.1 mm (95, 97, 99-101). Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743750" httpUri="https://zenodo.org/record/5743750/files/figure.png" pageId="5" pageNumber="279">Fig. 95</figureCitation>
) ...
<taxonomicName id="A2D44D02FF913F43FB4A226F6C418651" ID-CoL="999W7" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1268,1345,1889,1912]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="repetek" status="sp. nov.">
<emphasis id="57A0EA93FF913F43FB4A226F6C418651" box="[1268,1345,1889,1912]" italics="true" pageId="5" pageNumber="279">repetek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF913F43FAF6226F6C9B8651" box="[1352,1435,1889,1912]" pageId="5" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF913F43FF2922AF6C9A86F1" pageId="5" pageNumber="279">
<paragraph id="656B3681FF913F43FF2922AF6C9A8691" blockId="5.[151,1434,1953,2008]" box="[151,1434,1953,1976]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2922AF6C9A8691" box="[151,1434,1953,1976]" pageId="5" pageNumber="279">
14A Epigynal atrium markedly narrowed at its anterior end, pawn-shaped (
<figureCitation id="FDEF2A04FF913F43FC7122AF6D228691" box="[975,1058,1953,1976]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="5" pageNumber="279">Figs 27</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FB9322AF6D568691" box="[1069,1110,1953,1976]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="5" pageNumber="279">122</figureCitation>
) .......................................... 15
</keyLead>
</paragraph>
<paragraph id="656B3681FF913F43FF2922CF6C9A86F1" blockId="5.[151,1434,1953,2008]" box="[151,1434,1985,2008]" pageId="5" pageNumber="279">
<keyLead id="DE2595B4FF913F43FF2922CF6C9A86F1" box="[151,1434,1985,2008]" pageId="5" pageNumber="279">
14B Epigynal atrium much less narrower at its anterior end, with almost subparallel lateral edges (
<figureCitation id="FDEF2A04FF913F43FB7622CF6C2886F1" box="[1224,1320,1985,2008]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="5" pageNumber="279">Figs 167</figureCitation>
,
<figureCitation id="FDEF2A04FF913F43FA8A22CF6C5D86F1" box="[1332,1373,1985,2008]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="5" pageNumber="279">179</figureCitation>
) .. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF923F40FF29240C6C9A8050" pageId="6" pageNumber="280">
<paragraph id="656B3681FF923F40FF29240C6C9B8010" blockId="6.[151,1437,258,377]" pageId="6" pageNumber="280">
<keyLead id="DE2595B4FF923F40FF29240C6C9B8010" pageId="6" pageNumber="280">
15A Posterior transverse plate of epigyne in the shape of a low inverted triangle, epigynal edges strongly sigmoid (
<figureCitation id="FDEF2A04FF923F40FACF240C680F8010" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="6" pageNumber="280">Fig. 122</figureCitation>
); spermathecae markedly swollen in anterior portion (
<figureCitation id="FDEF2A04FF923F40FCF1242C6AAA8010" box="[847,938,290,313]" captionStart="Figs 115-125" captionStartId="24.[151,192,1685,1706]" captionTargetBox="[151,1435,257,1636]" captionTargetId="figure-9@24.[151,1436,257,1636]" captionTargetPageId="24" captionText="Figs 115-125. Karakumosa shmatkoi sp. nov.; male paratypes (115-120, 123-125) and female paratype (121-122); specimens from Tyub-Karagan Peninsula (115-116, 118, 120, 123, 125), Island Kulaly (117), Zhuzguntyube stow (121-122) and Tengiz (119, 124). (115, 118-119) Bulbus, ventral view. (116) Embolic division. (117) Left male palp, ventral view. (120) Bulbus, retrolateral view. (121) Vulva, dorsal view. (122) Epigyne, ventral view. (123-125) Median tooth of MA, posterior view. Scale bars 0.25 mm (115, 117, 120, 122), 0.2 mm (118-119), 0.1 mm (116, 121, 123-125).Abbreviations as explained in Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5743754" httpUri="https://zenodo.org/record/5743754/files/figure.png" pageId="6" pageNumber="280">Fig. 121</figureCitation>
) ...........................................
<taxonomicName id="A2D44D02FF923F40FB5F242C6C418010" ID-CoL="992FK" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1249,1345,290,313]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="280" phylum="Arthropoda" rank="species" species="shmatkoi" status="sp. nov.">
<emphasis id="57A0EA93FF923F40FB5F242C6C418010" box="[1249,1345,290,313]" italics="true" pageId="6" pageNumber="280">shmatkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF923F40FAF6242C6C9B8010" box="[1352,1435,290,313]" pageId="6" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="656B3681FF923F40FF29244C6C9A8050" blockId="6.[151,1437,258,377]" pageId="6" pageNumber="280">
<keyLead id="DE2595B4FF923F40FF29244C6C9A8050" pageId="6" pageNumber="280">
15B Posterior transverse plate of epigyne slightly procurved,epigynal edges indistinctly sigmoid (
<figureCitation id="FDEF2A04FF923F40FB13244C6DFA8070" box="[1197,1274,322,345]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="6" pageNumber="280">Fig. 27</figureCitation>
); spermathecae not swollen anteriorly (
<figureCitation id="FDEF2A04FF923F40FE65246C6B508050" box="[475,592,354,377]" captionStart="Figs 22-29" captionStartId="9.[157,198,1809,1830]" captionTargetBox="[154,1430,261,1756]" captionTargetId="figure-9@9.[151,1436,257,1764]" captionTargetPageId="9" captionText="Figs 22-29. Karakumosa alticeps (Kroneberg, 1875); male (22-23) and female (24-28) from Chu river valley, and female (29) from Karamola Mt. (22, 24) Body, dorsal view. (23, 25) Ditto, ventral view. (26) Ditto, lateral view. (27) Epigyne, ventral view. (28-29) Vulva, dorsal view. Scale bars 1 cm (22-26), 0.25 mm (27-29)." figureDoi="http://doi.org/10.5281/zenodo.5743722" httpUri="https://zenodo.org/record/5743722/files/figure.png" pageId="6" pageNumber="280">Figs 28-29</figureCitation>
) ..............................................................................................................
<taxonomicName id="A2D44D02FF923F40FAF9246C6C9A8050" ID-CoL="3M6FY" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[1351,1434,354,377]" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="280" phylum="Arthropoda" rank="species" species="alticeps">
<emphasis id="57A0EA93FF923F40FAF9246C6C9A8050" box="[1351,1434,354,377]" italics="true" pageId="6" pageNumber="280">alticeps</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE202E24FF923F40FF2924AC6C9B8330" pageId="6" pageNumber="280">
<paragraph id="656B3681FF923F40FF2924AC6C9B80F0" blockId="6.[151,1437,418,537]" pageId="6" pageNumber="280">
<keyLead id="DE2595B4FF923F40FF2924AC6C9B80F0" pageId="6" pageNumber="280">
16A Posterior transverse plate of epigyne with straight posterior margin (
<figureCitation id="FDEF2A04FF923F40FC7224AC6D2C8090" box="[972,1068,418,441]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="6" pageNumber="280">Fig. 179</figureCitation>
); spermathecae distinctly inclined towards each other (
<figureCitation id="FDEF2A04FF923F40FE0524CC6B1680F0" box="[443,534,450,473]" captionStart="Figs 173-186" captionStartId="34.[151,192,1877,1898]" captionTargetBox="[151,1435,257,1821]" captionTargetId="figure-9@34.[151,1436,257,1827]" captionTargetPageId="34" captionText="Figs 173-186. Karakumosa zyuzini sp. nov., male paratype (173-176, 178, 180-181, 183-186) and female paratype (177, 179, 182), in MMUE. (173-174, 176) Bulbus, ventral view. (175) Embolic division. (177) Left female chelicera, ventral view. (178) Bulbus, retrolateral view. (179) Epigyne, ventral view. (180-181, 183-186) Median tooth of MA, posterior view. (182) Vulva, dorsal view. Scale bars 0.5 mm (173, 176-179), 0.25 mm (182), 0.2 (174), 0.1 mm (175, 180-181, 183-186)." figureDoi="http://doi.org/10.5281/zenodo.5743772" httpUri="https://zenodo.org/record/5743772/files/figure.png" pageId="6" pageNumber="280">Fig. 182</figureCitation>
) ............................................................................................................
<taxonomicName id="A2D44D02FF923F40FB4924CC6C4180F0" ID-CoL="99C72" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1271,1345,450,473]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="280" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="57A0EA93FF923F40FB4924CC6C4180F0" box="[1271,1345,450,473]" italics="true" pageId="6" pageNumber="280">zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF923F40FAF624CC6C9B80F0" box="[1352,1435,450,473]" pageId="6" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="656B3681FF923F40FF2924EC6C9B8330" blockId="6.[151,1437,418,537]" pageId="6" pageNumber="280">
<keyLead id="DE2595B4FF923F40FF2924EC6C9B8330" pageId="6" pageNumber="280">
16B Posterior transverse plate of epigyne anchor-shaped, with convex posterior margin (
<figureCitation id="FDEF2A04FF923F40FBE024EC6DBA80D0" box="[1118,1210,482,505]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="6" pageNumber="280">Fig. 167</figureCitation>
); spermathecae more or less parallel to each other (
<figureCitation id="FDEF2A04FF923F40FDA3270C6B788330" box="[541,632,514,537]" captionStart="Figs 161-168" captionStartId="32.[151,192,1613,1634]" captionTargetBox="[159,1435,257,1543]" captionTargetId="figure-157@32.[151,1436,257,1551]" captionTargetPageId="32" captionText="Figs 161-168. Karakumosa turanica sp. nov., male holotype (161-164), female paratype (166-167) and paralectotype of Tarentula alticeps newly identified as K. turanica (165, 168). (161) Bulbus, ventral view. (162) Ditto, retrolateral view. (163) Synembolus, apical view. (164) Median tooth of MA, posterior view. (165-166) Vulva, dorsal view. (167-168) Epigyne, ventral view. Scale bars 0.25 mm (161-162, 166), 0.2 mm (165, 167-168), 0.1 mm (163-164)." figureDoi="http://doi.org/10.5281/zenodo.5743768" httpUri="https://zenodo.org/record/5743768/files/figure.png" pageId="6" pageNumber="280">Fig. 166</figureCitation>
) ..........................................................................................
<taxonomicName id="A2D44D02FF923F40FB59270C6C418330" ID-CoL="99B65" authority="Logunov &amp; Ponomarev, 2020" authorityName="Logunov &amp; Ponomarev" authorityYear="2020" box="[1255,1345,514,537]" class="Arachnida" family="Lycosidae" genus="Karakumosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="280" phylum="Arthropoda" rank="species" species="turanica" status="sp. nov.">
<emphasis id="57A0EA93FF923F40FB59270C6C418330" box="[1255,1345,514,537]" italics="true" pageId="6" pageNumber="280">turanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4C9357E8FF923F40FAF6270C6C9B8330" box="[1352,1435,514,537]" pageId="6" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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