296 lines
36 KiB
XML
296 lines
36 KiB
XML
<document ID-CLB-Dataset="9289" ID-DOI="http://dx.doi.org/10.3897/zookeys.1004.57526" ID-GBIF-Dataset="4e633bc1-16fc-4e83-b8a0-cb9289897616" ID-PMC="PMC7758311" ID-Pensoft-Pub="1313-2970-1004-27" ID-Pensoft-UUID="BEECC18115E95A798CA5250990A350E3" ID-PubMed="33384565" ID-ZooBank="320559CF19B5423CB7FB72555290241A" ModsDocID="1313-2970-1004-27" checkinTime="1608157838683" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. & Kunte, Krushnamegh" docDate="2020" docId="8CD7BE05252F53729494CDA75FA4DA80" docLanguage="en" docName="ZooKeys 1004: 27-97" docOrigin="ZooKeys 1004" docPubDate="2020-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1004.57526" docTitle="Maripanthus draconis Maddison, sp. nov." docType="treatment" docVersion="6" id="BEECC18115E95A798CA5250990A350E3" lastPageNumber="27" masterDocId="BEECC18115E95A798CA5250990A350E3" masterDocTitle="A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)" masterLastPageNumber="97" masterPageNumber="27" pageNumber="27" updateTime="1732936594770" updateUser="ExternalLinkService">
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<mods:title id="473AA475C7D9FF1B2ED80CA50E8B04D7">A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)</mods:title>
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<mods:namePart id="7721C5DFC5BB6D80354FD4CEED24EBBC">Maddison, Wayne P.</mods:namePart>
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<mods:affiliation id="156FA487226782AB857D7F30B0A242DF">Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:nameIdentifier id="ADB62B986D200DD04D07F6BB7286F73A" type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
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<mods:namePart id="8DF42D2A81E039CA0B9248E331D6AC57">Beattie, Imara</mods:namePart>
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<mods:affiliation id="2A33B095BFB6F825CE2413EFCED11136">Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:namePart id="09651BCE060E31CAB30E5B89407BFE5C">Marathe, Kiran</mods:namePart>
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<mods:affiliation id="58280C63CEF0D8D1121CA5460D519723">Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada & National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:namePart id="919F7114DBDCE6F8A4926CC527E43760">Ng, Paul Y. C.</mods:namePart>
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<mods:affiliation id="9CD3C14B13B196C0A07DC5592A84CE12">205 River Valley Road, # 16 - 53, Singapore 238274, Republic of Singapore</mods:affiliation>
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<mods:namePart id="9FAC9FDEA4965ADC422E2B6E1D831F7E">Kanesharatnam, Nilani</mods:namePart>
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<mods:nameIdentifier id="C3449E67244B38E6908DEA54F71AF081" type="ORCID">https://orcid.org/0000-0002-6070-8989</mods:nameIdentifier>
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<mods:affiliation id="1FF340AF821C993DCC7A580233F21BF0">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka & Department of Zoology, Faculty of Science, Eastern University, Vantharumoolai, Sri Lanka</mods:affiliation>
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<mods:namePart id="A3F4DC86EB5769B15883762E259662E7">Benjamin, Suresh P.</mods:namePart>
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<mods:affiliation id="283428E34A242091DEED015FC4610C05">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
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<mods:namePart id="1FE52C53F5A06897E62CF78D0A5E3628">Kunte, Krushnamegh</mods:namePart>
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<mods:affiliation id="2CF8237582FD4FB3AFEF24549F1C7127">National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:title id="ECF323F6C5CA7BD081DF014791D181F4">ZooKeys</mods:title>
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<mods:date id="9682DA711C8FC8A320EA1D385DEE7E96">2020</mods:date>
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<mods:number id="8CA135206AA1F1093FC5908FB8ADDFD6">2020-12-16</mods:number>
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<mods:number id="4C09DD507C199030792494115E24741B">1004</mods:number>
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<mods:start id="46DC69828C08EF6F246A82FD82D28B79">27</mods:start>
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<mods:end id="BB51CF12FE93C92F1D24165E8A16D32D">97</mods:end>
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<mods:url id="F74CA61A620181D51B29AEE6E933F60F">http://dx.doi.org/10.3897/zookeys.1004.57526</mods:url>
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<mods:classification id="9E9B48108EBFEFC5A13143C44AA7385F">journal article</mods:classification>
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<mods:identifier id="B6AF7CD158D541337CA7DCC0EBB8BFAC" type="DOI">http://dx.doi.org/10.3897/zookeys.1004.57526</mods:identifier>
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<treatment id="8CD7BE05252F53729494CDA75FA4DA80" ID-GBIF-Taxon="176088103" LSID="urn:lsid:zoobank.org:act:B511C74F-1343-48B8-ACDD-B01B20549324" httpUri="http://treatment.plazi.org/id/8CD7BE05252F53729494CDA75FA4DA80" lastPageNumber="27" pageId="0" pageNumber="27" scope_family="Salticidae" scope_order="Araneae" scope_tribe="Baviini">
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<subSubSection id="D1CE5CCD814E0110AC6B9C823BBF9F7F" pageId="0" pageNumber="27" type="nomenclature">
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<paragraph id="84C603F32ED5A26CFE28B599FE3D8426" pageId="0" pageNumber="27">
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<taxonomicName id="085FEF8608D58C4E36FEF93EAE4F1904" LSID="http://zoobank.org/B511C74F-1343-48B8-ACDD-B01B20549324" authority="Maddison, sp. nov." authorityName="Maddison, sp. nov." class="Magnoliopsida" family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus draconis" order="Apiales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="draconis">Maripanthus draconis Maddison, sp. nov.</taxonomicName>
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<figureCitation id="1B5DB5838A67C332D5756ACD4D888079" captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">Figs 27</figureCitation>
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<figureCitation id="B8AD8076325DBF86D16E75DE6F07B6C5" captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 34</figureCitation>
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<figureCitation id="F5A1850442E48BEAABC84A2E9C26BCD0" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">, 188-201</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="98AD36C61BCFB45BC73E80996538DC78" pageId="0" pageNumber="27" type="type material">
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<paragraph id="A774030AD62959E2366F342FBCD771FF" pageId="0" pageNumber="27">Type material.</paragraph>
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<paragraph id="BD6DD18166293635B4E6C02E78BEF608" pageId="0" pageNumber="27">
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<emphasis id="FFC92134CCA2A0F325E632989C505614" bold="true" italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
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male (specimen AS19.2232), two
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<emphasis id="8A919629C4961BF930E04E804C706539" bold="true" italics="true" pageId="0" pageNumber="27">paratype</emphasis>
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females (specimens AS19.2250 and d547), all in LKCNHM, from Singapore: Bukit Timah Nature Reserve, stream at Jungle Falls Path.
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<geoCoordinate id="BA7120312A4C211AC68E850C556B6318" degrees="1.3562" direction="north" orientation="latitude" precision="5" value="1.3562">1.3562°N</geoCoordinate>
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,
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<geoCoordinate id="66A2784F823B3006F4A18FE5591EFFE7" degrees="103.7748" direction="east" orientation="longitude" precision="5" value="103.7748">103.7748°E</geoCoordinate>
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to
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<geoCoordinate id="D0D365AD948CB94038A6872E9BABEADC" degrees="1.3572" direction="north" orientation="latitude" precision="5" value="1.3572">1.3572°N</geoCoordinate>
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,
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<geoCoordinate id="FD5849E6EFEE0079D0F55C75F5FDC285" degrees="103.7734" direction="east" orientation="longitude" precision="5" value="103.7734">103.7734°E</geoCoordinate>
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110-150 m elev. 12 June 2019 Maddison, Morehouse, & Marathe WPM#19-051.
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<emphasis id="FFADA10BCEF8C0E03A7B586E199D1A16" bold="true" italics="true" pageId="0" pageNumber="27">Paratype</emphasis>
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male (specimen WSG018) from Singapore: Nee Soon Swamp Forest. Beating vegetation.
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<geoCoordinate id="998A212281D43C9AFFD0FAC23680E9F0" degrees="1.39" direction="north" orientation="latitude" precision="555" value="1.39">1.39°N</geoCoordinate>
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,
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<geoCoordinate id="99C8A6A0BEC8924B61C60674AE5A178E" degrees="103.81" direction="east" orientation="longitude" precision="555" value="103.81">103.81°E</geoCoordinate>
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12 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-015.
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</paragraph>
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</subSubSection>
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<subSubSection id="C004DC55B21F3257C49105CBF2BD2D58" pageId="0" pageNumber="27" type="etymology">
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<paragraph id="9A9363BFAC3603EBC7EE1A5323799676" pageId="0" pageNumber="27">Etymology.</paragraph>
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<paragraph id="9E065419B41E7506D4F36E9B8A46F557" pageId="0" pageNumber="27">
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Greek,
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<emphasis id="F9FEC4C3ABC5CA5F35DECB54AB1165F2" italics="true" pageId="0" pageNumber="27">
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<normalizedToken id="6976AF4DED2AE3FB42665D5525EDC9A0" originalValue="δράκων">δράκων</normalizedToken>
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</emphasis>
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, referring to the fiery colours of the male.
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<emphasis id="67F9BE14766A7834983FD83CE1B209C5" italics="true" pageId="0" pageNumber="27">Other names</emphasis>
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: In
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<bibRefCitation id="230505AF85CB9B5C78253D4E9D0143D6" author="Maddison, WP" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="27" publicationUrl="http://salticidae.org/salticidImages/" refId="B23" refString="Maddison, WP, 2015b. Salticid images. http://salticidae.org/salticidImages/" title="Salticid images." url="http://salticidae.org/salticidImages/" year="2015 b">Maddison (2015b)</bibRefCitation>
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and
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<normalizedToken id="1447FBC64BB84F7AC117B009979B95C3" originalValue="WPM’s">WPM's</normalizedToken>
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field or lab notebooks the informal code for this species was
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<normalizedToken id="8D7F34770E13FA62AB6AB7A8D3AEDCDA" originalValue="“CFMAR”">"CFMAR"</normalizedToken>
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. A specimen that is likely a closely related but distinct species is shown in
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<bibRefCitation id="AA68B502A5C543439CC2DEE8D36BF1E1" author="Koh, JKH" journalOrPublisher="Sabah Forestry Department, city" pageId="0" pageNumber="27" refId="B18" refString="Koh, JKH, Bay, N, 2019. Borneo Spiders: A photographic field guide. Sabah Forestry Department, city" title="Borneo Spiders: A photographic field guide." year="2019">Koh and Bay 2019</bibRefCitation>
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as "
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<taxonomicName id="0AC39D040A476816B05E97FC923BAC57" authorityName="Simon" authorityYear="1877" class="Arachnida" family="Pittosporaceae" genus="Bavia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bavia" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="39AF71B7FC994C2894ED8E209864CCA2" italics="true" pageId="0" pageNumber="27">Bavia</emphasis>
|
||
</taxonomicName>
|
||
" sp. B Black-collared long-bellied jumping spider.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D59169309A7DA928CA7CCEB2E898F8CC" pageId="0" pageNumber="27" type="diagnosis">
|
||
<paragraph id="C31FB1DBD6D125A8191A17D8F8425F17" pageId="0" pageNumber="27">Diagnosis.</paragraph>
|
||
<paragraph id="B1FE24B7CC879F638D9D29B5122A686A" pageId="0" pageNumber="27">
|
||
Most similar to
|
||
<taxonomicName id="9BD14C98AE3BB49D918E16824896DA52" lsidName="M. smedleyi" pageId="0" pageNumber="27" rank="species" species="smedleyi">
|
||
<emphasis id="513785917560EC7892488C7731DAC69B" italics="true" pageId="0" pageNumber="27">M. smedleyi</emphasis>
|
||
</taxonomicName>
|
||
, of which only the female is known;
|
||
<taxonomicName id="3E4584E0387E1022875E52BEF4310DFD" lsidName="M. draconis" pageId="0" pageNumber="27" rank="species" species="draconis">
|
||
<emphasis id="5C7DA26C45ADDEBE908D04A7C72F0209" italics="true" pageId="0" pageNumber="27">M. draconis</emphasis>
|
||
</taxonomicName>
|
||
differs in having longer epigynal atria (greater than half of epigynal length) and less distinct atrial cliff (
|
||
<normalizedToken id="9ACB7E5E4F32F10B0E95E31FD8C3BDC0" originalValue="“ac”">"ac"</normalizedToken>
|
||
, Fig.
|
||
<figureCitation id="E5A4324B17CB1FB82FCC7722FB59C0D9" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">191</figureCitation>
|
||
). Similar to
|
||
<taxonomicName id="0EEB9FBA9CF6671EF0EF206CD9CBC485" lsidName="M. jubatus" pageId="0" pageNumber="27" rank="species" species="jubatus">
|
||
<emphasis id="8439E415F181C51DD8D0780D748A0F8C" italics="true" pageId="0" pageNumber="27">M. jubatus</emphasis>
|
||
</taxonomicName>
|
||
in being large bodied, but
|
||
<taxonomicName id="46E6335AB16A431828CD2E1FEACC8235" lsidName="M. draconis" pageId="0" pageNumber="27" rank="species" species="draconis">
|
||
<emphasis id="D58A306D9872EB2368C99CA8726A78E2" italics="true" pageId="0" pageNumber="27">M. draconis</emphasis>
|
||
</taxonomicName>
|
||
differs in having a honey-coloured translucent thorax, which in the female is speckled with black, and which in the male contrasts strongly against the black ocular area. Male front is impressively red-orange, with red-orange first femora, palps, and rings around eyes, and a golden face. Embolus slightly shorter than the other three species for which males are known. Ventral bump on palp femur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2E08D0FA9AA0C44157AE76B85E9758EC" pageId="0" pageNumber="27" type="description">
|
||
<paragraph id="0F2B5B5E9BB0B86E24A85A663C9EDC8A" pageId="0" pageNumber="27">Description.</paragraph>
|
||
<paragraph id="66C66E60E5A247962742CD325D080896" pageId="0" pageNumber="27">
|
||
<emphasis id="38FA1C14522163876F56ED4D68E5E681" bold="true" pageId="0" pageNumber="27">Male</emphasis>
|
||
(based on holotype, specimen AS19.2232). Carapace length 4.0; abdomen length 5.0.
|
||
<emphasis id="69D85F653BACC64573E60FB2AB20478E" bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
|
||
(Figs
|
||
<figureCitation id="B392C0BE3011F8A601E8CEA98027ED9C" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">196</figureCitation>
|
||
,
|
||
<figureCitation id="7297851CEF928C0EF6623D35403D9866" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">197</figureCitation>
|
||
) black in ocular area and in a thin line along ventral and posterior margin; thorax strongly contrasting honey coloured, glabrous except for a few vertical black setae.
|
||
<emphasis id="6AD79EABBEBE110C13612CAC15B3B68B" bold="true" italics="true" pageId="0" pageNumber="27">Clypeus</emphasis>
|
||
covered with yellow-cream setae.
|
||
<emphasis id="6924452CF292FA700DC724A2F1E3A109" bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
|
||
vertical, a soft orange-red, covered with yellow-cream setae. Three promarginal and five retromarginal teeth.
|
||
<emphasis id="96FA56D610B7D1C8D614EBBA82EDFF71" bold="true" italics="true" pageId="0" pageNumber="27">Palp</emphasis>
|
||
femur pale, darkening to red-brown distally. Bulb elongate (Figs
|
||
<figureCitation id="7DB377319203AF1DD49B0C43E4DF9F97" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">188</figureCitation>
|
||
,
|
||
<figureCitation id="E2789899D5F90146DBCB0E249FF9E538" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">189</figureCitation>
|
||
); embolus arising on the prolateral basal corner and proceeding distally. Femur with small but distinct distal ventral bump. Endite with small corner projection.
|
||
<emphasis id="2FB395B4735F134BD17E1649EAAFAE7F" bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
|
||
with pale femora and darker markings more distally. First femur striking orange-red, patella to tarsus dark brown to black. First patella through metatarsus with sparse ventral fringe of setae that are black at base, white at tip.
|
||
<emphasis id="E8032653A6278F7B1AC70315C38B3D08" bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
|
||
with brown longitudinal medial band dorsally, flanked by scattered red scales, with some scattered white scales basally.
|
||
</paragraph>
|
||
<paragraph id="024ED496A99C7B52A527A875D56D8A94" pageId="0" pageNumber="27">
|
||
<emphasis id="F15351BA63662E8399CCC36F52883CD7" bold="true" pageId="0" pageNumber="27">Female</emphasis>
|
||
(paratype, specimen AS19.2250). Carapace length 4.2; abdomen length 6.1.
|
||
<emphasis id="2C447E589651CECA185BD8E1A5F57A93" bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
|
||
(Fig.
|
||
<figureCitation id="F15A830624A174CFF82A0D69EA81DA16" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">199</figureCitation>
|
||
) integument black in ocular area, except for a pale spot on each side just medial to the PME (in other females, these pale spots form a transverse band that intersects with a longitudinal medial pale band, forming a pale cross in the ocular area). Thorax honey-coloured with distinct black speckles.
|
||
<emphasis id="EFBC2B369DB181D3A86E9F5ADB881BC8" bold="true" italics="true" pageId="0" pageNumber="27">Clypeus</emphasis>
|
||
with scattered white setae.
|
||
<emphasis id="378DDC714AAE6960C708D25600632A20" bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
|
||
brown with darker patches, with scattered white setae. Three promarginal and five retromarginal teeth.
|
||
<emphasis id="CA5C443ED30544702DDAC6AFF96374B2" bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
|
||
pale honey-coloured with various darker patches.
|
||
<emphasis id="5006294043831A9453456F04F399126F" bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
|
||
with central longitudinal pale chevroned band flanked by red scales.
|
||
<emphasis id="040B8B2C7AE2F226BC16F8106A1365CA" bold="true" italics="true" pageId="0" pageNumber="27">Epigyne</emphasis>
|
||
with large atria leading to broad ducts (Figs
|
||
<figureCitation id="CC5C879C71B7062599EB130858CD7B5E" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">191</figureCitation>
|
||
,
|
||
<figureCitation id="825D61B8A605E99DAD209D9A0978094C" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">192</figureCitation>
|
||
). Atria shallow laterally, declining rapidly at an "atrial cliff" (
|
||
<normalizedToken id="ACB756094F6D68AD0DD5BECC822AE13A" originalValue="“ac”">"ac"</normalizedToken>
|
||
, Fig.
|
||
<figureCitation id="D9075FFB8FDE552F1CC7DE575168A10C" captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">191</figureCitation>
|
||
) medial to which the surface is distinctly deeper. The ECP is apparently medial and small.
|
||
</paragraph>
|
||
<caption id="CDCC0D7E78693EC788C3A9A36AA53A8B" doi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27" start="Figures 188–201" startId="F21">
|
||
<paragraph id="70106FD6FE4497AA5710ABAE4C616BB0" pageId="0" pageNumber="27">
|
||
<emphasis id="A60175BEA4115E983C40027BE896A116" bold="true" pageId="0" pageNumber="27">Figures 188-201.</emphasis>
|
||
<taxonomicName id="DEC7B72180A01EC4C7C6BF3D7C1EB979" class="Magnoliopsida" family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus draconis" order="Apiales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="draconis">
|
||
<emphasis id="B694103DF51B23BA60EC123143A11869" italics="true" pageId="0" pageNumber="27">Maripanthus draconis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="2BA1D81A7CB62A78F9A6F0B600F506EB" bold="true" pageId="0" pageNumber="27">188</emphasis>
|
||
male left palp, prolateral view (holotype AS19.2232)
|
||
<emphasis id="FA0F41A9972A7D9663DDCD4821B55630" bold="true" pageId="0" pageNumber="27">189</emphasis>
|
||
same, ventral view
|
||
<emphasis id="5D290678D0E1C91FB88E7D3333492E57" bold="true" pageId="0" pageNumber="27">190</emphasis>
|
||
same, retrolateral view
|
||
<emphasis id="BF392C4083CB07E6AB2ADCBF79A14687" bold="true" pageId="0" pageNumber="27">191</emphasis>
|
||
epigyne, ventral (specimen AS19.2250)
|
||
<emphasis id="D266E8420F3C842559F8C705622CD6E4" bold="true" pageId="0" pageNumber="27">192</emphasis>
|
||
vulva, dorsal
|
||
<emphasis id="C0D5CB39CDC3AE833D543D268B7778C6" bold="true" pageId="0" pageNumber="27">193-197</emphasis>
|
||
holotype
|
||
<emphasis id="F632A3B393DB1771A8355A8CD1083E64" bold="true" pageId="0" pageNumber="27">198-201</emphasis>
|
||
female AS19.2250. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="85DEF85F5464C9F0AD33554595A3E7A8" pageId="0" pageNumber="27" type="natural history">
|
||
<paragraph id="B30DF0F174185E12A189D2DDF62E6517" pageId="0" pageNumber="27">Natural history.</paragraph>
|
||
<paragraph id="AA84E1A9896DD8AD65E7CB3BC7E04CED" pageId="0" pageNumber="27">In Singapore, beating vegetation in forest understory.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FD272AFC29FD51B12FF0C76447581DF" pageId="0" pageNumber="27" type="materials_examined">
|
||
<paragraph id="9A2D89C0EFBC850DA62233AD0F930E2B" pageId="0" pageNumber="27">Additional material examined.</paragraph>
|
||
<paragraph id="9F4CB1DF3B08C9AFD0649836D09620DF" pageId="0" pageNumber="27">
|
||
In UBCZ. Singapore: Bukit Timah Nature Reserve, stream at Jungle Falls Path.
|
||
<geoCoordinate id="00BCB193463D0C0C15E3ADB4B457EE8B" degrees="1.3562" direction="north" orientation="latitude" precision="5" value="1.3562">1.3562°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="80898B3CC8597A7790692026991E6BCB" degrees="103.7748" direction="east" orientation="longitude" precision="5" value="103.7748">103.7748°E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="58B56483906A2E70871FBDAEA87A2AF1" degrees="1.3572" direction="north" orientation="latitude" precision="5" value="1.3572">1.3572°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8E01DE8FFAEB2E5084666D61BBD6EB19" degrees="103.7734" direction="east" orientation="longitude" precision="5" value="103.7734">103.7734°E</geoCoordinate>
|
||
, 110-150 m elev. 12 June 2019 Maddison, Morehouse, & Marathe WPM#19-051 (one female); Bukit Timah Nature Reserve.
|
||
<geoCoordinate id="5B8ACF084A64DE4B54ADD6FF922F463A" degrees="1.355" direction="north" orientation="latitude" precision="55" value="1.355">1.355°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="86D2060AC27B13B1AFEC6A8A9A1B414A" degrees="103.78" direction="east" orientation="longitude" precision="555" value="103.78">103.78°E</geoCoordinate>
|
||
, 29 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-041 (1 female 1 juvenile). Malaysia: Johor: Gunung Belumut Recreational Forest.
|
||
<geoCoordinate id="129DE0FEACFC31B537443ADA1AAB3696" degrees="2.066" direction="north" orientation="latitude" precision="55" value="2.066">2.066°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A6B2E2D1F0C23DE41CE8B14FB0439F00" degrees="103.527" direction="east" orientation="longitude" precision="55" value="103.527">103.527°E</geoCoordinate>
|
||
, 60-100 m el. 24 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-038 (1 male, 1 female, 6 juveniles); near Kluang, Gunung Lambak.
|
||
<geoCoordinate id="E5005F472EB1D17A92F109DFE2BA69FA" degrees="2.025" direction="north" orientation="latitude" precision="55" value="2.025">2.025°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CEA18464ED3C6C376BBF5846324BE77B" degrees="103.344" direction="east" orientation="longitude" precision="55" value="103.344">103.344°E</geoCoordinate>
|
||
, 50-100 m el. 25 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-039 (1 male). Pahang: Tanah Rata. Jungle Trail 9 from Robinson Falls.
|
||
<geoCoordinate id="4DC4B2203719F4692D0390015935C24D" degrees="4.46" direction="north" orientation="latitude" precision="555" value="4.46">4.46°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F3871DE3362F1F4934A9AA59A8112C5D" degrees="101.40" direction="east" orientation="longitude" precision="555" value="101.4">101.40°E</geoCoordinate>
|
||
, 1200-1500 m el. 21-22 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-035 (1 female). Selangor: Ulu Gombak Field Station,
|
||
<geoCoordinate id="FB27348BBFCF13E0751B8EBB53372A01" degrees="3.325" direction="north" orientation="latitude" precision="55" value="3.325">3.325°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AE360AE17BC3DA0EBCB7D3053FC8DC57" degrees="101.753" direction="east" orientation="longitude" precision="55" value="101.753">101.753°E</geoCoordinate>
|
||
, 250 m el. 16-19 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-026 (1 female 1 juvenile); canyon near Ulu Gombak,
|
||
<geoCoordinate id="57A471CF4BE20E7E725FC29E5F1AC622" degrees="3.325" direction="north" orientation="latitude" precision="55" value="3.325">3.325°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0DCFF0C7DE0BE37E2F785C71EFB949DC" degrees="101.765" direction="east" orientation="longitude" precision="55" value="101.765">101.765°E</geoCoordinate>
|
||
, 275 m el. 17 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-027 (1 female 1 juvenile). Sarawak: Fairy Caves, near Kuching, 1.381-2°N, 110.117-9°E, 20 m el. 10 March 2012 Maddison/Piascik/Ang/Lee WPM#12-011 (1 female). The female from Sarawak is listed with some hesitation. It may be conspecific with a male
|
||
<taxonomicName id="307D1F32D70209A261A4853D2B54EF91" authorityName="Maddison" authorityYear="2020" class="Magnoliopsida" family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus" order="Apiales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="A5174C3924C438AF78CE8D70EEC649DC" italics="true" pageId="0" pageNumber="27">Maripanthus</emphasis>
|
||
</taxonomicName>
|
||
from Brunei, which appears to be a closely related but distinct species, with embolus initially directed distinctly more to the dorsal, and slight different carapace markings (specimen JK 08.08.23.0004, in LKCNHM, from Brunei: Ulu Temburong National Park, Ashton Trail
|
||
<geoCoordinate id="D8CDD6CCDBF7C5E172D78C0D99503769" degrees="4.5428" direction="north" orientation="latitude" precision="5" value="4.5428">4.5428°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4EE6E6B89A54C986C61F6CB3D71A11A3" degrees="115.1528" direction="east" orientation="longitude" precision="5" value="115.1528">115.1528°E</geoCoordinate>
|
||
J K H Koh 23 August 2008).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |