222 lines
25 KiB
XML
222 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.352.6294" ID-GBIF-Dataset="8de8ecaa-9c3f-45ca-820a-c9d7f93dd74d" ID-PMC="PMC3837397" ID-Pensoft-Pub="1313-2970-352-35" ID-PubMed="24294091" ID-ZBK="946C3E2D95DE4FD78613F46158017CB9" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-352-35" ModsDocOrigin="ZooKeys 352" ModsDocTitle="On five species of the tribes Abacetini and Pterostichini (Coleoptera, Carabidae)" checkinTime="1451246665382" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav V." docDate="2013" docId="219FB507732B14DF946772EE24F4E889" docLanguage="en" docName="ZooKeys 352: 35-50" docOrigin="ZooKeys 352" docSource="http://dx.doi.org/10.3897/zookeys.352.6294" docTitle="Rhytiferonia beroni Gueorguiev, 2013, sp. n." docType="treatment" docUuid="9DAC8D91-E7AE-4C20-BACB-A703133F19C7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="44" masterDocId="0E5D7D12FF8C417CFF86FFEBDF75BC16" masterDocTitle="On five species of the tribes Abacetini and Pterostichini (Coleoptera, Carabidae)" masterLastPageNumber="50" masterPageNumber="35" pageNumber="40" updateTime="1668156885337" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>On five species of the tribes Abacetini and Pterostichini (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gueorguiev, Borislav V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>352</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>35</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.352.6294</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.352.6294</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-352-35</mods:identifier>
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<mods:identifier type="ZBK">946C3E2D95DE4FD78613F46158017CB9</mods:identifier>
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<mods:identifier type="ZooBank">946C3E2D95DE4FD78613F46158017CB9</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152049641" LSID="urn:lsid:zoobank.org:act:9DAC8D91-E7AE-4C20-BACB-A703133F19C7" httpUri="http://treatment.plazi.org/id/219FB507732B14DF946772EE24F4E889" lastPageId="9" lastPageNumber="44" pageId="5" pageNumber="40">
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<subSubSection pageId="5" pageNumber="40" type="nomenclature">
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<paragraph pageId="5" pageNumber="40">
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<taxonomicName LSID="http://zoobank.org/9DAC8D91-E7AE-4C20-BACB-A703133F19C7" class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="40">sp. n.</taxonomicNameLabel>
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Figs 10-14
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="type material">
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<paragraph pageId="5" pageNumber="40">Type material.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
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Holotype
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<pageBreakToken pageId="6" pageNumber="41" start="start">♀</pageBreakToken>
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, "New Guinea, Bonforok bil, Tifalmin, 1600 m 17.10.75 leg. P. Beron" [typeset], "HOLOTYPE
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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spec. nov.
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<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
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des. 2012" [typeset, red label] (NMNHS). Paratypes 2♀♀, labeled as follows: 1♀, "Tifalmin W. Sepic Prov. IX.75" [handwritten], "British Speleological Expedition to Papua New Guinea 1975" [typeset], "
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
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sp. det. B. P. Moore
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<normalizedToken originalValue="’78”">'78"</normalizedToken>
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[handwritten & typeset] (NMNHS); 1♀, "Fimin tel 2300 m Western Prov. 30.viii.75 P. Beron" [handwritten], "British Speleological Expedition to Papua New Guinea 1975" [typeset], "
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
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sp. det. B. P. Moore
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<normalizedToken originalValue="’78”">'78"</normalizedToken>
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[handwritten & typeset], "UC Berkeley EMEC 345536" [typeset]; paratypes with subsequently added: "PARATYPE
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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spec. nov.
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<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
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des. 2012" [typeset, red label] (EMEC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="41" type="examined type material of other species">
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<paragraph pageId="6" pageNumber="41">Examined type material of other species.</paragraph>
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<paragraph pageId="6" pageNumber="41">
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia julianae" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="julianae">Rhytiferonia julianae</taxonomicName>
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Baehr, 2001, paratypes 2♂♂, "IRIAN JAYA: Mt. Juliana Gebiet 16.-17.9.1993 Sab-me Tal" [typeset], "ca.
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="140.28334">140°17'E</geoCoordinate>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-4.45">04°27'S</geoCoordinate>
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, 3400-3500m leg. M. Balke (14)" [typeset], "PARATYPE Rhytiferonia julianae, sp.nov. det. M. Baehr 2000" [typeset, red label] (NMW).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="41" type="diagnosis">
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<paragraph pageId="6" pageNumber="41">Diagnosis.</paragraph>
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<paragraph pageId="6" pageNumber="41">The new species is distinct from all other congeners in the following set of characters: 1) eyes moderately enclosed by temporae laterally; 2) pronotum with obtuse, somewhat perceptible basal angles, side without sinuation in front of angle; 3) posterolateral seta of pronotum slightly removed from angle; 4) elytra with parascutellar setigerous puncture, distinct parascutellar stria in interval 1, and angular base of stria 1 joining stria 2; 5) last three abdominal sterna with transverse sulci superficial, only laterally distinct.</paragraph>
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<paragraph pageId="6" pageNumber="41">
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It should be noted that the above diagnosis is based on the descriptions of the known species by
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr (2001)</bibRefCitation>
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as specimens of only
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
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species were used for comparison (see chapter "Examined type material of other species").
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42" type="description">
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<paragraph pageId="7" pageNumber="42">
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<pageBreakToken pageId="7" pageNumber="42" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="8" pageNumber="43">
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<pageBreakToken pageId="8" pageNumber="43" start="start">Habitus</pageBreakToken>
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. Moderately large-sized species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
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, with elongate, convex body and basal angles of pronotum rounded off (Fig. 10). Measurements. BL: 15.6-17.2 mm (17.1 mm in holotype); BW: 4.9-5.5 mm (5.5 mm in holotype). Ratios. PW/HW: 1.38-1.42 (1.42 in holotype); PW/PL: 1.10-1.16 (1.16 in holotype); PW/PbW: 1.35-1.37 (1.37 in holotype); PbW/PaW: 0.98-1.00 (0.98 in holotype); EW/PW: 1.20-1.23 (1.20 in holotype); EL/EW: 1.62-1.68 (1.62 in holotype). Color. Deep black on dorsal surface, mouthparts, antennae, legs, and ventral surface black brown to dark reddish. Microsculpture and lustre. Very fine, isodiametric, distinct on head and elytra, visible under magnification> 50 x, indistinct on pronotum; dorsal surface shiny. Head. Longer than wide, disc smooth, frontal furrows faintly impressed, oblique, divergent backward, hardly reach level of anterior supraorbital punctures; eyes small, modestly projecting laterally, as long as temporae; temporae slightly surpassing eyes laterally; paraorbital sulci moderately deep, reaching level of posterior supraorbital pore backward; labrum rectangular, anterior margin with six setigerous punctures; clypeus trapezoid, slightly emarginated anteriorly and laterally, with two setigerous punctures closer to lateral margins than to anterior one, clypeal suture faint; antennae filiform, pubescent from second fifth of segment 4, with terminal article not reaching base of pronotum; glossal sclerite of ligula with two long setae on anterior margin; maxillary palpomere 1 very massive, twice thicker than following two segments; mentum deeply emarginated, with tooth bifid at tip and pair of labial setae, epilobes large, significantly exceeding mentum tooth forward; submentum with two basal setae, without lateral ones. Pronotum (Fig. 11). Large, widest at middle, disc gently convex, smooth; midline very fine, distinct on medial half of pronotum, obsolescent apically and basally; anterior and posterior margins unbordered, almost of equal length, anterior margin slightly concave, fore angles moderately protruding forward; basal margin rather convex backward, hind angles subangular, incompletely round; lateral margins slightly convex to straight, without sinuation towards hind angles, marginal field narrow in apical half, widened and explanate towards base; anterolateral seta at apical third, posterolateral seta slightly in front of hind angles; posterolateral impressions faint. Elytra (Figs 12-13). Subelongate, oviform, convex dorsally, coalescent along suture, widest at third fourth; sides narrow basally, gradually widened apically; shoulders angulate, with minute teeth; lateral margins slightly sinuate before rounded apex; striae well impressed, complete, impunctate, parascutellar striae present, not anastomosing with stria 1 back, angular base of stria 1 present, joining stria 2 (in paratype) or reduced (in holotype), scutellar pores present, removed back from basal margin with distance from one to three diameters of pore, situated on angular base of stria 1, stria 7 deepened in apical third, with one setigerous puncture near to apex, discal setigerous punctures absent; intervals gently convex; umbilicate series of elytra entire, not interrupted in middle, consist of 19-21 setigerous punctures. Hind wings. Vestigial. Ventral surface (thorax and abdomen). Prosternum laterally with conspicuous longitudinal sulci; prosternal process unbordered; metaepisterna wider than long, with anterior margins longer than inner ones and as long as outer margins; apical three abdominal sternites with transverse sulci superficial, distinct laterally, indistinct in middle; last visible sternite quadrisetose in female. Legs. Fore and middle legs relatively short and massive, hind legs longer; fore and middle trochanteri with one seta, hind trochanteri asetose, with pointed apex, as long as half of hind femora; fore coxae asetose, middle coxae with two setae, hind coxae with three setae, including medial setae after meeting point of coxae; profemur posterior margin with four pores, mesofemur posterior margin with five pores, metafemur anterior margin with three pores; onychium setose ventrally. Female genitalia. Not studied.
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</paragraph>
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<caption pageId="8" pageNumber="43">
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<paragraph pageId="8" pageNumber="43">
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Figure 10.
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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sp. n., holotype.
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</paragraph>
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</caption>
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<caption pageId="8" pageNumber="43">
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<paragraph pageId="8" pageNumber="43">
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Figures 11-13.
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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sp. n. 11 pronotum, holotype 12-13 base of elytra (12 holotype 13 paratype). Scale line: 3 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="43" type="etymology">
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<paragraph pageId="8" pageNumber="43">Etymology.</paragraph>
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<paragraph pageId="8" pageNumber="43">A noun in the genitive case. Honour Dr. Petar Beron, a prominent Bulgarian zoologist, who first collected the new species.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="44" type="distribution">
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<paragraph pageId="9" pageNumber="44">
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<pageBreakToken pageId="9" pageNumber="44" start="start">Distribution</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="44">
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Papua New Guinea, Sandaun Province (= West Sepik Province), Telefomin District, Tifalmin env. (Fig. 14). For the time being, it is the first documented representative of
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
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Darlington, 1962 from Papua New Guinea.
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</paragraph>
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<caption pageId="9" pageNumber="44">
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<paragraph pageId="9" pageNumber="44">
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Figure 14. Localities of
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
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(grey circle) and
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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sp. n. (black circle).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="44" type="affinities">
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<paragraph pageId="9" pageNumber="44">Affinities.</paragraph>
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<paragraph pageId="9" pageNumber="44">
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The new species is provisionally placed in the nigra-group, which includes
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia nigra" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="nigra">Rhytiferonia nigra</taxonomicName>
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Darlington, 1962,
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia iebele" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="iebele">Rhytiferonia iebele</taxonomicName>
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Darlington, 1962, and
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
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Baehr, 2001.
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr (2001</bibRefCitation>
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: 43) associated the species from this complex due to the presence of: 1/ markedly enclosed eyes; 2/ basal angles of pronotum rounded off, without sinuation in front of them; 3/ posterolateral seta of pronotum far removed from angle; 4/ complete, deep, sharply impressed transverse sulci on three apical abdominal sterna; 5/ median lobe of aedeagus narrow, elongate, little curved medially, without spiniform sclerites near apex of internal sac. As the male genital characters of the new species are unknown, it possesses only two from the remaining four features: markedly enclosed eyes (i) and basal angles of pronotum rounded off, without sinuation in front of angles (ii). In contrast to that,
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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sp. n. possesses posterolateral seta of pronotum only slightly removed from basal angle and apical three abdominal sternites with superficial, distinct only laterally transverse sulci.
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</paragraph>
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<paragraph pageId="9" pageNumber="44">
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Except for the characters shared by the new species with the species from the nigra group,
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia beroni" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="beroni">Rhytiferonia beroni</taxonomicName>
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sp. n. most resembles
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
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in: 1) the pronotum with obtuse basal angles; 2) presence of parascutellar pore; 3) presence of parascutellar stria, which not anastomose with stria 1, and angular base of stria 1 joining stria 2. This set of shared traits places the last two species closer to each other than any of them to another species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
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. The new species can be distinguished from its closest congener in the following row:
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</paragraph>
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<paragraph pageId="9" pageNumber="44">
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1) temporae (
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<normalizedToken originalValue="“orbits”">"orbits"</normalizedToken>
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, after
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
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) laterally slightly surpassing eyes, vs. laterally perceptibly surpassing eyes in
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
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(
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
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: 45, 55, fig. 16);
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</paragraph>
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<paragraph pageId="9" pageNumber="44">
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2) pronotum widest in the middle, with fore angles moderately protruding (Fig. 11), vs. widest in anterior third, with fore angles little produced in
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
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(
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
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: 45, 54, fig. 9);
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</paragraph>
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<paragraph pageId="9" pageNumber="44">
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3) anterior and posterior margins of pronotum almost of equal length, PbW/PaW: 0.98, vs. "Base clearly narrower than apex." (
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<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
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: 45) in
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<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
|
||
;
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="44">
|
||
4) posterolateral seta of pronotum slightly removed from hind angle (Fig. 11), vs. far removed from hind angle in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
|
||
: 45, 54, fig. 9);
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="44">
|
||
5) transverse sulci on three apical abdominal sternites superficially impressed laterally, indistinct in middle, vs. complete and deep, sharply impressed in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
|
||
: 45);
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="44">
|
||
6) umbilicate series of 19-21 setigerous punctures, vs. umbilicate series of 18 setigerous punctures in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
|
||
: 45).
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="44">
|
||
In addition, several ratios with different values in the two species (
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia punctigera" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="punctigera">Rhytiferonia punctigera</taxonomicName>
|
||
in brackets): PW/HW: 1.38-1.42 (vs. 1.27); PW/PL: 1.10-1.16 (vs. 1.07); PW/PbW: 1.35-1.37 (vs. data questionable, 1.24, according to
|
||
<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
|
||
: 46, but 1.32, according to
|
||
<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="12" pageNumber="47" pagination="41 - 55" title="The genus Rhytiferonia Darlington in New Guinea (Carabidae: Pterostichinae)." volume="5" year="2001">Baehr 2001</bibRefCitation>
|
||
: 52); EL/EW: 1.62-1.68 (vs. 1.79).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |