432 lines
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432 lines
57 KiB
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<mods:title id="67B50931EF5202EA5A473B0129EBF7DF">Nuevos registros fósiles de Onthophagus pilauco Tello, Verdú, Rossini y Zunino, 2021 (Coleoptera: Scarabaeidae: Scarabaeinae), revelan un patrón morfológico único entre los Onthophagus americanos</mods:title>
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<mods:namePart id="C230297D65EE51311F851EE39C80FCE8">Tello, Francisco</mods:namePart>
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<mods:namePart id="E3058788C65151A73F7DE221593BDCEF">Rossini, Michele</mods:namePart>
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<mods:namePart id="C6569CB29F26D6F5C0F96C178CE6416B">Pino, Mario</mods:namePart>
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<mods:namePart id="E6AEC52109BFDDAF8E1C5AB0B7F9CC97">Verdú, José R.</mods:namePart>
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<treatment id="DE36F261A63FFFB7FEB5FFFF9A0650A9" ID-DOI="http://doi.org/10.5281/zenodo.13238777" ID-Zenodo-Dep="13238777" LSID="urn:lsid:plazi:treatment:DE36F261A63FFFB7FEB5FFFF9A0650A9" httpUri="http://treatment.plazi.org/id/DE36F261A63FFFB7FEB5FFFF9A0650A9" lastPageId="7" lastPageNumber="942" pageId="4" pageNumber="939">
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<subSubSection id="1E8510FCA63FFFB4FEB5FFFF9DD150C9" box="[349,1011,122,147]" pageId="4" pageNumber="939" type="nomenclature">
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<paragraph id="56204377A63FFFB4FEB5FFFF9DD150C9" blockId="4.[349,1011,122,178]" box="[349,1011,122,147]" pageId="4" pageNumber="939">
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<heading id="0D68F41BA63FFFB4FEB5FFFF9DD150C9" box="[349,1011,122,147]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="939" reason="4">
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<taxonomicName id="919F38F4A63FFFB4FEB5FFFF9DD150C9" ID-CoL="8KPR6" authority="Tello, Verdu, Rossini y Zunino, 2021" authorityName="Tello, Verdu, Rossini y Zunino" authorityYear="2021" box="[349,1011,122,147]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">
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<emphasis id="64EB9F65A63FFFB4FEB5FFFF9C6650C9" box="[349,580,122,146]" italics="true" pageId="4" pageNumber="939">Onthophagus pilauco</emphasis>
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<bibRefCitation id="320E3E86A63FFFB4FDA3FFFF9DD150C9" author="Tello, F. & Verdu, J. R. & Rossini, M. & Zunino, M." box="[587,1011,122,147]" pageId="4" pageNumber="939" pagination="133 - 145" refId="ref9640" refString="Tello, F., Verdu, J. R., Rossini, M. y Zunino, M. (2021) Onthophagus pilauco sp. nov. (Coleoptera, Scarabaeidae): evidence of beetle extinction in the Pleistocene - Holocene transition in Chilean Northern Patagonia. ZooKeys, 1043: 133 - 145." type="journal article" year="2021">Tello, Verdú, Rossini y Zunino, 2021</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="1E8510FCA63FFFB4FD80FF1F9CCB50E9" box="[616,745,154,178]" pageId="4" pageNumber="939" type="description">
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<paragraph id="56204377A63FFFB4FD80FF1F9CCB50E9" blockId="4.[349,1011,122,178]" box="[616,745,154,178]" pageId="4" pageNumber="939">
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(
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<figureCitation id="CEA45FF2A63FFFB4FD98FF1F9CC250E9" box="[624,736,154,178]" captionStart-0="Figuras 3-6" captionStart-1="Figuras 7-13" captionStart-2="Figuras 14-17" captionStartId-0="4.[151,232,1203,1225]" captionStartId-1="6.[151,232,1377,1399]" captionStartId-2="7.[91,172,1190,1212]" captionTargetBox-0="[214,1139,221,1180]" captionTargetBox-1="[201,1180,492,1357]" captionTargetBox-2="[148,1111,280,1174]" captionTargetId-0="figure-282@4.[151,1209,211,1199]" captionTargetId-1="figure-380@6.[151,1209,486,1371]" captionTargetId-2="figure-108@7.[91,1149,267,1191]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Figuras 3-6. Vista dorsal y laterodorsal de Onthophagus pilauco macho. 3, 5. Macho major. 4, 6. Macho minor. / Dorsal and laterodorsal view of male Onthophagus pilauco. 3, 5. Major male. 4, 6. Minor male." captionText-1="Figuras 7-13. Cuernos del clípeo presentes en los machos major y minor de Onthophagus pilauco. 7. Vista frontal del cuerno clipeal en la forma major. 8. Vista lateral del cuerno clipeal en la forma major. 9. Vista dorsal del cuerno clipeal en la forma major. 10. Reconstrucción fotográfica en vista frontal de la forma major. 11. Reconstrucción fotográfica en vista laterodorsal de la forma major. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma minor. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma minor. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both minor and major male forms of Onthophagus pilauco.7. Dorsal view of the clypeal horn in the major form. 8. Lateral view of the clypeal tubercle in the major form. 9. Dorsal view of the clypeal tubercle in the major form. 10. Photographic reconstruction of frontal view of the major form. 11. Photographic reconstruction in laterodorsal view of the major form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the minor form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the minor form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove." captionText-2="Figuras 14-17. 14-15. Vista dorsal de Onthophagus pilauco hembra. 16-17. Vista laterodorsal de Onthophagus pilauco hembra. / 14-15. Dorsal view of females Onthophagus pilauco. 16-17. Laterodorsal view of females Onthophagus pilauco." figureDoi-0="http://doi.org/10.5281/zenodo.13367592" figureDoi-1="http://doi.org/10.5281/zenodo.13367594" figureDoi-2="http://doi.org/10.5281/zenodo.13367596" httpUri-0="https://zenodo.org/record/13367592/files/figure.png" httpUri-1="https://zenodo.org/record/13367594/files/figure.png" httpUri-2="https://zenodo.org/record/13367596/files/figure.png" pageId="4" pageNumber="939">Figs. 3-17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<caption id="02E013FFA63FFFB4FF7FFB369A9854BD" ID-DOI="http://doi.org/10.5281/zenodo.13367592" ID-Zenodo-Dep="13367592" httpUri="https://zenodo.org/record/13367592/files/figure.png" pageId="4" pageNumber="939" startId="4.[151,232,1203,1225]" targetBox="[214,1139,221,1180]" targetPageId="4" targetType="figure">
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<paragraph id="56204377A63FFFB4FF7FFB369A9854BD" blockId="4.[151,1210,1203,1254]" pageId="4" pageNumber="939">
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<emphasis id="64EB9F65A63FFFB4FF7FFB369F365493" bold="true" box="[151,276,1203,1225]" pageId="4" pageNumber="939">Figuras 3-6.</emphasis>
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Vista dorsal y laterodorsal de
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<taxonomicName id="919F38F4A63FFFB4FDBFFB369D055493" authorityName="Tello, Verdu, Rossini y Zunino" authorityYear="2021" box="[599,807,1203,1224]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">
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<emphasis id="64EB9F65A63FFFB4FDBFFB369D055493" box="[599,807,1203,1224]" italics="true" pageId="4" pageNumber="939">Onthophagus pilauco</emphasis>
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macho. 3, 5. Macho
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<emphasis id="64EB9F65A63FFFB4FC15FB319A175492" box="[1021,1077,1204,1225]" italics="true" pageId="4" pageNumber="939">major</emphasis>
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. 4, 6. Macho
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<emphasis id="64EB9F65A63FFFB4FF7FFB549EF054BD" box="[151,210,1233,1254]" italics="true" pageId="4" pageNumber="939">minor</emphasis>
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. / Dorsal and laterodorsal view of male
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<emphasis id="64EB9F65A63FFFB4FD91FB559D6D54BD" box="[633,847,1232,1254]" italics="true" pageId="4" pageNumber="939">
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<taxonomicName id="919F38F4A63FFFB4FD91FB559D6B54BE" authorityName="Tello, Verdu, Rossini y Zunino" authorityYear="2021" box="[633,841,1232,1253]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">Onthophagus pilauco</taxonomicName>
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.
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</emphasis>
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3, 5. Major male. 4, 6. Minor male.
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</paragraph>
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</caption>
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<subSubSection id="1E8510FCA63FFFB7FF7FFA9A9A0650A9" lastPageId="7" lastPageNumber="942" pageId="4" pageNumber="939" type="materials_examined">
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<paragraph id="56204377A63FFFB5FF7FFA9A9FD352A9" blockId="4.[151,1210,1311,1751]" lastBlockId="5.[91,1151,122,754]" lastPageId="5" lastPageNumber="940" pageId="4" pageNumber="939">
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<emphasis id="64EB9F65A63FFFB4FF7FFA9A9CB2556C" bold="true" box="[151,656,1311,1335]" pageId="4" pageNumber="939">Material examinado (n=18): Machos (n=7)</emphasis>
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[N°1] a) “Cuad.: 17AA / N - / Prof.: 461 / Col.: F. Tello / Fecha:
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<date id="222165B7A63FFFB4FEB4FABA9FE9550C" box="[348,459,1343,1367]" pageId="4" pageNumber="939" value="2021-05-08">
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08.
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.2021
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/ NE –“, b)
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<taxonomicName id="919F38F4A63FFFB4FDB6FABA9D7E550C" authority="MHMOP" authorityName="MHMOP" box="[606,860,1343,1367]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO MHMOP</taxonomicName>
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/
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<collectionCode id="308EDBB2A63FFFB4FC8DFABA9D5D550C" box="[869,895,1343,1367]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
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/17AA-080521-
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”, c) “
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<taxonomicName id="919F38F4A63FFFB4FF4DFADA9D65552C" authority="Tello, Verdu, Rossini & Zunino, 2021" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[165,839,1375,1399]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">
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<emphasis id="64EB9F65A63FFFB4FF4DFADA9FAF552C" box="[165,397,1375,1399]" italics="true" pageId="4" pageNumber="939">Onthophagus pilauco</emphasis>
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<bibRefCitation id="320E3E86A63FFFB4FE7CFADA9D65552C" author="Tello, F. & Verdu, J. R. & Rossini, M. & Zunino, M." box="[404,839,1375,1399]" pageId="4" pageNumber="939" pagination="133 - 145" refId="ref9640" refString="Tello, F., Verdu, J. R., Rossini, M. y Zunino, M. (2021) Onthophagus pilauco sp. nov. (Coleoptera, Scarabaeidae): evidence of beetle extinction in the Pleistocene - Holocene transition in Chilean Northern Patagonia. ZooKeys, 1043: 133 - 145." type="journal article" year="2021">Tello, Verdú, Rossini & Zunino, 2021</bibRefCitation>
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<specimenCount id="409988FEA63FFFB4FCB8FADA9D415522" box="[848,867,1375,1401]" count="1" pageId="4" pageNumber="939" type="male">♂</specimenCount>
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”. [N°2] a) “Cuad.: 17AA / N - / Prof.: 461 / Col.: F. Tello / Fecha:
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<date id="222165B7A63FFFB4FDACFAFA9C9155CC" box="[580,691,1407,1431]" pageId="4" pageNumber="939" value="2021-05-08">
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08.
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<collectionCode id="308EDBB2A63FFFB4FD8BFAFA9C5A55CC" box="[611,632,1407,1431]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="939" type="Museum">V</collectionCode>
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.2021
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</date>
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/ NE –“, b)
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<taxonomicName id="919F38F4A63FFFB4FCACFAFA9A6155CC" authority="MHMOP" authorityName="MHMOP" box="[836,1091,1407,1431]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO MHMOP</taxonomicName>
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/
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<collectionCode id="308EDBB2A63FFFB4FBA3FAFA9A4755CC" box="[1099,1125,1407,1431]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
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/12AA-260321-
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08
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”, c) [mismos datos que N°1]. [N°3] a) “Cuad.: 16B / N - / Prof.: 441 / Col.: F. Tello / Fecha:
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<date id="222165B7A63FFFB4FEB6FA3A9FEF558C" box="[350,461,1471,1495]" pageId="4" pageNumber="939" value="2021-05-07">
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07.
