treatments-xml/data/03/BE/59/03BE5914116CFFE9E3DB60C3FA0E2C83.xml

<document id="57DB600BF88F38B2E1B2770B87C8C4D9" ID-CLB-Dataset="3788" ID-DOI="10.11646/zootaxa.5060.1.4" ID-GBIF-Dataset="0b735edc-d682-4b2e-9fb5-69de294147ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="5606771" ID-ZooBank="B96324E4-5651-472F-A443-31D9CF7109E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635406747450" checkinUser="plazi" docAuthor="Shy, Jhy-Yun, Shih, Hsi-Te &amp; Mao, Jean-Jay" docDate="2021" docId="03BE5914116CFFE9E3DB60C3FA0E2C83" docLanguage="en" docName="zootaxa.5060.1.4.pdf" docOrigin="Zootaxa 5060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geothelphusa boreas Shy &amp; Shih &amp; Mao 2021, sp. nov." docType="treatment" docUuid="6DC2B082-D052-49B3-B243-941D38306477" docUuidSource="ZooBank" docVersion="8" lastPageNumber="99" masterDocId="FF87216C116FFFEFE34C6304FF8A2D2E" masterDocTitle="Geothelphusa boreas, a new montane freshwater crab (Crustacea: Potamidae Geothelphusa) from northeastern Taiwan, and the identity of G. hirsuta Tan &amp; Liu, 1998" masterLastPageNumber="104" masterPageNumber="93" pageNumber="96" updateTime="1699467010177" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F4AC49E14C1350C64A94B56CB891872D">Geothelphusa boreas, a new montane freshwater crab (Crustacea: Potamidae Geothelphusa) from northeastern Taiwan, and the identity of G. hirsuta Tan &amp; Liu, 1998</mods:title>
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<mods:namePart id="E6ED21DDF6D92088DD22F2AC308DE8F9">Shy, Jhy-Yun</mods:namePart>
<mods:affiliation id="1955F930029CB6EBBED25F734278D13A">Department of Aquaculture, National Penghu University, 300 Liuhe Road, Magong City, Penghu 880, Taiwan.</mods:affiliation>
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<mods:namePart id="62BA6EACE8FA20E89491BA92AFF59B37">Shih, Hsi-Te</mods:namePart>
<mods:affiliation id="ACC4FFD4D3E0E7EEA003AC20728D54C8">Department of Life Science and Research Center for Global Change Biology, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<mods:namePart id="1B4C7D967592B9994450845F8EBF742B">Mao, Jean-Jay</mods:namePart>
<mods:affiliation id="1C871AA6D780D8755F170B42021020EC">Department of Forestry and Natural Resources, National Ilan University, 1, Sec. 1, Shen-Lung Road, Yilan 260, Taiwan.</mods:affiliation>
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<taxonomicName id="4C179381116CFFECE3DB60C3FE1C2ECC" ID-CoL="8RW7G" authority="Shy &amp; Shih &amp; Mao, 2021" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[151,406,967,994]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="boreas" status="sp. nov.">
<emphasis id="B9633410116CFFECE3DB60C3FE1C2ECC" bold="true" box="[151,406,967,994]" italics="true" pageId="3" pageNumber="96">Geothelphusa boreas</emphasis>
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<taxonomicNameLabel id="A250896B116CFFECE2D260CCFE712ECC" box="[414,507,968,994]" pageId="3" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BA8E802116CFFECE3DB60E8FE982929" blockId="3.[151,507,967,1031]" box="[151,274,1004,1031]" pageId="3" pageNumber="96">
(
<figureCitation id="132CF487116CFFECE3D360E8FE802929" box="[159,266,1004,1031]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1349,1373]" captionStartId-1="7.[151,250,1277,1301]" captionTargetBox-0="[279,1306,521,1284]" captionTargetBox-1="[172,1347,329,1244]" captionTargetId-0="figure-193@6.[166,1421,453,1329]" captionTargetId-1="graphics-93@7.[172,1075,333,1222]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Male right G1s of Geothelphusa takuan Shy, Yu &amp; Ng, 1994 (holotype, NTOU F10205; A, B) and G. hirsuta Tan &amp; Liu, 1998 (holotype, TMCD 3285; C, D). A, C, ventral view; B, D, dorsal view. Scale = 1 mm." captionText-1="FIGURE 5. A Bayesian Inference (BI) tree of Geothelphusa boreas sp. nov. and related montane species from northern Taiwan based on cytochrome oxidase I (COI) marker. Probability values at the nodes represent the support values for BI and maximum evolution (ME). For haplotype abbreviations, see Table 1. Corresponding numbers of localities in Fig. 1 are in parenthesis behind haplotype abbreviations." figureDoi-0="http://doi.org/10.5281/zenodo.5606781" figureDoi-1="http://doi.org/10.5281/zenodo.5606785" httpUri-0="https://zenodo.org/record/5606781/files/figure.png" httpUri-1="https://zenodo.org/record/5606785/files/figure.png" pageId="3" pageNumber="96">Figs. 4–5</figureCitation>
)
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urn:lsid:zoobank.org:act: 
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<taxonomicName id="4C179381116CFFECE3DB6750FED12943" box="[151,347,1108,1133]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B9633410116CFFECE3DB6750FEAE2942" box="[151,292,1108,1132]" italics="true" pageId="3" pageNumber="96">Geothelphusa</emphasis>
sp. 1
</taxonomicName>
. — Shih 
<emphasis id="B9633410116CFFECE2F76751FE622942" box="[443,488,1108,1132]" italics="true" pageId="3" pageNumber="96">et al</emphasis>
. 2011: 461.
</paragraph>
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<paragraph id="8BA8E802116CFFECE3DB679EFC4528C3" blockId="3.[151,1437,1178,2038]" pageId="3" pageNumber="96">
<emphasis id="B9633410116CFFECE3DB679EFEF0299A" bold="true" box="[151,378,1178,1204]" pageId="3" pageNumber="96">Material examined</emphasis>
. 
<materialsCitation id="3B7FE25F116CFFECE2C5679FFDE429D4" ID-GBIF-Occurrence="3395042310" collectingDate="1994-02-16" collectorName="H. - T. Shih" county="Hapen River" elevation="740" latitude="24.758333" location="Fushan Botanical Garden area" longLatPrecision="21" longitude="121.576385" municipality="River" pageId="3" pageNumber="96" specimenCount="1" specimenCount-male="1" stateProvince="New Taipei" typeStatus="holotype">
<emphasis id="B9633410116CFFECE2C5679FFE7C299B" bold="true" box="[393,502,1179,1205]" pageId="3" pageNumber="96">
<typeStatus id="54AC56A0116CFFECE2C5679FFE7C299B" box="[393,502,1179,1205]" pageId="3" pageNumber="96" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<specimenCount id="9D11238B116CFFECE149679FFDBA299B" box="[517,560,1179,1205]" pageId="3" pageNumber="96" type="male">1 ♂</specimenCount>
(21.2 × 
<quantity id="4CEF45E7116CFFECE1DD679FFD7C299B" box="[657,758,1179,1205]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="3" pageNumber="96" unit="mm" value="16.4">16.4 mm</quantity>
) (NCHUZOOL 13615), 
<collectingCounty id="62C9908E116CFFECE742679FFB14299B" box="[1038,1182,1179,1205]" pageId="3" pageNumber="96">Hapen River</collectingCounty>
, head branch of Danshuei (= Tansui) 
<collectingMunicipality id="6BCC7278116CFFECE20767B9FE0129F9" box="[331,395,1213,1239]" pageId="3" pageNumber="96">River</collectingMunicipality>
, the 
<location id="8EC8BED9116CFFECE28D67B9FC9329F6" LSID="urn:lsid:plazi:treatment:03BE5914116CFFE9E3DB60C3FA0E2C83:8EC8BED9116CFFECE28D67B9FC9329F6" box="[449,793,1213,1240]" county="Hapen River" latitude="24.758333" longLatPrecision="21" longitude="121.576385" municipality="River" name="Fushan Botanical Garden area" pageId="3" pageNumber="96" stateProvince="New Taipei">Fushan Botanical Garden area</location>
, 
<collectingRegion id="49D326E0116CFFECE06967BAFC5529F9" box="[805,991,1213,1240]" country="Taiwan" name="New Taipei" pageId="3" pageNumber="96">New Taipei City</collectingRegion>
(
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; 
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), elevation of 
<quantity id="4CEF45E7116CFFECE3F467E4FE8929D5" box="[184,259,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="96" unit="m" value="740.0">
<elevation id="003A0F31116CFFECE3F467E4FE8929D5" box="[184,259,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="96" unit="m" value="740.0">740 m</elevation>
</quantity>
, coll. 
<collectorName id="26E28DD4116CFFECE20767E5FE4F29D4" box="[331,453,1248,1275]" pageId="3" pageNumber="96">H.-T. Shih</collectorName>
, 
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<collectingDate id="EFED372A116CFFECE29F67E4FDE429D4" box="[467,622,1248,1274]" pageId="3" pageNumber="96" value="1994-02-16">16 Feb. 1994</collectingDate>
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. 
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<typeStatus id="54AC56A0116CFFECE13767E4FD7929D4" box="[635,755,1248,1274]" pageId="3" pageNumber="96" type="paratype">Paratypes</typeStatus>
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: 
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(15.7 × 12.4–19.0 × 
<quantity id="4CEF45E7116CFFECE77167E4FB2229D5" box="[1085,1192,1248,1275]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="3" pageNumber="96" unit="mm" value="14.7">14.7 mm</quantity>
), 
<specimenCount id="9D11238B116CFFECE7F167E4FB7429D5" box="[1213,1278,1248,1275]" pageId="3" pageNumber="96" type="female">2 ♀♀</specimenCount>
(15.3 × 12.1, 17.1 × 
<quantity id="4CEF45E7116CFFECE3AB6607FEC72833" box="[231,333,1283,1309]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="96" unit="mm" value="13.1">13.1 mm</quantity>
) (NCHUZOOL 13429), 
<materialsCitation id="3B7FE25F116CFFECE1286607FA1E2832" ID-GBIF-Occurrence="3395042301" box="[612,1428,1282,1310]" collectingDate="1994-02-16" collectionCode="ZRC" collectorName="Others" county="Hapen River" elevation="740" latitude="24.758333" location="Fushan Botanical Garden area" longLatPrecision="21" longitude="121.576385" municipality="River" pageId="3" pageNumber="96" specimenCode="ZRC 2021.0648" specimenCount="1" specimenCount-male="1" stateProvince="New Taipei" typeStatus="paratype">
<specimenCount id="9D11238B116CFFECE1286607FD042833" box="[612,654,1283,1309]" pageId="3" pageNumber="96" type="male">1 ♂</specimenCount>
(16.2 × 
<quantity id="4CEF45E7116CFFECE1A36607FCDF2833" box="[751,853,1283,1309]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="3" pageNumber="96" unit="mm" value="13.2">13.2 mm</quantity>
) (
<specimenCode id="DBB14079116CFFECE0206607FBA22830" box="[876,1064,1283,1310]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="96">ZRC 2021.0648</specimenCode>
); same data as holotype. 
<emphasis id="B9633410116CFFECE60E6606FA1E2832" bold="true" box="[1346,1428,1282,1308]" pageId="3" pageNumber="96">
<collectorName id="26E28DD4116CFFECE60E6606FA1E2832" box="[1346,1428,1282,1308]" pageId="3" pageNumber="96">Others</collectorName>
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</materialsCitation>
: 
<specimenCount id="9D11238B116CFFECE3DB6622FF53286E" box="[151,217,1318,1344]" pageId="3" pageNumber="96" type="male">7 ♂♂</specimenCount>
(11.4 × 9.2–15.3 × 
<quantity id="4CEF45E7116CFFECE28F6621FDA4286E" box="[451,558,1317,1344]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="3" pageNumber="96" unit="mm" value="12.2">12.2 mm</quantity>
), 
<materialsCitation id="3B7FE25F116CFFECE10F6621FB08284C" ID-GBIF-Occurrence="3395042305" collectingDate="2010-09-08" collectorName="Y. - H. Wang" elevation="280" latitude="24.530083" location="Suao" longLatPrecision="1" longitude="121.85361" municipality="Dakeng Bridge" pageId="3" pageNumber="96" specimenCount="1" specimenCount-female="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCount id="9D11238B116CFFECE10F6621FDFA286E" box="[579,624,1317,1344]" pageId="3" pageNumber="96" type="female">1 ♀</specimenCount>
(16.6 × 
<quantity id="4CEF45E7116CFFECE19A6621FCB4286E" box="[726,830,1317,1344]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="96" unit="mm" value="12.9">12.9 mm</quantity>
) (NCHUZOOL 13616), 
<collectingMunicipality id="6BCC7278116CFFECE7176621FA802811" box="[1115,1290,1317,1343]" pageId="3" pageNumber="96">Dakeng Bridge</collectingMunicipality>
, 
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, 
<collectingRegion id="49D326E0116CFFECE62C6621FA172811" box="[1376,1437,1317,1343]" country="Taiwan" name="Yilan" pageId="3" pageNumber="96">Yilan</collectingRegion>
(
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; 
<geoCoordinate id="EE238EC5116CFFECE202664CFE70284D" box="[334,506,1352,1379]" degrees="121" direction="east" minutes="51" orientation="longitude" pageId="3" pageNumber="96" precision="1" seconds="13.0" value="121.85361">121°51’13.0”E</geoCoordinate>
), elevation of 
<quantity id="4CEF45E7116CFFECE1D7664CFD6E284D" box="[667,740,1352,1379]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="96" unit="m" value="280.0">
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</quantity>
, coll. 
<collectorName id="26E28DD4116CFFECE06B664DFC21284D" box="[807,939,1353,1379]" pageId="3" pageNumber="96">Y.-H. Wang</collectorName>
<emphasis id="B9633410116CFFECE0FE664DFC69284C" box="[946,995,1352,1378]" italics="true" pageId="3" pageNumber="96">et al</emphasis>
., 
<date id="FFA9CEC2116CFFECE0B4664CFB08284C" box="[1016,1154,1352,1378]" pageId="3" pageNumber="96" value="2010-09-08">
<collectingDate id="EFED372A116CFFECE0B4664CFB08284C" box="[1016,1154,1352,1378]" pageId="3" pageNumber="96" value="2010-09-08">8 Sep. 2010</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B7FE25F116CFFECE7C1664CFD272889" ID-GBIF-Occurrence="3395042312" collectingDate="2018-03-10" collectorName="J. - J. Mao" elevation="1100" latitude="24.37886" location="Nan-ao" longLatPrecision="1" longitude="121.74963" municipality="Shenmihu" pageId="3" pageNumber="96" specimenCount="2" specimenCount-female="2" stateProvince="Yilan" typeStatus="paratype">
<specimenCount id="9D11238B116CFFECE7C1664CFB46284D" box="[1165,1228,1352,1379]" pageId="3" pageNumber="96" type="female">2 ♀♀</specimenCount>
(15.1 × 10.9, 16.7 × 
<quantity id="4CEF45E7116CFFECE3E1666FFE9B28AB" box="[173,273,1387,1413]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="3" pageNumber="96" unit="mm" value="12.8">12.8 mm</quantity>
) (NCHUZOOL 16436), around the lake of 
<collectingMunicipality id="6BCC7278116CFFECE1B8666FFCE228AB" box="[756,872,1387,1413]" pageId="3" pageNumber="96">Shenmihu</collectingMunicipality>
, 
<location id="8EC8BED9116CFFECE03E666FFC4D28AB" LSID="urn:lsid:plazi:treatment:03BE5914116CFFE9E3DB60C3FA0E2C83:8EC8BED9116CFFECE03E666FFC4D28AB" box="[882,967,1387,1413]" latitude="24.37886" longLatPrecision="1" longitude="121.74963" municipality="Shenmihu" name="Nan-ao" pageId="3" pageNumber="96" stateProvince="Yilan">Nan-ao</location>
, 
<collectingRegion id="49D326E0116CFFECE09D666FFB8428AB" box="[977,1038,1387,1413]" country="Taiwan" name="Yilan" pageId="3" pageNumber="96">Yilan</collectingRegion>
(
<geoCoordinate id="EE238EC5116CFFECE750666FFB3328AB" box="[1052,1209,1387,1413]" degrees="24" direction="north" minutes="22" orientation="latitude" pageId="3" pageNumber="96" precision="1" seconds="43.9" value="24.37886">24°22'43.9&quot;N</geoCoordinate>
; 
<geoCoordinate id="EE238EC5116CFFECE788666EFAE128AB" box="[1220,1387,1386,1413]" degrees="121" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="96" precision="1" seconds="58.7" value="121.74963">121°44'58.7&quot;E</geoCoordinate>
), elevation of 
<quantity id="4CEF45E7116CFFECE2416689FEE82886" box="[269,354,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="96" unit="m" value="1100.0">
<elevation id="003A0F31116CFFECE2416689FEE82886" box="[269,354,1421,1448]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="96" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, coll. 
<collectorName id="26E28DD4116CFFECE2EF668AFD862886" box="[419,524,1422,1448]" pageId="3" pageNumber="96">J.-J. Mao</collectorName>
, 
<date id="FFA9CEC2116CFFECE15A6689FD272889" box="[534,685,1421,1448]" pageId="3" pageNumber="96" value="2018-03-10">
<collectingDate id="EFED372A116CFFECE15A6689FD272889" box="[534,685,1421,1448]" pageId="3" pageNumber="96" value="2018-03-10">10 Mar. 2018</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="9D11238B116CFFECE1F46689FD6B2886" box="[696,737,1421,1448]" pageId="3" pageNumber="96" type="male">1 ♂</specimenCount>
(12.7 × 
<quantity id="4CEF45E7116CFFECE0716689FC1C2886" box="[829,918,1421,1448]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="96" unit="mm" value="9.7">9.7 mm</quantity>
), 
<specimenCount id="9D11238B116CFFECE0EB6689FC5A2886" box="[935,976,1421,1448]" pageId="3" pageNumber="96" type="female">1 ♀</specimenCount>
(13.9 × 
<quantity id="4CEF45E7116CFFECE7606689FB1A2886" box="[1068,1168,1421,1448]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="96" unit="mm" value="10.5">10.5 mm</quantity>
) (NCHUZOOL 14998), 
<materialsCitation id="3B7FE25F116CFFECE3DB66B4FC4128C3" ID-GBIF-Occurrence="3395042308" collectingDate="2009-09-26" collectionCode="ZRC" collectorName="J. - J. Mao" elevation="1808" latitude="24.500973" location="Nan-ao" longLatPrecision="1" longitude="121.63616" municipality="Tongshan" pageId="3" pageNumber="96" specimenCode="ZRC 2021.0649" specimenCount="1" specimenCount-female="1" stateProvince="Yilan" typeStatus="paratype">
<specimenCount id="9D11238B116CFFECE3DB66B4FF4B28E5" box="[151,193,1456,1483]" pageId="3" pageNumber="96" type="female">1 ♀</specimenCount>
(13.5 × 
<quantity id="4CEF45E7116CFFECE26E66B4FE0D28E5" box="[290,391,1456,1483]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="96" unit="mm" value="10.3">10.3 mm</quantity>
) (
<specimenCode id="DBB14079116CFFECE2D266B4FDD328E5" box="[414,601,1456,1483]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="96">ZRC 2021.0649</specimenCode>
), 
<collectingMunicipality id="6BCC7278116CFFECE12766B4FD5028E4" box="[619,730,1456,1482]" pageId="3" pageNumber="96">Tongshan</collectingMunicipality>
, 
<location id="8EC8BED9116CFFECE1AA66B5FCB128E5" LSID="urn:lsid:plazi:treatment:03BE5914116CFFE9E3DB60C3FA0E2C83:8EC8BED9116CFFECE1AA66B5FCB128E5" box="[742,827,1457,1483]" latitude="24.500973" longLatPrecision="1" longitude="121.63616" municipality="Tongshan" name="Nan-ao" pageId="3" pageNumber="96" stateProvince="Yilan">Nan-ao</location>
, 
<collectingRegion id="49D326E0116CFFECE00A66B4FC0928E4" box="[838,899,1456,1482]" country="Taiwan" name="Yilan" pageId="3" pageNumber="96">Yilan</collectingRegion>
(
<geoCoordinate id="EE238EC5116CFFECE0DE66B4FBA528E5" box="[914,1071,1456,1483]" degrees="24" direction="north" minutes="30" orientation="latitude" pageId="3" pageNumber="96" precision="1" seconds="15.5" value="24.504305">24°30'15.5&quot;N</geoCoordinate>
; 
<geoCoordinate id="EE238EC5116CFFECE77766B4FB5528E4" box="[1083,1247,1456,1482]" degrees="121" direction="east" minutes="37" orientation="longitude" pageId="3" pageNumber="96" precision="1" seconds="41.8" value="121.62828">121°37'41.8&quot;E</geoCoordinate>
– 24°30'03.5&quot;N; 121°38'10.2&quot;E), elevation of 
<quantity id="4CEF45E7116CFFECE29266D6FDF128C3" box="[478,635,1490,1517]" metricMagnitude="3" metricUnit="m" metricValue="1.8074999999999999" metricValueMax="1.88" metricValueMin="1.735" pageId="3" pageNumber="96" unit="m" value="1807.5" valueMax="1880.0" valueMin="1735.0">
<elevation id="003A0F31116CFFECE29266D6FDF128C3" box="[478,635,1490,1517]" metricMagnitude="3" metricUnit="m" metricValue="1.8074999999999999" metricValueMax="1.88" metricValueMin="1.735" pageId="3" pageNumber="96" unit="m" value="1807.5" valueMax="1880.0" valueMin="1735.0">1735–1880 m</elevation>
</quantity>
, coll. 
<collectorName id="26E28DD4116CFFECE1F366D7FCA328C3" box="[703,809,1491,1517]" pageId="3" pageNumber="96">J.-J. Mao</collectorName>
, 
<date id="FFA9CEC2116CFFECE07866D7FC4128C3" box="[820,971,1491,1517]" pageId="3" pageNumber="96" value="2009-09-26">
<collectingDate id="EFED372A116CFFECE07866D7FC4128C3" box="[820,971,1491,1517]" pageId="3" pageNumber="96" value="2009-09-26">26 Sep. 2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA8E802116CFFECE38B66F1FD402B7B" blockId="3.[151,1437,1178,2038]" pageId="3" pageNumber="96">
<emphasis id="B9633410116CFFECE38B66F1FE582B21" bold="true" box="[199,466,1525,1551]" pageId="3" pageNumber="96">Comparative material</emphasis>
. 
<taxonomicName id="4C179381116CFFECE2AD66F1FC4F2B21" authority="Shy, Ng &amp; Yu, 1994" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[481,965,1525,1552]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B9633410116CFFECE2AD66F1FD5B2B21" box="[481,721,1525,1551]" italics="true" pageId="3" pageNumber="96">Geothelphusa takuan</emphasis>
<bibRefCitation id="EF8695F3116CFFECE19566F1FC4F2B21" author="Shy, J. - Y. &amp; Ng, P. K. L. &amp; Yu, H. - P." box="[729,965,1525,1552]" pageId="3" pageNumber="96" pagination="781 - 846" refId="ref8705" refString="Shy, J. - Y., Ng, P. K. L. &amp; Yu, H. - P. (1994) Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42, 781 - 846." type="journal article" year="1994">Shy, Ng &amp; Yu, 1994</bibRefCitation>
</taxonomicName>
: see “Material examined” under 
<taxonomicName id="4C179381116CFFECE67666F1FEA42B1C" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B9633410116CFFECE67666F1FEA42B1C" italics="true" pageId="3" pageNumber="96">Geothelphusa takuan</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C179381116CFFECE272651CFC982B1C" authority="Shy, Ng &amp; Yu, 1994: 1" authorityName="Shy, Ng &amp; Yu" authorityPageNumber="1" authorityYear="1994" box="[318,786,1560,1587]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="tali">
<emphasis id="B9633410116CFFECE272651CFD8C2B1C" box="[318,518,1560,1586]" italics="true" pageId="3" pageNumber="96">Geothelphusa tali</emphasis>
<bibRefCitation id="EF8695F3116CFFECE143651CFC982B1C" author="Shy, J. - Y. &amp; Ng, P. K. L. &amp; Yu, H. - P." box="[527,786,1560,1587]" pageId="3" pageNumber="96" pagination="781 - 846" refId="ref8705" refString="Shy, J. - Y., Ng, P. K. L. &amp; Yu, H. - P. (1994) Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42, 781 - 846." type="journal article" year="1994">
Shy, Ng &amp; Yu, 1994: 
<specimenCount id="9D11238B116CFFECE048651CFC982B1C" box="[772,786,1560,1586]" pageId="3" pageNumber="96" type="male" typeStatus="holotype">1</specimenCount>
</bibRefCitation>
</taxonomicName>
holotype male (
<collectionCode id="ED0670C7116CFFECE085651CFB922B1C" box="[969,1048,1560,1586]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="3" pageNumber="96">NTOU</collectionCode>
F10187), Dali (= Tali), 
<collectingRegion id="49D326E0116CFFECE665651CFAEC2B1C" box="[1321,1382,1560,1586]" country="Taiwan" name="Yilan" pageId="3" pageNumber="96">Yilan</collectingRegion>
Co., 
<collectingCountry id="F300A892116CFFECE3DB653FFF662B7B" box="[151,236,1595,1621]" name="Taiwan" pageId="3" pageNumber="96">Taiwan</collectingCountry>
, coll. J.-Y. Shy &amp; W.-L. Tsay, 
<date id="FFA9CEC2116CFFECE173653FFD4C2B7B" box="[575,710,1595,1621]" pageId="3" pageNumber="96" value="1993-04-04">4 Apr. 1993</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C30DBB89116CFFECE38B6559FDE92BCE" pageId="3" pageNumber="96" type="diagnosis">
<paragraph id="8BA8E802116CFFECE38B6559FDE92BCE" blockId="3.[151,1437,1178,2038]" pageId="3" pageNumber="96">
<emphasis id="B9633410116CFFECE38B6559FEB02B59" bold="true" box="[199,314,1629,1655]" pageId="3" pageNumber="96">Diagnosis</emphasis>
. Carapace length, width 1.7-, 2.3-times carapace height, respectively. Anterolateral margins distinct, lined with inconspicuous granules, without epibranchial tooth. Subterminal segment of G1 (
<figureCitation id="132CF487116CFFECE7F66584FABF2BB4" box="[1210,1333,1664,1690]" captionStart="FIGURE 3" captionStartId="5.[151,250,1208,1232]" captionTargetBox="[237,1350,189,1172]" captionTargetId="figure-16@5.[225,1363,181,1184]" captionTargetPageId="5" captionText="FIGURE 3. Male right G1 and G2 of Geothelphusa boreas sp. nov. (holotype, NCHUZOOL 13615). A, D, ventral view of G1; B, dorsal view of G1; C, terminal segment of G1; D, ventral view of G2. Scale = 2.5 mm for A, B, D; scale = 0.5 mm for C." figureDoi="http://doi.org/10.5281/zenodo.5606777" httpUri="https://zenodo.org/record/5606777/files/figure.png" pageId="3" pageNumber="96">Fig. 3A–C</figureCitation>
) curving inwards, outer proximal margin without tooth; terminal segment slightly curving outwards to almost straight; total length of G1 5.8 times terminal segment.
</paragraph>
</subSubSection>
<subSubSection id="C30DBB89116CFFEAE38B65ECFCDF2B17" lastPageId="5" lastPageNumber="98" pageId="3" pageNumber="96" type="description">
<paragraph id="8BA8E802116CFFECE38B65ECFD5E2AD8" blockId="3.[151,1437,1178,2038]" pageId="3" pageNumber="96">
<emphasis id="B9633410116CFFECE38B65ECFEDB2A2C" bold="true" box="[199,337,1768,1794]" pageId="3" pageNumber="96">Description</emphasis>
. Carapace (
<figureCitation id="132CF487116CFFECE29A65ECFDC72A2D" box="[470,589,1768,1795]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="3" pageNumber="96">Fig. 2A, B</figureCitation>
) swollen longitudinally, transversely; dorsal surface smooth, glabrous, with fine pits. Carapace length, width 1.7-, 2.3-times carapace height, respectively. Frontal margin slightly dived into 2 lobes, without tooth. Postorbital cristae distinct, supraorbital margin smooth, without granules; infraorbital margin smooth to almost smooth, lined with inconspicuous granules. External orbital angles stout, external orbital regions concave. Anterolateral margins distinct, lined with inconspicuous granules, without epibranchial tooth. Postorbital cristae faint, smooth. Gastric, cardiac, intestinal regions smooth. H-shaped groove distinct. Tip of medium lobe of epistome stout. Distance between tip of sternopleonal cavity (
<figureCitation id="132CF487116CFFECE01764BCFC392AFC" box="[859,947,1976,2002]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="3" pageNumber="96">Fig. 2C</figureCitation>
) and anterior margin of thoracic sternite 3 about 2.2 times length of thoracic sternites 1 and 2.
</paragraph>
<caption id="DF68B88A116BFFEBE3DB642CFE832A82" ID-DOI="http://doi.org/10.5281/zenodo.5606775" ID-Zenodo-Dep="5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="4" pageNumber="97" startId="4.[151,250,1832,1856]" targetBox="[184,1403,182,1808]" targetPageId="4">
<paragraph id="8BA8E802116BFFEBE3DB642CFE832A82" blockId="4.[151,1436,1832,1964]" pageId="4" pageNumber="97">
<emphasis id="B9633410116BFFEBE3DB642CFE992A6E" bold="true" box="[151,275,1832,1856]" pageId="4" pageNumber="97">FIGURE 2.</emphasis>
<taxonomicName id="4C179381116BFFEBE255642CFE7A2A6E" authority="Shy &amp; Shih &amp; Mao, 2021" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[281,496,1832,1856]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="boreas" status="sp. nov.">
<emphasis id="B9633410116BFFEBE255642CFE7A2A6E" box="[281,496,1832,1856]" italics="true" pageId="4" pageNumber="97">Geothelphusa boreas</emphasis>
</taxonomicName>
<emphasis id="B9633410116BFFEBE2BB642CFDCD2A6E" bold="true" box="[503,583,1832,1856]" pageId="4" pageNumber="97">
<taxonomicNameLabel id="A250896B116BFFEBE2BB642CFDCD2A6E" box="[503,583,1832,1856]" pageId="4" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat.
</paragraph>
</caption>
<caption id="DF68B88A116AFFEAE3DB67BCFAF529DB" ID-DOI="http://doi.org/10.5281/zenodo.5606777" ID-Zenodo-Dep="5606777" httpUri="https://zenodo.org/record/5606777/files/figure.png" pageId="5" pageNumber="98" startId="5.[151,250,1208,1232]" targetBox="[237,1350,189,1172]" targetPageId="5">
<paragraph id="8BA8E802116AFFEAE3DB67BCFAF529DB" blockId="5.[151,1436,1208,1269]" pageId="5" pageNumber="98">
<emphasis id="B9633410116AFFEAE3DB67BCFE9829FE" bold="true" box="[151,274,1208,1232]" pageId="5" pageNumber="98">FIGURE 3.</emphasis>
Male right G1 and G2 of 
<taxonomicName id="4C179381116AFFEAE15B67BDFD6429FF" authority="Shy &amp; Shih &amp; Mao, 2021" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[535,750,1209,1233]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="boreas" status="sp. nov.">
<emphasis id="B9633410116AFFEAE15B67BDFD6429FF" box="[535,750,1209,1233]" italics="true" pageId="5" pageNumber="98">Geothelphusa boreas</emphasis>
</taxonomicName>
<emphasis id="B9633410116AFFEAE1B867BDFCCE29FF" bold="true" box="[756,836,1209,1233]" pageId="5" pageNumber="98">
<taxonomicNameLabel id="A250896B116AFFEAE1B867BDFCCE29FF" box="[756,836,1209,1233]" pageId="5" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, NCHUZOOL 13615). A, D, ventral view of G1; B, dorsal view of G1; C, terminal segment of G1; D, ventral view of G2. Scale = 2.5 mm for A, B, D; scale = 0.5 mm for C.
</paragraph>
</caption>
<paragraph id="8BA8E802116AFFEAE38B6627FACA28AB" blockId="5.[151,1437,1315,2025]" pageId="5" pageNumber="98">
Chelipeds (
<figureCitation id="132CF487116AFFEAE2046627FE4B2813" box="[328,449,1315,1341]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="5" pageNumber="98">Fig. 2A, B</figureCitation>
) of adult male unequal, fingers of larger chela forming obliquely triangular gape when closed. Ambulatory legs (
<figureCitation id="132CF487116AFFEAE2F86643FDA1284F" box="[436,555,1351,1377]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="5" pageNumber="98">Fig. 2A, C</figureCitation>
) smooth, dorsal, ventral margins of dactyli with 2 rows of small spines, respectively. Second leg about 1.8 carapace length. Telson bell-shaped, moderately short, width about 1.5 length.
</paragraph>
<paragraph id="8BA8E802116AFFEAE38B668BFCDF2B17" blockId="5.[151,1437,1315,2025]" pageId="5" pageNumber="98">
Subterminal segment of G1 (
<figureCitation id="132CF487116AFFEAE142668BFD0D2887" box="[526,647,1423,1449]" captionStart="FIGURE 3" captionStartId="5.[151,250,1208,1232]" captionTargetBox="[237,1350,189,1172]" captionTargetId="figure-16@5.[225,1363,181,1184]" captionTargetPageId="5" captionText="FIGURE 3. Male right G1 and G2 of Geothelphusa boreas sp. nov. (holotype, NCHUZOOL 13615). A, D, ventral view of G1; B, dorsal view of G1; C, terminal segment of G1; D, ventral view of G2. Scale = 2.5 mm for A, B, D; scale = 0.5 mm for C." figureDoi="http://doi.org/10.5281/zenodo.5606777" httpUri="https://zenodo.org/record/5606777/files/figure.png" pageId="5" pageNumber="98">Fig. 3A–C</figureCitation>
) curving inwards, outer proximal margin without tooth in most specimen, some smaller specimen (&lt;
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) with low tooth, inner proximal margin slightly dilated; terminal segment slightly curving outwards to almost straight; total length of G1 5.8 times terminal segment; length of synovial membrane about 4.6 times maximum width. Outer proximal margin of basal segment of G2 (
<figureCitation id="132CF487116AFFEAE72766FFFB4B2B3B" box="[1131,1217,1531,1557]" captionStart="FIGURE 3" captionStartId="5.[151,250,1208,1232]" captionTargetBox="[237,1350,189,1172]" captionTargetId="figure-16@5.[225,1363,181,1184]" captionTargetPageId="5" captionText="FIGURE 3. Male right G1 and G2 of Geothelphusa boreas sp. nov. (holotype, NCHUZOOL 13615). A, D, ventral view of G1; B, dorsal view of G1; C, terminal segment of G1; D, ventral view of G2. Scale = 2.5 mm for A, B, D; scale = 0.5 mm for C." figureDoi="http://doi.org/10.5281/zenodo.5606777" httpUri="https://zenodo.org/record/5606777/files/figure.png" pageId="5" pageNumber="98">Fig. 3D</figureCitation>
) dilated, showing a single lobe; distal segment short, about 0.16 times total length.
</paragraph>
</subSubSection>
<subSubSection id="C30DBB89116AFFEAE38B6547FB732AC7" pageId="5" pageNumber="98" type="etymology">
<paragraph id="8BA8E802116AFFEAE38B6547FDD42BAF" blockId="5.[151,1437,1315,2025]" pageId="5" pageNumber="98">
<emphasis id="B9633410116AFFEAE38B6547FEC42B73" bold="true" box="[199,334,1603,1629]" pageId="5" pageNumber="98">Etymology.</emphasis>
The species is derived from the Greek “ 
<taxonomicName id="4C179381116AFFEAE0586547FCD52B73" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[788,863,1603,1629]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="boreas">
<emphasis id="B9633410116AFFEAE0586547FCD52B73" box="[788,863,1603,1629]" italics="true" pageId="5" pageNumber="98">boreas</emphasis>
</taxonomicName>
” (for north), alluding to the northernmost distribution of this species in the Central Range.
</paragraph>
<paragraph id="8BA8E802116AFFEAE38B658FFB732AC7" blockId="5.[151,1437,1315,2025]" pageId="5" pageNumber="98">
<emphasis id="B9633410116AFFEAE38B658FFE042B8B" bold="true" box="[199,398,1675,1701]" pageId="5" pageNumber="98">Ecological notes.</emphasis>
The specimens were collected from the montane area with an elevation of 
<quantity id="4CEF45E7116AFFEAE781658FFA992B8B" box="[1229,1299,1675,1701]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="98" unit="m" value="740.0">740 m</quantity>
(the Fushan Botanical Garden area) to 
<quantity id="4CEF45E7116AFFEAE28C65ABFD9E2BE7" box="[448,532,1711,1737]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="98" unit="m" value="1100.0">1100 m</quantity>
(Shenmihu) and about 
<quantity id="4CEF45E7116AFFEAE05065ABFCF82BE7" box="[796,882,1711,1737]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="98" unit="m" value="1800.0">1800 m</quantity>
(Tonghsan), but also extended to the coastal cliff area of Suao (elevation of 
<quantity id="4CEF45E7116AFFEAE28965D7FD842BC3" box="[453,526,1747,1773]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="98" unit="m" value="280.0">280 m</quantity>
) (
<figureCitation id="132CF487116AFFEAE16B65D7FDE32BC3" box="[551,617,1747,1773]" captionStart="FIGURE 1" captionStartId="1.[151,250,1662,1686]" captionTargetBox="[200,1368,750,1619]" captionTargetPageId="1" captionText="FIGURE 1. Collection sites (empty circles) for the Geothelphusa species from northern Taiwan. For locality names, see Table 1. R. = river." figureDoi="http://doi.org/10.5281/zenodo.5606773" httpUri="https://zenodo.org/record/5606773/files/figure.png" pageId="5" pageNumber="98">Fig. 1</figureCitation>
). The distributional range of this species covers an area of 
<quantity id="4CEF45E7116AFFEAE64065D7FADF2BC3" box="[1292,1365,1747,1774]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="98" unit="km" value="45.0">45 km</quantity>
(from Fushan to Shenmihu; 
<figureCitation id="132CF487116AFFEAE2DF65F3FE5F2A3F" box="[403,469,1783,1809]" captionStart="FIGURE 1" captionStartId="1.[151,250,1662,1686]" captionTargetBox="[200,1368,750,1619]" captionTargetPageId="1" captionText="FIGURE 1. Collection sites (empty circles) for the Geothelphusa species from northern Taiwan. For locality names, see Table 1. R. = river." figureDoi="http://doi.org/10.5281/zenodo.5606773" httpUri="https://zenodo.org/record/5606773/files/figure.png" pageId="5" pageNumber="98">Fig. 1</figureCitation>
). There was running water in most habitats (
<figureCitation id="132CF487116AFFEAE09265F3FBD22A3F" box="[990,1112,1783,1809]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="5" pageNumber="98">Fig. 2D–F</figureCitation>
), but some crabs were more semiterrestrial, hiding under fallen leaves with only some seepage in Tongshan (
<figureCitation id="132CF487116AFFEAE761641FFB272A1B" box="[1069,1197,1819,1845]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="5" pageNumber="98">Fig. 2G, H</figureCitation>
). Other species sympatric with this species include members of the 
<taxonomicName id="4C179381116AFFEAE1F7643BFCFE2A77" authorityName="Shy, P. K. L. Ng &amp; Yu" authorityYear="1994" box="[699,884,1855,1881]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="eucrinodonta">
<emphasis id="B9633410116AFFEAE1F7643BFCFE2A77" box="[699,884,1855,1881]" italics="true" pageId="5" pageNumber="98">G. eucrinodonta</emphasis>
</taxonomicName>
species complex, 
<taxonomicName id="4C179381116AFFEAE70A643BFB052A77" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1094,1167,1855,1881]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="tali">
<emphasis id="B9633410116AFFEAE70A643BFB052A77" box="[1094,1167,1855,1881]" italics="true" pageId="5" pageNumber="98">G. tali</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C179381116AFFEAE786643BFF762A53" authority="Shy, Ng &amp; Yu, 1994" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="ilan">
<emphasis id="B9633410116AFFEAE786643BFA902A77" box="[1226,1306,1855,1881]" italics="true" pageId="5" pageNumber="98">G. ilan</emphasis>
<bibRefCitation id="EF8695F3116AFFEAE66E643BFF762A53" author="Shy, J. - Y. &amp; Ng, P. K. L. &amp; Yu, H. - P." pageId="5" pageNumber="98" pagination="781 - 846" refId="ref8705" refString="Shy, J. - Y., Ng, P. K. L. &amp; Yu, H. - P. (1994) Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42, 781 - 846." type="journal article" year="1994">
Shy, Ng &amp; Yu, 
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</bibRefCitation>
</taxonomicName>
in Fushan area (
<bibRefCitation id="EF8695F3116AFFEAE2F86467FDE02A53" author="Shih, H. - T. &amp; Shy, J. - Y. &amp; Lee, J. - H." box="[436,618,1891,1917]" pageId="5" pageNumber="98" pagination="661 - 675" refId="ref8603" refString="Shih, H. - T., Shy, J. - Y., Lee, J. - H. (2010) A new freshwater crab of the genus Geothelphusa (Brachyura, Potamidae) from southwestern Taiwan. Crustaceana Monographs, 14, 661 - 675. https: // doi. org / 10.1163 / 9789047427759 _ 048" type="journal article" year="2010">
Shih 
<emphasis id="B9633410116AFFEAE2A36460FDAA2A53" box="[495,544,1891,1917]" italics="true" pageId="5" pageNumber="98">et al</emphasis>
. 2010
</bibRefCitation>
). In the higher altitude (Tongshan and Shenmihu), the habitats of the new species are near the 
<taxonomicName id="4C179381116AFFEAE235648CFD952A8F" box="[377,543,1927,1953]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="98" phylum="Tracheophyta" rank="species" species="hayatae">
<emphasis id="B9633410116AFFEAE235648CFD952A8F" box="[377,543,1927,1953]" italics="true" pageId="5" pageNumber="98">Fagus hayatae</emphasis>
</taxonomicName>
(
<collectingCountry id="F300A892116AFFEAE1616483FD0B2A8F" box="[557,641,1927,1953]" name="Taiwan" pageId="5" pageNumber="98">Taiwan</collectingCountry>
beech) forest (
<figureCitation id="132CF487116AFFEAE0646483FC282A8F" box="[808,930,1927,1953]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="5" pageNumber="98">Fig. 2E–H</figureCitation>
), with the mean (and standard deviation) water temperature 17.66 ± 2.12 
<superScript id="7C62454A116AFFEAE29164AFFE722AEB" attach="right" box="[477,504,1963,1989]" fontSize="6" pageId="5" pageNumber="98">oC</superScript>
(
<date id="FFA9CEC2116AFFEAE14A64AFFD0E2AEB" box="[518,644,1963,1989]" pageId="5" pageNumber="98" value="2018-04">April 2018</date>
) and 23.25 ± 1.78 
<superScript id="7C62454A116AFFEAE01B64AFFCF82AEB" attach="right" box="[855,882,1963,1989]" fontSize="6" pageId="5" pageNumber="98">oC</superScript>
(
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), pH 7.4 ± 0.3 (
<date id="FFA9CEC2116AFFEAE60E64AFFEB82AC7" pageId="5" pageNumber="98" value="2018-04" valueMax="2018-10-31" valueMin="2018-04-00">April to October 2018</date>
) and dissolved oxygen 8.7 ± 
<quantity id="4CEF45E7116AFFEAE13764CBFD422AC7" box="[635,712,1999,2025]" metricMagnitude="-7" metricUnit="kg" metricValue="9.0" pageId="5" pageNumber="98" unit="mg" value="0.9">0.9 mg</quantity>
/L (
<date id="FFA9CEC2116AFFEAE1A364CBFC612AC7" box="[751,1003,1999,2025]" pageId="5" pageNumber="98" value="2018-04" valueMax="2018-10-31" valueMin="2018-04-00">April to October 2018</date>
) in Shenmihu (
<figureCitation id="132CF487116AFFEAE7DB64CBFB662AC7" box="[1175,1260,1999,2025]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[184,1403,182,1808]" captionTargetId="figure-21@4.[182,1405,181,1808]" captionTargetPageId="4" captionText="FIGURE 2. Geothelphusa boreas sp. nov. A, B, C, dorsal, frontal and ventral views of the holotype (NCHUZOOL 13615); D, habitat in Dakeng Bridge, Suao, Yilan; E, F, habitat in Shenmihu, Nan-ao, Yilan; G, H, habitat in Tongshan, Nan-ao, Yilan. The arrows in E and G indicated the localities with crabs observed. The two rulers in H were used for a separate ecological study of the habitat." figureDoi="http://doi.org/10.5281/zenodo.5606775" httpUri="https://zenodo.org/record/5606775/files/figure.png" pageId="5" pageNumber="98">Fig. 2F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30DBB891169FFE9E38B6393FA0E2C83" pageId="6" pageNumber="99" type="discussion">
<paragraph id="8BA8E8021169FFE9E38B6393FA0E2C83" blockId="6.[151,1437,151,429]" pageId="6" pageNumber="99">
<emphasis id="B96334101169FFE9E38B6393FEBE2D9F" bold="true" box="[199,308,151,177]" pageId="6" pageNumber="99">Remarks</emphasis>
. 
<taxonomicName id="4C1793811169FFE9E20D6393FDA42D9F" authority="Shy &amp; Shih &amp; Mao, 2021" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[321,558,151,177]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="boreas" status="sp. nov.">
<emphasis id="B96334101169FFE9E20D6393FDA42D9F" box="[321,558,151,177]" italics="true" pageId="6" pageNumber="99">Geothelphusa boreas</emphasis>
</taxonomicName>
<emphasis id="B96334101169FFE9E1796393FD072D9F" bold="true" box="[565,653,151,177]" pageId="6" pageNumber="99">
<taxonomicNameLabel id="A250896B1169FFE9E1796393FD072D9F" box="[565,653,151,177]" pageId="6" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically most similar to 
<taxonomicName id="4C1793811169FFE9E7556393FB0C2D9F" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1049,1158,151,177]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101169FFE9E7556393FB0C2D9F" box="[1049,1158,151,177]" italics="true" pageId="6" pageNumber="99">G. takuan</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1793811169FFE9E7F76393FA882D9F" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1211,1282,151,177]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="tali">
<emphasis id="B96334101169FFE9E7F76393FA882D9F" box="[1211,1282,151,177]" italics="true" pageId="6" pageNumber="99">G. tali</emphasis>
</taxonomicName>
which occurs in adjacent regions in northern 
<collectingCountry id="F300A8921169FFE9E2B663BFFDDA2DFB" box="[506,592,187,213]" name="Taiwan" pageId="6" pageNumber="99">Taiwan</collectingCountry>
. The G1 structures remain the best way to separate them. The subterminal segment of the G1 of 
<taxonomicName id="4C1793811169FFE9E2C463DBFE452DD7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[392,463,223,249]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="tali">
<emphasis id="B96334101169FFE9E2C463DBFE452DD7" box="[392,463,223,249]" italics="true" pageId="6" pageNumber="99">G. tali</emphasis>
</taxonomicName>
is straight (
<bibRefCitation id="EF8695F31169FFE9E11E63DBFD752DD7" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[594,767,223,249]" pageId="6" pageNumber="99" refId="ref8837" refString="Shy, J. - Y., Shih, H. - T. &amp; Ng, P. K. L. (2020) Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. III. Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, 26 + 232 pp." type="book" year="2020">
Shy 
<emphasis id="B96334101169FFE9E1C963DBFD3C2DD7" box="[645,694,223,249]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2020
</bibRefCitation>
: fig. 124), while those of present new species and 
<taxonomicName id="4C1793811169FFE9E66263DBFA162DD7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1326,1436,223,249]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101169FFE9E66263DBFA162DD7" box="[1326,1436,223,249]" italics="true" pageId="6" pageNumber="99">G. takuan</emphasis>
</taxonomicName>
are distinctly curved inwards (
<figureCitation id="132CF4871169FFE9E2A66207FDB22C33" box="[490,568,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1208,1232]" captionTargetBox="[237,1350,189,1172]" captionTargetId="figure-16@5.[225,1363,181,1184]" captionTargetPageId="5" captionText="FIGURE 3. Male right G1 and G2 of Geothelphusa boreas sp. nov. (holotype, NCHUZOOL 13615). A, D, ventral view of G1; B, dorsal view of G1; C, terminal segment of G1; D, ventral view of G2. Scale = 2.5 mm for A, B, D; scale = 0.5 mm for C." figureDoi="http://doi.org/10.5281/zenodo.5606777" httpUri="https://zenodo.org/record/5606777/files/figure.png" pageId="6" pageNumber="99">Figs. 3</figureCitation>
, 
<figureCitation id="132CF4871169FFE9E1096207FD012C33" box="[581,651,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1349,1373]" captionTargetBox="[279,1306,521,1284]" captionTargetId="figure-193@6.[166,1421,453,1329]" captionTargetPageId="6" captionText="FIGURE 4. Male right G1s of Geothelphusa takuan Shy, Yu &amp; Ng, 1994 (holotype, NTOU F10205; A, B) and G. hirsuta Tan &amp; Liu, 1998 (holotype, TMCD 3285; C, D). A, C, ventral view; B, D, dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5606781" httpUri="https://zenodo.org/record/5606781/files/figure.png" pageId="6" pageNumber="99">4A, B</figureCitation>
; 
<bibRefCitation id="EF8695F31169FFE9E1DB6207FCCE2C33" author="Shy, J. - Y. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[663,836,259,285]" pageId="6" pageNumber="99" refId="ref8837" refString="Shy, J. - Y., Shih, H. - T. &amp; Ng, P. K. L. (2020) Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. III. Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, 26 + 232 pp." type="book" year="2020">
Shy 
<emphasis id="B96334101169FFE9E1856207FD702C33" box="[713,762,259,285]" italics="true" pageId="6" pageNumber="99">et al</emphasis>
. 2020
</bibRefCitation>
: fig. 121). In addition, the G1 subterminal segment in 
<taxonomicName id="4C1793811169FFE9E3DB6223FE8C2C6F" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[151,262,295,321]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101169FFE9E3DB6223FE8C2C6F" box="[151,262,295,321]" italics="true" pageId="6" pageNumber="99">G. takuan</emphasis>
</taxonomicName>
has a distinct tooth on the outer proximal margin (
<figureCitation id="132CF4871169FFE9E00D6223FC112C6F" box="[833,923,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1349,1373]" captionTargetBox="[279,1306,521,1284]" captionTargetId="figure-193@6.[166,1421,453,1329]" captionTargetPageId="6" captionText="FIGURE 4. Male right G1s of Geothelphusa takuan Shy, Yu &amp; Ng, 1994 (holotype, NTOU F10205; A, B) and G. hirsuta Tan &amp; Liu, 1998 (holotype, TMCD 3285; C, D). A, C, ventral view; B, D, dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5606781" httpUri="https://zenodo.org/record/5606781/files/figure.png" pageId="6" pageNumber="99">Fig. 4A</figureCitation>
), while in 
<taxonomicName id="4C1793811169FFE9E75E6223FBF52C6F" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[1042,1151,295,321]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="boreas">
<emphasis id="B96334101169FFE9E75E6223FBF52C6F" box="[1042,1151,295,321]" italics="true" pageId="6" pageNumber="99">G. boreas</emphasis>
</taxonomicName>
, there is usually no tooth present on the structure (or at most with a faint knob in some small individuals) (
<figureCitation id="132CF4871169FFE9E75B624FFBFA2C4B" box="[1047,1136,331,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1208,1232]" captionTargetBox="[237,1350,189,1172]" captionTargetId="figure-16@5.[225,1363,181,1184]" captionTargetPageId="5" captionText="FIGURE 3. Male right G1 and G2 of Geothelphusa boreas sp. nov. (holotype, NCHUZOOL 13615). A, D, ventral view of G1; B, dorsal view of G1; C, terminal segment of G1; D, ventral view of G2. Scale = 2.5 mm for A, B, D; scale = 0.5 mm for C." figureDoi="http://doi.org/10.5281/zenodo.5606777" httpUri="https://zenodo.org/record/5606777/files/figure.png" pageId="6" pageNumber="99">Fig. 3A</figureCitation>
). The distance between the tip of the sternopleonal cavity and the anterior margin of thoracic sternite 
<quantity id="4CEF45E71169FFE9E082626BFC732CA7" box="[974,1017,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="99" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C1793811169FFE9E0B3626BFBE72CA7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1023,1133,367,393]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="takuan">
<emphasis id="B96334101169FFE9E0B3626BFBE72CA7" box="[1023,1133,367,393]" italics="true" pageId="6" pageNumber="99">G. takuan</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1793811169FFE9E7EF626BFB602CA7" authorityName="Shy, Ng &amp; Yu" authorityYear="1994" box="[1187,1258,367,393]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="tali">
<emphasis id="B96334101169FFE9E7EF626BFB602CA7" box="[1187,1258,367,393]" italics="true" pageId="6" pageNumber="99">G. tali</emphasis>
</taxonomicName>
being 1.0 times and 1.2 times the length of sternites 1 and 2, respectively, which are shorter than the ratio 2.2 times in 
<emphasis id="B96334101169FFE9E65C6297FA0E2C83" box="[1296,1412,403,429]" italics="true" pageId="6" pageNumber="99">
<taxonomicName id="4C1793811169FFE9E65C6297FA0B2C83" authorityName="Shy &amp; Shih &amp; Mao" authorityYear="2021" box="[1296,1409,403,429]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="99" phylum="Arthropoda" rank="species" species="boreas">G. boreas</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>