291 lines
37 KiB
XML
291 lines
37 KiB
XML
<document id="ACE8805E9479EB770A0E7A325FB22379" ID-DOI="10.11646/zootaxa.5032.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5495554" ID-ZooBank="65894101-2D60-426D-B149-4CF6E1C561BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631115156823" checkinUser="plazi" docAuthor="Enuschenko, Ilya V. & Shavrin, Alexey V." docDate="2021" docId="0C1987ABFFE3297EFF7EA464B38EF853" docLanguage="en" docName="zootaxa.5032.3.2.pdf" docOrigin="Zootaxa 5032 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyrophaena (Gyrophaena) furcata" docType="treatment" docVersion="3" lastPageNumber="336" masterDocId="F020FFD3FFE6297BFFE9A646B131FFA5" masterDocTitle="On some species of Gyrophaenina Kraatz 1856 of Sri Lanka and India (Coleoptera: Staphylinidae: Aleocharinae: Homalotini)" masterLastPageNumber="356" masterPageNumber="331" pageNumber="336" updateTime="1699405676542" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6D6A978DFBF998635F29C1E1336107B8">On some species of Gyrophaenina Kraatz 1856 of Sri Lanka and India (Coleoptera: Staphylinidae: Aleocharinae: Homalotini)</mods:title>
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<mods:namePart id="8529BB4CEAEBA6AF78028A3DE416B9D4">Enuschenko, Ilya V.</mods:namePart>
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<mods:affiliation id="943F1CE2868B52E97E7FAC368B21C7E0">Limnological Institute SB RAS, Ulan-Batorskaya str., 3, Irkutsk, 664033, Russia.</mods:affiliation>
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<mods:namePart id="736E49F3EC3BBEFD37C77CBC2B50FB03">Shavrin, Alexey V.</mods:namePart>
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<mods:affiliation id="65A3C23D99472C97501BB315EDB39346">Institute of Life Sciences and Technologies, Daugavpils Universit, Vienibas 13, Daugavpils, LV- 5401, Latvia. ashavrin @ hotmail. com</mods:affiliation>
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<mods:title id="F9EF55B0EA8AE42B38941E0D33B535B8">Zootaxa</mods:title>
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<mods:date id="F04EBE28D290764BF4CF70B04FF75006">2021</mods:date>
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<paragraph id="840F36BDFFE3297EFF7EA464B20EFD99" blockId="5.[151,831,546,609]" box="[151,831,546,573]" pageId="5" pageNumber="336">
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<heading id="DF4781D1FFE3297EFF7EA464B20EFD99" bold="true" box="[151,831,546,573]" fontSize="11" level="1" pageId="5" pageNumber="336" reason="1">
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<taxonomicName id="43B04D3EFFE3297EFF7EA464B20EFD99" ID-CoL="8Y463" authority="(Motschulsky, 1858)" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[151,831,546,573]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcata" subGenus="Gyrophaena">
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<emphasis id="B6C4EAAFFFE3297EFF7EA464B01CFD98" bold="true" box="[151,301,546,573]" italics="true" pageId="5" pageNumber="336">Gyrophaena</emphasis>
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(
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<emphasis id="B6C4EAAFFFE3297EFED7A464B0E5FD98" bold="true" box="[318,468,546,573]" italics="true" pageId="5" pageNumber="336">Gyrophaena</emphasis>
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)
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<emphasis id="B6C4EAAFFFE3297EFE0CA464B30CFD99" bold="true" box="[485,573,546,572]" italics="true" pageId="5" pageNumber="336">furcata</emphasis>
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(
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<bibRefCitation id="E0214B4CFFE3297EFDA5A464B206FD99" author="Motschulsky, V. I." box="[588,823,546,572]" pageId="5" pageNumber="336" pagination="204 - 264" refId="ref14608" refString="Motschulsky, V. I. (1858) Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. Bulletin de la Societe. Imperiale des Naturalistes de Moscou, 31 (3), 204 - 264." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
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)
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</emphasis>
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<subSubSection id="CCAA6536FFE3297EFF7EA401B016FDC4" box="[151,295,583,609]" pageId="5" pageNumber="336" type="description">