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<collectionCode id="308EDBB2A63FFFB4FE95FA3A9FB0558C" box="[381,402,1471,1495]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="939" type="Museum">V</collectionCode>
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.2021
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</date>
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/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63FFFB4FD8AFA3A9D7C558C" authority="MHMOP" authorityName="MHMOP" box="[610,862,1471,1495]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO MHMOP</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FC8FFA3A9DA2558C" box="[871,896,1471,1495]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
|
|
/16B-070521-
|
|
<collectionCode id="308EDBB2A63FFFB4FBFDFA3A9AA7558C" box="[1045,1157,1471,1495]" pageId="4" pageNumber="939">IMZCWS</collectionCode>
|
|
”, c) [mismos datos que N°1]. [N°4] a) “Cuad.: 17AA / N - / Prof.: 431 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63FFFB4FF7FFA7A9F24564C" box="[151,262,1535,1559]" pageId="4" pageNumber="939" value="2021-05-28">
|
|
28.
|
|
<collectionCode id="308EDBB2A63FFFB4FF5EFA7A9EE9564C" box="[182,203,1535,1559]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="939" type="Museum">V</collectionCode>
|
|
.2021
|
|
</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63FFFB4FE74FA7A9C01564C" box="[412,547,1535,1559]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FDD6FA7A9C88564C" box="[574,682,1535,1559]" pageId="4" pageNumber="939">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FD5BFA7A9CEE564C" box="[691,716,1535,1559]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
|
|
/16B-070521- Q6B2
|
|
<collectionCode id="308EDBB2A63FFFB4FC47FA7A9DF4564C" box="[943,982,1535,1559]" country="Austria" name="Zoological Collection, University of Vienna" pageId="4" pageNumber="939" type="University or college">ZC</collectionCode>
|
|
”, c) [mismos datos que N°1]. [N°5] a) “Cuad.: 12AA / N°247 / Prof.: 421 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63FFFB4FBEDF99A9A5F566C" box="[1029,1149,1567,1591]" pageId="4" pageNumber="939" value="2021-03-04">04.III.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63FFFB4FF35F9BA9F47560C" box="[221,357,1599,1623]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FE93F9BA9FCA560C" box="[379,488,1599,1623]" pageId="4" pageNumber="939">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FE19F9BA9C28560C" box="[497,522,1599,1623]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
|
|
/12AA-030421- ZTI80D”, c) [mismos datos que N°1]. [N°6] a) “Cuad.: 15A / N° - / Prof.: 420 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63FFFB4FCC4F9DA9D86562C" box="[812,932,1631,1655]" pageId="4" pageNumber="939" value="2021-04-28">28.IV.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63FFFB4FBD9F9DA9A9B562C" box="[1073,1209,1631,1655]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FF40F9FA9F3556CC" box="[168,279,1663,1687]" pageId="4" pageNumber="939">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FEF7F9FA9F1B56CC" box="[287,313,1663,1687]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
|
|
/15A-280421-
|
|
<collectionCode id="308EDBB2A63FFFB4FE3DF9FA9C3756CC" box="[469,533,1663,1687]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="4" pageNumber="939">MTD</collectionCode>
|
|
0
|
|
<collectionCode id="308EDBB2A63FFFB4FDCAF9FA9C6956CC" box="[546,587,1663,1687]" pageId="4" pageNumber="939">VH</collectionCode>
|
|
”, c) [mismos datos que N°1]. [N°7] a) “Cuad.: 16AA / N° 21 / Prof.: 410 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63FFFB4FD7DF91A9D2F56EC" box="[661,781,1695,1719]" pageId="4" pageNumber="939" value="2021-04-13">13.IV.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63FFFB4FC49F91A9A0B56EC" box="[929,1065,1695,1719]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="939" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FBABF91A9A9356EC" box="[1091,1201,1695,1719]" pageId="4" pageNumber="939">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63FFFB4FF7FF93A9E92568C" box="[151,176,1727,1751]" country="Russia" name="Paleontological Institute" pageId="4" pageNumber="939">PI</collectionCode>
|
|
/16AA-130421-
|
|
<collectionCode id="308EDBB2A63FFFB4FEB5F93A9FF5568C" box="[349,471,1727,1751]" pageId="4" pageNumber="939">NANQYE</collectionCode>
|
|
“, c) [mismos datos que N°1].
|
|
<emphasis id="64EB9F65A63FFFB4FCD0F93A9DD4568C" bold="true" box="[824,1014,1727,1751]" pageId="4" pageNumber="939">Hembras (n = 6)</emphasis>
|
|
[N°8] a) “Cuad.: 14A / N° - / Prof.: 397 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FD8EFFFF9CFC50C9" box="[614,734,122,146]" pageId="5" pageNumber="940" value="2021-03-24">24.III.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FC81FFFF9DD250C9" box="[873,1008,122,146]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FBEFFFFF9A5750C9" box="[1031,1141,122,146]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FFB3FF1F9E5750E9" box="[91,117,154,178]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/14A-240321-D5
|
|
<collectionCode id="308EDBB2A63EFFB5FED9FF1F9F7950E9" box="[305,347,154,178]" pageId="5" pageNumber="940">XA</collectionCode>
|
|
39”, c) “
|
|
<taxonomicName id="919F38F4A63EFFB5FE5CFF1F9A6C50E9" authority="Tello, Verdu, Rossini & Zunino, 2021" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[436,1102,154,178]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">
|
|
<emphasis id="64EB9F65A63EFFB5FE5CFF1F9CB950E9" box="[436,667,154,178]" italics="true" pageId="5" pageNumber="940">Onthophagus pilauco</emphasis>
|
|
<bibRefCitation id="320E3E86A63EFFB5FD49FF1F9A6C50E9" author="Tello, F. & Verdu, J. R. & Rossini, M. & Zunino, M." box="[673,1102,154,178]" pageId="5" pageNumber="940" pagination="133 - 145" refId="ref9640" refString="Tello, F., Verdu, J. R., Rossini, M. y Zunino, M. (2021) Onthophagus pilauco sp. nov. (Coleoptera, Scarabaeidae): evidence of beetle extinction in the Pleistocene - Holocene transition in Chilean Northern Patagonia. ZooKeys, 1043: 133 - 145." type="journal article" year="2021">Tello, Verdú, Rossini & Zunino, 2021</bibRefCitation>
|
|
</taxonomicName>
|
|
<specimenCount id="409988FEA63EFFB5FBBDFF1E9A4B50EE" box="[1109,1129,155,181]" count="1" pageId="5" pageNumber="940" type="female">♀</specimenCount>
|
|
”. [N°9] a) “Cuad.: 12AA / N°247 / Prof.: 241 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FC91FF3F9DD35089" box="[889,1009,186,210]" pageId="5" pageNumber="940" value="2021-03-04">04.III.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FFB3FF5F9EC050A9" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[91,226,218,242]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FF12FF5F9F4A50A9" box="[250,360,218,242]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FE99FF5F9FA950A9" box="[369,395,218,242]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/12AA-030421-7
|
|
<collectionCode id="308EDBB2A63EFFB5FDA1FF5F9C9150A9" box="[585,691,218,242]" pageId="5" pageNumber="940">VMHZV</collectionCode>
|
|
”, c) [mismos datos que N°8]. [N°10] a) “Cuad.: 16AA / N°- / Prof.: 460 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FD37FF7F9D7A5149" box="[735,856,250,274]" pageId="5" pageNumber="940" value="2021-04-30">30.IV.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FC0DFF7F9A4F5149" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[997,1133,250,274]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FFB3FE9F9EE85169" box="[91,202,282,306]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FF3AFE9F9ECE5169" box="[210,236,282,306]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/16AA-300421-
|
|
<collectionCode id="308EDBB2A63EFFB5FE76FE9F9FF35169" box="[414,465,282,306]" pageId="5" pageNumber="940">OUI</collectionCode>
|
|
12F”, c) [mismos datos que N°8]. [N°11] a) “Cuad.: 17A / N°- / Prof.: 407 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FDFAFEBF9CA85109" box="[530,650,314,338]" pageId="5" pageNumber="940" value="2021-04-07">07.IV.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FCF8FEBF9DBA5109" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[784,920,314,338]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FC45FEBF9A3E5109" box="[941,1052,314,338]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FBCCFEBF9A1C5109" box="[1060,1086,314,338]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/17A-070421-
|
|
<collectionCode id="308EDBB2A63EFFB5FF5EFEDF9F2A5129" box="[182,264,346,370]" pageId="5" pageNumber="940">RIVST</collectionCode>
|
|
3”, c) [mismos datos que N°8]. [N°12] a) “Cuad.: 12AA / N°- / Prof.: 241 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FEF5FEFF9FB751C9" box="[285,405,378,402]" pageId="5" pageNumber="940" value="2021-03-04">04.III.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FDCEFEFF9C8C51C9" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[550,686,378,402]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FD2FFEFF9D1651C9" box="[711,820,378,402]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FCD4FEFF9D7451C9" box="[828,854,378,402]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/12AA-040321-
|
|
<collectionCode id="308EDBB2A63EFFB5FBEDFEFF9A7151C9" box="[1029,1107,378,402]" pageId="5" pageNumber="940">OYCE</collectionCode>
|
|
76”, c) [mismos datos que N°8]. [N°13] a) “Cuad.: 16AA / N°- / Prof.: 460 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FFB3FE3F9EF15189" box="[91,211,442,466]" pageId="5" pageNumber="940" value="2021-04-30">30.IV.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FEB3FE3F9FC15189" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[347,483,442,466]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FE11FE3F9C445189" box="[505,614,442,466]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FD87FE3F9CAB5189" box="[623,649,442,466]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/16AA-301521-
|
|
<collectionCode id="308EDBB2A63EFFB5FCD0FE3F9DA45189" box="[824,902,442,466]" pageId="5" pageNumber="940">OYCE</collectionCode>
|
|
76”, c) [mismos datos que N°8].
|
|
<emphasis id="64EB9F65A63EFFB5FF3DFE5F9DBD51A9" bold="true" box="[213,927,474,498]" pageId="5" pageNumber="940">
|
|
Otro material examinado. Macho
|
|
<emphasis id="64EB9F65A63EFFB5FD98FE5F9C9551A9" bold="true" box="[624,695,474,498]" italics="true" pageId="5" pageNumber="940">minor</emphasis>
|
|
con cuerno (n = 1).
|
|
</emphasis>
|
|
[N° 14] a) “Cuad.: 17AA / N°- / Prof.: 431-430 / Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FD38FE7F9D6A5249" box="[720,840,506,530]" pageId="5" pageNumber="940" value="2021-05-28">28.05.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FC0BFE7F9A495249" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[995,1131,506,530]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FFB3FD9F9EEA5269" box="[91,200,538,562]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FF38FD9F9ECB5269" box="[208,233,538,562]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/12AA-040321- N4
|
|
<collectionCode id="308EDBB2A63EFFB5FE2FFD9F9C025269" box="[455,544,538,562]" pageId="5" pageNumber="940">MZXN</collectionCode>
|
|
”.
|
|
<emphasis id="64EB9F65A63EFFB5FDDFFD9F9E885209" bold="true" pageId="5" pageNumber="940">
|
|
Cuernos del clípeo de ejemplares macho
|
|
<emphasis id="64EB9F65A63EFFB5FBDEFD9F9A5F5269" bold="true" box="[1078,1149,538,562]" italics="true" pageId="5" pageNumber="940">major</emphasis>
|
|
(n = 4).
|
|
</emphasis>
|
|
[N° 15] a) “Cuad.: 10AA / N°- / Prof.: 490/ Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FC6FFDBF9DDD5209" box="[903,1023,570,594]" pageId="5" pageNumber="940" value="2021-05-25">25.05.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FFB3FDDF9EC05229" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[91,226,602,626]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FF11FDDF9F445229" box="[249,358,602,626]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FE86FDDF9FAA5229" box="[366,392,602,626]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/10AA-250521-
|
|
<collectionCode id="308EDBB2A63EFFB5FDD7FDDF9C8A5229" box="[575,680,602,626]" pageId="5" pageNumber="940">DNPDV</collectionCode>
|
|
9”. [N° 16] a) “Cuad.: 15AA / N°- / Prof.: 470/ Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FE65FDFF9C3252C9" box="[397,528,634,658]" pageId="5" pageNumber="940" value="2021-07-05">05.VII.2021</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FD72FDFF9D0052C9" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[666,802,634,658]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FCD7FDFF9D8E52C9" box="[831,940,634,658]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FC5DFDFF9DED52C9" box="[949,975,634,658]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/15AA-050721-
|
|
<collectionCode id="308EDBB2A63EFFB5FFB3FD1F9E9752E9" box="[91,181,666,690]" pageId="5" pageNumber="940">AGCM</collectionCode>
|
|
74”. [N° 17] a) “Cuad.: 16AA / N°- / Prof.: 410-419/ Col.: F. Tello / Fecha:
|
|
<date id="222165B7A63EFFB5FBEBFD1F9A5F52E9" box="[1027,1149,666,690]" pageId="5" pageNumber="940" value="2021-06-13">
|
|
13.
|
|
<collectionCode id="308EDBB2A63EFFB5FBC8FD1F9A1D52E9" box="[1056,1087,666,690]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="940">VI</collectionCode>
|
|
.2021
|
|
</date>
|
|
/ NE –“, b)
|
|
<taxonomicName id="919F38F4A63EFFB5FF0AFD3F9F4B5289" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[226,361,698,722]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="940" phylum="Arthropoda" rank="species" species="pilauco">“PILAUCO</taxonomicName>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FE6AFD3F9FD25289" box="[386,496,698,722]" pageId="5" pageNumber="940">MHMOP</collectionCode>
|
|
/
|
|
<collectionCode id="308EDBB2A63EFFB5FE10FD3F9C305289" box="[504,530,698,722]" country="Russia" name="Paleontological Institute" pageId="5" pageNumber="940">PI</collectionCode>
|
|
/16AA-130621- A8
|
|
<collectionCode id="308EDBB2A63EFFB5FD02FD3F9D135289" box="[746,817,698,722]" pageId="5" pageNumber="940">JRPD</collectionCode>
|
|
”. [N°18] [Mismos datos que N°3, añadido a la misma puntilla].
|
|
</paragraph>
|
|
<paragraph id="56204377A63EFFB5FFB3FC9C9E895569" blockId="5.[89,1151,793,1330]" pageId="5" pageNumber="940">
|
|
<emphasis id="64EB9F65A63EFFB5FFB3FC9C9CE15369" bold="true" box="[91,707,793,818]" pageId="5" pageNumber="940">
|
|
Descripción del macho (n=12). Cabeza (
|
|
<figureCitation id="CEA45FF2A63EFFB5FDA8FC9F9C945369" box="[576,694,794,818]" captionStart-0="Figuras 3-6" captionStart-1="Figuras 7-13" captionStartId-0="4.[151,232,1203,1225]" captionStartId-1="6.[151,232,1377,1399]" captionTargetBox-0="[214,1139,221,1180]" captionTargetBox-1="[201,1180,492,1357]" captionTargetId-0="figure-282@4.[151,1209,211,1199]" captionTargetId-1="figure-380@6.[151,1209,486,1371]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Figuras 3-6. Vista dorsal y laterodorsal de Onthophagus pilauco macho. 3, 5. Macho major. 4, 6. Macho minor. / Dorsal and laterodorsal view of male Onthophagus pilauco. 3, 5. Major male. 4, 6. Minor male." captionText-1="Figuras 7-13. Cuernos del clípeo presentes en los machos major y minor de Onthophagus pilauco. 7. Vista frontal del cuerno clipeal en la forma major. 8. Vista lateral del cuerno clipeal en la forma major. 9. Vista dorsal del cuerno clipeal en la forma major. 10. Reconstrucción fotográfica en vista frontal de la forma major. 11. Reconstrucción fotográfica en vista laterodorsal de la forma major. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma minor. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma minor. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both minor and major male forms of Onthophagus pilauco.7. Dorsal view of the clypeal horn in the major form. 8. Lateral view of the clypeal tubercle in the major form. 9. Dorsal view of the clypeal tubercle in the major form. 10. Photographic reconstruction of frontal view of the major form. 11. Photographic reconstruction in laterodorsal view of the major form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the minor form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the minor form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove." figureDoi-0="http://doi.org/10.5281/zenodo.13367592" figureDoi-1="http://doi.org/10.5281/zenodo.13367594" httpUri-0="https://zenodo.org/record/13367592/files/figure.png" httpUri-1="https://zenodo.org/record/13367594/files/figure.png" pageId="5" pageNumber="940">Figs. 3-13</figureCitation>
|
|
):
|
|
</emphasis>
|
|
Macho
|
|
<emphasis id="64EB9F65A63EFFB5FCCDFC9E9D415369" box="[805,867,795,818]" italics="true" pageId="5" pageNumber="940">major</emphasis>
|
|
de color predominante verde, cobrizo o negro, brillante o semiopaco; microescultura del tegumento lisa; clípeo subtrapezoidal, notoriamente curvado hacia arriba; disco del clípeo y frente con puntuación fina en la porción central, profunda y redondeada hacia el margen que divide el clípeo de la gena; genas claramente expandidas más allá del clípeo (margen de la cabeza con una muesca en la unión clípeogenal), notoriamente o moderadamente curvadas, sin puntuación aparente o con pocos puntos (~4) en la porción anterior; sutura clípeogenal entera hasta los cuernos frontales; carenas frontoclipeal y frontal ausentes; presenta un cuerno largo en el disco del clípeo, más ancho en la porción media y superior en vista frontal (
|
|
<figureCitation id="CEA45FF2A63EFFB5FBC3FB9F9A4D5469" box="[1067,1135,1050,1074]" captionStart="Figuras 7-13" captionStartId="6.[151,232,1377,1399]" captionTargetBox="[201,1180,492,1357]" captionTargetId="figure-380@6.[151,1209,486,1371]" captionTargetPageId="6" captionText="Figuras 7-13. Cuernos del clípeo presentes en los machos major y minor de Onthophagus pilauco. 7. Vista frontal del cuerno clipeal en la forma major. 8. Vista lateral del cuerno clipeal en la forma major. 9. Vista dorsal del cuerno clipeal en la forma major. 10. Reconstrucción fotográfica en vista frontal de la forma major. 11. Reconstrucción fotográfica en vista laterodorsal de la forma major. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma minor. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma minor. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both minor and major male forms of Onthophagus pilauco.7. Dorsal view of the clypeal horn in the major form. 8. Lateral view of the clypeal tubercle in the major form. 9. Dorsal view of the clypeal tubercle in the major form. 10. Photographic reconstruction of frontal view of the major form. 11. Photographic reconstruction in laterodorsal view of the major form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the minor form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the minor form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove." figureDoi="http://doi.org/10.5281/zenodo.13367594" httpUri="https://zenodo.org/record/13367594/files/figure.png" pageId="5" pageNumber="940">Fig. 7</figureCitation>
|
|
), cuerno con disco elipsoidal prominente en vista ventral (
|
|
<figureCitation id="CEA45FF2A63EFFB5FCE7FBBF9D545409" box="[783,886,1082,1106]" captionStart="Figuras 7-13" captionStartId="6.[151,232,1377,1399]" captionTargetBox="[201,1180,492,1357]" captionTargetId="figure-380@6.[151,1209,486,1371]" captionTargetPageId="6" captionText="Figuras 7-13. Cuernos del clípeo presentes en los machos major y minor de Onthophagus pilauco. 7. Vista frontal del cuerno clipeal en la forma major. 8. Vista lateral del cuerno clipeal en la forma major. 9. Vista dorsal del cuerno clipeal en la forma major. 10. Reconstrucción fotográfica en vista frontal de la forma major. 11. Reconstrucción fotográfica en vista laterodorsal de la forma major. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma minor. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma minor. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both minor and major male forms of Onthophagus pilauco.7. Dorsal view of the clypeal horn in the major form. 8. Lateral view of the clypeal tubercle in the major form. 9. Dorsal view of the clypeal tubercle in the major form. 10. Photographic reconstruction of frontal view of the major form. 11. Photographic reconstruction in laterodorsal view of the major form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the minor form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the minor form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove." figureDoi="http://doi.org/10.5281/zenodo.13367594" httpUri="https://zenodo.org/record/13367594/files/figure.png" pageId="5" pageNumber="940">Figs. 8-9</figureCitation>
|
|
); cuernos de la frente notoriamente achatados, expandidos hasta la base (carenados), notoriamente avanzados, es decir, ubicados en la posición medial del disco de la frente formando una línea con el ápice de la foseta del ojo, ambos cuernos de la frente forman una “
|
|
<collectionCode id="308EDBB2A63EFFB5FC88FB1F9D5554E9" box="[864,887,1178,1202]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="940" type="Museum">V</collectionCode>
|
|
” expandida sin unirse en la frente; ojos pequeños en vista dorsal, separados por al menos 12 veces el ancho de la foseta ocular, foseta más larga que ancha; quillas paraocular y postoccipital enteramente conectadas (sin división en la base de la frente), bien desarrolladas y sobresalientes en vista lateral.
|
|
</paragraph>
|
|
<paragraph id="56204377A63EFFB5FFB3FADC9EF75609" blockId="5.[90,1150,1369,1618]" pageId="5" pageNumber="940">
|
|
<emphasis id="64EB9F65A63EFFB5FFB3FADC9EF4552A" bold="true" box="[91,214,1369,1393]" pageId="5" pageNumber="940">Variación.</emphasis>
|
|
Los machos
|
|
<emphasis id="64EB9F65A63EFFB5FE86FADE9F8D5529" box="[366,431,1371,1394]" italics="true" pageId="5" pageNumber="940">minor</emphasis>
|
|
se diferencian de los
|
|
<emphasis id="64EB9F65A63EFFB5FD45FADE9CC95529" box="[685,747,1371,1394]" italics="true" pageId="5" pageNumber="940">major</emphasis>
|
|
por el tamaño y forma del cuerno del clípeo, siendo más largo en las formas
|
|
<emphasis id="64EB9F65A63EFFB5FD88FAFE9CBC55C9" box="[608,670,1403,1426]" italics="true" pageId="5" pageNumber="940">major</emphasis>
|
|
(
|
|
<figureCitation id="CEA45FF2A63EFFB5FD45FAFF9D0355C9" box="[685,801,1402,1426]" captionStart="Figuras 7-13" captionStartId="6.[151,232,1377,1399]" captionTargetBox="[201,1180,492,1357]" captionTargetId="figure-380@6.[151,1209,486,1371]" captionTargetPageId="6" captionText="Figuras 7-13. Cuernos del clípeo presentes en los machos major y minor de Onthophagus pilauco. 7. Vista frontal del cuerno clipeal en la forma major. 8. Vista lateral del cuerno clipeal en la forma major. 9. Vista dorsal del cuerno clipeal en la forma major. 10. Reconstrucción fotográfica en vista frontal de la forma major. 11. Reconstrucción fotográfica en vista laterodorsal de la forma major. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma minor. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma minor. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both minor and major male forms of Onthophagus pilauco.7. Dorsal view of the clypeal horn in the major form. 8. Lateral view of the clypeal tubercle in the major form. 9. Dorsal view of the clypeal tubercle in the major form. 10. Photographic reconstruction of frontal view of the major form. 11. Photographic reconstruction in laterodorsal view of the major form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the minor form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the minor form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove." figureDoi="http://doi.org/10.5281/zenodo.13367594" httpUri="https://zenodo.org/record/13367594/files/figure.png" pageId="5" pageNumber="940">Figs. 7-11</figureCitation>
|
|
), mientras que en las formas
|
|
<emphasis id="64EB9F65A63EFFB5FFB5FA1E9EBC55E9" box="[93,158,1435,1458]" italics="true" pageId="5" pageNumber="940">minor</emphasis>
|
|
el cuerno es reducido y tan ancho como largo, sin el disco elipsoidal prominente (
|
|
<figureCitation id="CEA45FF2A63EFFB5FF8BFA3F9EC05589" box="[99,226,1466,1490]" captionStart="Figuras 7-13" captionStartId="6.[151,232,1377,1399]" captionTargetBox="[201,1180,492,1357]" captionTargetId="figure-380@6.[151,1209,486,1371]" captionTargetPageId="6" captionText="Figuras 7-13. Cuernos del clípeo presentes en los machos major y minor de Onthophagus pilauco. 7. Vista frontal del cuerno clipeal en la forma major. 8. Vista lateral del cuerno clipeal en la forma major. 9. Vista dorsal del cuerno clipeal en la forma major. 10. Reconstrucción fotográfica en vista frontal de la forma major. 11. Reconstrucción fotográfica en vista laterodorsal de la forma major. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma minor. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma minor. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both minor and major male forms of Onthophagus pilauco.7. Dorsal view of the clypeal horn in the major form. 8. Lateral view of the clypeal tubercle in the major form. 9. Dorsal view of the clypeal tubercle in the major form. 10. Photographic reconstruction of frontal view of the major form. 11. Photographic reconstruction in laterodorsal view of the major form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the minor form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the minor form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove." figureDoi="http://doi.org/10.5281/zenodo.13367594" httpUri="https://zenodo.org/record/13367594/files/figure.png" pageId="5" pageNumber="940">Figs. 12-13</figureCitation>
|
|
); la cabeza de los machos
|
|
<emphasis id="64EB9F65A63EFFB5FDFCFA3E9C775589" box="[532,597,1467,1490]" italics="true" pageId="5" pageNumber="940">minor</emphasis>
|
|
es de menor tamaño, siendo estos ~0,8 veces el tamaño de uno
|
|
<emphasis id="64EB9F65A63EFFB5FEFCFA5E9F7055A9" box="[276,338,1499,1522]" italics="true" pageId="5" pageNumber="940">major</emphasis>
|
|
; tegumento opaco; clípeo levemente curvado hacia arriba; y cuernos de la frente notoriamente separados entre sí 0,5 veces el largo de los mismos, mientras que la separación de los cuernos de la frente en los machos
|
|
<emphasis id="64EB9F65A63EFFB5FD04F99E9D085669" box="[748,810,1563,1586]" italics="true" pageId="5" pageNumber="940">major</emphasis>
|
|
es significativamente menor (
|
|
<figureCitation id="CEA45FF2A63EFFB5FF8BF9BF9EE55609" box="[99,199,1594,1618]" captionStart="Figuras 3-6" captionStartId="4.[151,232,1203,1225]" captionTargetBox="[214,1139,221,1180]" captionTargetId="figure-282@4.[151,1209,211,1199]" captionTargetPageId="4" captionText="Figuras 3-6. Vista dorsal y laterodorsal de Onthophagus pilauco macho. 3, 5. Macho major. 4, 6. Macho minor. / Dorsal and laterodorsal view of male Onthophagus pilauco. 3, 5. Major male. 4, 6. Minor male." figureDoi="http://doi.org/10.5281/zenodo.13367592" httpUri="https://zenodo.org/record/13367592/files/figure.png" pageId="5" pageNumber="940">Figs. 3-6</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="56204377A63EFFB6FFB3F9FC9EC55189" blockId="5.[91,1149,1657,1747]" lastBlockId="6.[150,1211,122,466]" lastPageId="6" lastPageNumber="941" pageId="5" pageNumber="940">
|
|
<emphasis id="64EB9F65A63EFFB5FFB3F9FC9D0456C9" bold="true" box="[91,806,1657,1682]" pageId="5" pageNumber="940">
|
|
Redescripción de la hembra (n=6). Cabeza (
|
|
<figureCitation id="CEA45FF2A63EFFB5FD79F9FF9D3B56C9" box="[657,793,1658,1682]" captionStart="Figuras 14-17" captionStartId="7.[91,172,1190,1212]" captionTargetBox="[148,1111,280,1174]" captionTargetId="figure-108@7.[91,1149,267,1191]" captionTargetPageId="7" captionText="Figuras 14-17. 14-15. Vista dorsal de Onthophagus pilauco hembra. 16-17. Vista laterodorsal de Onthophagus pilauco hembra. / 14-15. Dorsal view of females Onthophagus pilauco. 16-17. Laterodorsal view of females Onthophagus pilauco." figureDoi="http://doi.org/10.5281/zenodo.13367596" httpUri="https://zenodo.org/record/13367596/files/figure.png" pageId="5" pageNumber="940">Figs. 14-17</figureCitation>
|
|
):
|
|
</emphasis>
|
|
Color predominante verde brillante o semi-opaco; microescultura del tegumento lisa; clípeo subtrapezoidal, leve o notoriamente curvado hacia arriba en su extremo; disco del clípeo y frente con puntuación fina en la porción central, profunda y redondeada llegando hasta la carena que divide el clípeo de la mejilla; porción central del surco del clípeo con una hendidura profunda y ovalada; genas claramente expandidas más allá del clípeo (margen de la cabeza con una muesca en la unión clípeogenal), notoriamente o moderadamente curvadas; carenas frontoclipeal y frontal ausentes; cuernos de la frente bien desarrollados, forma semitriangular o cónica, aguzado o redondeado en su ápice, notoriamente avanzados, es decir, ubicados en la posición medial del disco de la frente formando una línea con el ápice de la foseta del ojo; ojos pequeños en vista dorsal, foseta más larga que ancha, separados por al menos 10 veces el ancho de la foseta; quillas paraocular y postoccipital enteramente conectadas (sin división en la base de la frente), bien desarrolladas y sobresalientes en vista lateral.
|
|
</paragraph>
|
|
<caption id="02E013FFA63DFFB6FF7FFAE49C87568D" ID-DOI="http://doi.org/10.5281/zenodo.13367594" ID-Zenodo-Dep="13367594" httpUri="https://zenodo.org/record/13367594/files/figure.png" pageId="6" pageNumber="941" startId="6.[151,232,1377,1399]" targetBox="[201,1180,492,1357]" targetPageId="6" targetType="figure">
|
|
<paragraph id="56204377A63DFFB6FF7FFAE49C87568D" blockId="6.[150,1210,1377,1751]" pageId="6" pageNumber="941">
|
|
<emphasis id="64EB9F65A63DFFB6FF7FFAE49F01552D" bold="true" box="[151,291,1377,1399]" pageId="6" pageNumber="941">Figuras 7-13.</emphasis>
|
|
Cuernos del clípeo presentes en los machos
|
|
<emphasis id="64EB9F65A63DFFB6FCE4FAE79D66552C" box="[780,836,1378,1399]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
y
|
|
<emphasis id="64EB9F65A63DFFB6FC8AFAE79DBF552C" box="[866,925,1378,1399]" italics="true" pageId="6" pageNumber="941">minor</emphasis>
|
|
de
|
|
<emphasis id="64EB9F65A63DFFB6FC2FFAE49A82552D" box="[967,1184,1377,1398]" italics="true" pageId="6" pageNumber="941">
|
|
<taxonomicName id="919F38F4A63DFFB6FC2FFAE49ABE552D" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[967,1180,1377,1398]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="941" phylum="Arthropoda" rank="species" species="pilauco">Onthophagus pilauco</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
7. Vista frontal del cuerno clipeal en la forma
|
|
<emphasis id="64EB9F65A63DFFB6FDB6FAFA9CB455CF" box="[606,662,1407,1428]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
. 8. Vista lateral del cuerno clipeal en la forma
|
|
<emphasis id="64EB9F65A63DFFB6FB93FAFA9A9155CF" box="[1147,1203,1407,1428]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
. 9. Vista dorsal del cuerno clipeal en la forma
|
|
<emphasis id="64EB9F65A63DFFB6FD6DFA199C9F55EA" box="[645,701,1436,1457]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
. 10. Reconstrucción fotográfica en vista frontal de la forma
|
|
<emphasis id="64EB9F65A63DFFB6FEF6FA3F9F745594" box="[286,342,1466,1487]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
. 11. Reconstrucción fotográfica en vista laterodorsal de la forma
|
|
<emphasis id="64EB9F65A63DFFB6FBF5FA3F9A775594" box="[1053,1109,1466,1487]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
. 12 Vista frontal de un fragmento del clípeo y cuerno del clípeo de la forma
|
|
<emphasis id="64EB9F65A63DFFB6FCBAFA529DAF55B7" box="[850,909,1495,1516]" italics="true" pageId="6" pageNumber="941">minor</emphasis>
|
|
. 13. Vista laterodorsal de un fragmento del clípeo y cuerno del clípeo de la forma
|
|
<emphasis id="64EB9F65A63DFFB6FD27FA719D2B5652" box="[719,777,1524,1545]" italics="true" pageId="6" pageNumber="941">minor</emphasis>
|
|
. MC: margen del clípeo. TM: cuerno del disco del clípeo. SC: surco del clípeo. / Clypeal horns observed in both
|
|
<emphasis id="64EB9F65A63DFFB6FC95F9979D9A567C" box="[893,952,1554,1575]" italics="true" pageId="6" pageNumber="941">minor</emphasis>
|
|
and
|
|
<emphasis id="64EB9F65A63DFFB6FC02F9979A00567C" box="[1002,1058,1554,1575]" italics="true" pageId="6" pageNumber="941">major</emphasis>
|
|
male forms of
|
|
<emphasis id="64EB9F65A63DFFB6FF7FF9AB9F4E5618" box="[151,364,1582,1603]" italics="true" pageId="6" pageNumber="941">
|
|
<taxonomicName id="919F38F4A63DFFB6FF7FF9AB9F4A5618" authorityName="Tello, Verdu, Rossini & Zunino" authorityYear="2021" box="[151,360,1582,1603]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="941" phylum="Arthropoda" rank="species" species="pilauco">Onthophagus pilauco</taxonomicName>
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.
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</emphasis>
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7. Dorsal view of the clypeal horn in the
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<emphasis id="64EB9F65A63DFFB6FCFAF9AA9D68561F" box="[786,842,1583,1604]" italics="true" pageId="6" pageNumber="941">major</emphasis>
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form. 8. Lateral view of the clypeal tubercle in the
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<emphasis id="64EB9F65A63DFFB6FEDEF9C99F4C563A" box="[310,366,1612,1633]" italics="true" pageId="6" pageNumber="941">major</emphasis>
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form. 9. Dorsal view of the clypeal tubercle in the
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<emphasis id="64EB9F65A63DFFB6FC60F9C99DE2563A" box="[904,960,1612,1633]" italics="true" pageId="6" pageNumber="941">major</emphasis>
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form. 10. Photographic reconstruction of frontal view of the
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<emphasis id="64EB9F65A63DFFB6FDFDF9EF9C6F5624" box="[533,589,1642,1663]" italics="true" pageId="6" pageNumber="941">major</emphasis>
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form. 11. Photographic reconstruction in laterodorsal view of the
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<emphasis id="64EB9F65A63DFFB6FF34F9029F3656C7" box="[220,276,1671,1692]" italics="true" pageId="6" pageNumber="941">major</emphasis>
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form. 12. Frontal view of a fragment of the clypeus and clypeal horn of the
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<emphasis id="64EB9F65A63DFFB6FBD7F9029A5856C7" box="[1087,1146,1671,1692]" italics="true" pageId="6" pageNumber="941">minor</emphasis>
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form. 13. Laterodorsal view of a fragment of the clypeus and clypeal horn of the
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<emphasis id="64EB9F65A63DFFB6FC5FF9219DD056E2" box="[951,1010,1700,1721]" italics="true" pageId="6" pageNumber="941">minor</emphasis>
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form. MC: margin of clypeus. TM: Clypeal horn. SC: clypeal groove.
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</paragraph>
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</caption>
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<paragraph id="56204377A63CFFB7FFB3FFFC9A0650A9" blockId="7.[88,1150,121,243]" pageId="7" pageNumber="942">
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<emphasis id="64EB9F65A63CFFB7FFB3FFFC9F7C50C9" bold="true" box="[91,350,121,146]" pageId="7" pageNumber="942">Variación en hembras</emphasis>
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. Largo máximo de la cabeza: 2,0-
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<quantity id="9167EE92A63CFFB7FD3EFFFF9D0C50C9" box="[726,814,122,146]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="942" unit="mm" value="2.5">2,5 mm</quantity>
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. Ancho máximo de la cabeza: 1,8-3,0 mm. Se observa una leve variación entre hembras; aquellos de aspecto brillante presentan cuernos aguzados en su ápice y de tamaño mayor (
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<figureCitation id="CEA45FF2A63CFFB7FCC0FF3F9D885089" box="[808,938,186,210]" captionStart="Figuras 14-17" captionStartId="7.[91,172,1190,1212]" captionTargetBox="[148,1111,280,1174]" captionTargetId="figure-108@7.[91,1149,267,1191]" captionTargetPageId="7" captionText="Figuras 14-17. 14-15. Vista dorsal de Onthophagus pilauco hembra. 16-17. Vista laterodorsal de Onthophagus pilauco hembra. / 14-15. Dorsal view of females Onthophagus pilauco. 16-17. Laterodorsal view of females Onthophagus pilauco." figureDoi="http://doi.org/10.5281/zenodo.13367596" httpUri="https://zenodo.org/record/13367596/files/figure.png" pageId="7" pageNumber="942">Figs. 14, 16</figureCitation>
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), mientras que las de aspecto semiopaco presentan cuernos cónicos y de menor tamaño (
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<figureCitation id="CEA45FF2A63CFFB7FC7BFF5F9A3550A9" box="[915,1047,218,242]" captionStart="Figuras 14-17" captionStartId="7.[91,172,1190,1212]" captionTargetBox="[148,1111,280,1174]" captionTargetId="figure-108@7.[91,1149,267,1191]" captionTargetPageId="7" captionText="Figuras 14-17. 14-15. Vista dorsal de Onthophagus pilauco hembra. 16-17. Vista laterodorsal de Onthophagus pilauco hembra. / 14-15. Dorsal view of females Onthophagus pilauco. 16-17. Laterodorsal view of females Onthophagus pilauco." figureDoi="http://doi.org/10.5281/zenodo.13367596" httpUri="https://zenodo.org/record/13367596/files/figure.png" pageId="7" pageNumber="942">Figs. 15, 17</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |