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<mods:title id="B6E53D2C22DFA006511EDAAA747642A5">A new icriodontid conodont cluster with specific mesowear supports an alternative apparatus motion model for Icriodontidae</mods:title>
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<mods:namePart id="BF9D4107C52CFE1728219051A090A02D">Suttner, Thomas J.</mods:namePart>
<mods:affiliation id="C9A03F4FD15526F287BAC4B936167A73">University of Graz, Institute for Earth Sciences, Heinrichstrasse 26, 8010 Graz, Austria; &amp; Geological-Palaeontological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria;</mods:affiliation>
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<mods:namePart id="208A04F6C4EEF170EEE9B8A455BC680D">Kido, Erika</mods:namePart>
<mods:affiliation id="C6E3E18C9FB0443FDF161646DEF12291">University of Graz, Institute for Earth Sciences, Heinrichstrasse 26, 8010 Graz, Austria;</mods:affiliation>
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<mods:namePart id="329AC758D4069E43922AC132559A412D">Briguglio, Antonino</mods:namePart>
<mods:affiliation id="3824FEB3C4815327E3FC4B04323251FE">Faculty of Science, Universiti Brunei Darussalam, Jalan Tungku Link, Gadong BE 1410, Brunei Darussalam</mods:affiliation>
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<subSubSection id="54E2654AFFAC0637AFA7F81FFAAFF821" box="[959,1352,1942,1966]" pageId="2" pageNumber="910" type="nomenclature">
<paragraph id="1C4736C1FFAC0637AFA7F81FFAAFF821" blockId="2.[882,1430,1783,1966]" box="[959,1352,1942,1966]" pageId="2" pageNumber="910">
Genus
<taxonomicName id="DBF84D42FFAC0637A812F81EFAAFF821" authority="Bultynck, 1976" authorityName="Bultynck" authorityYear="1976" box="[1034,1352,1942,1966]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prioniodontida" pageId="2" pageNumber="910" phylum="Chordata" rank="genus">
<emphasis id="2E8CEAD3FFAC0637A812F81EFB79F821" bold="true" box="[1034,1182,1942,1966]" italics="true" pageId="2" pageNumber="910">Caudicriodus</emphasis>
<bibRefCitation id="78694B30FFAC0637A8BCF81FFAAFF821" author="Bultynck, P." box="[1188,1352,1943,1966]" pageId="2" pageNumber="910" pagination="1 - 73" refId="ref9131" refString="Bultynck, P. 1976. Le Silurien Superieur et le Devonien Inferieur de la Sierra de Guadarrama (Espagne Centrale). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, 49 (5), 1 - 73." type="journal article" year="1976">Bultynck, 1976</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<caption id="48876649FFAD0636AC87FAFBFE49FA2C" ID-DOI="http://doi.org/10.5281/zenodo.10883200" ID-Zenodo-Dep="10883200" httpUri="https://zenodo.org/record/10883200/files/figure.png" pageId="3" pageNumber="911" startId="3.[159,227,1395,1416]" targetBox="[173,1453,183,1363]" targetPageId="3" targetType="figure">
<paragraph id="1C4736C1FFAD0636AC87FAFBFE49FA2C" blockId="3.[159,1466,1395,1443]" pageId="3" pageNumber="911">
<emphasis id="2E8CEAD3FFAD0636AC87FAFBFF1BFA07" bold="true" box="[159,252,1395,1416]" pageId="3" pageNumber="911">Figure 1.</emphasis>
Locality map and section log from the Kottwitz quarry (southern Burgenland, Austria), where the
<taxonomicName id="DBF84D42FFAD0636A8DDFAFBFA5DFA07" baseAuthorityName="Serpagli" baseAuthorityYear="1983" box="[1221,1466,1395,1416]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="woschmidti">
<emphasis id="2E8CEAD3FFAD0636A8DDFAFBFA5DFA07" box="[1221,1466,1395,1416]" italics="true" pageId="3" pageNumber="911">Caudicriodus woschmidti</emphasis>
</taxonomicName>
conodont cluster was found.
</paragraph>
</caption>
<subSubSection id="54E2654AFFAD0636AC87F99FFEEDF9E1" pageId="3" pageNumber="911" type="type_taxon">
<paragraph id="1C4736C1FFAD0636AC87F99FFEEDF9E1" blockId="3.[159,781,1559,1646]" pageId="3" pageNumber="911">
<materialsCitation id="AC903C9CFFAD0636AC87F99FFEE1F9E1" country="Germany" county="Rhenish Slate Mountains" location="lower Lochkovian" municipality="Untenruden" pageId="3" pageNumber="911" specimenCount="1" typeStatus="holotype">
<emphasis id="2E8CEAD3FFAD0636AC87F99FFEADF9A1" bold="true" box="[159,330,1559,1582]" pageId="3" pageNumber="911">
<typeStatus id="C3438863FFAD0636AC87F99FFF3EF9A1" box="[159,217,1559,1582]" pageId="3" pageNumber="911">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="DBF84D42FFAD0636AD40F99FFCE2F9A1" authority="Ziegler, 1960" authorityName="Ziegler" authorityYear="1960" box="[344,773,1559,1582]" family="Icriodontidae" genus="Icriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="woschmidti">
<emphasis id="2E8CEAD3FFAD0636AD40F99FFDA2F9A1" box="[344,581,1559,1582]" italics="true" pageId="3" pageNumber="911">Icriodus woschmidti</emphasis>
<bibRefCitation id="78694B30FFAD0636AE79F99FFCE2F9A1" author="Ziegler, W." box="[609,773,1559,1582]" pageId="3" pageNumber="911" pagination="169 - 201" refId="ref12391" refString="Ziegler, W. 1960. Conodonten aus dem Rheinischen Unterdevon (Gedinnium) des Remscheider Sattels (Rheinisches Schiefergebirge). Pala ¨ ontologische Zeitschrift, 34, 169 - 201." type="journal article" year="1960">Ziegler, 1960</bibRefCitation>
</taxonomicName>
;
<location id="1927601AFFAD0636AC87F9BFFE82F9C1" LSID="urn:lsid:plazi:treatment:945187D7FFAC0636AFA7F81FFA6FF803:1927601AFFAD0636AC87F9BFFE82F9C1" box="[159,357,1591,1614]" country="Germany" county="Rhenish Slate Mountains" municipality="Untenruden" name="lower Lochkovian" pageId="3" pageNumber="911">lower Lochkovian</location>
;
<collectingMunicipality id="FC23ACBBFFAD0636AD68F9BFFE17F9C1" box="[368,496,1591,1614]" pageId="3" pageNumber="911">Untenruden</collectingMunicipality>
,
<collectingCounty id="F5264E4DFFAD0636ADE3F9BFFCEEF9C1" box="[507,777,1591,1614]" pageId="3" pageNumber="911">Rhenish Slate Mountains</collectingCounty>
,
<collectingCountry id="64EF7651FFAD0636AC87F9DFFEE1F9E1" box="[159,262,1623,1646]" name="Germany" pageId="3" pageNumber="911">Germany</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection id="54E2654AFFAD0636AC87F910FA6FF803" pageId="3" pageNumber="911" type="discussion">
<paragraph id="1C4736C1FFAD0636AC87F910FB99F9E1" blockId="3.[159,781,1688,1934]" lastBlockId="3.[845,1467,1559,1932]" pageId="3" pageNumber="911">
<emphasis id="2E8CEAD3FFAD0636AC87F910FEE9F920" bold="true" box="[159,270,1688,1711]" pageId="3" pageNumber="911">Remarks.</emphasis>
The generic diagnosis follows
<bibRefCitation id="78694B30FFAD0636AEB1F910FF0DF940" author="Bultynck, P." pageId="3" pageNumber="911" pagination="1 - 73" refId="ref9131" refString="Bultynck, P. 1976. Le Silurien Superieur et le Devonien Inferieur de la Sierra de Guadarrama (Espagne Centrale). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, 49 (5), 1 - 73." type="journal article" year="1976">Bultynck (1976)</bibRefCitation>
. The outline of the widely opened basal cavity (termed platform in earlier publications, but not homologous to the platform in polygnathids, for example) is identical in shape to that of other species of
<taxonomicName id="DBF84D42FFAD0636AC87F8BFFF1DF8C1" authorityName="Branson &amp; Mehl" authorityYear="1938" box="[159,250,1847,1870]" family="Icriodontidae" genus="Icriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="genus">
<emphasis id="2E8CEAD3FFAD0636AC87F8BFFF1DF8C1" box="[159,250,1847,1870]" italics="true" pageId="3" pageNumber="911">Icriodus</emphasis>
</taxonomicName>
. Features which discriminate
<taxonomicName id="DBF84D42FFAD0636AE61F8BFFCEAF8C1" authorityName="Bultynck" authorityYear="1976" box="[633,781,1847,1870]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="genus">
<emphasis id="2E8CEAD3FFAD0636AE61F8BFFCEAF8C1" box="[633,781,1847,1870]" italics="true" pageId="3" pageNumber="911">Caudicriodus</emphasis>
</taxonomicName>
from
<taxonomicName id="DBF84D42FFAD0636ACFCF8DFFED8F8E1" authorityName="Branson &amp; Mehl" authorityYear="1938" box="[228,319,1879,1902]" family="Icriodontidae" genus="Icriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="genus">
<emphasis id="2E8CEAD3FFAD0636ACFCF8DFFED8F8E1" box="[228,319,1879,1902]" italics="true" pageId="3" pageNumber="911">Icriodus</emphasis>
</taxonomicName>
are the lateral process (denticulate or adenticulate) that extends posterior of the cusp at a specific angle, and a spur developed between the cusp and posterior-most transverse denticle row on the inner side of the element.
</paragraph>
<paragraph id="1C4736C1FFAD0636AF7FF9FEFA6FF803" blockId="3.[845,1467,1559,1932]" pageId="3" pageNumber="911">
<bibRefCitation id="78694B30FFAD0636AF7FF9FEFBFEF902" author="Bultynck, P." box="[871,1049,1654,1677]" pageId="3" pageNumber="911" pagination="1 - 73" refId="ref9131" refString="Bultynck, P. 1976. Le Silurien Superieur et le Devonien Inferieur de la Sierra de Guadarrama (Espagne Centrale). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, 49 (5), 1 - 73." type="journal article" year="1976">Bultynck (1976)</bibRefCitation>
included the following species within
<taxonomicName id="DBF84D42FFAD0636AF55F91EFC39F922" authorityName="Bultynck" authorityYear="1976" box="[845,990,1686,1709]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="genus">
<emphasis id="2E8CEAD3FFAD0636AF55F91EFC39F922" box="[845,990,1686,1709]" italics="true" pageId="3" pageNumber="911">Caudicriodus</emphasis>
</taxonomicName>
:
<taxonomicName id="DBF84D42FFAD0636AFEAF91EFAEEF922" baseAuthorityName="Serpagli" baseAuthorityYear="1983" box="[1010,1289,1686,1709]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="woschmidti">
<emphasis id="2E8CEAD3FFAD0636AFEAF91EFAEEF922" box="[1010,1289,1686,1709]" italics="true" pageId="3" pageNumber="911">Caudicriodus woschmidti</emphasis>
</taxonomicName>
,
<taxonomicName id="DBF84D42FFAD0636A905F91EFC63F942" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="postwoschmidti">
<emphasis id="2E8CEAD3FFAD0636A905F91EFC63F942" italics="true" pageId="3" pageNumber="911">C. postwoschmidti</emphasis>
</taxonomicName>
,
<taxonomicName id="DBF84D42FFAD0636AF8BF93EFBD3F942" box="[915,1076,1718,1741]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="angustoides">
<emphasis id="2E8CEAD3FFAD0636AF8BF93EFBD3F942" box="[915,1076,1718,1741]" italics="true" pageId="3" pageNumber="911">C. angustoides</emphasis>
</taxonomicName>
,
<taxonomicName id="DBF84D42FFAD0636A85CF93EFB3AF942" box="[1092,1245,1718,1741]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="curvicauda">
<emphasis id="2E8CEAD3FFAD0636A85CF93EFB3AF942" box="[1092,1245,1718,1741]" italics="true" pageId="3" pageNumber="911">C. curvicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="DBF84D42FFAD0636A8F5F93EFA6CF942" box="[1261,1419,1718,1741]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="celtibericus">
<emphasis id="2E8CEAD3FFAD0636A8F5F93EFA6CF942" box="[1261,1419,1718,1741]" italics="true" pageId="3" pageNumber="911">C. celtibericus</emphasis>
</taxonomicName>
and
<taxonomicName id="DBF84D42FFAD0636AF55F95EFC0FF962" box="[845,1000,1750,1773]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="sigmoidalis">
<emphasis id="2E8CEAD3FFAD0636AF55F95EFC0FF962" box="[845,1000,1750,1773]" italics="true" pageId="3" pageNumber="911">C. sigmoidalis</emphasis>
</taxonomicName>
. Later,
<bibRefCitation id="78694B30FFAD0636A825F95EFB03F962" author="Drygant, D. M." box="[1085,1252,1750,1773]" pageId="3" pageNumber="911" refId="ref9670" refString="Drygant, D. M. 2010. [Devonian Conodonts from South - West Margin of the East European Platform (Volyn' - Podolian Ukraine]. Academperiodyka, Kyiv, 156 pp. [In Ukrainian.]" type="book" year="2010">Drygant (2010)</bibRefCitation>
included four additional species,
<taxonomicName id="DBF84D42FFAD0636AFF4F97EFB0DF882" box="[1004,1258,1782,1805]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="2E8CEAD3FFAD0636AFF4F97EFB0DF882" box="[1004,1258,1782,1805]" italics="true" pageId="3" pageNumber="911">Caudicriodus hesperius</emphasis>
</taxonomicName>
,
<taxonomicName id="DBF84D42FFAD0636A8E0F97EFA54F882" box="[1272,1459,1782,1805]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="ruthmawsonae">
<emphasis id="2E8CEAD3FFAD0636A8E0F97EFA54F882" box="[1272,1459,1782,1805]" italics="true" pageId="3" pageNumber="911">C. ruthmawsonae</emphasis>
</taxonomicName>
,
<taxonomicName id="DBF84D42FFAD0636AF55F89EFC33F8A2" box="[845,980,1814,1837]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="transiens">
<emphasis id="2E8CEAD3FFAD0636AF55F89EFC33F8A2" box="[845,980,1814,1837]" italics="true" pageId="3" pageNumber="911">C. transiens</emphasis>
</taxonomicName>
and
<taxonomicName id="DBF84D42FFAD0636A80DF89EFB94F8A2" box="[1045,1139,1814,1837]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="serus">
<emphasis id="2E8CEAD3FFAD0636A80DF89EFB94F8A2" box="[1045,1139,1814,1837]" italics="true" pageId="3" pageNumber="911">C. serus</emphasis>
</taxonomicName>
, and
<bibRefCitation id="78694B30FFAD0636A8A2F89EFC73F8C2" author="Drygant, D. &amp; Szaniawski, H." pageId="3" pageNumber="911" pagination="833 - 861" refId="ref9710" refString="Drygant, D. &amp; Szaniawski, H. 2012. Lochkovian conodonts from Podolia, Ukraine and their stratigraphic significance. Acta Palaeontologica Polonica, 57, 833 - 861." type="journal article" year="2012">Drygant &amp; Szaniawski (2012)</bibRefCitation>
described
<taxonomicName id="DBF84D42FFAD0636A829F8BEFAB8F8C2" box="[1073,1375,1846,1869]" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="schoenlaubi">
<emphasis id="2E8CEAD3FFAD0636A829F8BEFAB8F8C2" box="[1073,1375,1846,1869]" italics="true" pageId="3" pageNumber="911">Caudicriodus schoenlaubi</emphasis>
</taxonomicName>
. More recently,
<bibRefCitation id="78694B30FFAD0636AFA0F8DEFB4AF8E2" author="Murphy, M. A. &amp; Carls, P. &amp; J. I" box="[952,1197,1878,1901]" pageId="3" pageNumber="911" pagination="4 - 10" refId="ref10920" refString="Murphy, M. A., Carls, P. &amp; Valenzuela-R´ios, J. I. 2016. Cypricriodus new genus. University of California, Riverside Campus Museum Contribution, 8, 4 - 10." type="journal article" year="2016">
Murphy
<emphasis id="2E8CEAD3FFAD0636A803F8DEFBBEF8E2" box="[1051,1113,1878,1901]" italics="true" pageId="3" pageNumber="911">et al.</emphasis>
(2016)
</bibRefCitation>
excluded
<taxonomicName id="DBF84D42FFAD0636A931F8DEFC54F803" family="Icriodontidae" genus="Caudicriodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="911" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="2E8CEAD3FFAD0636A931F8DEFC54F803" italics="true" pageId="3" pageNumber="911">Caudicriodus hesperius</emphasis>
</taxonomicName>
, placing it in their new genus
<taxonomicName id="DBF84D42FFAD0636A8ECF8FDFA67F803" box="[1268,1408,1909,1932]" genus="Cypricriodus" pageId="3" pageNumber="911" rank="genus">
<emphasis id="2E8CEAD3FFAD0636A8ECF8FDFA67F803" box="[1268,1408,1909,1932]" italics="true" pageId="3" pageNumber="911">Cypricriodus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>