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<paragraph id="840F36BDFFE3297EFF7EA401B016FDC4" blockId="5.[151,831,546,609]" box="[151,295,583,609]" pageId="5" pageNumber="336">
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(
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<figureCitation id="1C8B2A38FFE3297EFF76A401B011FDC4" box="[159,288,583,609]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1135,1159]" captionTargetBox="[168,1427,194,1098]" captionTargetId="figure-24@6.[151,1436,181,1112]" captionTargetPageId="6" captionText="FIGURES 17–21. Gyrophaena furcata (holotype): 17–habitus, 18–aedeagus, lateral view, 19–male abdominal tergite VIII, dorsal view, 20–male abdominal sternite VIII, ventral view, 21–labels of the holotype. Scale bars: 1.0 mm (Fig. 17), 0.10 mm (Figs 18–20)." figureDoi="http://doi.org/10.5281/zenodo.5495636" httpUri="https://zenodo.org/record/5495636/files/figure.png" pageId="5" pageNumber="336">Figs 17–20</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="CCAA6536FFE3297EFF7EA4CDB56EFD67" pageId="5" pageNumber="336" type="reference_group">
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<paragraph id="840F36BDFFE3297EFF7EA4CDB261FD06" blockId="5.[151,1119,651,706]" box="[151,848,651,675]" pageId="5" pageNumber="336">
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<treatmentCitation id="051110ACFFE3297EFF7EA4CDB352FD06" author="Motschulsky, V. I." box="[151,611,651,675]" page="227" pageId="5" pageNumber="336" year="1858">
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<taxonomicName id="43B04D3EFFE3297EFF7EA4CDB352FD06" authority="Motschulsky, 1858: 227" authorityName="Motschulsky" authorityPageNumber="227" authorityYear="1858" box="[151,611,651,675]" class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcatus">
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<emphasis id="B6C4EAAFFFE3297EFF7EA4CDB057FD06" box="[151,358,651,675]" italics="true" pageId="5" pageNumber="336">Encephalus furcatus</emphasis>
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<bibRefCitation id="E0214B4CFFE3297EFE85A4CDB352FD06" author="Motschulsky, V. I." box="[364,611,651,675]" pageId="5" pageNumber="336" pagination="204 - 264" refId="ref14608" refString="Motschulsky, V. I. (1858) Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. Bulletin de la Societe. Imperiale des Naturalistes de Moscou, 31 (3), 204 - 264." type="journal article" year="1858">Motschulsky, 1858: 227</bibRefCitation>
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;
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<treatmentCitation id="051110ACFFE3297EFD84A4CDB261FD06" author="Motschulsky, V. I." box="[621,848,651,675]" page="51" pageId="5" pageNumber="336" year="1859">
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<bibRefCitation id="E0214B4CFFE3297EFD84A4CDB261FD06" author="Motschulsky, V. I." box="[621,848,651,675]" pageId="5" pageNumber="336" pagination="25 - 118" refId="ref14648" refString="Motschulsky, V. I. (1859) Insectes des Indes orientales et de contrees analogues. Etudes Entomologiques, 8, 25 - 118." type="journal article" year="1859">Motschulsky 1859: 51</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="840F36BDFFE3297EFF7EA4ECB56EFD67" blockId="5.[151,1119,651,706]" box="[151,1119,682,706]" pageId="5" pageNumber="336">
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<treatmentCitationGroup id="A4A01193FFE3297EFF7EA4ECB56EFD67" box="[151,1119,682,706]" pageId="5" pageNumber="336">
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<treatmentCitation id="051110ACFFE3297EFF7EA4ECB217FD67" author="Bernhauer, M. & Scheerpeltz, O." box="[151,806,682,706]" page="530" pageId="5" pageNumber="336" year="1926">
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<taxonomicName id="43B04D3EFFE3297EFF7EA4ECB217FD67" ID-CoL="8Y463" authority=": Bernhauer & Scheerpeltz 1926: 530" authorityName="Bernhauer & Scheerpeltz" authorityPageNumber="530" authorityYear="1926" box="[151,806,682,706]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcata">
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<emphasis id="B6C4EAAFFFE3297EFF7EA4ECB024FD67" box="[151,277,682,706]" italics="true" pageId="5" pageNumber="336">Gyrophaena</emphasis>
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(s.str.)
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||
<emphasis id="B6C4EAAFFFE3297EFE88A4ECB098FD67" box="[353,425,682,706]" italics="true" pageId="5" pageNumber="336">furcata</emphasis>
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:
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<bibRefCitation id="E0214B4CFFE3297EFE5FA4ECB217FD67" author="Bernhauer, M. & Scheerpeltz, O." box="[438,806,682,706]" pageId="5" pageNumber="336" pagination="499 - 988" refId="ref13956" refString="Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. (Pars 82). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Volumen 5. Staphylinidae. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer & Scheerpeltz 1926: 530</bibRefCitation>
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;
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<treatmentCitation id="051110ACFFE3297EFCD9A4ECB2CDFD67" author="Cameron, M." box="[816,1020,682,706]" page="219" pageId="5" pageNumber="336" year="1933">
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<bibRefCitation id="E0214B4CFFE3297EFCD9A4ECB2CDFD67" author="Cameron, M." box="[816,1020,682,706]" pageId="5" pageNumber="336" pagination="219 - 220" refId="ref14132" refString="Cameron, M. (1933) Remarks on some of Motschoulsky's types of Staphylinidae (Col.). The Entomologist ' sMonthly Magazine, 69, 219 - 220." type="journal article" year="1933">Cameron 1933: 219</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="051110ACFFE3297EFBEFA4ECB56EFD67" author="Cameron, M." box="[1030,1119,682,706]" page="89" pageId="5" pageNumber="336" year="1939">
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<bibRefCitation id="E0214B4CFFE3297EFBEFA4ECB56EFD67" author="Cameron, M." box="[1030,1119,682,706]" pageId="5" pageNumber="336" pagination="1 - 3" refId="ref14209" refString="Cameron, M. (1939) Coleoptera, Staphylinidae. Vol. IV. Parts I & II. In: Sewell, R. B. S. (Ed.), The Fauna of British India, including Ceylon and Burma. Taylor & Francis, London, pp. i - xviii + 1 - 691, pls. 1 - 3." type="book chapter" year="1939">1939: 89</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCAA6536FFE3297EFF7EA4B6B0ABFCD3" pageId="5" pageNumber="336" type="materials_examined">
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<paragraph id="840F36BDFFE3297EFF7EA4B6B0ABFCD3" blockId="5.[151,1437,752,2039]" pageId="5" pageNumber="336">
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<emphasis id="B6C4EAAFFFE3297EFF7EA4B6B086FCAF" bold="true" box="[151,439,752,778]" pageId="5" pageNumber="336">Type material examined</emphasis>
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:
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<typeStatus id="5B0B881FFFE3297EFE2CA4B7B304FCAE" box="[453,565,753,779]" pageId="5" pageNumber="336" type="lectotype">Lectotype</typeStatus>
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(
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<emphasis id="B6C4EAAFFFE3297EFDACA4B6B231FCAF" bold="true" box="[581,768,752,778]" pageId="5" pageNumber="336">here designated</emphasis>
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) of
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<taxonomicName id="43B04D3EFFE3297EFCC4A4B6B5DBFCAE" authority="Motschulsky 1859" authorityName="Motschulsky" authorityYear="1859" box="[813,1258,752,779]" class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcatus">
|
||
<emphasis id="B6C4EAAFFFE3297EFCC4A4B6B520FCAF" box="[813,1041,752,778]" italics="true" pageId="5" pageNumber="336">Encephalus furcatus</emphasis>
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<bibRefCitation id="E0214B4CFFE3297EFBF1A4B6B5DBFCAE" author="Motschulsky, V. I." box="[1048,1258,752,779]" pageId="5" pageNumber="336" pagination="25 - 118" refId="ref14648" refString="Motschulsky, V. I. (1859) Insectes des Indes orientales et de contrees analogues. Etudes Entomologiques, 8, 25 - 118." type="journal article" year="1859">Motschulsky 1859</bibRefCitation>
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</taxonomicName>
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||
,
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||
<specimenCount id="92B6FD34FFE3297EFB1DA4B6B42FFCAE" box="[1268,1310,752,779]" pageId="5" pageNumber="336" type="male">1 ♂</specimenCount>
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[dissected; specimen without left antennomeres 9–11 and left protarsus; labels as in
|
||
<figureCitation id="1C8B2A38FFE3297EFC3AA552B519FC8B" box="[979,1064,787,814]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1135,1159]" captionTargetBox="[168,1427,194,1098]" captionTargetId="figure-24@6.[151,1436,181,1112]" captionTargetPageId="6" captionText="FIGURES 17–21. Gyrophaena furcata (holotype): 17–habitus, 18–aedeagus, lateral view, 19–male abdominal tergite VIII, dorsal view, 20–male abdominal sternite VIII, ventral view, 21–labels of the holotype. Scale bars: 1.0 mm (Fig. 17), 0.10 mm (Figs 18–20)." figureDoi="http://doi.org/10.5281/zenodo.5495636" httpUri="https://zenodo.org/record/5495636/files/figure.png" pageId="5" pageNumber="336">Fig. 21</figureCitation>
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||
]: ‘
|
||
<taxonomicName id="43B04D3EFFE3297EFBAFA552B5F8FC8B" authorityName="Kirby" authorityYear="1832" box="[1094,1225,788,814]" class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B6C4EAAFFFE3297EFBAFA552B5F8FC8B" box="[1094,1225,788,814]" italics="true" pageId="5" pageNumber="336">Encephalus</emphasis>
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</taxonomicName>
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|
|
||
<taxonomicName id="43B04D3EFFE3297EFB37A552B1F5FCF7" authority="Motsch. Ind." authorityName="Motsch. Ind." class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcatus">
|
||
<emphasis id="B6C4EAAFFFE3297EFB37A552B408FC8B" box="[1246,1337,788,814]" italics="true" pageId="5" pageNumber="336">furcatus</emphasis>
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||
Motsch. Ind.
