treatments-xml/data/A3/88/A6/A388A615077953D8889F95AD493D28E9.xml

<document ID-CLB-Dataset="229646" ID-DOI="http://dx.doi.org/10.3897/zse.99.95923" ID-GBIF-Dataset="b0059ad9-fa19-4539-821a-0bec19b167c9" ID-Pensoft-Pub="1860-0743-1-77" ID-Pensoft-UUID="F3CD023B35EE51769693A41205F7F028" ID-ZooBank="4D1F8139D4204E6E9B8E2401EF6232DA" ModsDocID="1860-0743-99-1-77" checkinTime="1674309841633" checkinUser="pensoft" docAuthor="Makino, Tomohisa, Nakano, Takafumi, Okamoto, Taku &amp; Hikida, Tsutomu" docDate="2023" docId="A388A615077953D8889F95AD493D28E9" docLanguage="en" docName="ZoosystEvol 99(1): 77-91" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/zse.99.95923" docTitle="Ateuchosaurus pellopleurus" docType="treatment" docVersion="6" id="F3CD023B35EE51769693A41205F7F028" lastPageNumber="77" masterDocId="F3CD023B35EE51769693A41205F7F028" masterDocTitle="Taxonomic revision and re-description of Ateuchosaurus pellopleurus (Hallowell, 1861) (Reptilia, Squamata, Scincidae) with resurrection of A. okinavensis (Thompson, 1912)" masterLastPageNumber="91" masterPageNumber="77" pageNumber="77" updateTime="1732849130218" updateUser="ExternalLinkService">
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<mods:title id="EC0DDAAF48784C9070D7EFCF25EF6791">Taxonomic revision and re-description of Ateuchosaurus pellopleurus (Hallowell, 1861) (Reptilia, Squamata, Scincidae) with resurrection of A. okinavensis (Thompson, 1912)</mods:title>
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<mods:namePart id="F49D399B01F4AAEF0A3DD7EB897AF867">Makino, Tomohisa</mods:namePart>
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<mods:affiliation id="A7554C238BDC9953F2D436528AFC0885">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
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<mods:namePart id="294469909063BDA51DC1FDBAEB707C33">Nakano, Takafumi</mods:namePart>
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<mods:affiliation id="8CF177EAB36B25C3C6EE986BC21905AE">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
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<mods:namePart id="1F17A03EA4C3F07DDAB51338E250126E">Okamoto, Taku</mods:namePart>
<mods:affiliation id="D5CF45A0D6525B25D6752C4E2BE440C5">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
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<mods:namePart id="FD700CC74D79CE2573D936E0134F4197">Hikida, Tsutomu</mods:namePart>
<mods:affiliation id="943C0E9271E748CC81F73718802147D5">Department of Zoology, Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto 606 - 8502, Japan</mods:affiliation>
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<mods:date id="001F0EB47C670FD57B58B05264B9DD9B">2023</mods:date>
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<subSubSection id="88B85DE0B99C55A8FCB00DA43BE78C64" pageId="0" pageNumber="77" type="nomenclature">
<paragraph id="74CA745910A167E132B5450E1EB4E24A" pageId="0" pageNumber="77">
<taxonomicName id="D83ED26BE4188101C819040CF2FD4A15" ID-CoL="67VM7" ID-ENA="316453" LSID="A388A615-0779-53D8-889F-95AD493D28E9" authority="(Hallowell, 1861)" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Ateuchosaurus pellopleurus (Hallowell, 1861)</taxonomicName>
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<paragraph id="4B2205BB8DE0944EFA03A78F5F7DD9DC" pageId="0" pageNumber="77">
<figureCitation id="057EEBADF1A5526CD0FCFB193B2AA6F8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (A) and ventral (B) views of the lectotype of Ateuchosaurus pellopleurus (USNM 42110). These photographs were cited from the collection database of the Division of Amphibians and Reptiles in the National Museum of Natural History at the Smithsonian Institution (EZID of the dorsal view: http: // n 2 t. net / ark: / 65665 / m 3 c 86 ab 460 - 1 e 75 - 4 bb 1 - a 19 b- 89 e 8439 f 129 c, EZID of the ventral view: http: // n 2 t. net / ark: / 65665 / m 390991 fa 3 - b 108 - 4 c 49 - bdb 2 - a 4 e 443 ac 1 cfe)." figureDoi="10.3897/zse.99.95923.figure3" httpUri="https://binary.pensoft.net/fig/798124" pageId="0" pageNumber="77">Figs 3</figureCitation>
<figureCitation id="FD772D243FC67E4F9A5B07BD9A0E5720" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal, lateral, and ventral views of head scalation of Ateuchosaurus pellopleurus (KUZ R 77703). CS, chinshield; EcP, ectoparietal; F, frontal; FrNa, frontonasal; FrP, frontoparietal; InLa, infralabial; IP, interparietal; Lo, loreal; M, mental; Na, nasal; P, parietal; PE, parietal eye; PoG, postgenial; PoLa, postlabial; PoM, postmental; PoO, postocular; PoSO, postsubocular; PrO, preocular; PrSO, presubocular; PrT, pretemporal; R, rostral; SuCi, supraciliary; SuLa, supralabial; SuLo, supraloreal; SuNa, supranasal; SuO, supraocular; T, temporal. The dashed line on the fused scale of Na and first SuLa in the lateral view indicates the line of a shallow groove on the scale." figureDoi="10.3897/zse.99.95923.figure4" httpUri="https://binary.pensoft.net/fig/798125" pageId="0" pageNumber="77">, 4 Japanese name: Heriguro-Hime-Tokage</figureCitation>
</paragraph>
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<paragraph id="063525F23B86805D9214837893DDD3F0" pageId="0" pageNumber="77">
<taxonomicName id="4B6059A8D976994DAFB30E6B77DAD8AD" authorityName="Hallowell" authorityYear="1861" family="Scincidae" genus="Lygosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Lygosaurus pellopleurus</taxonomicName>
Hallowell, 1861: 496-497 (part); 
<bibRefCitation id="756B10EBC1F042BB7CBEC4BA5827D913" author="Stejneger, L" journalOrPublisher="Bulletin - United States National Museum" pageId="0" pageNumber="77" pagination="1 - 577" refId="B36" refString="Stejneger, L, 1907. Herpetology of Japan and adjacent territory. Bulletin - United States National Museum 58: 1 - 577" title="Herpetology of Japan and adjacent territory." volume="58" year="1907">Stejneger 1907</bibRefCitation>
: 222-224 (part); 1927: 2; 
<bibRefCitation id="D7807A5E9FEE1E2BFA1808FB710CF524" author="Sclater, JA" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="77" refId="B32" refString="Sclater, JA, 1950. Bionomic notes on some reptiles form Okinawa Shima, Ryukyu Islands, Japan. The Annals &amp; magazine of natural history. 12th series, Zoology, Botany, and Geology 3: 113-116." title="Bionomic notes on some reptiles form Okinawa Shima, Ryukyu Islands, Japan. The Annals &amp; magazine of natural history. 12 th series, Zoology, Botany, and Geology 3: 113 - 116." year="1950">Sclater 1950</bibRefCitation>
: 114.
</paragraph>
<paragraph id="F3B07BEE23F4558C58890C77CEC9D741" pageId="0" pageNumber="77">
<taxonomicName id="02C34F3A18BF92F38ED4A8DE0520793C" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma pellopleurum" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurum">Lygosoma pellopleurum</taxonomicName>
: 
<bibRefCitation id="D77691A85839151EF613786638D6A41F" author="Boulenger, GA" journalOrPublisher="British Museum (Natural history), London" pageId="0" pageNumber="77" refId="B3" refString="Boulenger, GA, 1887. Catalogue of the lizards in the British Museum (Natural History). 2nd edn. Volume III. British Museum (Natural history), London" title="Catalogue of the lizards in the British Museum (Natural History). 2 nd edn. Volume III." year="1887">Boulenger 1887</bibRefCitation>
: 319; 
<bibRefCitation id="A6E66E99890449154FF879B5D496D0D6" author="Okada, S" journalOrPublisher="Kinkōdō, Tokyo" pageId="0" pageNumber="77" refId="B23" refString="Okada, S, 1891. Catalogue of Vertebrated Animals of Japan. Kinkōdō, Tokyo" title="Catalogue of Vertebrated Animals of Japan." year="1891">Okada 1891</bibRefCitation>
: 70 (part).
</paragraph>
<paragraph id="7BFD28A667A491410607752316F2718F" pageId="0" pageNumber="77">
<taxonomicName id="441CB578C049B621FB084D2296B4343B" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma (Homolepida) pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus" subGenus="Homolepida">Lygosoma (Homolepida) pellopleurus</taxonomicName>
: 
<bibRefCitation id="8C29F2E5205F6CBDA0FC640A54E87A9F" author="Boettger, O" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="77" refId="B2" refString="Boettger, O, 1895. Neue Froesche und Schlangen von den Liukiu-Inseln. Offenbacker Vereins fur Naturkunde. Bericht 33-36: 101-117." title="Neue Froesche und Schlangen von den Liukiu-Inseln. Offenbacker Vereins fur Naturkunde. Bericht 33 - 36: 101 - 117." year="1895">Boettger 1895</bibRefCitation>
: 107.
</paragraph>
<paragraph id="E5B2D9D3E1F9E7ACFE7D434DFD0279BC" pageId="0" pageNumber="77">
<taxonomicName id="6F7D49BB56DEEF36AD4520A2804D2D73" authorityName="Van Denburgh" authorityYear="1912" family="Scincidae" genus="Lygosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosaurus pellopleurus subsp. browni" order="Squamata" pageId="0" pageNumber="77" rank="subSpecies" species="pellopleurus" subSpecies="browni">Lygosaurus pellopleurus browni</taxonomicName>
Van Denburgh, 1912a: 7; 
<bibRefCitation id="EDBF2BBD09147CB920D4C9F67B769B19" author="Van Denburgh, J" journalOrPublisher="Private publication, San Francisco" pageId="0" pageNumber="77" refId="B42" refString="Van Denburgh, J, 1912b. Concerning certain species of reptiles and amphibians from China, Japan, the Loo Choo Islands and Formosa. Proceedings of the California Academy of Sciences, 4th Series 3: 187-258." title="Concerning certain species of reptiles and amphibians from China, Japan, the Loo Choo Islands and Formosa. Proceedings of the California Academy of Sciences, 4 th Series 3: 187 - 258." year="1912 b">Van Denburgh 1912b</bibRefCitation>
: 240-241.
</paragraph>
<paragraph id="D879262AF70E92F37DEC4103A9BFA5E0" pageId="0" pageNumber="77">
<taxonomicName id="8F3A9621BC32FC3687984AFF2ABD019F" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Ateuchosaurus pellopleurus</taxonomicName>
: 
<bibRefCitation id="1735B3A9CAFB08C9D1156366FD9C6C54" author="Smith, MA" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="77" pagination="213 - 234" refId="B34" refString="Smith, MA, 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39: 213 - 234" title="A review of the genus Lygosoma (Scincidae: Reptilia) and its allies." volume="39" year="1937">Smith 1937</bibRefCitation>
: 231; 
<bibRefCitation id="9194B97B85ECFB324E7C949EF770AB25" author="Tanaka, S" journalOrPublisher="Ienohikari Kyokai, Tokyo" pageId="0" pageNumber="77" refId="B38" refString="Tanaka, S, 1979. Heriguro Himetokage. In: Sengoku S (Ed.) Gensyoku Ryosei Hachurui. Ienohikari Kyokai, Tokyo" title="Heriguro Himetokage. In: Sengoku S (Ed.) Gensyoku Ryosei Hachurui." year="1979">Tanaka 1979</bibRefCitation>
: 37 (part); 
<bibRefCitation id="F1804C26A38F33BA7A50EE1ED6A0856B" author="Hikida, T" editor="Sengoku, S" journalOrPublisher="Heibonsha Limited, Publishers, Tokyo" pageId="0" pageNumber="77" pagination="80 - 82" refId="B13" refString="Hikida, T, 1996. Scincidae. In: Sengoku, S, Hikida, T, Matsui, M, Nakaya, K, Eds., The Encyclopedia of Animals in Japan, Volume 5: Amphibians, Reptiles, Chondrichthyes. Heibonsha Limited, Publishers, Tokyo: 80 - 82" title="Scincidae." volumeTitle="The Encyclopedia of Animals in Japan, Volume 5: Amphibians, Reptiles, Chondrichthyes." year="1996">Hikida 1996</bibRefCitation>
: 81 (part); 
<bibRefCitation id="FC8E1B53FADE1B7994A8C4770B0A83A0" DOI="https://doi.org/10.2307/1565549" author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="77" pagination="106 - 118" refId="B30" refString="Ota, H, Miyaguni, H, Hikida, T, 1999. Geographic variation in the endemic skink, Ateuchosaurus pellopleurus from the Ryukyu Archipelago. Journal of Herpetology 33 (1): 106 - 118, DOI: https://doi.org/10.2307/1565549" title="Geographic variation in the endemic skink, Ateuchosaurus pellopleurus from the Ryukyu Archipelago." url="https://doi.org/10.2307/1565549" volume="33" year="1999">Ota et al. 1999</bibRefCitation>
: 106 (part); 
<bibRefCitation id="F8D93B40E6E8ECD5A9DAE59FAE7B91CB" author="Goris, RC" journalOrPublisher="Krieger, Malabar" pageId="0" pageNumber="77" refId="B4" refString="Goris, RC, Maeda, N, 2004. Guide to the Amphibians and Reptiles of Japan. Krieger, Malabar" title="Guide to the Amphibians and Reptiles of Japan." year="2004">Goris and Maeda 2004</bibRefCitation>
: 155-156 (part); 
<bibRefCitation id="976A48ADAF1648129244CAA8526938F4" DOI="https://doi.org/10.1016/j.ympev.2005.12.011" author="Austin, JJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="77" pagination="503 - 511" refId="B1" refString="Austin, JJ, Arnold, EN, 2006. Using ancient and recent DNA to explore relationships of extinct and endangered Leiolopisma skinks (Reptilia: Scincidae) in the Mascarene islands. Molecular Phylogenetics and Evolution 39 (2): 503 - 511, DOI: https://doi.org/10.1016/j.ympev.2005.12.011" title="Using ancient and recent DNA to explore relationships of extinct and endangered Leiolopisma skinks (Reptilia: Scincidae) in the Mascarene islands." url="https://doi.org/10.1016/j.ympev.2005.12.011" volume="39" year="2006">Austin and Arnold 2006</bibRefCitation>
: fig. 2, table 1; 
<bibRefCitation id="45283FC41492D866F42B329DBE9C104D" DOI="https://doi.org/10.1186/1471-2148-13-93" author="Pyron, RA" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="77" pagination=": 93" refId="B31" refString="Pyron, RA, Burbrink, FT, Wiens, JJ, 2013. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology 13 (1): 93, DOI: https://doi.org/10.1186/1471-2148-13-93" title="A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes." url="https://doi.org/10.1186/1471-2148-13-93" volume="13" year="2013">Pyron et al. 2013</bibRefCitation>
: fig. 7; 
<bibRefCitation id="9A028F95F77F7DCF79996FAFEA73B2E4" DOI="https://doi.org/10.11646/zootaxa.3765.4.2" author="Hedges, SB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="77" pagination="317 - 338" refId="B12" refString="Hedges, SB, 2014. The high-level classification of skinks (Reptilia, Squamata, Scincomorpha). Zootaxa 3765 (4): 317 - 338, DOI: https://doi.org/10.11646/zootaxa.3765.4.2" title="The high-level classification of skinks (Reptilia, Squamata, Scincomorpha)." url="https://doi.org/10.11646/zootaxa.3765.4.2" volume="3765" year="2014">Hedges 2014</bibRefCitation>
: 322; 
<bibRefCitation id="1465CCF906AD0B04065ADD393EC295E9" DOI="https://doi.org/10.1016/j.ympev.2015.10.009" author="Zheng, Y" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="77" pagination="537 - 547" refId="B44" refString="Zheng, Y, Wiens, JJ, 2016. Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species. Molecular Phylogenetics and Evolution 94: 537 - 547, DOI: https://doi.org/10.1016/j.ympev.2015.10.009" title="Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species." url="https://doi.org/10.1016/j.ympev.2015.10.009" volume="94" year="2016">Zheng and Wiens 2016</bibRefCitation>
: figs 1-2; 
<bibRefCitation id="4D3F28FD35026A3E5C808A433A6BD2CB" DOI="https://doi.org/10.1007/978-4-431-56432-4_5" author="Okamoto, T" editor="Motokawa, M" journalOrPublisher="Springer Japan KK, Tokyo" pageId="0" pageNumber="77" pagination="135 - 163" refId="B26" refString="Okamoto, T, 2017. Historical biogeography of the terrestrial reptiles of Japan: a comparative analysis of geographic ranges and molecular phylogenies. In: Motokawa, M, Kajihara, H, Eds., Species Diversity of Animals in Japan. Springer Japan KK, Tokyo: 135 - 163, DOI: https://doi.org/10.1007/978-4-431-56432-4_5" title="Historical biogeography of the terrestrial reptiles of Japan: a comparative analysis of geographic ranges and molecular phylogenies." url="https://doi.org/10.1007/978-4-431-56432-4_5" volumeTitle="Species Diversity of Animals in Japan." year="2017">Okamoto 2017</bibRefCitation>
: table 5.2; 
<bibRefCitation id="570578283FCCE6B04800219AE83657BB" DOI="https://doi.org/10.1016/j.jcz.2020.06.008" author="Makino, T" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="77" pagination="1 - 10" refId="B17" refString="Makino, T, Okamoto, T, Kurita, K, Nakano, T, Hikida, T, 2020. Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan. Zoologischer Anzeiger 288: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.06.008" title="Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan." url="https://doi.org/10.1016/j.jcz.2020.06.008" volume="288" year="2020">Makino et al. 2020</bibRefCitation>
: 6 (&quot;northern lineage&quot;); 
<bibRefCitation id="08892941E9A36BE81C617D96308AF860" author="Okamoto, T" journalOrPublisher="Sunrise Publishing, Hikone" pageId="0" pageNumber="77" pagination="142 - 143" refId="B27" refString="Okamoto, T, Kurita, K, 2021. Ateuchosaurus pellopleurus (Hallowell, 1861). In: Amphibians and Reptiles of Japan. Sunrise Publishing, Hikone: 142 - 143" title="Ateuchosaurus pellopleurus (Hallowell, 1861)." volumeTitle="Amphibians and Reptiles of Japan." year="2021">Okamoto and Kurita 2021</bibRefCitation>
: 142-143 (part).
</paragraph>
<paragraph id="8A9F0365AD9CC77209882322F60587C1" pageId="0" pageNumber="77">
<taxonomicName id="9DD1E48896EA5C7E8F1214B3181D7BB1" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">Lygosoma pellopleurus</taxonomicName>
: 
<bibRefCitation id="875658B5BE950A140E87A067A83DC21E" author="Okada, Y" journalOrPublisher="Section B" pageId="0" pageNumber="77" pagination="159 - 214" refId="B24" refString="Okada, Y, 1939. Studies on the lizards of Japan. Contribution III. Scincidae. Science Reports of the Tokyo Bunrika Daigaku. Section B 4: 159 - 214" title="Studies on the lizards of Japan. Contribution III. Scincidae. Science Reports of the Tokyo Bunrika Daigaku." volume="4" year="1939">Okada 1939</bibRefCitation>
: 206-209 (part).
</paragraph>
<paragraph id="59A3C89A9E4483E741D3B94D797C4D81" pageId="0" pageNumber="77">
<taxonomicName id="16AD0707C2F693BC2118A3C297E24D50" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lygosoma (Ateuchosaurus) pellopleurum" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurum" subGenus="Ateuchosaurus">Lygosoma (Ateuchosaurus) pellopleurum</taxonomicName>
: 
<bibRefCitation id="C97F6B23FE6ABADA35281865AF56B6A5" author="Nakamura, K" journalOrPublisher="Hoikusha Publishing, Osaka" pageId="0" pageNumber="77" refId="B19" refString="Nakamura, K, Ueno, S, 1963. Japanese reptiles and amphibians in colour. Hoikusha Publishing, Osaka" title="Japanese reptiles and amphibians in colour." year="1963">
Nakamura and 
<normalizedToken id="D8376AF20607CDE0F58C9FC0622811B9" originalValue="Uéno">Ueno</normalizedToken>
1963
</bibRefCitation>
: 123-125 (part).
</paragraph>
</subSubSection>
<subSubSection id="5F4B7FC7A72820DE6B97DAF86D4D3868" pageId="0" pageNumber="77" type="lectotype designation">
<paragraph id="0133388CAF56260506DBC529BD765AEC" pageId="0" pageNumber="77">Lectotype designation.</paragraph>
<paragraph id="9DE0B88E5F2A8E7995C973A5059005A8" pageId="0" pageNumber="77">
One of the existing syntypes, USNM 42110, was designated as the lectotype for 
<taxonomicName id="804BBBB8BD751FE8D1CC22D1FA771187" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="8B11CD79CF4047A1EB8EC7EBDD901F76" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="3BEF68F2524A576626971743874078E3" doi="10.3897/zse.99.95923.figure3" httpUri="https://binary.pensoft.net/fig/798124" pageId="0" pageNumber="77" start="Figure 3" startId="F3">
<paragraph id="80AB534D4EC5310A74B6213273F8430B" pageId="0" pageNumber="77">
<emphasis id="E3F2837F48AD89B7740614E0E636E712" bold="true" pageId="0" pageNumber="77">Figure 3.</emphasis>
Dorsal (
<emphasis id="456CAF4E7B0BAA3E7B2B43C12B3B9C0E" bold="true" pageId="0" pageNumber="77">A</emphasis>
) and ventral (
<emphasis id="AFA5CC55B0867545B5D8265E65186491" bold="true" pageId="0" pageNumber="77">B</emphasis>
) views of the lectotype of 
<taxonomicName id="DA84CA53FA42BFEEA31033AD11B15A67" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="67D7B6C23EE59CE0B29B7387691B6B67" italics="true" pageId="0" pageNumber="77">Ateuchosaurus pellopleurus</emphasis>
</taxonomicName>
(USNM 42110). These photographs were cited from the collection database of the Division of Amphibians and Reptiles in the National Museum of Natural History at the Smithsonian Institution (EZID of the dorsal view: http://n2t.net/ark:/65665/m3c86ab460-1e75-4bb1-a19b-89e8439f129c, EZID of the ventral view: http://n2t.net/ark:/65665/m390991fa3-b108-4c49-bdb2-a4e443ac1cfe).
</paragraph>
</caption>
<paragraph id="2590B9F23C665893B24B992D87A6E4AA" pageId="0" pageNumber="77">
<emphasis id="7A99591E1FAF901C9E6BD14EC4DF4357" bold="true" pageId="0" pageNumber="77">
<emphasis id="A052F4C8D5605EB8447F5BE510F38932" italics="true" pageId="0" pageNumber="77">Lectotype</emphasis>
.
</emphasis>
USNM 42110 collected from Amamioshima Island, Amami Group, Ryukyu Archipelago, Japan, in the United States North Pacific Surveying and Exploring Expedition in 1853-1856.
</paragraph>
<caption id="E49713A7CA9778D7360A8A909E995EF5" doi="10.3897/zse.99.95923.figure4" httpUri="https://binary.pensoft.net/fig/798125" pageId="0" pageNumber="77" start="Figure 4" startId="F4">
<paragraph id="C8FB53D3DE8CEAF1A330B39CF47D995A" pageId="0" pageNumber="77">
<emphasis id="5477FA51EEAAC7BE9D58CFFDC7DA6ACF" bold="true" pageId="0" pageNumber="77">Figure 4.</emphasis>
Dorsal, lateral, and ventral views of head scalation of 
<taxonomicName id="E4343C71EC44E24AB2C4C34E04C61D8E" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="632B0052C7D98B3FF4D6F922258F17CB" italics="true" pageId="0" pageNumber="77">Ateuchosaurus pellopleurus</emphasis>
</taxonomicName>
(KUZ R77703). CS, chinshield; EcP, ectoparietal; F, frontal; FrNa, frontonasal; FrP, frontoparietal; InLa, infralabial; IP, interparietal; Lo, loreal; M, mental; Na, nasal; P, parietal; PE, parietal eye; PoG, postgenial; PoLa, postlabial; PoM, postmental; PoO, postocular; PoSO, postsubocular; PrO, preocular; PrSO, presubocular; PrT, pretemporal; R, rostral; SuCi, supraciliary; SuLa, supralabial; SuLo, supraloreal; SuNa, supranasal; SuO, supraocular; T, temporal. The dashed line on the fused scale of Na and first SuLa in the lateral view indicates the line of a shallow groove on the scale.
</paragraph>
</caption>
<paragraph id="43749DA4EBFF67E758444561D5537E80" pageId="0" pageNumber="77">
<emphasis id="4D0666CBBF40079317D1179912306A25" bold="true" pageId="0" pageNumber="77">
<emphasis id="552DA8FD19945E7AFEC1BFE0FBD0D48B" italics="true" pageId="0" pageNumber="77">Paralectotype</emphasis>
.
</emphasis>
USNM 42114. The locality of this specimen is the same as USNM 42110. Missing syntype(s) collected from Loo-Choo is also automatically fixed as the paralectotype(s) of 
<taxonomicName id="C454758B52BB4E8EDEAC68A8B64FFCDA" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="46619E0205CB5DCE6BA724C4BF91348A" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
. However, the specimen(s) may belong to 
<taxonomicName id="68CB0683D162C3855535CFD4ED5C0796" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="CE7D0100033B80AE08665CE01804A4FB" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F0031A9B89F0CC39B7361C7EAB17D9E7" pageId="0" pageNumber="77" type="materials_examined">
<paragraph id="DE2B8A19850C0B74F6F4CD9554911334" pageId="0" pageNumber="77">Material examined.</paragraph>
<paragraph id="068CBEB6A7E6AF4240A1F6E783E92720" pageId="0" pageNumber="77">
<materialsCitation id="962D16FEA9A15B1124BF5E7D759BE510" accessionNumber="R54830" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Takeshima Island" municipality="Mishima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingCountry id="3E7E3A02977934ED18508CF6474D16F6" name="Japan">Japan</collectingCountry>
• 
<collectingRegion id="C901FFDBFD14729EE53B67EE407491F0" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="67EA9726D5852C06F56C02FE9D5C01FF">Mishima Village</collectingMunicipality>
, 
<location id="76325607F8BB77FD8DCB1F74A7ADDC61" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:76325607F8BB77FD8DCB1F74A7ADDC61" country="Japan" municipality="Mishima Village" name="Takeshima Island" stateProvince="Kagoshima Prefecture">Takeshima Island</location>
; KUZ 
<accessionNumber id="119EF81A8B5D4AD2FCB6851B1FECA7EC" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R54830">R54830</accessionNumber>
to 54834, 70451, 70453 to 70456, 70458, 70461
</materialsCitation>
• 
<materialsCitation id="7D88E08EED9432873D921FA693726083" accessionNumber="R70413" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Iojima Island" municipality="Mishima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="BC4FFC4F33A8497B50F3B5B1E34EA11A" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="2214BF9B216B58873C6712496DEA0C1F">Mishima Village</collectingMunicipality>
, 
<location id="0747185A0F5C883E1A7B8A3895244C02" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:0747185A0F5C883E1A7B8A3895244C02" country="Japan" municipality="Mishima Village" name="Iojima Island" stateProvince="Kagoshima Prefecture">Iojima Island</location>
; KUZ 
<accessionNumber id="686EBDEAEA8DFE81CDEFC1C8EBCE5AC1" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70413">R70413</accessionNumber>
to 70417, 70420, 70422 to 70425, 70442 to 70443, 70445 to 70450
</materialsCitation>
• 
<materialsCitation id="658AC104C5CCA8BE5473F3C3B9F1B218" accessionNumber="R47185" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Kuroshima Island" municipality="Mishima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="F7D6C4702EFEED7CAFDACE076BE719FA" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="820063754BD09E862FDBEE46B4A343FC">Mishima Village</collectingMunicipality>
, 
<location id="5133D13283BDF5E1545A2D222F8EDE2E" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:5133D13283BDF5E1545A2D222F8EDE2E" country="Japan" municipality="Mishima Village" name="Kuroshima Island" stateProvince="Kagoshima Prefecture">Kuroshima Island</location>
; KUZ 
<accessionNumber id="249E915BC95228971EA38289A32F7CE9" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R47185">R47185</accessionNumber>
to 47202
</materialsCitation>
• 
<materialsCitation id="3A0771A3B3CE97DFC0AEA94DB8E8B5E0" accessionNumber="R66811" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Kuchinoshima Island" municipality="Toshima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="1C639FE1631B6B83E6E14A79508DB7E3" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="2611C18730A7B17B91A2B2B0D155D04C">Toshima Village</collectingMunicipality>
, 
<location id="7A26559482D2FECDFBBDDDFCD9DBE84A" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:7A26559482D2FECDFBBDDDFCD9DBE84A" country="Japan" municipality="Toshima Village" name="Kuchinoshima Island" stateProvince="Kagoshima Prefecture">Kuchinoshima Island</location>
; KUZ 
<accessionNumber id="07A630F694B61FBA04AE4E759F10F9FF" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R66811">R66811</accessionNumber>
to 66812, 68678
</materialsCitation>
• 
<materialsCitation id="A3C02B2472BCC60DB235B6DFDD2CAD57" accessionNumber="R66815" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Nakanoshima Island" municipality="Toshima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="345F627915DDBC551C4B3CBE59B0845B" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="B076BA57900609C11CFF5BEE498EAE37">Toshima Village</collectingMunicipality>
, 
<location id="88940A7A2AFBE96D04C96D770C41A224" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:88940A7A2AFBE96D04C96D770C41A224" country="Japan" municipality="Toshima Village" name="Nakanoshima Island" stateProvince="Kagoshima Prefecture">Nakanoshima Island</location>
; KUZ 
<accessionNumber id="B3E794DEFC5F9EFD07D63928F61D6839" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R66815">R66815</accessionNumber>
, 66834 to 66835, OMNH R3024 to 3028, 6880
</materialsCitation>
• 
<materialsCitation id="A64DB25B07862343AA1B1EA8AE4E4A48" accessionNumber="R69905" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Suwanosejima Island" municipality="Toshima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="95CB4EFC023B219DB284362DDBA2B63F" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="2858531BB6A87E69D2970B0DEADF54EA">Toshima Village</collectingMunicipality>
, 
<location id="2EEF5A4174C0ACEDA8D8CEFABC93984F" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:2EEF5A4174C0ACEDA8D8CEFABC93984F" country="Japan" municipality="Toshima Village" name="Suwanosejima Island" stateProvince="Kagoshima Prefecture">Suwanosejima Island</location>
; KUZ 
<accessionNumber id="E1C1CFA4C976B80035089806A49E18A4" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R69905">R69905</accessionNumber>
to 69910, OMNH R3022 to 3023, 6886, 6889
</materialsCitation>
• 
<materialsCitation id="25D3A064E1AB290F01260998A43585DC" accessionNumber="R66781" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Akusekijima Island" municipality="Toshima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="AF4D17C5495EA019FEDA4F8961377B89" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="42AF318990B935FE825B7BCD3798AB62">Toshima Village</collectingMunicipality>
, 
<location id="3E89AC77115F55DFED531406D4AAAE30" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:3E89AC77115F55DFED531406D4AAAE30" country="Japan" municipality="Toshima Village" name="Akusekijima Island" stateProvince="Kagoshima Prefecture">Akusekijima Island</location>
; KUZ 
<accessionNumber id="16D2BEEA9909F099E586B69E83E5B544" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R66781">R66781</accessionNumber>
to 66782, OMNH R3018 to 3019
</materialsCitation>
• 
<materialsCitation id="2B863E0F5051E89C8177D5C54396ECE7" accessionNumber="R34983" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Kodakarajima Island" municipality="Toshima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="F69B638586B47BB2D4969EF49B7BF30A" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="C3B2ED7C3FDD933ED44B59C4C710456B">Toshima Village</collectingMunicipality>
, 
<location id="D2961CF0380E2C61C0DF48313FBC04C8" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:D2961CF0380E2C61C0DF48313FBC04C8" country="Japan" municipality="Toshima Village" name="Kodakarajima Island" stateProvince="Kagoshima Prefecture">Kodakarajima Island</location>
; KUZ 
<accessionNumber id="E79B5BE570EAEAAD448302042756CC79" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R34983">R34983</accessionNumber>
to 34986, 70147 to 70151, 70153 to 70155, 70162
</materialsCitation>
• 
<materialsCitation id="DE42ABC777C3B6B0DAC788C13C7661B7" accessionNumber="R13169" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Takarajima Island" municipality="Toshima Village" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="F4C0257EAC45C54CE6FBC6E6FEE73A8A" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="69FA556D8D8802CA5122619482711CD0">Toshima Village</collectingMunicipality>
, 
<location id="84586F399B13E0AC0FFC7AAB57D05B26" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:84586F399B13E0AC0FFC7AAB57D05B26" country="Japan" municipality="Toshima Village" name="Takarajima Island" stateProvince="Kagoshima Prefecture">Takarajima Island</location>
; KUZ 
<accessionNumber id="D20CA5203663601EC7B8A00989908826" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R13169">R13169</accessionNumber>
to 13207, OMNH R1 to 4, 2217, 3014 to 3016, 6890
</materialsCitation>
• 
<materialsCitation id="CAAD583EA3B4F08D82CE33DF9A2952EA" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Kikaijima Island" municipality="Kikai Town" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="BAFBE9B2E4DA5D96F729EE8052D9BD35" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<collectingMunicipality id="143502C0D289CEC52BE57B31E7013BC4">Kikai Town</collectingMunicipality>
, 
<location id="3964FB55FD8C431A72018262BC21BD97" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:3964FB55FD8C431A72018262BC21BD97" country="Japan" municipality="Kikai Town" name="Kikaijima Island" stateProvince="Kagoshima Prefecture">Kikaijima Island</location>
; OMNH R3029 to 3036
</materialsCitation>
• 
<materialsCitation id="44B8F9764AC907DA65B5DD9FD0830E24" accessionNumber="R62708" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Amamioshima Island" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="E13AEF0D19DC4B2EBCE7809E9C30DD44" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<location id="5D1A90DD23114CA84F7949B9D9919887" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:5D1A90DD23114CA84F7949B9D9919887" country="Japan" name="Amamioshima Island" stateProvince="Kagoshima Prefecture">Amamioshima Island</location>
; KUZ 
<accessionNumber id="4DCA2C1FE658CFFC01EA4C9490515661" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R62708">R62708</accessionNumber>
, 66187 to 66190, 69075, 69901 to 69904, 70700, 72543, 72545, 77679 to 77682, 77686, 77689 to 77690, 77692, 77694 to 77698, 77700, 77702 to 77708, OMNH R245 to 246, 248 to 249, 260, 621 to 623, 2777, 3077 to 3094, 3096 to 3100, 3102, 3105 to 3112, 6876, 6878 to 6879, 6881 to 6883, 6885, 6891 to 6892
</materialsCitation>
• 
<materialsCitation id="626E550B584CDEFCA4B8F0B5762724A2" accessionNumber="R70680" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Tokunoshima Island" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="8D828A79E2DAE5CA7BD88471383FAB46" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<location id="1292DAE42CEE5D68CD553A47B6F70D7C" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:1292DAE42CEE5D68CD553A47B6F70D7C" country="Japan" name="Tokunoshima Island" stateProvince="Kagoshima Prefecture">Tokunoshima Island</location>
; KUZ 
<accessionNumber id="C50F4AA90E96F55D706AA491D22178C9" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R70680">R70680</accessionNumber>
, 77672, 77674, OMNH R3039 to 3042, 3044 to 3060, 6877, 6888
</materialsCitation>
• 
<materialsCitation id="4E6225CB5909829383590670C835CEEF" accessionNumber="R34987" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Japan" location="Okinoerabujima Island" specimenCount="1" stateProvince="Kagoshima Prefecture">
<collectingRegion id="E9A7D752952E2357409C3F96B19C9C79" country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
, 
<location id="96358D6112047CDCA2EEAEC8AE1C8412" LSID="urn:lsid:plazi:treatment:A388A615077953D8889F95AD493D28E9:96358D6112047CDCA2EEAEC8AE1C8412" country="Japan" name="Okinoerabujima Island" stateProvince="Kagoshima Prefecture">Okinoerabujima Island</location>
; KUZ 
<accessionNumber id="C78EE4EC70760396270DBB35B4DF46B0" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R34987">R34987</accessionNumber>
to 34996, 66740, 66780, 70675 to 70679, 70681 to 70682, 77637 to 77640, 77642 to 77644, 77647, 77649 to 77650, 77652 to 77661, 77664 to 77665, 77667 to 77671, OMNH R262 to 266, 3037
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DF486736E60F98ADC9BF5E8B50475E25" pageId="0" pageNumber="77" type="emended diagnosis">
<paragraph id="094B2A7D78E8E060697D8AC57E93954B" pageId="0" pageNumber="77">Emended diagnosis.</paragraph>
<paragraph id="8FA58F3F66E2FE4776864069C2B16B7E" pageId="0" pageNumber="77">
An 
<taxonomicName id="3060FE8F21CCD200875BA904F6E56EFC" authorityName="Gray" authorityYear="1845" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus" order="Squamata" pageId="0" pageNumber="77" rank="genus">
<emphasis id="9556CB3724AA841D3665507079DDF676" italics="true" pageId="0" pageNumber="77">Ateuchosaurus</emphasis>
</taxonomicName>
species characterized by the following characters: FrNa and frontal usually distinct; a pair of frontoparietals that do not contact each other; eight SuCis; anteroposteriorly reduced parietals separated from SuO and pretemporal by EcP; no distinct nuchals; usually six InLas; body size medium (SVL ca. 42-70 mm); widely separated forelimb and hindlimb when appressed; usually 26 or 28 MSRs (mode = 26, 24-30); 53-70 (mode = 62) and 56-69 DMSs (mode = 63) in male and female, respectively; 8-14 TIVs (mode = 11); 10 preanals not enlarged; usually 2-3 pairs of DSRBS.
</paragraph>
</subSubSection>
<subSubSection id="22C1C9AA716E971B80B8BD92F3A29815" pageId="0" pageNumber="77" type="comparisons">
<paragraph id="D1C6DC6DA1ABAC0662A46B4F0D35B6D9" pageId="0" pageNumber="77">Comparison.</paragraph>
<paragraph id="5FE0679B56DC0838B737304B205CE1D7" pageId="0" pageNumber="77">
<taxonomicName id="5CE7962AD6F6C11F3825C11791D9CDF5" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="E3A6F39AA302D851434E3168EBB25E1E" italics="true" pageId="0" pageNumber="77">Ateuchosaurus pellopleurus</emphasis>
</taxonomicName>
, together with 
<taxonomicName id="F1DEB406C8C2E18929DB62B7E0D4EE07" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="6F5C7A5176BF140BA00FB6C8FF482A11" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
(see below), is distinguished from 
<taxonomicName id="BB4DB3524F789E783AFB0FE6888290A4" lsidName="A. chinensis" pageId="0" pageNumber="77" rank="species" species="chinensis">
<emphasis id="619BC77E4318A295D8785948F490A51D" italics="true" pageId="0" pageNumber="77">A. chinensis</emphasis>
</taxonomicName>
by having the following characters (
<bibRefCitation id="B33C088DDF15CAE979A5BC2447E2CE85" author="Nguyen, QT" journalOrPublisher="Herpetology Notes" pageId="0" pageNumber="77" pagination="17 - 21" refId="B21" refString="Nguyen, QT, Tung, TT, Ngoc, HV, Boehme, W, Ziegler, T, 2008. Rediscovery and redescription of Ateuchosaurus chinensis Gray, 1845 (Squamata: Sauria: Scincidae) from northeastern Vietnam. Herpetology Notes 1: 17 - 21" title="Rediscovery and redescription of Ateuchosaurus chinensis Gray, 1845 (Squamata: Sauria: Scincidae) from northeastern Vietnam." volume="1" year="2008">Nguyen et al. 2008</bibRefCitation>
): usually six InLas (vs. seven InLas); smaller body size (SVL ca. 42-70 mm vs. 70.0-83.8 mm; see also 
<bibRefCitation id="D940984EB46862E10F5BCAC701BE9E24" author="Okamoto, T" journalOrPublisher="Sunrise Publishing, Hikone" pageId="0" pageNumber="77" pagination="142 - 143" refId="B27" refString="Okamoto, T, Kurita, K, 2021. Ateuchosaurus pellopleurus (Hallowell, 1861). In: Amphibians and Reptiles of Japan. Sunrise Publishing, Hikone: 142 - 143" title="Ateuchosaurus pellopleurus (Hallowell, 1861)." volumeTitle="Amphibians and Reptiles of Japan." year="2021">Okamoto and Kurita 2021</bibRefCitation>
); usually 26 or 28 MSRs (vs. 30 scale rows); usually 10-14 TIVs (vs. usually 16-18 TIVs); 10 preanals (vs. usually six preanals); blackish line on dorsolateral surface from snout to midbody (vs. no such line). This species resembles 
<taxonomicName id="CA4D2F3D4640FA0C28BA75BE96A68DDC" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="B716D881015751DEAA3B545BFF33F413" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
in MSR and body size but differs in usually having the separated state of FrNa and frontal (vs. the fused state), 53-70 and 56-69 DMSs in male and female (vs. 54-64 and 55-67 DMSs in male and female), 8-14 TIVs (vs. 10-16 TIVs), and usually 2-3 pairs of DSRBS (vs. usually 0-1 pair).
</paragraph>
<paragraph id="91158720C4C29BAB0E761ACD4FAFC228" pageId="0" pageNumber="77">
The mean K2P genetic distance 
<normalizedToken id="E684C8331AA181E2189C21E1045A6FB0" originalValue="±">+/-</normalizedToken>
standard deviation (range in parenthesis) based on cytochrome 
<emphasis id="A432362ADB8CD4EEDADA2D4FCAE00442" italics="true" pageId="0" pageNumber="77">b</emphasis>
sequences between 
<taxonomicName id="14DCB03DF083C89F01907674527A9956" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="BBA86C7198445F8E0192B88DF0ACB466" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="B51ED4A83F8C11455A50CFBBCE3AFA81" lsidName="A. chinensis" pageId="0" pageNumber="77" rank="species" species="chinensis">
<emphasis id="D58CFBD47BE509803D86C972C518E130" italics="true" pageId="0" pageNumber="77">A. chinensis</emphasis>
</taxonomicName>
was 24.6% 
<normalizedToken id="6CA544AE7321484053E8BC48427465FC" originalValue="±">+/-</normalizedToken>
0.3% (24.2%-25.5%); the distance between 
<taxonomicName id="4F1C7B662633E2DB086C289EA3E8B995" lsidName="A. pellopleurus" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="4888DF69052E44D5F46ACFD62CD70A8B" italics="true" pageId="0" pageNumber="77">A. pellopleurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="256FBECD2FF21239F7C44A50BE9685EF" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="1969EBEBFEF17D2E25A27333D0B0EA3F" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
was 13.1% 
<normalizedToken id="13A439B8588C77EA6770E2A5DE115496" originalValue="±">+/-</normalizedToken>
0.9% (11.3%-16.0%).
</paragraph>
</subSubSection>
<subSubSection id="A8F21F0F112E124DBA9233D7DF4BF470" pageId="0" pageNumber="77" type="description">
<paragraph id="A1225A7785DBD18927BBF59099EA8328" pageId="0" pageNumber="77">Description.</paragraph>
<paragraph id="0BC1DAD34CC70F12A746108EF5695D0C" pageId="0" pageNumber="77">
Mainly based on a topotype (KUZ R77703), an adult male collected from Amami City in Amamioshima Island (
<geoCoordinate id="65E7904569BD0333EDC68E0217F9DC07" degrees="28" direction="north" minutes="23" orientation="latitude" precision="1" seconds="44.29" value="28.395636">28°23'44.29&quot;N</geoCoordinate>
, 
<geoCoordinate id="29EDFD83B448C5740FFF2E3251B886D4" degrees="129" direction="east" minutes="28" orientation="longitude" precision="1" seconds="04.32" value="129.46786">129°28'04.32&quot;E</geoCoordinate>
, 2 m above sea level) (followed by ranges of several morphometric and meristic characters in parenthesis; 
<emphasis id="0D21EC97F121FBBBF2042FA0EC624BEE" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 23, topotypes, KUZ R66188, 72543, 77679-77682, 77686, 77689-77690, 77692, 77694-77698, 77700, 77702-77708); SVL 53.5 mm (53.5 
<normalizedToken id="03C3F71756598009EFC849E5A0CE98CD" originalValue="±">+/-</normalizedToken>
2.3, 49.0-57.0); HL 6.7 mm (6.5 
<normalizedToken id="7ED20CEC2D70F9C7B0B23E2F0F70CDC7" originalValue="±">+/-</normalizedToken>
0.3, 6.0-7.0); HW 5.0 mm (5.4 
<normalizedToken id="A2F4AEAF5ADA70E55DF2954260C79A2F" originalValue="±">+/-</normalizedToken>
0.3, 4.8-6.0); HH 4.2 mm (4.2 
<normalizedToken id="E5FEE74CF13369BEF4C91D0A16631261" originalValue="±">+/-</normalizedToken>
0.3, 3.7-5.0); SEyL 3.3 mm (3.2 
<normalizedToken id="E0147FC39431969BE2B0E34C69B0959E" originalValue="±">+/-</normalizedToken>
0.2, 2.8-3.6); EyL 2.1 mm (2.3 
<normalizedToken id="8A1E8ADE3DD3C192683A98C2DC109408" originalValue="±">+/-</normalizedToken>
0.1, 2.1-2.5); EyEaL 3.1 mm (3.2 
<normalizedToken id="2B3981DD150BAB4AE13E9EFC9CF2DA78" originalValue="±">+/-</normalizedToken>
0.2, 2.9-3.5); EaD 1.1 mm (1.0 
<normalizedToken id="CE57BF2F05ED0254DDA805E32515D87E" originalValue="±">+/-</normalizedToken>
0.1, 0.8-1.2); SAL 18.1 mm (17.2 
<normalizedToken id="3F8DED90FBC2A210409BDABE5A6CD404" originalValue="±">+/-</normalizedToken>
0.9, 15.5-18.6); AGL 31.4 mm (32.2 
<normalizedToken id="3ED8B32895857B27EDB8B003CC4B256F" originalValue="±">+/-</normalizedToken>
1.7, 28.9-34.5); TaL 56.3 mm (original tail 32.6 mm; regenerated tail 23.7 mm; 66.1 
<normalizedToken id="CC58798BEDA3E32DE6BE32681BDC663B" originalValue="±">+/-</normalizedToken>
0.9, 65.5-66.8, 
<emphasis id="D12EC19D4EEC5F2F60D7C4A0158EBC98" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 2 with undamaged original tail); FlL 9.0 mm (8.8 
<normalizedToken id="3047B4A996108BF71DD4F4E77107F369" originalValue="±">+/-</normalizedToken>
0.6, 7.8-9.8); HlL 15.9 mm (14.8 
<normalizedToken id="A724740F39FE0E96AB2E5AC5F040443D" originalValue="±">+/-</normalizedToken>
1.0, 12.5-16.5); FIIIL 1.8 mm (1.6 
<normalizedToken id="68E05B0F6ECAC92D86DCABB781E95D18" originalValue="±">+/-</normalizedToken>
0.2, 1.3-2.0); TIVL 4.3 mm (4.0 
<normalizedToken id="180955AE1DF8C9FA6201EBB9E0471583" originalValue="±">+/-</normalizedToken>
0.3, 3.5-4.6).
</paragraph>
<paragraph id="E162437CA20316AE71D5685D68E09DD1" pageId="0" pageNumber="77">Snout obtusely pointed; rostral visible from above, overlapping supranasal, nasal and first supralabial; nasal fused with first supralabial, with shallow groove on boundary between nasal and labial parts; nostril at center of nasal; supranasal unpaired with posterior edge concaved, overlapped by nasal; supraloreal overlapped by supranasal and anterior and posterior loreals, overlapping frontonasal and first supraciliary; frontonasal not fused with frontal, as large as supranasal, with flat posterior edge, overlapped by supranasal, overlapping first supraocular and frontal; no prefrontal; frontal large, shorter than distance from it to snout, longer than frontonasal, a part between first supraoculars narrower anteriorly, overlapping frontoparietals and interparietal; four supraoculars (no variation), anterior ones overlapping posterior ones, first one overlapped by first, second and third supraciliaries, first and second ones overlapping frontal, third and fourth ones overlapping frontoparietal; a pair of frontoparietals as long as width, separated from each other by interparietal, overlapping interparietal, parietal and ectoparietal; interparietal with short arrowhead-like tetragonal shape, similar length and width of frontoparietal, overlapping parietals; ectoparietal pentagonal and similar size to fourth supraocular, overlapped by fourth supraocular and upper pretemporal, overlapping parietal; parietals shorter and 1.5 times wider than interparietal, left one overlapped by right one; no distinct nuchal.</paragraph>
<paragraph id="EA86DBDCA49A8859FAACBB13FC0962BE" pageId="0" pageNumber="77">Two loreals with a half length and similar height of second supralabial, anterior one overlapped by nasaland second supralabial, overlapping supranasal and posterior one, posterior one overlapped by second supralabial, overlapping first supraciliary and upper and lower preoculars; eight supraciliaries (mode = 8, 7-8), anterior ones overlapping posterior ones, first one overlapped by upper preocular, third and fourth ones overlapping second supraocular, fourth to sixth ones overlapping third supraocular, sixth and seventh ones overlapping fourth supraocular, eighth one overlapping upper pretemporal and upper postocular; two preoculars, upper one similar size to first supraciliary, overlapped by lower one, lower one three times larger than upper one, overlapping third supralabial and anterior presubocular; two presuboculars overlapped by third supralabial, anterior one overlapping posterior one, posterior one overlapping fourth supralabial; three postsuboculars, lower ones overlapping upper ones, lowermost one largest, overlapped by fourth supralabial, lowermost and middle ones overlapping fifth supralabial, middle and uppermost ones overlapping primary temporal, uppermost one overlapping lower postocular; two postoculars, upper one overlapping lower one (usually three postoculars, anterior one overlapped by uppermost postsubocular, overlapping upper and lower ones); two pretemporals, upper one overlapped by fourth supraocular and upper postocular, overlapping lower one and uppermost secondary temporal, lower one smaller than upper one, overlapped by upper and lower postoculars, overlapping first to third secondary temporals; temporals with similar shape and size of body scales; single primary temporal overlapped by lower postocular and fifth supralabial, overlapping sixth supralabial; four secondary temporals, uppermost one overlapped by ectoparietal, second one overlapped by uppermost one, third one overlapped by primary temporal, overlapping second one, lowermost one overlapped by primary temporal, third one and sixth supralabial, overlapping upper postlabial; six supralabials (no variation), anterior ones overlapping posterior ones, fourth and fifth ones largely interrupted by lower-anterior postsubocular, fifth one longest, third to fifth ones under eye; two postlabials overlapped by sixth supralabial, upper one overlapping lower one.</paragraph>
<paragraph id="336C2072025789772483CBB050F2A378" pageId="0" pageNumber="77">Mental wider than and as high as rostral, overlapping postmental and first infralabial; single postmental, right part enlarged relative to left part, overlapped by first infralabial, overlapping chinshields; six infralabials on left side (five on right side) (mode = 6, 5-6), anterior ones overlapping posterior ones, first one overlapping chinshield and postgenial, sixth one smallest; a pair of chinshields, separated from each other, left one enlarged toward right side, right one distorted and smaller than left one, overlapping postgenial; a postgenial only in left side, smaller than chinshields, overlapped by second infralabial, no right postgenial.</paragraph>
<paragraph id="D3046A74A15D71B763659243E9A725E7" pageId="0" pageNumber="77">Ear opening with oval shape, no ear lobule; forelimbs with five short but distinct fingers, third one longest; third finger with two rows of supradigitals at base and seven subdigital scales (mode = 7, 5-7); 25 scale rows around midbody (mode = 26, 24-26); 61 pairs of dorsal median scales from posterior side of parietal to position of posterior margin of preanal (mode = 57, 53-62); usually three keels on a dorsal scale; hindlimbs with five short but distinct toes, fourth one longest; fourth toe with two rows of supradigitals at base and 13 subdigital scales (mode = 11, 10-14); 10 preanals, central ones with shorter length and similar width of midbody scales, overlapped by outer ones, right-central one overlapped by left-central one, outer ones small; subcaudal not enlarged; 15 scale rows around tail at 10th subcaudal position.</paragraph>
<paragraph id="73D3EE66C2613B65249816EABD1C80B3" pageId="0" pageNumber="77">
In alcohol, dorsal surface of head to tail light brown, first and second scale rows from dorsal median line with many blackish spots; no distinct blackish stripe on dorsal scale row (mode = 2, 0-3, 
<emphasis id="589829D2465F50B721E39A73F266912B" italics="true" pageId="0" pageNumber="77">N</emphasis>
= 21); first scale row from dorsal median line on original tail with a pale blackish stripe; upper and lower eyelids whitish with blackish edge; blackish line on dorsolateral surface from snout to midbody interrupting dorsal and lateral sides, with narrower width of second supralabial on snout, interrupted by eye, with similar width of dorsolateral scale from eye to neck, 1.0-1.5 times as wide as dorsolateral scale at position above forelimb, gradually obscure from that position to midbody; upper margin of blackish line distinct from posterior corner of eye to midbody, located on middle position of scales on fourth scale row from dorsal median line; lateral surface of head to posterior position of forelimb whitish with small blackish spots; lateral surface of midbody to tail dark brown with small blackish spots; ventral surface of head to tail whitish, with a small number of blackish spots on head, small gray spots on midbody, rough blackish stripe on each trunk scale row around hindlimb including preanal scales, and a blackish spot on each subcaudal scale; forelimb and hindlimb light brown on dorsal surface and whitish on ventral surface with a blackish speckle on each scale row.
</paragraph>
</subSubSection>
<subSubSection id="4C08D5D4E8BDC19B61C5C3308C6D8C0D" pageId="0" pageNumber="77" type="variation">
<paragraph id="A406CE8384F6E0BB0F16E5B43C43A891" pageId="0" pageNumber="77">Variation.</paragraph>
<paragraph id="86B288A18F17B1907CABFE73648F4439" pageId="0" pageNumber="77">
<taxonomicName id="FE2903508BB3DB0967071B0326EDB02F" baseAuthorityName="Hallowell" baseAuthorityYear="1861" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus pellopleurus" order="Squamata" pageId="0" pageNumber="77" rank="species" species="pellopleurus">
<emphasis id="2E33C01EBEA324BBDBD8D68874BDE52D" italics="true" pageId="0" pageNumber="77">Ateuchosaurus pellopleurus</emphasis>
</taxonomicName>
usually has a single SuNa with a concave posterior edge (248/299 specimens), but sometimes has a SuNa with a straight posterior margin (28/299 specimens) or two laterally separated SuNas (23/299 specimens).
</paragraph>
<paragraph id="322F38DB7C0D56BE2D59FDA5316475D7" pageId="0" pageNumber="77">
This species usually has distinct FrNa and frontal, but sometimes these are fused (Table 
<tableCitation id="E5421BF30A2255146E717A8394697C32" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Variation of the condition of frontonasal-frontal and the number of subdigital scales of the fourth toe (TIV) in Ateuchosaurus pellopleurus and A. okinavensis." httpUri="http://table.plazi.org/id/4D51506529DC5DA604A7AEAEFFC59096" pageId="0" pageNumber="77" tableUuid="4D51506529DC5DA604A7AEAEFFC59096">1</tableCitation>
). The Takeshima and Iojima populations (sites 1 and 2) also show comparable frequencies for the separated and fused types.
</paragraph>
<paragraph id="D0F7B4235B69786C3A874420827B1961" pageId="0" pageNumber="77">
This species has 8-14 TIVs (mode = 11) (Table 
<tableCitation id="2464E16103BB3EBBB797440B25312E53" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Variation of the condition of frontonasal-frontal and the number of subdigital scales of the fourth toe (TIV) in Ateuchosaurus pellopleurus and A. okinavensis." httpUri="http://table.plazi.org/id/4D51506529DC5DA604A7AEAEFFC59096" pageId="0" pageNumber="77" tableUuid="4D51506529DC5DA604A7AEAEFFC59096">1</tableCitation>
). This character shows clinal geographic variation from northern to southern populations: 8-12 (mode = 10, sites 1-7), 10-12 (mode = 11, sites 8 and 9), 10-14 (mode = 11, sites 10 and 11), and 9-14 TIVs (mode = 12, sites 12 and 13).
</paragraph>
<paragraph id="AA7E323D96FB33528712F6693293F2CB" pageId="0" pageNumber="77">The Amamioshima population has smaller MSRs, especially in males (mode = 24, 24-28 in males; mode = 26, 24-26 in females), than the other populations, which usually have 26 or 28 MSRs (mode = 26, 24-30 in males; mode = 26, 26-30 in females) with few sexual differences (Suppl. material 2).</paragraph>
<paragraph id="D74A455A1F862C2B5779693A98DFB385" pageId="0" pageNumber="77">Males have smaller DMSs (mode = 62, 53-70) than females (mode = 63, 56-69) (Suppl. material 3). This species usually has 2-3 pairs of DSRBS, but sometimes has no stripe on the dorsum or one pair of DSRBS.</paragraph>
<paragraph id="A30A228F187A3BD71965CDB7F6C8D78E" pageId="0" pageNumber="77">
The populations of the Osumi and Tokara Groups (sites 1-9), Kikaijima and Amamioshima Islands (sites 10 and 11), and Tokunoshima and Okinoerabujima Islands (sites 12 and 13) have different mitochondrial DNA (mtDNA) sequences (
<bibRefCitation id="1CF7C3EBC62BF90397B4BFDBCDADB255" DOI="https://doi.org/10.1016/j.jcz.2020.06.008" author="Makino, T" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="77" pagination="1 - 10" refId="B17" refString="Makino, T, Okamoto, T, Kurita, K, Nakano, T, Hikida, T, 2020. Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan. Zoologischer Anzeiger 288: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.06.008" title="Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan." url="https://doi.org/10.1016/j.jcz.2020.06.008" volume="288" year="2020">Makino et al. 2020</bibRefCitation>
). The Osumi-Tokara populations have extremely low genetic diversity in the partial sequences of mtDNA and recombination activating gene 1 (
<emphasis id="8D2FF863FF5D2FABC4AC2D38D75F0F46" italics="true" pageId="0" pageNumber="77">RAG1</emphasis>
) (
<bibRefCitation id="73FDA2D37A09AB80A844F19BAD8D72FB" DOI="https://doi.org/10.1016/j.jcz.2020.06.008" author="Makino, T" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="77" pagination="1 - 10" refId="B17" refString="Makino, T, Okamoto, T, Kurita, K, Nakano, T, Hikida, T, 2020. Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan. Zoologischer Anzeiger 288: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.06.008" title="Origin and intraspecific diversification of the scincid lizard Ateuchosaurus pellopleurus with implications for historical island biogeography of the Central Ryukyus of Japan." url="https://doi.org/10.1016/j.jcz.2020.06.008" volume="288" year="2020">Makino et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B298199A76C18E0D7DEAD547FCDFC53C" pageId="0" pageNumber="77" type="distribution">
<paragraph id="AFF7140436188EA03A6E95A1D25C766E" pageId="0" pageNumber="77">Distribution.</paragraph>
<paragraph id="5A6F2436FB7F950B64F306B02DA73A2A" pageId="0" pageNumber="77">
Mishima (sites 1-3 in Fig. 
<figureCitation id="7D1F3571818C5BB7E80928C1E7212015" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the geographic range of Ateuchosaurus pellopleurus and A. okinavensis in the Ryukyu Archipelago, and the sampling sites of the specimens examined in this study. A. The geographic location of the Ryukyu Archipelago; B. Sampling sites in the northern and central parts of the Ryukyu Archipelago. Each solid line indicates the geographic range of the Ateuchosaurus species revealed by Makino et al. (2020). Site numbers correspond to the locality numbers in Tables 1 - 2 and Suppl. materials 1 - 3." figureDoi="10.3897/zse.99.95923.figure1" httpUri="https://binary.pensoft.net/fig/798122" pageId="0" pageNumber="77">1</figureCitation>
) of the Osumi Group, and the Tokara (sites 4-9 and surrounding islets) and Amami Groups (sites 10-13 and surrounding islets) of the Ryukyu Archipelago.
</paragraph>
<paragraph id="FE6CE7B6AB9A28AE424D430C250F2B31" pageId="0" pageNumber="77">
A skeletal remain, probably dating from the late 19th century, is known from Yoronjima Island (the island between sites 13 and 15) (
<bibRefCitation id="4314EB3BF139387D4836B13E0F488742" DOI="https://doi.org/10.13128/Acta_Herpetol-11924" author="Nakamura, Y" journalOrPublisher="Acta Herpetologica" pageId="0" pageNumber="77" pagination="19 - 34" refId="B20" refString="Nakamura, Y, Takahashi, A, Ota, H, 2013. Recent cryptic extinction of squamate reptiles on Yoronjima Island of the Ryukyu Archipelago, Japan, inferred from garbage dump remains. Acta Herpetologica 8 (1): 19 - 34, DOI: https://doi.org/10.13128/Acta_Herpetol-11924" title="Recent cryptic extinction of squamate reptiles on Yoronjima Island of the Ryukyu Archipelago, Japan, inferred from garbage dump remains." url="https://doi.org/10.13128/Acta_Herpetol-11924" volume="8" year="2013">Nakamura et al. 2013</bibRefCitation>
), although it remains unknown whether the specimen represents this species or 
<taxonomicName id="77BAD1280FC08E8CD2701B22DCEBC54D" lsidName="A. okinavensis" pageId="0" pageNumber="77" rank="species" species="okinavensis">
<emphasis id="1F8AAF4477DF29287AF3172B2D072305" italics="true" pageId="0" pageNumber="77">A. okinavensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22F0FC92A7FC539BD81E9F3152CB8C1D" pageId="0" pageNumber="77" type="natural history">
<paragraph id="1666A6E79936077DE3F89E4D110D5B5F" pageId="0" pageNumber="77">Natural history.</paragraph>
<paragraph id="F4A78E5FDE1FE750AD3193CCCB5AA3F4" pageId="0" pageNumber="77">
This species occurs in leaf litter of grasslands and forest floors, including small vegetation around urban areas. Its reproductive season may range from May to August (
<bibRefCitation id="D393AC6AE29AECD1553A27BDB3AEE56B" author="Okada, S" journalOrPublisher="Natural History Research" pageId="0" pageNumber="77" pagination="43 - 52" refId="B25" refString="Okada, S, Ota, H, Hasegawa, T, Hikida, T, Miyaguni, H, Kato, J, 1992. Reproductive traits of seven species of lygosominae skinks (Squamata: Reptilia) from East Asia. Natural History Research 2 (1): 43 - 52" title="Reproductive traits of seven species of lygosominae skinks (Squamata: Reptilia) from East Asia." volume="2" year="1992">Okada et al. 1992</bibRefCitation>
).
</paragraph>
<paragraph id="B9375681782E4048EF6CCCF91EDFE97C" pageId="0" pageNumber="77">
This skink is more active on sunny days (56/75 specimens) than on cloudy (11/75 specimens) or rainy days (8/75 specimens). This species is usually seen in the daytime (10:03-17:52, 79/84 specimens) in spring to autumn, but is sometimes seen in the evening to nighttime (18:11-21:48, 5/84 specimens) on Takeshima, Iojima, Kodakarajima, Amamioshima, Tokunoshima and Okinoerabujima Islands (sites 1, 2, 8, and 11-13 in Fig. 
<figureCitation id="EC85E3B347704178BF5687A459B17F94" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the geographic range of Ateuchosaurus pellopleurus and A. okinavensis in the Ryukyu Archipelago, and the sampling sites of the specimens examined in this study. A. The geographic location of the Ryukyu Archipelago; B. Sampling sites in the northern and central parts of the Ryukyu Archipelago. Each solid line indicates the geographic range of the Ateuchosaurus species revealed by Makino et al. (2020). Site numbers correspond to the locality numbers in Tables 1 - 2 and Suppl. materials 1 - 3." figureDoi="10.3897/zse.99.95923.figure1" httpUri="https://binary.pensoft.net/fig/798122" pageId="0" pageNumber="77">1</figureCitation>
). It is found in warm microhabitats with temperatures of more than 20.0 °C (25.2 
<normalizedToken id="843181739A45A54BBCB10546CD27FFA3" originalValue="±">+/-</normalizedToken>
2.5 °C, 20.8-30.2 °C, 84 specimens) (T.M., personal observation).
</paragraph>
</subSubSection>
<subSubSection id="C5B5DE61332C80F23EAB9AD2AECBAFD5" pageId="0" pageNumber="77" type="conservation">
<paragraph id="74CC03725CA1FBCA0778DA53B007AC23" pageId="0" pageNumber="77">Conservation.</paragraph>
<paragraph id="C31761A050707EC6F18042DFF1C44CF5" pageId="0" pageNumber="77">
The island population of Iojima (site 2) is endangered by predation from the non-native Indian Peafowl 
<taxonomicName id="3C5F7A0FB32D2D145E3AFD145AE59E3A" authorityName="Linnaeus" authorityYear="1758" class="Aves" family="Phasianidae" genus="Pavo" kingdom="Animalia" lsidName="Pavo cristatus" order="Galliformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="cristatus">
<emphasis id="EA8BE4C083F3D17FEBD9C815CB37170A" italics="true" pageId="0" pageNumber="77">Pavo cristatus</emphasis>
</taxonomicName>
Linnaeus, 1758, and the populations of Mishima (sites 1-3) are designated as Threatened Local Population (LP) in the Red Data Book of Japan (
<bibRefCitation id="233A808A745966586CCAECD5A36A5F03" author="Ota, H" journalOrPublisher="- Threatened Wildlife of Japan- Volume 3, Reptilia / Amphibia. GYOSEI Corporation, Tokyo" pageId="0" pageNumber="77" pagination="89 - 90" refId="B28" refString="Ota, H, 2014. Ryukyu short-legged skink (populations in Mishima Islands). In: Red Data Book 2014. -Threatened Wildlife of Japan- Volume 3, Reptilia/Amphibia. GYOSEI Corporation, Tokyo: 89 - 90" title="Ryukyu short-legged skink (populations in Mishima Islands)." volumeTitle="Red Data Book 2014." year="2014">Ota 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>