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<document id="DB8313A87EB86B3C4B5EF8C2EDA88B61" ID-CLB-Dataset="25961" ID-DOI="10.11646/zootaxa.4658.1.1" ID-GBIF-Dataset="31d6e081-95c3-4086-a45a-c11555dda3aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373040" ID-ZooBank="ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566364910789" checkinUser="plazi" docAuthor="Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed &amp; Barbero, Enrico" docDate="2019" docId="8861532B885E182FFF7608C2FE66C334" docLanguage="en" docName="zootaxa.4658.1.1.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus (Furconthophagus) frontalis Raffray 1877" docType="treatment" docVersion="12" lastPageNumber="14" masterDocId="74582B5388531822FFE10957FFA3C74B" masterDocTitle="An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="14" updateTime="1698751869822" updateUser="ExternalLinkService">
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<mods:title id="3DB73C4AFFDC73C71CCE84CEA2E1CF08">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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<mods:namePart id="75CDC3847E0809177A6AA6B89C4816E5">Ziani, Stefano</mods:namePart>
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<treatment id="8861532B885E182FFF7608C2FE66C334" ID-DOI="http://doi.org/10.5281/zenodo.5667073" ID-GBIF-Taxon="159217226" ID-Zenodo-Dep="5667073" LSID="urn:lsid:plazi:treatment:8861532B885E182FFF7608C2FE66C334" httpUri="http://treatment.plazi.org/id/8861532B885E182FFF7608C2FE66C334" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="48D2B1B6885E182FFF7608C2FCF3C6E4" box="[151,848,405,432]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="0077E23D885E182FFF7608C2FCF3C6E4" blockId="13.[151,848,405,467]" box="[151,848,405,432]" pageId="13" pageNumber="14">
<heading id="5B3F5551885E182FFF7608C2FCF3C6E4" bold="true" box="[151,848,405,432]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="C7C899BE885E182FFF7608C2FCF3C6E4" ID-CoL="74MKP" authority="Raffray, 1877" authorityName="Raffray" authorityYear="1877" box="[151,848,405,432]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frontalis" subGenus="Furconthophagus">
<emphasis id="32BC3E2F885E182FFF7608C2FCF3C6E4" bold="true" box="[151,848,405,432]" pageId="13" pageNumber="14">
<emphasis id="32BC3E2F885E182FFF7608C2FE9FC6FB" bold="true" box="[151,316,405,432]" italics="true" pageId="13" pageNumber="14">Onthophagus</emphasis>
(
<emphasis id="32BC3E2F885E182FFEAC08C2FD84C6FB" bold="true" box="[333,551,405,432]" italics="true" pageId="13" pageNumber="14">Furconthophagus</emphasis>
)
<emphasis id="32BC3E2F885E182FFDD808C2FD03C6FB" bold="true" box="[569,672,405,432]" italics="true" pageId="13" pageNumber="14">frontalis</emphasis>
Raffray, 1877
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="48D2B1B6885E182FFF7608EEFF4FC698" box="[151,236,440,467]" pageId="13" pageNumber="14" type="description">
<paragraph id="0077E23D885E182FFF7608EEFF4FC698" blockId="13.[151,848,405,467]" box="[151,236,440,467]" pageId="13" pageNumber="14">
[
<figureCitation id="98F3FEB8885E182FFF7F08EEFF47C698" box="[158,228,440,467]" captionStart="FIGURES 813" captionStartId="18.[151,264,1861,1886]" captionTargetBox="[315,1265,200,1801]" captionTargetId="figure@18.[280,1306,166,1842]" captionTargetPageId="18" captionText="FIGURES 813. Onthophagini of Arabian Peninsula. 8, Onthophagus (Furconthophagus) aethiopicus dOrbigny, 1902: YE (Sanaa) Alharf; 9, Onthophagus (Furconthophagus) frontalis Raffray, 1877: YE (Sanaa) Wadi Dhahr; 10, Onthophagus (Furconthophagus) rugulipennis Fairmaire, 1887: YE (Sanaa) Wadi Dhahar; 11, Onthophagus (Furconthophagus) sellatus Klug, 1845: SA (Riyadh) Deirab; 12, Onthophagus (Furconthophagus) variegatus (Fabricius, 1798): YE (Sanaa) Wadi Dhahr; 13, Onthophagus (Indonthophagus) nitidulus Klug, 1856: OM (Dhofar) Salalah. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373047/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
]
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B6885E182FFF760B57FE66C334" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="0077E23D885E182FFF760B57FE19C550" blockId="13.[151,1437,512,1151]" box="[151,442,512,539]" pageId="13" pageNumber="14">
<emphasis id="32BC3E2F885E182FFF760B57FECBC550" bold="true" box="[151,360,512,539]" pageId="13" pageNumber="14">Literature quotes</emphasis>
. None.
</paragraph>
<paragraph id="0077E23D885E182FFF260B73FE6AC528" blockId="13.[151,1437,512,1151]" pageId="13" pageNumber="14">
<emphasis id="32BC3E2F885E182FFF260B73FE0BC574" bold="true" box="[199,424,548,575]" pageId="13" pageNumber="14">Material examined</emphasis>
.
<materialsCitation id="B0A0E860885E182FFE540B72FE65C528" ID-GBIF-Occurrence="2350486362" collectingDate="2003-11-17" collectionCode="NMPC" collectorName="D. Kral" country="Yemen" elevation="2240" location="Wadi Dhahr" pageId="13" pageNumber="14" specimenCount="1" stateProvince="San'a'">
<collectingCountry id="78DFA2AD885E182FFE540B72FDA2C577" box="[437,513,549,575]" name="Yemen" pageId="13" pageNumber="14">
<emphasis id="32BC3E2F885E182FFE540B72FDA2C577" bold="true" box="[437,513,549,575]" pageId="13" pageNumber="14">
<subScript id="9C4CE078885E182FFE540B72FDA2C577" attach="left" box="[437,513,549,575]" fontSize="7" pageId="13" pageNumber="14">YEMEN</subScript>
</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF885E182FFDEF0B72FD4CC574" box="[526,751,549,575]" country="Yemen" name="San'a'" pageId="13" pageNumber="14">Sanaa Governorate</collectingRegion>
,
<location id="0517B4E6885E182FFD180B72FC22C574" LSID="urn:lsid:plazi:treatment:8861532B885E182FFF7608C2FE66C334:0517B4E6885E182FFD180B72FC22C574" box="[761,897,549,575]" country="Yemen" name="Wadi Dhahr" pageId="13" pageNumber="14" stateProvince="San'a'">Wadi Dhahr</location>
,
<quantity id="C7304FD8885E182FFC6B0B72FC7DC574" box="[906,990,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="13" pageNumber="14" unit="m" value="2240.0">
<elevation id="8BE5050E885E182FFC6B0B72FC7DC574" box="[906,990,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="13" pageNumber="14" unit="m" value="2240.0">2240 m</elevation>
</quantity>
(152643″N 446734″E), 15/
<date id="7476C4FD885E182FFAFE0B73FA34C574" box="[1311,1431,548,575]" pageId="13" pageNumber="14" value="2003-11-17">
<collectingDate id="64323D15885E182FFAFE0B73FA34C574" box="[1311,1431,548,575]" pageId="13" pageNumber="14" value="2003-11-17">17.xi.2003</collectingDate>
</date>
,
<collectorName id="AD3D87EB885E182FFF760B1EFF48C528" box="[151,235,585,611]" pageId="13" pageNumber="14">D. Král</collectorName>
leg.
<specimenCount id="16CE29B4885E182FFEC30B1EFEFAC528" box="[290,345,585,611]" pageId="13" pageNumber="14" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885E182FFE890B1EFE1DC528" box="[360,446,585,611]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="14">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0077E23D885E182FFF260B3BFAA7C5CC" blockId="13.[151,1437,512,1151]" box="[199,1284,620,647]" pageId="13" pageNumber="14">
<emphasis id="32BC3E2F885E182FFF260B3BFEFAC5CC" bold="true" box="[199,345,620,647]" pageId="13" pageNumber="14">Distribution</emphasis>
.
<emphasis id="32BC3E2F885E182FFE860B3BFE10C5CC" bold="true" box="[359,435,620,647]" pageId="13" pageNumber="14">Africa</emphasis>
:
<collectingCountry id="78DFA2AD885E182FFE230B3AFDB2C5CC" box="[450,529,621,647]" name="Eritrea" pageId="13" pageNumber="14">Eritrea</collectingCountry>
,
<collectingCountry id="78DFA2AD885E182FFDFC0B3AFDD9C5CC" box="[541,634,621,647]" name="Djibouti" pageId="13" pageNumber="14">Djibouti</collectingCountry>
(dOrbigny 1913).
<collectingCountry id="78DFA2AD885E182FFCB40B3AFC12C5CC" box="[853,945,621,647]" name="Somalia" pageId="13" pageNumber="14">Somalia</collectingCountry>
(
<bibRefCitation id="64599FCC885E182FFC5E0B3AFB54C5CC" author="Carpaneto, G. M. &amp; Piattella, E." box="[959,1271,621,647]" pageId="13" pageNumber="14" pagination="265 - 292" refId="ref37075" refString="Carpaneto, G. M. &amp; Piattella, E. (1990) Analisi zoogeografica preliminare dei Coleotteri Scarabeidi della Somalia (Coleoptera, Scarabaeidae s. str.). Biogeographia, 14 [1988], 265 - 292. https: // doi. org / 10.21426 / B 61411034 7" type="journal article" year="1990">Carpaneto &amp; Piattella 1990</bibRefCitation>
).
</paragraph>
<paragraph id="0077E23D885E182FFF260BC6FAD1C358" blockId="13.[151,1437,512,1151]" pageId="13" pageNumber="14">
<emphasis id="32BC3E2F885E182FFF260BC6FE97C5E0" bold="true" box="[199,308,657,683]" pageId="13" pageNumber="14">Remarks</emphasis>
. First record for the Arabian Peninsula, notably
<collectingCountry id="78DFA2AD885E182FFC830BC6FC17C5E0" box="[866,948,657,683]" name="Yemen" pageId="13" pageNumber="14">Yemen</collectingCountry>
. The sole specimen examined, a male, has an apical reddish yellow spot on 3
<superScript id="F7BD4F75885E182FFDC00BE3FD8CC589" attach="none" box="[545,559,692,706]" fontSize="6" pageId="13" pageNumber="14">rd</superScript>
/4
<superScript id="F7BD4F75885E182FFDA40BE3FDF1C589" attach="left" box="[581,594,692,706]" fontSize="6" pageId="13" pageNumber="14">th</superScript>
elytral interstria. DOrbigny (1904) writes: “Dans mon Synopsis des Onthophagides paléarctiques et dans le Supplément (in LAbeille, XXIX, p. 158 et 294), jai confondu cette espèce [
<taxonomicName id="C7C899BE885E182FFF400BAAFDE0C45C" authority="Raffray, 1877" authorityName="Raffray" authorityYear="1877" box="[161,579,764,791]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="32BC3E2F885E182FFF400BAAFE3EC45C" box="[161,413,765,791]" italics="true" pageId="13" pageNumber="14">Onthophagus frontalis</emphasis>
Raffray, 1877
</taxonomicName>
] avec
<taxonomicName id="C7C899BE885E182FFD6D0BAAFC93C45C" authority="Fabr." authorityName="Fabr." box="[652,816,765,791]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="32BC3E2F885E182FFD6D0BAAFD4DC45C" box="[652,750,765,791]" italics="true" pageId="13" pageNumber="14">parvulus</emphasis>
Fabr.
</taxonomicName>
qui lui ressemble beaucoup et est décrit de lInde. Les exemplaires provenant du Sind: Kurrachee (Maindron) sont bien des parvulus, mais ceux que jai signalés dObock (Maindron, coll. Muséum de Paris), et de
<collectingCountry id="78DFA2AD885E182FFD8F0A12FD68C414" box="[622,715,837,863]" name="Djibouti" pageId="13" pageNumber="14">Djibouti</collectingCountry>
(H. Coutière, id.), sont des
<taxonomicName id="C7C899BE885E182FFBE50A12FBC1C414" authorityName="Raffray" authorityYear="1877" box="[1028,1122,837,863]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="32BC3E2F885E182FFBE50A12FBC1C414" box="[1028,1122,837,863]" italics="true" pageId="13" pageNumber="14">frontalis</emphasis>
</taxonomicName>
. Le
<taxonomicName id="C7C899BE885E182FFB740A12FB54C414" box="[1173,1271,837,863]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="32BC3E2F885E182FFB740A12FB54C414" box="[1173,1271,837,863]" italics="true" pageId="13" pageNumber="14">parvulus</emphasis>
</taxonomicName>
est donc rayer de la liste des espèces africaines” [In my “Synopsis des Onthophagides paléarctiques and in the “Supplementum (in LAbeille,XXIX, p. 158 et 294), I confounded this species [
<taxonomicName id="C7C899BE885E182FFCB70ADAFB56C4EC" authority="Raffray, 1877" authorityName="Raffray" authorityYear="1877" box="[854,1269,908,935]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="32BC3E2F885E182FFCB70ADAFBF1C4EC" box="[854,1106,909,935]" italics="true" pageId="13" pageNumber="14">Onthophagus frontalis</emphasis>
Raffray, 1877
</taxonomicName>
] with
<taxonomicName id="C7C899BE885E182FFADB0ADAFF72C480" authority="Fabr." authorityName="Fabr." class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="32BC3E2F885E182FFADB0ADAFA3FC4EC" box="[1338,1436,909,935]" italics="true" pageId="13" pageNumber="14">parvulus</emphasis>
Fabr.
</taxonomicName>
which is very similar to it and is described from
<collectingCountry id="78DFA2AD885E182FFD060AE6FC87C480" box="[743,804,945,971]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
. Specimens from
<collectingRegion id="C20C2CDF885E182FFC080AE6FB82C480" box="[1001,1057,945,971]" country="Pakistan" name="Sindh" pageId="13" pageNumber="14">Sind</collectingRegion>
: Kurrachee (Maindron) [
<collectingCountry id="78DFA2AD885E182FFAD00AE7FA35C480" box="[1329,1430,944,971]" name="Pakistan" pageId="13" pageNumber="14">Pakistan</collectingCountry>
] are indeed
<taxonomicName id="C7C899BE885E182FFEF20A82FED6C4A4" box="[275,373,981,1007]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="32BC3E2F885E182FFEF20A82FED6C4A4" box="[275,373,981,1007]" italics="true" pageId="13" pageNumber="14">parvulus</emphasis>
</taxonomicName>
, whereas the ones I have recorded from
<collectingRegion id="C20C2CDF885E182FFCDE0A82FC29C4A4" box="[831,906,981,1007]" country="Djibouti" name="Obock" pageId="13" pageNumber="14">Obock</collectingRegion>
(Maindron, Paris Museum collection) and
<collectingCountry id="78DFA2AD885E182FFA8E0A82FF72C358" name="Djibouti" pageId="13" pageNumber="14">Djibouti</collectingCountry>
(H. Coutiér, id.) are
<taxonomicName id="C7C899BE885E182FFE5B0AAEFDBBC358" authorityName="Raffray" authorityYear="1877" box="[442,536,1017,1043]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="32BC3E2F885E182FFE5B0AAEFDBBC358" box="[442,536,1017,1043]" italics="true" pageId="13" pageNumber="14">frontalis</emphasis>
</taxonomicName>
.
<taxonomicName id="C7C899BE885E182FFDC70AAEFD08C358" box="[550,683,1017,1043]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="32BC3E2F885E182FFDC70AAEFD08C358" box="[550,683,1017,1043]" italics="true" pageId="13" pageNumber="14">O. parvulus</emphasis>
</taxonomicName>
is therefore to be removed from the list of the African species].
</paragraph>
<paragraph id="0077E23D885E182FFF260D4AFE66C334" blockId="13.[151,1437,512,1151]" pageId="13" pageNumber="14">
After having examined the aedeagus and the internal sac, specifically the lamella copulatrix, of
<taxonomicName id="C7C899BE885E182FFB0F0D4AFACDC37C" authorityName="Raffray" authorityYear="1877" box="[1262,1390,1053,1079]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="32BC3E2F885E182FFB0F0D4AFACDC37C" box="[1262,1390,1053,1079]" italics="true" pageId="13" pageNumber="14">O. frontalis</emphasis>
</taxonomicName>
and compared them with those of
<taxonomicName id="C7C899BE885E182FFE070D16FDC7C310" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[486,612,1089,1115]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="furcatus">
<emphasis id="32BC3E2F885E182FFE070D16FDC7C310" box="[486,612,1089,1115]" italics="true" pageId="13" pageNumber="14">O. furcatus</emphasis>
</taxonomicName>
, the
<typeStatus id="DF735C9F885E182FFD7D0D16FD6FC310" box="[668,716,1089,1115]" pageId="13" pageNumber="14">type</typeStatus>
species of
<taxonomicName id="C7C899BE885E182FFCAB0D16FBB3C310" authorityName="Zunino" authorityYear="1979" box="[842,1040,1089,1115]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="31" phylum="Arthropoda" rank="subGenus" subGenus="Furconthophagus">
<emphasis id="32BC3E2F885E182FFCAB0D16FBB3C310" box="[842,1040,1089,1115]" italics="true" pageId="13" pageNumber="14">Furconthophagus</emphasis>
</taxonomicName>
, we believe that the species should be placed in this subgenus.
</paragraph>
</subSubSection>
</treatment>
</document>