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||
</taxonomicName>
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||
or.’, ‘Type’, [small round yellow label], ‘
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||
<typeStatus id="5B0B881FFFE3297EFD7AA57EB209FCF7" box="[659,824,824,850]" pageId="5" pageNumber="336" type="lectotype">LECTOTYPE</typeStatus>
|
||
<taxonomicName id="43B04D3EFFE3297EFCA9A57EB1CFFCD3" authority="Motschulsky, 1858 Enushchenko I. V. 2019" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcata">
|
||
<emphasis id="B6C4EAAFFFE3297EFCA9A57EB513FCF7" box="[832,1058,824,850]" italics="true" pageId="5" pageNumber="336">Gyrophaena furcata</emphasis>
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<bibRefCitation id="E0214B4CFFE3297EFBC0A57EB5CFFCF6" author="Motschulsky, V. I." box="[1065,1278,824,851]" pageId="5" pageNumber="336" pagination="204 - 264" refId="ref14608" refString="Motschulsky, V. I. (1858) Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. Bulletin de la Societe. Imperiale des Naturalistes de Moscou, 31 (3), 204 - 264." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
|
||
Enushchenko I.V. 2019
|
||
</taxonomicName>
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||
det.’ (
|
||
<collectionCode id="E2A1AE78FFE3297EFEACA51AB0BFFCD3" box="[325,398,860,886]" pageId="5" pageNumber="336">ZMM</collectionCode>
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).
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</paragraph>
|
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</subSubSection>
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<subSubSection id="CCAA6536FFE3297EFF2EA5C6B0B8F9CE" pageId="5" pageNumber="336" type="description">
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<paragraph id="840F36BDFFE3297EFF2EA5C6B011FA83" blockId="5.[151,1437,752,2039]" pageId="5" pageNumber="336">
|
||
<emphasis id="B6C4EAAFFFE3297EFF2EA5C6B05CFC3F" bold="true" box="[199,365,896,922]" pageId="5" pageNumber="336">Redescription</emphasis>
|
||
. Body length
|
||
<quantity id="43489B58FFE3297EFDE7A5C6B344FC3E" box="[526,629,896,923]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="336" unit="mm" value="2.05">2.05 mm</quantity>
|
||
. Body wide, robust (
|
||
<figureCitation id="1C8B2A38FFE3297EFCB6A5C6B282FC3F" box="[863,947,896,922]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1135,1159]" captionTargetBox="[168,1427,194,1098]" captionTargetId="figure-24@6.[151,1436,181,1112]" captionTargetPageId="6" captionText="FIGURES 17–21. Gyrophaena furcata (holotype): 17–habitus, 18–aedeagus, lateral view, 19–male abdominal tergite VIII, dorsal view, 20–male abdominal sternite VIII, ventral view, 21–labels of the holotype. Scale bars: 1.0 mm (Fig. 17), 0.10 mm (Figs 18–20)." figureDoi="http://doi.org/10.5281/zenodo.5495636" httpUri="https://zenodo.org/record/5495636/files/figure.png" pageId="5" pageNumber="336">Fig. 17</figureCitation>
|
||
); surface of forebody shiny, without reticulate microsculpture. Head and pronotum black; elytra and abdomen red-brown (apical portion of elytra blackish); antennomeres 5–11 dark brown; mouthparts, antennomeres 1–4 and legs yellow-brown. Head 1.5 times as wide as long, with two moderately large punctures in middle and a few distinct, sparse and fine punctures behind eyes; vertex impunctate. Basal antennomere elliptic, twice as long as wide, antennomere 2 1.6 times as long as wide, 3 slightly longer than 2, 1.4 times as long as wide, 4 transverse, as long as wide, 5–9 distinctly transverse, twice as wide as long, 10 1.2–1.3 times as wide as long, apical antennomere oblong, 1.4 times as long as wide, about twice as long as two preceding antennomeres. Pronotum strongly convex, transverse, 1.5 times as wide as long, with two median rows consisting of three large and deep punctures; basal portion of pronotum without punctation. Elytra slightly longer than pronotum, 1.6 times as wide as long, surface covered with moderately large tubercles (ca.
|
||
<quantity id="43489B58FFE3297EFA82A282B1F2FAA6" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="336" unit="mm" value="0.03">0.03 mm</quantity>
|
||
diam.), denser on shoulders, along suture and posterior margins. Abdomen almost parallel-sided, slightly tapering apically.
|
||
</paragraph>
|
||
<paragraph id="840F36BDFFE3297EFF2EA376B42EFA13" blockId="5.[151,1437,752,2039]" pageId="5" pageNumber="336">
|
||
Male. Median lobe long, narrow, with subacute apex (
|
||
<figureCitation id="1C8B2A38FFE3297EFCF0A376B25DFAEF" box="[793,876,1328,1354]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1135,1159]" captionTargetBox="[168,1427,194,1098]" captionTargetId="figure-24@6.[151,1436,181,1112]" captionTargetPageId="6" captionText="FIGURES 17–21. Gyrophaena furcata (holotype): 17–habitus, 18–aedeagus, lateral view, 19–male abdominal tergite VIII, dorsal view, 20–male abdominal sternite VIII, ventral view, 21–labels of the holotype. Scale bars: 1.0 mm (Fig. 17), 0.10 mm (Figs 18–20)." figureDoi="http://doi.org/10.5281/zenodo.5495636" httpUri="https://zenodo.org/record/5495636/files/figure.png" pageId="5" pageNumber="336">Fig. 18</figureCitation>
|
||
); apical projection of internal sac wider and longer than median lobe, with very long, spirally folded flagellum. Abdominal tergite VII with lateral irregular rows of small, oblong tubercles and very wide oval impression. Apical margin of abdominal tergite VIII with very deep and moderately wide excision (
|
||
<figureCitation id="1C8B2A38FFE3297EFE2FA3DAB32BFA13" box="[454,538,1436,1462]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1135,1159]" captionTargetBox="[168,1427,194,1098]" captionTargetId="figure-24@6.[151,1436,181,1112]" captionTargetPageId="6" captionText="FIGURES 17–21. Gyrophaena furcata (holotype): 17–habitus, 18–aedeagus, lateral view, 19–male abdominal tergite VIII, dorsal view, 20–male abdominal sternite VIII, ventral view, 21–labels of the holotype. Scale bars: 1.0 mm (Fig. 17), 0.10 mm (Figs 18–20)." figureDoi="http://doi.org/10.5281/zenodo.5495636" httpUri="https://zenodo.org/record/5495636/files/figure.png" pageId="5" pageNumber="336">Fig. 19</figureCitation>
|
||
). Apical margin of abdominal sternite VIII widely rounded (
|
||
<figureCitation id="1C8B2A38FFE3297EFB57A3DAB423FA13" box="[1214,1298,1436,1462]" captionStart="FIGURES 17–21" captionStartId="6.[151,264,1135,1159]" captionTargetBox="[168,1427,194,1098]" captionTargetId="figure-24@6.[151,1436,181,1112]" captionTargetPageId="6" captionText="FIGURES 17–21. Gyrophaena furcata (holotype): 17–habitus, 18–aedeagus, lateral view, 19–male abdominal tergite VIII, dorsal view, 20–male abdominal sternite VIII, ventral view, 21–labels of the holotype. Scale bars: 1.0 mm (Fig. 17), 0.10 mm (Figs 18–20)." figureDoi="http://doi.org/10.5281/zenodo.5495636" httpUri="https://zenodo.org/record/5495636/files/figure.png" pageId="5" pageNumber="336">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="840F36BDFFE3297EFF2EA386B0A0FA7F" blockId="5.[151,1437,752,2039]" box="[199,401,1472,1498]" pageId="5" pageNumber="336">Female unknown.</paragraph>
|
||
<paragraph id="840F36BDFFE3297EFF2EA3A2B0B8F9CE" blockId="5.[151,1437,752,2039]" pageId="5" pageNumber="336">
|
||
<emphasis id="B6C4EAAFFFE3297EFF2EA3A2B09DFA5B" bold="true" box="[199,428,1508,1534]" pageId="5" pageNumber="336">Comparative notes</emphasis>
|
||
. Regarding the absence of microsculpture on the pronotum and the elytra, and general shape of the apical portion of the aedeagus,
|
||
<taxonomicName id="43B04D3EFFE3297EFD6EA04EB3CBF987" box="[647,762,1544,1571]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcata">
|
||
<emphasis id="B6C4EAAFFFE3297EFD6EA04EB3AAF987" box="[647,667,1544,1570]" italics="true" pageId="5" pageNumber="336">G</emphasis>
|
||
.
|
||
<emphasis id="B6C4EAAFFFE3297EFD43A04EB3CBF987" box="[682,762,1544,1570]" italics="true" pageId="5" pageNumber="336">furcata</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="43B04D3EFFE3297EFC67A04EB5FCF986" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[910,1229,1544,1571]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="livida">
|
||
<emphasis id="B6C4EAAFFFE3297EFC67A04EB2C0F987" box="[910,1009,1544,1570]" italics="true" pageId="5" pageNumber="336">G. livida</emphasis>
|
||
<bibRefCitation id="E0214B4CFFE3297EFC11A04EB5FCF986" author="Motschulsky, V. I." box="[1016,1229,1544,1571]" pageId="5" pageNumber="336" pagination="204 - 264" refId="ref14608" refString="Motschulsky, V. I. (1858) Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. Bulletin de la Societe. Imperiale des Naturalistes de Moscou, 31 (3), 204 - 264." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
(see below), from which it can be distinguished by its significantly larger and wider body and different morphology of the aedeagus and male tergite VIII.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CCAA6536FFE3297EFF2EA032B27CF92B" box="[199,845,1652,1678]" pageId="5" pageNumber="336" type="distribution">
|
||
<paragraph id="840F36BDFFE3297EFF2EA032B27CF92B" blockId="5.[151,1437,752,2039]" box="[199,845,1652,1678]" pageId="5" pageNumber="336">
|
||
<emphasis id="B6C4EAAFFFE3297EFF2EA032B050F92B" bold="true" box="[199,353,1652,1678]" pageId="5" pageNumber="336">Distribution.</emphasis>
|
||
<collectingCountry id="FCA7762DFFE3297EFE8EA032B090F92B" box="[359,417,1652,1678]" name="India" pageId="5" pageNumber="336">India</collectingCountry>
|
||
orientale;
|
||
<collectingCountry id="FCA7762DFFE3297EFDF0A032B3B7F92B" box="[537,646,1652,1678]" name="Sri Lanka" pageId="5" pageNumber="336">Sri Lanka</collectingCountry>
|
||
(
|
||
<bibRefCitation id="E0214B4CFFE3297EFD7CA032B271F92B" author="Cameron, M." box="[661,832,1652,1678]" pageId="5" pageNumber="336" pagination="1 - 3" refId="ref14209" refString="Cameron, M. (1939) Coleoptera, Staphylinidae. Vol. IV. Parts I & II. In: Sewell, R. B. S. (Ed.), The Fauna of British India, including Ceylon and Burma. Taylor & Francis, London, pp. i - xviii + 1 - 691, pls. 1 - 3." type="book chapter" year="1939">Cameron 1939</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CCAA6536FFE3297EFF2EA0DEB38EF853" pageId="5" pageNumber="336" type="discussion">
|
||
<paragraph id="840F36BDFFE3297EFF2EA0DEB1C1F8C3" blockId="5.[151,1437,752,2039]" pageId="5" pageNumber="336">
|
||
<emphasis id="B6C4EAAFFFE3297EFF2EA0DEB005F917" bold="true" box="[199,308,1688,1714]" pageId="5" pageNumber="336">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="43B04D3EFFE3297EFEA8A0DEB315F917" box="[321,548,1688,1714]" class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcatus">
|
||
<emphasis id="B6C4EAAFFFE3297EFEA8A0DEB315F917" box="[321,548,1688,1714]" italics="true" pageId="5" pageNumber="336">Encephalus furcatus</emphasis>
|
||
</taxonomicName>
|
||
was originally described from “Indes orientales”.A male from ZMM was designated as the
|
||
<typeStatus id="5B0B881FFFE3297EFEC8A0FAB0B9F973" box="[289,392,1724,1750]" pageId="5" pageNumber="336" type="lectotype">lectotype</typeStatus>
|
||
in order to fix the identity of the name.
|
||
<bibRefCitation id="E0214B4CFFE3297EFCACA0FAB580F972" author="Bernhauer, M. & Scheerpeltz, O." box="[837,1201,1724,1751]" pageId="5" pageNumber="336" pagination="499 - 988" refId="ref13956" refString="Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. (Pars 82). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Volumen 5. Staphylinidae. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer & Scheerpeltz (1926)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="E0214B4CFFE3297EFB0EA0FAB4A6F973" author="Cameron, M." box="[1255,1431,1724,1750]" pageId="5" pageNumber="336" pagination="219 - 220" refId="ref14132" refString="Cameron, M. (1933) Remarks on some of Motschoulsky's types of Staphylinidae (Col.). The Entomologist ' sMonthly Magazine, 69, 219 - 220." type="journal article" year="1933">Cameron (1933</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E0214B4CFFE3297EFF7EA0A6B1E1F95F" author="Cameron, M." box="[151,208,1760,1786]" pageId="5" pageNumber="336" pagination="1 - 3" refId="ref14209" refString="Cameron, M. (1939) Coleoptera, Staphylinidae. Vol. IV. Parts I & II. In: Sewell, R. B. S. (Ed.), The Fauna of British India, including Ceylon and Burma. Taylor & Francis, London, pp. i - xviii + 1 - 691, pls. 1 - 3." type="book chapter" year="1939">1939</bibRefCitation>
|
||
) cited
|
||
<taxonomicName id="43B04D3EFFE3297EFECAA0A6B33AF95F" box="[291,523,1760,1786]" class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcatus">
|
||
<emphasis id="B6C4EAAFFFE3297EFECAA0A6B33AF95F" box="[291,523,1760,1786]" italics="true" pageId="5" pageNumber="336">Encephalus furcatus</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="43B04D3EFFE3297EFDDFA0A6B3CDF95F" box="[566,764,1760,1786]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcata">
|
||
<emphasis id="B6C4EAAFFFE3297EFDDFA0A6B363F95F" box="[566,594,1760,1786]" italics="true" pageId="5" pageNumber="336">G.</emphasis>
|
||
(s.str.)
|
||
<emphasis id="B6C4EAAFFFE3297EFD45A0A6B3CDF95F" box="[684,764,1760,1786]" italics="true" pageId="5" pageNumber="336">furcata</emphasis>
|
||
</taxonomicName>
|
||
. Among synonyms of
|
||
<taxonomicName id="43B04D3EFFE3297EFBEFA0A6B54DF95F" box="[1030,1148,1760,1786]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcata">
|
||
<emphasis id="B6C4EAAFFFE3297EFBEFA0A6B54DF95F" box="[1030,1148,1760,1786]" italics="true" pageId="5" pageNumber="336">G. furcata</emphasis>
|
||
</taxonomicName>
|
||
they listed
|
||
<taxonomicName id="43B04D3EFFE3297EFAEEA0A6B018F8BA" authority="Kraatz, 1859" authorityName="Kraatz" authorityYear="1859" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="B6C4EAAFFFE3297EFAEEA0A6B4ADF95F" box="[1287,1436,1760,1786]" italics="true" pageId="5" pageNumber="336">G. humeralis</emphasis>
|
||
<bibRefCitation id="E0214B4CFFE3297EFF7EA143B018F8BA" author="Kraatz, G." box="[151,297,1796,1823]" pageId="5" pageNumber="336" pagination="1 - 193" refId="ref14558" refString="Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25 (1), 1 - 193, pls. 1 - 3. https: // doi. org / 10.5962 / bhl. title. 66002" type="journal article" year="1859">Kraatz, 1859</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="43B04D3EFFE3297EFE8BA142B39CF8BA" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[354,685,1796,1823]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="indica">
|
||
<emphasis id="B6C4EAAFFFE3297EFE8BA142B0FAF8BB" box="[354,459,1796,1822]" italics="true" pageId="5" pageNumber="336">G. indica</emphasis>
|
||
<bibRefCitation id="E0214B4CFFE3297EFE3DA142B39CF8BA" author="Motschulsky, V. I." box="[468,685,1796,1823]" pageId="5" pageNumber="336" pagination="204 - 264" refId="ref14608" refString="Motschulsky, V. I. (1858) Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. Bulletin de la Societe. Imperiale des Naturalistes de Moscou, 31 (3), 204 - 264." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
. The senior author studied types of both
|
||
<taxonomicName id="43B04D3EFFE3297EFB96A143B5CBF8BB" box="[1151,1274,1796,1823]" class="Insecta" family="Staphylinidae" genus="Encephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="furcatus">
|
||
<emphasis id="B6C4EAAFFFE3297EFB96A143B5A1F8BB" box="[1151,1168,1797,1822]" italics="true" pageId="5" pageNumber="336">E</emphasis>
|
||
.
|
||
<emphasis id="B6C4EAAFFFE3297EFB76A142B5CBF8BB" box="[1183,1274,1796,1822]" italics="true" pageId="5" pageNumber="336">furcatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="43B04D3EFFE3297EFADAA142B4ADF8BB" authorityName="Motschulsky" authorityYear="1858" box="[1331,1436,1796,1823]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="indica">
|
||
<emphasis id="B6C4EAAFFFE3297EFADAA142B476F8BB" box="[1331,1351,1796,1822]" italics="true" pageId="5" pageNumber="336">G</emphasis>
|
||
.
|
||
<emphasis id="B6C4EAAFFFE3297EFABFA142B4ADF8BB" box="[1366,1436,1796,1822]" italics="true" pageId="5" pageNumber="336">indica</emphasis>
|
||
</taxonomicName>
|
||
deposited in ZMM, and based on external and internal morphological differences revealed that they are different species.
|
||
</paragraph>
|
||
<paragraph id="840F36BDFFE3297EFF2EA136B38EF853" blockId="5.[151,1437,752,2039]" pageId="5" pageNumber="336">
|
||
Based on the general shape of the aedeagus and some details of the arrangement of pronotal punctures and elytral tubercles,
|
||
<taxonomicName id="43B04D3EFFE3297EFEB3A1D2B332F80B" box="[346,515,1940,1967]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="331" phylum="Arthropoda" rank="species" species="furcata" subGenus="Gyrophaena">
|
||
<emphasis id="B6C4EAAFFFE3297EFEB3A1D2B05FF80B" box="[346,366,1940,1966]" italics="true" pageId="5" pageNumber="336">G</emphasis>
|
||
. (
|
||
<emphasis id="B6C4EAAFFFE3297EFE6EA1D2B0AAF80B" box="[391,411,1940,1966]" italics="true" pageId="5" pageNumber="336">G</emphasis>
|
||
.)
|
||
<emphasis id="B6C4EAAFFFE3297EFE5AA1D2B332F80B" box="[435,515,1940,1966]" italics="true" pageId="5" pageNumber="336">furcata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="43B04D3EFFE3297EFDD2A1D2B3E2F80B" authorityName="Motschulsky" authorityYear="1858" box="[571,723,1940,1967]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="331" phylum="Arthropoda" rank="species" species="livida" subGenus="Gyrophaena">
|
||
<emphasis id="B6C4EAAFFFE3297EFDD2A1D2B37EF80B" box="[571,591,1940,1966]" italics="true" pageId="5" pageNumber="336">G</emphasis>
|
||
. (
|
||
<emphasis id="B6C4EAAFFFE3297EFD8EA1D2B34AF80B" box="[615,635,1940,1966]" italics="true" pageId="5" pageNumber="336">G</emphasis>
|
||
.)
|
||
<emphasis id="B6C4EAAFFFE3297EFD7AA1D2B3E2F80B" box="[659,723,1940,1966]" italics="true" pageId="5" pageNumber="336">livida</emphasis>
|
||
</taxonomicName>
|
||
(see below) somewhat resemble some species of the American genus
|
||
<taxonomicName id="43B04D3EFFE3297EFF09A1FFB0D4F877" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[224,485,1976,2002]" class="Insecta" family="Staphylinidae" genus="Eumicrota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B6C4EAAFFFE3297EFF09A1FFB064F877" box="[224,341,1977,2002]" italics="true" pageId="5" pageNumber="336">Eumicrota</emphasis>
|
||
Casey, 1906
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="43B04D3EFFE3297EFDCCA1FFB261F877" authority="(Erichson, 1840)" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[549,848,1976,2002]" class="Insecta" family="Staphylinidae" genus="Eumicrota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="socia">
|
||
<emphasis id="B6C4EAAFFFE3297EFDCCA1FFB3BCF877" box="[549,653,1977,2002]" italics="true" pageId="5" pageNumber="336">Eu. socia</emphasis>
|
||
(Erichson, 1840)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="43B04D3EFFE3297EFC6EA1FFB581F876" authority="Seevers, 1951" authorityName="Seevers" authorityYear="1951" box="[903,1200,1976,2003]" class="Insecta" family="Staphylinidae" genus="Eumicrota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="spinosa">
|
||
<emphasis id="B6C4EAAFFFE3297EFC6EA1FFB53BF877" box="[903,1034,1977,2002]" italics="true" pageId="5" pageNumber="336">Eu. spinosa</emphasis>
|
||
<bibRefCitation id="E0214B4CFFE3297EFBF8A1FEB581F876" author="Seevers, C. H." box="[1041,1200,1976,2003]" pageId="5" pageNumber="336" pagination="1 - 762" refId="ref15634" refString="Seevers, C. H. (1951) A revision of the North American and European staphylinid beetles of the subtribe Gyrophaenina (Aleocharinae, Bolitocharini). Fieldiana Zoology, 32 (10), 1 - 762. https: // doi. org / 10.5962 / bhl. title. 2816" type="journal article" year="1951">Seevers, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
)), distributed in both American continents (
|
||
<bibRefCitation id="E0214B4CFFE3297EFE67A19AB33DF853" author="Ashe, J. S." box="[398,524,2012,2038]" pageId="5" pageNumber="336" pagination="129 - 349" refId="ref13828" refString="Ashe, J. S. (1984) Generic revision of the subtribe Gyrophaenina (Coleoptera: Staphylinidae: Aleocharinae) with a review of the described subgenera and major features of evolution. Quaestiones Entomologicae, 20, 129 - 349." type="journal article" year="1984">Ashe 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E0214B4CFFE3297EFDFEA19BB382F853" author="Newton, A. F." box="[535,691,2012,2039]" pageId="5" pageNumber="336" pagination="2000" refId="ref14740" refString="Newton, A. F. (2019) StaphBase: Staphyliniformia world catalog database (version Nov 2018): Staphylinoidea, Hydrophiloidea, Synteliidae. In: Roskov, Y., Ower, G., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, E., Decock, W., van Nieukerken, E., Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life. Species 2000: Naturalis, Leiden. Updated to 17 May 2021 by A. Newton (unpublished database). Availabe from: http: // www. catalogueoflife. org / col (accessed 18 May 2021)" type="book chapter" year="2019">Newton 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |