201 lines
25 KiB
XML
201 lines
25 KiB
XML
<document id="8679D963F126EC3D489E0C707B772B6D" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.24505" ID-GBIF-Dataset="4d1c8337-e338-4fc8-aeae-16e9f0f2ef6d" ID-Pensoft-Pub="2535-0730-1-45" ID-ZooBank="2949085F15034A9A9791EAC7B4E87FDF" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-1-45" ModsDocOrigin="Evolutionary Systematics 2 (1)" ModsDocTitle="A new species of Charinus (Amblypygi: Charinidae) from Ghana, with notes on West African whip spiders" checkinTime="1558621786533" checkinUser="pensoft" docAuthor="Harms, Danilo" docDate="2018" docId="C6CC7B938328430B7B9966218F23541D" docLanguage="en" docName="EvolutSyst 2(1): 45-53" docOrigin="Evolutionary Systematics 2 (1)" docSource="http://dx.doi.org/10.3897/evolsyst.2.24505" docTitle="Charinus kakum Harms, 2018, sp. n." docType="treatment" docUuid="FFC44C02-FB47-4B54-BE2B-25A07EEEFF95" docUuidSource="ZooBank" docVersion="5" lastPageNumber="45" masterDocId="A813FFBAFFC48D7AFF82FFF1FF81FFC5" masterDocTitle="A new species of Charinus (Amblypygi: Charinidae) from Ghana, with notes on West African whip spiders" masterLastPageNumber="53" masterPageNumber="45" pageNumber="45" updateTime="1701316493799" updateUser="plazi">
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<mods:title id="04EFC7ADC9B1F6A5C027B82A1A107FE6">A new species of Charinus (Amblypygi: Charinidae) from Ghana, with notes on West African whip spiders</mods:title>
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<mods:name id="355BEDD8831DC3490AB98014A0C927C7" type="personal">
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<mods:roleTerm id="57719B765BD61B0FC96ED87C4356533A">Author</mods:roleTerm>
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<mods:namePart id="0761ED7A5F33DFA4E0E8CF2BA19D0CAD">Harms, Danilo</mods:namePart>
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<mods:typeOfResource id="185C755834E9281DA915B778FAE8DAA5">text</mods:typeOfResource>
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<mods:titleInfo id="CF6BDC13598FEE22434D37E85A813D13">
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<mods:title id="6FA4A5A91989B90F711A730382F0D23A">Evolutionary Systematics</mods:title>
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<mods:part id="E8C9B186CC96FD059687AC6465EC1C9B">
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<mods:date id="0BC44327360C0033674182345E2DA946">2018</mods:date>
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<mods:number id="286CCE9519D080D9E6CF27F46E4FC2AE">2</mods:number>
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<mods:detail id="BDC596206687CE8084D4AB9159B2A799" type="issue">
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<mods:number id="65ED427DF0183350BFCB0948B3784B83">1</mods:number>
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<mods:start id="04CE17F432802B35ACEC68E441EC02C8">45</mods:start>
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<mods:url id="FC036E8C61713B5ED59D4FBFF3BE4CA3">http://dx.doi.org/10.3897/evolsyst.2.24505</mods:url>
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<mods:classification id="A737C3E414B533AF473E2F0D8940329E">journal article</mods:classification>
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<mods:identifier id="155512E0AA7881490A9EDAB8A2B131FC" type="DOI">http://dx.doi.org/10.3897/evolsyst.2.24505</mods:identifier>
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<treatment id="C6CC7B938328430B7B9966218F23541D" ID-GBIF-Taxon="157251974" LSID="urn:lsid:zoobank.org:act:FFC44C02-FB47-4B54-BE2B-25A07EEEFF95" httpUri="http://treatment.plazi.org/id/C6CC7B938328430B7B9966218F23541D" lastPageNumber="45" pageId="0" pageNumber="45">
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<subSubSection id="BF4E40966395525AA13226D600348BBB" pageId="0" pageNumber="45" type="nomenclature">
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<paragraph id="C8A7CA7AA2ABA625F1B91E3271B6DBD9" pageId="0" pageNumber="45">
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<taxonomicName id="9263AEA4F74FC383DE0D02552E13BD1B" ID-CoL="TNVF" LSID="http://zoobank.org/FFC44C02-FB47-4B54-BE2B-25A07EEEFF95" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
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<taxonomicNameLabel id="4042C38F7E2807E7977AF2B33D8ECD53" pageId="0" pageNumber="45">sp. n.</taxonomicNameLabel>
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(Figs 1, 2, 3, 4, 5)
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</paragraph>
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</subSubSection>
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<subSubSection id="3786C45C240BD539BBA85A1822F54141" pageId="0" pageNumber="45" type="type material">
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<paragraph id="136FFFBE55D74B2F48425D1AB82388A1" pageId="0" pageNumber="45">Type material.</paragraph>
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<paragraph id="EC92EACE4265F395913C02FACA2DC009" pageId="0" pageNumber="45">
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Female holotype: GHANA, Central Province, Kakum National Park, Track to Treehouse (
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<geoCoordinate id="49B6832C1630577522FA20744359A300" direction="north" orientation="latitude" precision="1" value="5.355897">
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5°21
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<normalizedToken id="E3C4429E60E419BBCC090543115B9C48" originalValue="’21.23”">'21.23"</normalizedToken>
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N
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</geoCoordinate>
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;
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<geoCoordinate id="D689A575B2EB1F6470578D6A51D4CC98" direction="north" orientation="latitude" precision="1" value="1.3821862">
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1°22
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<normalizedToken id="93A60111DA963792C12F56939481984F" originalValue="’55.87”">'55.87"</normalizedToken>
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N
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</geoCoordinate>
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), under flat rock near forest floor, 13.XII. 2017, coll. D. Harms and B.K. Williams (ZMH-A0000893).
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</paragraph>
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</subSubSection>
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<subSubSection id="7B36C3E653175C025E4634E558387EBB" pageId="0" pageNumber="45" type="diagnosis">
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<paragraph id="7C3C5BED0C5F5E12D85D94D069C7526D" pageId="0" pageNumber="45">Diagnosis.</paragraph>
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<paragraph id="87B0048916C12489917923DBC312F9B2" pageId="0" pageNumber="45">
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<taxonomicName id="5CF8535FE6BE6B4F5984D61E15E69E83" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
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sp. n. differs from the cave-inhabiting species
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<taxonomicName id="D5F25910ECF516971B925EED1AFEA159" lsidName="C. milloti" pageId="0" pageNumber="45" rank="species" species="milloti">C. milloti</taxonomicName>
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Fage, 1939 and
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<taxonomicName id="1B8C42A2AB2027998BED965CD5F576ED" lsidName="C. fagei" pageId="0" pageNumber="45" rank="species" species="fagei">C. fagei</taxonomicName>
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Weygoldt, 1972 in low number of tibial segments on leg IV (5 in
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<taxonomicName id="CA9A55EACB521F05738A99C9387F29D2" lsidName="C. millotei" pageId="0" pageNumber="45" rank="species" species="millotei">C. millotei</taxonomicName>
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and
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<taxonomicName id="A0B01089CF0AF8A070F966FF84083802" lsidName="C. fagei" pageId="0" pageNumber="45" rank="species" species="fagei">C. fagei</taxonomicName>
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vs. 3 in
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<taxonomicName id="668F70C03053B7DFAC1BC207BAF4CE55" lsidName="C. kakum" pageId="0" pageNumber="45" rank="species" species="kakum">C. kakum</taxonomicName>
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), trichobothria patterns of leg IV (compare with
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<bibRefCitation id="12A2067AA82E8BFA307035B10DBE95EE" author="Weygoldt, P" journalOrPublisher="Zoologische Jahrbuecher, Abteilung fuer Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="45" pagination="107 - 132" title="Charontidae (Amblypygi) aus Brasilien." volume="99" year="1972">Weygoldt 1972</bibRefCitation>
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,
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<bibRefCitation id="380BE5F32DCA632509B27D4DD4FCA757" pageId="0" pageNumber="45">2000a</bibRefCitation>
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), small body size (prosoma length 2.6 in
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<taxonomicName id="49806FBEF11D44DF176ED5363AE20907" lsidName="C. kakum" pageId="0" pageNumber="45" rank="species" species="kakum">C. kakum</taxonomicName>
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), and shorter legs; from the island species
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<taxonomicName id="700CC837219C8B99FC83888ABF653FBA" lsidName="C. africanus" pageId="0" pageNumber="45" rank="species" species="africanus">C. africanus</taxonomicName>
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<bibRefCitation id="167F9709EF670E59611AB941BA2C406C" pageId="0" pageNumber="45">Hansen 1921</bibRefCitation>
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through the female genital operculum which has a steep ventral flexure at about two third of its length in
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<taxonomicName id="FBA617E559CCCF9E9A6648B63FD4A4C4" lsidName="C. africanus" pageId="0" pageNumber="45" rank="species" species="africanus">C. africanus</taxonomicName>
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(
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<bibRefCitation id="B982CF362883358E13B9A7F70DB0F1F1" author="Weygoldt, P" journalOrPublisher="Zoologische Jahrbuecher, Abteilung fuer Systematik, Oekologie und Geographie der Tiere" pageId="0" pageNumber="45" pagination="107 - 132" title="Charontidae (Amblypygi) aus Brasilien." volume="99" year="1972">Weygoldt 1972</bibRefCitation>
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;
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<bibRefCitation id="211D423599419F4E110B529191FBBED5" author="Miranda, GS" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="45" pagination="1 - 17" title="A new species of Charinus Simon, 1892 (Arachnida: Amblypygi: Charinidae) from Israel and new records of C. ioanniticus (Kritscher, 1959)." url="https://doi.org/10.5852/ejt.2016.234" volume="234" year="2016 b">Miranda et al. 2016b</bibRefCitation>
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) but is uniform in
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<taxonomicName id="C378CDA1FCEE1A6E6889E926EC06382B" lsidName="C. kakum" pageId="0" pageNumber="45" rank="species" species="kakum">C. kakum</taxonomicName>
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, and body size (ca. 8.0-8.5 mm in
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<taxonomicName id="61240483CD0613B76F3975E484D9D0AF" lsidName="C. africanus" pageId="0" pageNumber="45" rank="species" species="africanus">C. africanus</taxonomicName>
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and 5.8 in
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<taxonomicName id="404D872F4F092CF3BD7DAADBEE34DE66" lsidName="C. kakum" pageId="0" pageNumber="45" rank="species" species="kakum">C. kakum</taxonomicName>
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;
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<bibRefCitation id="4F0249F8A29394EA7B9C015707DECF5A" pageId="0" pageNumber="45">Hansen 1921</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="646574F9A5F53BD9F83E87A85DFDF974" pageId="0" pageNumber="45" type="description">
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<paragraph id="5F57EAA997D96737FA4172C6D8A4660C" pageId="0" pageNumber="45">Description.</paragraph>
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<paragraph id="52C008B6F5FB86D5C965E00717DF2796" pageId="0" pageNumber="45">
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Carapace: flattened and wider than long (Figs 3A, C), uniformly brown except for the margins that are pale. Ratio length/width approximately
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<normalizedToken id="E4486D8FEC69DA4C012E5BE53835BF0B" originalValue="¾">3/4</normalizedToken>
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. Anterior margin rounded and with six spines situated anterior to the ME, individual spines projecting upwards. ME well developed, projecting forwards and situated on a common dark tubercle. LE well developed and on a common tubercle, projecting sideward. Fovea well developed and rounded, followed by a triangular depression at the posterior margin of the carapace. Three additional depressions in lateral position: i) smallest situated posterior to the LE; ii) a second larger depression in mediolateral position; and iii) a posterior-lateral position. Carina runs from the corners of the anterior margin and extends from the coxae of legs I to the corners of the posterior margin. Small granules and punctuations present, denser at the anterior region and pars cephalica. Sternum: tri-segmented but individual segments weakly sclerotized and rounded (Fig. 3B, D), pale except for the reddish-brown sternites. Tritosternum projected anteriorly and reaching far into the coxae of the pedipalps, elongate and cone-shaped, with one apical pair of spines, one medial pair, and ca. 6 spinelets at the base, distal margins serrate, base more or less triangular. Tetrasternum (second segment) rounded and hard to discern, with a pair of spinelets. Pentasternum (third segment) reduced and hard to discern, with a pair of setae (or small spinelets). Two additional setae between coxae of leg IV. The segments separated from each other by ca. 1.5 times the diameter of the tetrasternum. Opisthosoma: ovate and light brown, except for the tergites that are brown (Figs 2 C; 3A, B). Tergites with punctuations, in particular the distal positions. Tergites distally also with a small longitudinal row of up to 6 tiny setae. Chelicera: Cheliceral furrow with 4 teeth of the basal segment (Figs 4D, E; 5B). Proximal tooth largest, size range: IV>I>II>III. Distal tooth bifid and distal cusp larger than proximal cusp. Claw with 5 basal teeth, the 3 proximal teeth on a common base; some fine hairs at the retrolateral margin, condyle strongly sclerotized and reddish-brown, fang brown. Basal segment dorsally with four setae: first one situated distally near fang condyle, two medio-distal, and one in medial position. Both sides of basal segment with a longitudinal band of five or six fine setae. Ventral side of basal segment setose and reddish. Pedipalp: Trochanter (Fig. 4
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<normalizedToken id="C3136DEABB362C305F1B4BEF890E07EC" originalValue="A–C">A-C</normalizedToken>
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): Ventral apophysis sharply pointed and with a series of 14 strong spines (Fig. 4C); distal three largest, proximally decreasing in size; prolateral side with one spine and 3 spinelets (Fig. 4A). Femur: with three rows of spines, prolateral row with 5 setiferous tubercles, median row with large 3 spines (size ranges distal to proximal II>III>I), and retrolateral row with 4 spines (size range distal to proximal II>III>I>IV). Spines of retrolateral row largest. Size distal to proximal II>III>I>IV. Some setae present distally in pro- and retrolateral position (Fig. 4A, B; 5A). Tibia: with typical charinid spine armature (
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<bibRefCitation id="624028DA85F45F955229E2173064DFAD" pageId="0" pageNumber="45">Weygoldt 2000a</bibRefCitation>
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). Retrolateral side with 3 spines and one distal setiferous tubercle; distal spine largest and and spines proximally decreasing in length (III>II>I), distal setiferous tubercle near base of basitarsus in front of largest spine; Prolateral side with 2 spines and one proximal spinelet, distal spine largest. Dorsal side of tibia adorned with setae that distally decrease in length. Basitarsus: broadly flattened and each side with 2 spines, distal ones largest but proximal spine in retrolateral position much smaller (Fig. 4A). Retrolateral surface with several spinelets, prolateral surface smooth and with a median row of three trichobothria-like setae plus three additional setae more retrolaterally. Distitarsus: with 2 small spines above the cleaning organ, distal one about twice the size of proximal spine. Dorsal side setose, ventral side with ca. 10 long setae which are distally serrate. Cleaning organ occupies about 1/2 of the article length. Claw: long and with sharp curved tip, ca. 2/3 the length of distitarsus, divided from distitarsus and not fused. Legs: All moderately setose. Femur lengths: I>III>II>IV. Leg I: tibia with 24 articles and tarsus with 41 articles. Leg IV: Barsitibia with 3 pseudoarticles and distitarsus undivided. Trichobothria pattern: pseudosegment one: 1, two: 1, three: 11 (Fig. 5D). All claws smooth and not serrate. All walking legs with pulvilli that are slightly smaller than the claws. Genitalia: Genital operculum rounded but distolateral margins concave (Fig. 2B). Sparsely setose in the proximal section but distal margins with ca. 22 setae. Genital opening rounded and slightly concave in distal view. Ventral sac covers visible and medially fused. Gonopods longer than wide and with rounded openings (Fig. 5C), separated by ca. the diameter of their base; otherwise indistinct and comparable to other species of the genus (e.g. compare
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<bibRefCitation id="D572CAE5FFC899596809B38E3AB51421" author="Vasconcelos, ACO" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="277 - 292" title="Two new species of cave-dwelling Charinus Simon, 1892 from Brazil (Arachnida: Amblypygi: Charinidae)." url="https://doi.org/10.11646/zootaxa.4312.2.4" volume="4312" year="2017">Vasconcelos and Ferreira 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="AE56971CBAECD6763D5FF8563F432C1F" pageId="0" pageNumber="45" type="colour">
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<paragraph id="F7F5F5EAE2BEE1CBE6036AE9A47C426C" pageId="0" pageNumber="45">Colour.</paragraph>
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<paragraph id="423739E9F76AC063F99B926583D339B1" pageId="0" pageNumber="45">Body chestnut brown (dark brown in the life animal), abdomen yellowish except for the tergites. Chelicerae and distal sections of the pedipalp reddish (Figs 2A; 3A, B).</paragraph>
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</subSubSection>
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<subSubSection id="FFEA993EA051B3AAAA9D5E6C8AB6BFDE" pageId="0" pageNumber="45" type="description">
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<paragraph id="BC08734414D9793CE70BC5DF4C7E2D9E" pageId="0" pageNumber="45">Measurements.</paragraph>
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<paragraph id="5B219F91A0B3554257DBC56C2A516EE2" pageId="0" pageNumber="45">Total length 5.8: Carapace: Length: 2.6; Width: 1.9; Opisthosoma Length: 3.3; Opisthosoma Width: 2.25. Left Pedipalp: Femur: 1.35; Tibia: 1.66; Basitarsus: 0.57; Distitarsus: 0.58; Claw 0.43. Chelicera (basal segment): Length: 0.85. Width: 0.53.</paragraph>
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</subSubSection>
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<subSubSection id="DE06C34D0897B46A541B5AEC35F36181" pageId="0" pageNumber="45" type="etymology">
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<paragraph id="5AA4E9DDD476726B1CC35A9AB49456E7" pageId="0" pageNumber="45">Etymology.</paragraph>
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<paragraph id="200CDDE1C41A1221F6413B1B268BA5D8" pageId="0" pageNumber="45">The specific epithet refers to the type locality, Kakum National Park in Ghana.</paragraph>
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</subSubSection>
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<subSubSection id="3AC0FF87F3407440F4AE355CF9077953" pageId="0" pageNumber="45" type="co1 gene sequence. genbank submssion no. mh107031">
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<paragraph id="CE1E9530721FD9958FC7554046B74976" pageId="0" pageNumber="45">CO1 gene sequence. GenBank Submssion No. MH107031</paragraph>
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<paragraph id="F812D370576D3DDD538A58274C5FAB85" pageId="0" pageNumber="45">.</paragraph>
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</subSubSection>
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<subSubSection id="33F1195790CE583442937649766965FF" pageId="0" pageNumber="45" type="distribution">
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<paragraph id="718E1E18F6BDCE13BF2873DD2C65F5C7" pageId="0" pageNumber="45">Distribution.</paragraph>
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<paragraph id="557B0CFC68E965E40FF5971DBF7EF164" pageId="0" pageNumber="45">Only known from the type locality by a single female individual. The wider distribution remains unknown and unfortunately no other specimens could be collected, despite targeted searches in the vicinity of the type locality.</paragraph>
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</subSubSection>
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<subSubSection id="5789CCB2F8004628D587601F20E2F74F" pageId="0" pageNumber="45" type="habitat">
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<paragraph id="B475EEE861F58E38056D1F744525145D" pageId="0" pageNumber="45">Habitat.</paragraph>
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<paragraph id="67C6054165B8D58E401AC8AE598BCF34" pageId="0" pageNumber="45">
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The species was collected in closed primary rainforest in Kakum National Park, Ghana (Fig. 1). It was found in a small rocky outcrop close to a dry creek-bed under a rock which was positioned directly on top soil. Other arachnids that were seen at this outcrop included
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<taxonomicName id="A2EA589EB3605EAD96749DAC305045DF" class="Arachnida" family="Ricinoididae" genus="Ricinoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ricinoides" order="Ricinulei" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Ricinoides</taxonomicName>
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sp. (
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<taxonomicName id="A2C391F9F2B359E5A34E73AE1917AB9C" lsidName="" order="Ricinulei" pageId="0" pageNumber="45" rank="order">Ricinulei</taxonomicName>
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) and schizomids (
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<taxonomicName id="5249F9C2FEB30FDCACC9F8BC6D243F78" lsidName="" order="Uropygi" pageId="0" pageNumber="45" rank="order">Uropygi</taxonomicName>
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:
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<taxonomicName id="65B8C34FC791DB28B50F905553203A04" lsidName="" order="Schizomida" pageId="0" pageNumber="45" rank="order">Schizomida</taxonomicName>
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). No other specimens could be found under adjacent rocks or tree bark. Kakum National Park comprises moist evergreen (where the holotype was collected) and swamp or riverine forests. Average annual rainfall is ca. 1380 mm and the elevation range is 135-250 meters.
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</paragraph>
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</subSubSection>
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<subSubSection id="2DC389187412C3304D6E36078B1941FB" pageId="0" pageNumber="45" type="notes on biology">
|
||
<paragraph id="86BE2F3D532A3C61B9CE63BAA5E89367" pageId="0" pageNumber="45">Notes on biology.</paragraph>
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<paragraph id="2B20345363B903665121FE4ECBEFE0B8" pageId="0" pageNumber="45">The collected female was brooding and carried a total of seven eggs in a brood sac. These eggs were large, ca. 1.25 mm in diameter (Fig. 2D). Considering the low number of eggs and their large size, the reproductive potential of this species is considered to be low. Nothing else is known about the biology and distribution of this species. So far, it is the smallest whip spider in western Africa and may have a preference for rocky boulders in densely vegetated areas, under which it retreats.</paragraph>
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<caption id="6DF4DFE0744BC7309081B1ACE18F8F10" pageId="0" pageNumber="45">
|
||
<paragraph id="2F12B98F8DF2B780CD180C6D1DBBBBB3" pageId="0" pageNumber="45">
|
||
Figure 1. Distribution map showing the type localities of the described species of
|
||
<taxonomicName id="04CFEDBB9D567549DCD4BCF3E79EC392" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Charinus</taxonomicName>
|
||
in West Africa according to
|
||
<bibRefCitation id="FDD9B0F0AE7165A9F2570AE6F3FF6A42" author="Harvey, MS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" url="http://www.museum.wa.gov.au/catalogues/whip-spiders" year="2013">Harvey (2013)</bibRefCitation>
|
||
. The type locality of
|
||
<taxonomicName id="C0C19B992D4B57F503F2C73DB44004A8" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
|
||
sp. n. is highlighted in red. Note that the localities for
|
||
<taxonomicName id="9C8657B03590627546CB75E235F566E0" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus milloti" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="milloti">Charinus milloti</taxonomicName>
|
||
and
|
||
<taxonomicName id="F63A31284D6D41F3A1AF426C10EFBD3C" lsidName="C. fagei" pageId="0" pageNumber="45" rank="species" species="fagei">C. fagei</taxonomicName>
|
||
may contain some error because both species were originally described from caves with an unknown location. The symbols indicate the regions in which the caves may be located.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E4E15671BBC984A2391D1C9A866F7B21" pageId="0" pageNumber="45">
|
||
<paragraph id="65CDED4252ECBEDD1AEFBF22D73F6F92" pageId="0" pageNumber="45">
|
||
Figure 2. Female holotype of
|
||
<taxonomicName id="BA53F04825A9AE10BE683CD50C9CD921" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
|
||
sp. n.: A, habitus dorsal; B, genital operculum in ventral view. Arrow indicates the position of the ventral sac covers; C, opisthosoma in dorsal view; D, eggs carried by the female (removed from the genital operculum for imaging).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="30908B3F62DD4D4D6E4A10721C62609F" pageId="0" pageNumber="45">
|
||
<paragraph id="A45EDF7E98BD07531F82534337E80C95" pageId="0" pageNumber="45">
|
||
Figure 3. Female holotype of
|
||
<taxonomicName id="B05EDAF51EB11842E9D040DC33E5EC5E" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
|
||
sp. n.: A, details of habitus in dorsal view; B, details of habitus in ventral view; C, carapace in dorsal view; D, prosomal sternites in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9588C5A0377BE250E19B66E25A6E3B99" pageId="0" pageNumber="45">
|
||
<paragraph id="15DBD7A5DB5B081E1FE2F90969F8EA5B" pageId="0" pageNumber="45">
|
||
Figure 4. Female holotype of
|
||
<taxonomicName id="E8188D438AF3CDD846644B8517C34CCA" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
|
||
sp. n.: A, left pedipalp in prolateral view. Arrow points to the position of the cleaning organ; B, left pedipalp in retrolateral view; C, ventral spiniform apophysis, retrolateral view; D, right chelicera in retrolateral view; E, same in prolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="91F5CA29F211601E6CEB7C078A434A4B" pageId="0" pageNumber="45">
|
||
<paragraph id="0632685B8727891DA6C72E410509B9F9" pageId="0" pageNumber="45">
|
||
Figure 5. Female holotype of
|
||
<taxonomicName id="74261706C3F7D5EAAC437654E1A40927" class="Arachnida" family="Charinidae" genus="Charinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charinus kakum" order="Amblypygi" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="kakum">Charinus kakum</taxonomicName>
|
||
sp. n.: A, left pedipalp in prolateral view; B, right chelicera in retrolateral view; C, female gonopods, dorsal view; and D, trichobothria patterns on leg IV.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |