216 lines
33 KiB
XML
216 lines
33 KiB
XML
<document id="5EFD365C90342C1E5C68CC3B2AA69AA1" ID-DOI="10.11646/zootaxa.2765.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5290957" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629712688807" checkinUser="felipe" docAuthor="Prieto-Márquez, Albert" docDate="2011" docId="E34487A0FF8FE7372C86FDC1FB56F84B" docLanguage="en" docName="zt02765p068.pdf" docOrigin="Zootaxa 2765 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2765.1.6" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Claosaurus agilis Marsh 1872" docType="treatment" docVersion="4" lastPageNumber="62" masterDocId="1F7DFFD8FF8EE7362C11FFD4FF9DFF97" masterDocTitle="Revised diagnoses of Hadrosaurus foulkii Leidy, 1858 (the type genus and species of Hadrosauridae Cope, 1869) and Claosaurus agilis Marsh, 1872 (Dinosauria: Ornithopoda) from the Late Cretaceous of North America" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1699100317202" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C51868052CD6983FCF40A2CD8FC67BB2">Revised diagnoses of Hadrosaurus foulkii Leidy, 1858 (the type genus and species of Hadrosauridae Cope, 1869) and Claosaurus agilis Marsh, 1872 (Dinosauria: Ornithopoda) from the Late Cretaceous of North America</mods:title>
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<mods:namePart id="DBE99FAEC71015391AF3DD569E3B6D1F">Prieto-Márquez, Albert</mods:namePart>
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<mods:date id="89E5B34561496EC64825DCBA0879E51B">2011</mods:date>
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<mods:number id="0648804A9A2D388CFE26F6B95BE5400F">2011-02-15</mods:number>
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<mods:number id="034FE00A50453FBC8259FC39632193FC">2765</mods:number>
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<treatment id="E34487A0FF8FE7372C86FDC1FB56F84B" ID-DOI="http://doi.org/10.5281/zenodo.5294866" ID-Zenodo-Dep="5294866" LSID="urn:lsid:plazi:treatment:E34487A0FF8FE7372C86FDC1FB56F84B" httpUri="http://treatment.plazi.org/id/E34487A0FF8FE7372C86FDC1FB56F84B" lastPageNumber="62" pageId="1" pageNumber="62">
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<subSubSection id="23F7653DFF8FE7372C86FDC1FE76FDBB" box="[151,491,532,557]" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="6B5236B6FF8FE7372C86FDC1FE76FDBB" blockId="1.[151,491,532,591]" box="[151,491,532,557]" pageId="1" pageNumber="62">
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<heading id="301A81DAFF8FE7372C86FDC1FE76FDBB" bold="true" box="[151,491,532,557]" fontSize="10" level="2" pageId="1" pageNumber="62" reason="2">
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<taxonomicName id="ACED4D35FF8FE7372C86FDC1FE76FDBB" authority="Marsh, 1872" authorityName="Marsh" authorityYear="1872" box="[151,491,532,557]" class="Reptilia" family="Hadrosauridae" genus="Claosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="agilis">
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<emphasis id="5999EAA4FF8FE7372C86FDC1FE76FDBB" bold="true" box="[151,491,532,557]" pageId="1" pageNumber="62">
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<emphasis id="5999EAA4FF8FE7372C86FDC1FECAFDBA" bold="true" box="[151,343,533,557]" italics="true" pageId="1" pageNumber="62">Claosaurus agilis</emphasis>
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<bibRefCitation id="0F7C4B47FF8FE7372D4CFDC0FE76FDBB" author="Marsh, O. C." box="[349,491,532,556]" pageId="1" pageNumber="62" pagination="301" refId="ref3959" refString="Marsh, O. C. (1872) Notice on a new species of Hadrosaurus. American Journal of Science, 3, 301." type="journal article" year="1872">Marsh, 1872</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="23F7653DFF8FE7372C86FDECFF48FDD8" box="[151,213,568,591]" pageId="1" pageNumber="62" type="description">
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<paragraph id="6B5236B6FF8FE7372C86FDECFF48FDD8" blockId="1.[151,491,532,591]" box="[151,213,568,591]" pageId="1" pageNumber="62">
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<figureCitation id="F3D62A33FF8FE7372C86FDECFF48FDD8" box="[151,213,568,591]" captionStart="FIGURE 2" captionStartId="4.[151,250,1095,1118]" captionTargetBox="[173,1416,221,1068]" captionTargetId="figure-269@4.[151,1436,195,1071]" captionTargetPageId="4" captionText="FIGURE 2. (a) Right ilium of the holotype specimen of Claosaurus agilis (YPM 1190) in lateral view. The horizontal and double-arrowed white lines indicate the breadth of the supraacetabular crest being approximately 75 per cent of the length of the central plate of the ilium. The vertical white arrow indicates the position of the ventral apex of the supraacetabular crest relative to the posteroventral corner (black arrow) of the posterior tuberosity of the ischial peduncle. (b) Line drawing of the right ilium of YPM 1190 showing in gray the areas with plaster reconstruction." figureDoi="http://doi.org/10.5281/zenodo.5290961" httpUri="https://zenodo.org/record/5290961/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="23F7653DFF8FE7372C86FDA8FACFFB86" pageId="1" pageNumber="62" type="materials_examined">
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<paragraph id="6B5236B6FF8FE7372C86FDA8FC9FFC88" blockId="1.[151,1437,636,2012]" pageId="1" pageNumber="62">
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<materialsCitation id="DB853CEBFF8FE7372C86FDA8FC9FFC88" collectionCode="YPM" pageId="1" pageNumber="62" specimenCode="YPM 1190" specimenCount="1" typeStatus="holotype">
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<emphasis id="5999EAA4FF8FE7372C86FDA8FE98FD03" bold="true" box="[151,261,636,660]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">
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<typeStatus id="B4568814FF8FE7372C86FDA8FE9CFD03" box="[151,257,636,660]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="3B4B9ECDFF8FE7372D1FFDA9FE1AFD03" box="[270,391,637,660]" collectionCode="YPM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="1" pageNumber="62" type="Museum">YPM 1190</specimenCode>
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, a partial skeleton consisting of a fragment of coronoid process from the left dentary, maxillary fragments with several teeth exposed, partial left postorbital, cervical through caudal vertebral series missing the neural spines, right scapula, partial left and right sternals, left and right humeri, proximal and distal segments of both ulnae and radii, nearly complete right ilium, distal-most end of the right ischium, nearly complete left and right femora, partial tibiae missing their shafts, and partial left and right pedes.
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</materialsCitation>
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</paragraph>
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<paragraph id="6B5236B6FF8FE7372CD6FCFEFB4FFC10" blockId="1.[151,1437,636,2012]" pageId="1" pageNumber="62">
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<emphasis id="5999EAA4FF8FE7372CD6FCFEFE68FCD5" bold="true" box="[199,501,810,834]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">Type locality and horizon.</emphasis>
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The
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<typeStatus id="B4568814FF8FE7372E20FCFEFD13FCD6" box="[561,654,810,833]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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and only known exemplar of
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<taxonomicName id="ACED4D35FF8FE7372FCAFCFEFB07FCD6" authorityName="Marsh" authorityYear="1872" box="[987,1178,810,833]" class="Reptilia" family="Hadrosauridae" genus="Claosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="agilis">
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<emphasis id="5999EAA4FF8FE7372FCAFCFEFB07FCD6" box="[987,1178,810,833]" italics="true" pageId="1" pageNumber="62">Claosaurus agilis</emphasis>
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</taxonomicName>
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was collected from the Smoky Hill Chalk Member of the Niobrara Chalk Formation (near Smoky Hill River, Logan County,
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<collectingRegion id="A929F854FF8FE7372911FC99FAD2FCF3" box="[1280,1359,845,868]" country="United States of America" name="Kansas" pageId="1" pageNumber="62">Kansas</collectingRegion>
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,
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<collectingCountry id="13FA7626FF8FE737294FFC99FA08FCF3" box="[1374,1429,845,868]" name="United States of America" pageId="1" pageNumber="62">USA</collectingCountry>
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) (
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<bibRefCitation id="0F7C4B47FF8FE7372C8FFCA4FE18FC10" author="Carpenter, K. & Dilkes, D. & Weishampel, D. B." box="[158,389,880,903]" pageId="1" pageNumber="62" pagination="275 - 297" refId="ref3635" refString="Carpenter, K., Dilkes, D. & Weishampel, D. B. (1995) The dinosaurs of the Niobrara Chalk Formation (Upper Cretaceous, Kansas). Journal of Vertebrate Paleontology, 15, 275 - 297." type="journal article" year="1995">
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Carpenter
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<emphasis id="5999EAA4FF8FE7372D1EFCA4FEA3FC10" box="[271,318,880,903]" italics="true" pageId="1" pageNumber="62">et al</emphasis>
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. 1995
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</bibRefCitation>
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); these beds are Late Coniacian in age according to
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<bibRefCitation id="0F7C4B47FF8FE7372FBEFCA4FB53FC10" author="Everhart, M. J. & Ewell, K." box="[943,1230,880,903]" pageId="1" pageNumber="62" pagination="27 - 35" refId="ref3738" refString="Everhart, M. J. & Ewell, K. (2006) Shark-bitten dinosaur (Hadrosauridae) caudal vertebrae from the Niobrara Chalk (Upper Coniacian) of western Kansas. Transactions of the Kansas Academy of Science, 109, 27 - 35." type="journal article" year="2006">Everhart and Ewell (2006)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="6B5236B6FF8FE7372CD6FC46FACFFB86" blockId="1.[151,1437,636,2012]" pageId="1" pageNumber="62">
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<emphasis id="5999EAA4FF8FE7372CD6FC46FE36FC3D" bold="true" box="[199,427,914,938]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">Amended diagnosis.</emphasis>
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Hadrosauroid dinosaur characterized by the unique combination of the following humeral and iliac characters: deltopectoral crest length less than 48% of humerus length and with wide arcuate laterodistal corner; anteroposterior width of supraacetabular crest being approximately 75 per cent of length of central plate of ilium; and ventral apex of supraacetabular crest positioned anterior to the posteroventral corner of ischial peduncle of ilium..
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</paragraph>
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</subSubSection>
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<subSubSection id="23F7653DFF8FE7372CD6FBC8FB56F84B" pageId="1" pageNumber="62" type="discussion">
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<paragraph id="6B5236B6FF8FE7372CD6FBC8FAC6FB76" blockId="1.[151,1437,636,2012]" pageId="1" pageNumber="62">
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<emphasis id="5999EAA4FF8FE7372CD6FBC8FEABFBA3" bold="true" box="[199,310,1052,1076]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">Remarks.</emphasis>
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<bibRefCitation id="0F7C4B47FF8FE7372D2DFBC9FDADFBA3" author="Carpenter, K. & Dilkes, D. & Weishampel, D. B." box="[316,560,1053,1076]" pageId="1" pageNumber="62" pagination="275 - 297" refId="ref3635" refString="Carpenter, K., Dilkes, D. & Weishampel, D. B. (1995) The dinosaurs of the Niobrara Chalk Formation (Upper Cretaceous, Kansas). Journal of Vertebrate Paleontology, 15, 275 - 297." type="journal article" year="1995">
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Carpenter
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<emphasis id="5999EAA4FF8FE7372DBDFBC9FE46FBA3" box="[428,475,1053,1076]" italics="true" pageId="1" pageNumber="62">et al</emphasis>
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. (1995)
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</bibRefCitation>
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tentatively diagnosed
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<taxonomicName id="ACED4D35FF8FE7372F33FBC9FC42FBA3" authorityName="Marsh" authorityYear="1872" box="[802,991,1053,1076]" class="Reptilia" family="Hadrosauridae" genus="Claosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="agilis">
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<emphasis id="5999EAA4FF8FE7372F33FBC9FC42FBA3" box="[802,991,1053,1076]" italics="true" pageId="1" pageNumber="62">Claosaurus agilis</emphasis>
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</taxonomicName>
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on the basis of the relatively short length of the preacetabular process in YPM 1190. As preserved in this specimen, the process is certainly proximodistally shorter (in relation to the anteroposterior length of the iliac central plate) than in any other known iguanodontian ornithopod (Prieto-Márquez unpublished data). However, the distal-most tip of the preacetabular process in YPM 1190 is reconstructed, as evidenced by the presence of plaster (
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<figureCitation id="F3D62A33FF8FE7372EE6FB7CFCD4FB28" box="[759,841,1192,1215]" captionStart="FIGURE 2" captionStartId="4.[151,250,1095,1118]" captionTargetBox="[173,1416,221,1068]" captionTargetId="figure-269@4.[151,1436,195,1071]" captionTargetPageId="4" captionText="FIGURE 2. (a) Right ilium of the holotype specimen of Claosaurus agilis (YPM 1190) in lateral view. The horizontal and double-arrowed white lines indicate the breadth of the supraacetabular crest being approximately 75 per cent of the length of the central plate of the ilium. The vertical white arrow indicates the position of the ventral apex of the supraacetabular crest relative to the posteroventral corner (black arrow) of the posterior tuberosity of the ischial peduncle. (b) Line drawing of the right ilium of YPM 1190 showing in gray the areas with plaster reconstruction." figureDoi="http://doi.org/10.5281/zenodo.5290961" httpUri="https://zenodo.org/record/5290961/files/figure.png" pageId="1" pageNumber="62">Fig. 2b</figureCitation>
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). This indicates that the preacetabular process is most probably incomplete and, thus,
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<taxonomicName id="ACED4D35FF8FE7372DF6FB1EFDDCFB76" authorityName="Marsh" authorityYear="1872" box="[487,577,1226,1249]" class="Reptilia" family="Hadrosauridae" genus="Claosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="agilis">
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<emphasis id="5999EAA4FF8FE7372DF6FB1EFDDCFB76" box="[487,577,1226,1249]" italics="true" pageId="1" pageNumber="62">C. agilis</emphasis>
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</taxonomicName>
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cannot be diagnosed based on the possession of a relatively short process.
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</paragraph>
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<paragraph id="6B5236B6FF8FE7372CD6FB39FBA1F9AB" blockId="1.[151,1437,636,2012]" pageId="1" pageNumber="62">
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As shown in
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<tableCitation id="266F030DFF8FE7372D40FB39FE02FA93" box="[337,415,1261,1284]" captionStart="TABLE 1" captionStartId="2.[151,235,151,174]" captionText="TABLE 1. Distribution of four humeral and iliac characters relevant to the diagnoses of Hadrosaurus foulkii and Claosaurus agilis in a sample of iguanodontian ornithopods. DPCL: proximodistal length of the deltopectoral crest relative to the total length of the humerus (0 = less than 48% of humeral length; 1 = between 48 and 55%; 2 = over 55%). DPCC: morphology of the laterodistal corner of the deltopectoral crest of the humerus. SAPL: anteroposterior width of the supraacetabular crest relative to the anteroporterior length of the central plate of the ilium (0 = over 85% of the length of the iliac central plate; 1 = between 85 and 70%; 2 = between 69 and 55%; 3 = less 55%). SAPP: position of the ventral apex of the supraacetabular crest of the ilium relative to the posteroventral corner of the tuberosity of the ischial peduncle. The data displayed in this table are based on observation of actual specimens, except in the following cases in which they are based on literature sources: (1) Norman (1998); (2) Gilpin et al. (2007); (3) McDonald et al. (2010); (4) Xu et al. (2000); (5) Norman (2002). Character coding and definitions for DPCL and SAPL are from Prieto-Márquez (2010)." pageId="1" pageNumber="62">Table 1</tableCitation>
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,
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<taxonomicName id="ACED4D35FF8FE7372DBDFB39FD9BFA93" authorityName="Marsh" authorityYear="1872" box="[428,518,1261,1284]" class="Reptilia" family="Hadrosauridae" genus="Claosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="agilis">
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<emphasis id="5999EAA4FF8FE7372DBDFB39FD9BFA93" box="[428,518,1261,1284]" italics="true" pageId="1" pageNumber="62">C. agilis</emphasis>
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</taxonomicName>
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is the only non-hadrosaurid hadrosauroid species that combines such a relatively wide supraacetabular crest with its ventral apex being located anterior to the posteroventral corner of the ischial peduncle of the ilium. All other non-hadrosaurid hadrosauroid taxa that have a supraacetabular crest that is 70 per cent or more of the length of the iliac central plate show a ventral apex of this crest positioned posterior to the ischial peduncle (
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<figureCitation id="F3D62A33FF8FE737290CFA81FAC1FAFB" box="[1309,1372,1365,1388]" captionStart="FIGURE 3" captionStartId="5.[151,250,1320,1343]" captionTargetBox="[152,1428,307,1288]" captionTargetId="figure-412@5.[151,1436,297,1299]" captionTargetPageId="5" captionText="FIGURE 3. Distribution, in a phylogenetic context, of the relative length and profile of the laterodistal corner of the deltopectoral crest of the humerus, and the anteroposterior width of the supraacetabular crest and position of its apex relative to the ischial peduncle of the ilium in Hadrosaurus foulkii and selected major groups of iguanodontian dinosaurs. The cladogram has been simplified from the strict reduced consensus tree presented by Prieto-Márquez (2010). From left to right, specimens used as clade or taxon representatives are as follows. Non-hadrosauroid Iguanodontia: skull of Iguanodon bernissartensis (IRSNB 1731), and right humerus (NHMUK R6616; reversed) and left ilium (NHMUK 11521) of Mantellisaurus atherfieldensis. Nonhadrosaurid Hadrosauroidea: skull of Equijubus normani (IVPP V12534), and right humerus (AMNH 30728; reversed) and right ilium (AMNH 30735; reversed) of Gilmoreosaurus mongoliensis. Hadrosaurus foulkii: line drawing of the historical skull model sculpted by Waterhouse Hawkins, and left humerus and left ilium of ANSP 10005. Saurolophinae: skull of Edmontosaurus annectens (SM R4036) and left humerus of Gryposaurus latidens (AMNH 5465). Lambeosaurinae: skull of Parasaurolophus walkeri (ROM 768) and left humerus of P. cyrtocristatus (FMNH P27393). The right ilium (reversed) representing both saurolophines and lambeosaurines (i.e., Saurolophidae) is that of Brachylophosaurus canadensis (MOR 794). Skulls and ilia are shown in left lateral view; humeri are displayed in anteromedial view." figureDoi="http://doi.org/10.5281/zenodo.5290965" httpUri="https://zenodo.org/record/5290965/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
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). The validity of the diagnosis proposed here depends on interpretation of the actual position of the ventral apex of the supraacetabular crest in YPM 1190. In this exemplar, the lateroventral margin of the crest is not completely preserved. However, the preserved lateroventral border indicates that the apex must have been located at least slightly anterior to the level of the ischial peduncle. Other possible interpretations of the position of the apex would place it even more anteriorly. The most conservative estimate is considered here (
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<figureCitation id="F3D62A33FF8FE7372F6CF9D6FC48F98E" box="[893,981,1538,1561]" captionStart="FIGURE 2" captionStartId="4.[151,250,1095,1118]" captionTargetBox="[173,1416,221,1068]" captionTargetId="figure-269@4.[151,1436,195,1071]" captionTargetPageId="4" captionText="FIGURE 2. (a) Right ilium of the holotype specimen of Claosaurus agilis (YPM 1190) in lateral view. The horizontal and double-arrowed white lines indicate the breadth of the supraacetabular crest being approximately 75 per cent of the length of the central plate of the ilium. The vertical white arrow indicates the position of the ventral apex of the supraacetabular crest relative to the posteroventral corner (black arrow) of the posterior tuberosity of the ischial peduncle. (b) Line drawing of the right ilium of YPM 1190 showing in gray the areas with plaster reconstruction." figureDoi="http://doi.org/10.5281/zenodo.5290961" httpUri="https://zenodo.org/record/5290961/files/figure.png" pageId="1" pageNumber="62">Fig. 2a</figureCitation>
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). An anteriorly-positioned apex of the supraacetabular crest is convergent with
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<taxonomicName id="ACED4D35FF8FE7372E58F9F1FC43F9AB" authority="(Prieto-Marquez 2010)" baseAuthorityName="Prieto-Marquez" baseAuthorityYear="2010" box="[585,990,1573,1596]" class="Reptilia" family="Saurolophidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="family">
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Saurolophidae (
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<bibRefCitation id="0F7C4B47FF8FE7372EE1F9F1FC4AF9AB" author="Prieto-Marquez, A." box="[752,983,1573,1596]" pageId="1" pageNumber="62" pagination="435 - 502" refId="ref4248" refString="Prieto-Marquez, A. (2010) Global phylogeny of Hadrosauridae (Dinosauria: Ornithopoda) using parsimony and Bayesian methods. Zoological Journal of the Linnean Society, 159, 435 - 502." type="journal article" year="2010">Prieto-Márquez 2010</bibRefCitation>
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)
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</taxonomicName>
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(
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<figureCitation id="F3D62A33FF8FE7372FFCF9F1FBB3F9AB" box="[1005,1070,1573,1596]" captionStart="FIGURE 3" captionStartId="5.[151,250,1320,1343]" captionTargetBox="[152,1428,307,1288]" captionTargetId="figure-412@5.[151,1436,297,1299]" captionTargetPageId="5" captionText="FIGURE 3. Distribution, in a phylogenetic context, of the relative length and profile of the laterodistal corner of the deltopectoral crest of the humerus, and the anteroposterior width of the supraacetabular crest and position of its apex relative to the ischial peduncle of the ilium in Hadrosaurus foulkii and selected major groups of iguanodontian dinosaurs. The cladogram has been simplified from the strict reduced consensus tree presented by Prieto-Márquez (2010). From left to right, specimens used as clade or taxon representatives are as follows. Non-hadrosauroid Iguanodontia: skull of Iguanodon bernissartensis (IRSNB 1731), and right humerus (NHMUK R6616; reversed) and left ilium (NHMUK 11521) of Mantellisaurus atherfieldensis. Nonhadrosaurid Hadrosauroidea: skull of Equijubus normani (IVPP V12534), and right humerus (AMNH 30728; reversed) and right ilium (AMNH 30735; reversed) of Gilmoreosaurus mongoliensis. Hadrosaurus foulkii: line drawing of the historical skull model sculpted by Waterhouse Hawkins, and left humerus and left ilium of ANSP 10005. Saurolophinae: skull of Edmontosaurus annectens (SM R4036) and left humerus of Gryposaurus latidens (AMNH 5465). Lambeosaurinae: skull of Parasaurolophus walkeri (ROM 768) and left humerus of P. cyrtocristatus (FMNH P27393). The right ilium (reversed) representing both saurolophines and lambeosaurines (i.e., Saurolophidae) is that of Brachylophosaurus canadensis (MOR 794). Skulls and ilia are shown in left lateral view; humeri are displayed in anteromedial view." figureDoi="http://doi.org/10.5281/zenodo.5290965" httpUri="https://zenodo.org/record/5290965/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
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).
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</paragraph>
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<paragraph id="6B5236B6FF8FE7372CD6F99CFB56F84B" blockId="1.[151,1437,636,2012]" pageId="1" pageNumber="62">
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The proposed iliac characters for diagnosing
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<taxonomicName id="ACED4D35FF8FE7372EBBF99CFCFAF9C8" authorityName="Marsh" authorityYear="1872" box="[682,871,1608,1631]" class="Reptilia" family="Hadrosauridae" genus="Claosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="agilis">
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<emphasis id="5999EAA4FF8FE7372EBBF99CFCFAF9C8" box="[682,871,1608,1631]" italics="true" pageId="1" pageNumber="62">Claosaurus agilis</emphasis>
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</taxonomicName>
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(and
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<taxonomicName id="ACED4D35FF8FE7372FB4F99CFBE6F9C8" authorityName="Leidy" authorityYear="1858" box="[933,1147,1608,1631]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="foulkii">
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<emphasis id="5999EAA4FF8FE7372FB4F99CFBE6F9C8" box="[933,1147,1608,1631]" italics="true" pageId="1" pageNumber="62">Hadrosaurus foulkii</emphasis>
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</taxonomicName>
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; see below) do not display intraspecific variation such as that due to ontogeny. In those hadrosauroid taxa in which different ontogenetic stages are known preserving appendicular elements (e.g.,
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<taxonomicName id="ACED4D35FF8FE7372EA3F959FC37F933" authorityName="Gilmore" authorityYear="1933" box="[690,938,1677,1700]" class="Reptilia" family="Hadrosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="johnsoni">
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<emphasis id="5999EAA4FF8FE7372EA3F959FC37F933" box="[690,938,1677,1700]" italics="true" pageId="1" pageNumber="62">Bactrosaurus johnsoni</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ACED4D35FF8FE7372FAFF959FA8FF933" authorityName="Sternberg" authorityYear="1953" box="[958,1298,1677,1700]" class="Reptilia" family="Hadrosauridae" genus="Brachylophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="canadensis">
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<emphasis id="5999EAA4FF8FE7372FAFF959FA8FF933" box="[958,1298,1677,1700]" italics="true" pageId="1" pageNumber="62">Brachylophosaurus canadensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ACED4D35FF8FE7372936F959FE82F950" authorityName="Horner & Makela" authorityYear="1979" class="Reptilia" family="Hadrosauridae" genus="Maiasaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="peeblesorum">
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<emphasis id="5999EAA4FF8FE7372936F959FE82F950" italics="true" pageId="1" pageNumber="62">Maiasaura peeblesorum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ACED4D35FF8FE7372D2DF964FDF7F950" authorityName="Rozhdestvensky" authorityYear="1952" box="[316,618,1712,1735]" class="Reptilia" family="Hadrosauridae" genus="Saurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="angustirostris">
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<emphasis id="5999EAA4FF8FE7372D2DF964FDF7F950" box="[316,618,1712,1735]" italics="true" pageId="1" pageNumber="62">Saurolophus angustirostris</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ACED4D35FF8FE7372E96F964FCEEF950" box="[647,883,1712,1735]" class="Reptilia" family="Hadrosauridae" genus="Edmontosaurus" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="5999EAA4FF8FE7372E96F964FCB3F950" box="[647,814,1712,1735]" italics="true" pageId="1" pageNumber="62">Edmontosaurus</emphasis>
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spp.
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</taxonomicName>
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,
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|
<taxonomicName id="ACED4D35FF8FE7372F9CF964FBEDF950" box="[909,1136,1712,1735]" class="Reptilia" family="Lambeosauridae" genus="Corythosaurus" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="5999EAA4FF8FE7372F9CF964FBB1F950" box="[909,1068,1712,1735]" italics="true" pageId="1" pageNumber="62">Corythosaurus</emphasis>
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spp.
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="ACED4D35FF8FE737289BF964FAF6F950" box="[1162,1387,1712,1735]" class="Reptilia" family="Lambeosauridae" genus="Lambeosaurus" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="5999EAA4FF8FE737289BF964FABAF950" box="[1162,1319,1712,1735]" italics="true" pageId="1" pageNumber="62">Lambeosaurus</emphasis>
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spp.
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</taxonomicName>
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, or
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<taxonomicName id="ACED4D35FF8FE7372C86F906FE33F97E" box="[151,430,1746,1769]" class="Reptilia" family="Hadrosauridae" genus="Hypacrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="stebingeri">
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<emphasis id="5999EAA4FF8FE7372C86F906FE33F97E" box="[151,430,1746,1769]" italics="true" pageId="1" pageNumber="62">Hypacrosaurus stebingeri</emphasis>
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</taxonomicName>
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|
) the anteroposterior breadth of the supraacetabular crest and the position of its ventral apex relative to the posterior tuberosity of the ischial peduncle are maintained throughout ontogeny (Prieto-Márquez in press and unpublished data). More specifically, this is exemplified when comparing the
|
|
<taxonomicName id="ACED4D35FF8FE7372FEFF8CCFB0CF8B8" authorityName="Sternberg" authorityYear="1953" box="[1022,1169,1816,1839]" class="Reptilia" family="Hadrosauridae" genus="Brachylophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="canadensis">
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<emphasis id="5999EAA4FF8FE7372FEFF8CCFB0CF8B8" box="[1022,1169,1816,1839]" italics="true" pageId="1" pageNumber="62">B. canadensis</emphasis>
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</taxonomicName>
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|
juvenile specimen MOR
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|
<date id="1F531076FF8FE7372C86F8EEFE9EF8C6" box="[151,259,1850,1873]" pageId="1" pageNumber="62" value="1071-07-15">1071-7-15</date>
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|
-98-215 with the adults of the same species MOR 794 or MOR MOR 8-2-98-469; the juvenile ZPAL MgD-I 159 with the adult MPC-D 100/706, both referable to
|
|
<taxonomicName id="ACED4D35FF8FE7372EDEF889FCE1F8E3" authorityName="Rozhdestvensky" authorityYear="1952" box="[719,892,1885,1908]" class="Reptilia" family="Hadrosauridae" genus="Saurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="angustirostris">
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|
<emphasis id="5999EAA4FF8FE7372EDEF889FCE1F8E3" box="[719,892,1885,1908]" italics="true" pageId="1" pageNumber="62">S. angustirostris</emphasis>
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</taxonomicName>
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|
; the adult
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|
<taxonomicName id="ACED4D35FF8FE7372FFDF889FB24F8E3" authority="MOR" authorityName="MOR" box="[1004,1209,1885,1908]" class="Reptilia" family="Hadrosauridae" genus="Hypacrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="stebingeri">
|
|
<emphasis id="5999EAA4FF8FE7372FFDF889FBEAF8E3" box="[1004,1143,1885,1908]" italics="true" pageId="1" pageNumber="62">H. stebingeri</emphasis>
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MOR
|
|
</taxonomicName>
|
|
549 with the neonate ilia from MOR 548; or the juvenile
|
|
<taxonomicName id="ACED4D35FF8FE7372E3AF854FD6CF800" box="[555,753,1920,1943]" class="Reptilia" family="Lambeosauridae" genus="Lambeosaurus" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="undetermined">
|
|
<emphasis id="5999EAA4FF8FE7372E3AF854FD55F800" box="[555,712,1920,1943]" italics="true" pageId="1" pageNumber="62">Lambeosaurus</emphasis>
|
|
sp.
|
|
</taxonomicName>
|
|
specimen AMNH 5340 with the adult
|
|
<taxonomicName id="ACED4D35FF8FE73728B6F854FA04F800" authority="ROM 1218" authorityName="ROM" authorityYear="1218" box="[1191,1433,1920,1943]" class="Reptilia" family="Lambeosauridae" genus="Lambeosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="lambei">
|
|
<emphasis id="5999EAA4FF8FE73728B6F854FA92F800" box="[1191,1295,1920,1943]" italics="true" pageId="1" pageNumber="62">L. lambei</emphasis>
|
|
ROM 1218
|
|
</taxonomicName>
|
|
. Among basal hadrosauroids, the fact that those characters are maintained throughout ontogeny is documented in
|
|
<taxonomicName id="ACED4D35FF8FE7372994F876FF6EF84B" authorityName="Gilmore" authorityYear="1933" class="Reptilia" family="Hadrosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="62" phylum="Chordata" rank="species" species="johnsoni">
|
|
<emphasis id="5999EAA4FF8FE7372994F876FF6EF84B" italics="true" pageId="1" pageNumber="62">B. johnsoni</emphasis>
|
|
</taxonomicName>
|
|
; for example, in the juvenile AMNH 6577 and the adult or larger subadult IRSNB 95E5/25.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |