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<document id="97A7D15B4D844013CA1E91AF5C4327A0" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E35BE22FFE6AA3B85E88FDCF" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Mulinoides plenicostatus" docType="treatment" docVersion="22" lastPageNumber="1552" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1551" updateTime="1698783606523" updateUser="ExternalLinkService">
<mods:mods id="8EC510201FCBB19E42B5B27B13236FD9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="042367B0C210342C827A40062F47D671">
<mods:title id="E3B269123B9AC51BD4206DFD51F945C0">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
</mods:titleInfo>
<mods:name id="1AA0B6DE4FAB3E8F2615DF584F80D3FE" type="personal">
<mods:role id="02D50DC2F54DA1BF5CDA2CDED430C8E2">
<mods:roleTerm id="975C02D300B509CB5900155D37D31D18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E721C7ED78E15420147377D4569C1D2">DeVries, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7781F3340CED76BA7698816CB49DC4EE">text</mods:typeOfResource>
<mods:relatedItem id="E438D70DECC3471CD4D316CDB16AB480" type="host">
<mods:titleInfo id="757910105205A68788A549A215C17AB5">
<mods:title id="5DE32EDE9DF432E74FF3671DE832495A">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="8D1D0BD64A21DDA56F7D858F49F2127F">
<mods:date id="403403075A2AB41A85BC34E73DB47A0A">2019</mods:date>
<mods:detail id="24B3B388E54B4098A844EF69982D2227" type="pubDate">
<mods:number id="153A08F50B38443526046CA4FA4AB2E2">2018-12-17</mods:number>
</mods:detail>
<mods:detail id="B7C79E3DEADFABB75E4084C0EC62036F" type="volume">
<mods:number id="6EF398F49D04CC46992426A710E359DF">53</mods:number>
</mods:detail>
<mods:detail id="20D1F4DFC7E526BFB43DFFB8EC5DCC3B" type="issue">
<mods:number id="4F3CD76B1C3C2AD5C244D7F1EFDA6C5D">25</mods:number>
</mods:detail>
<mods:extent id="DEB2EB32EF66B554730451C749DB0DEE" unit="page">
<mods:start id="B21AF3E431804F383541B11A81882465">1533</mods:start>
<mods:end id="F307C31731AF856789EF88F906D06CF1">1584</mods:end>
</mods:extent>
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<mods:classification id="775676DAFE030B46ED24B42CB18CD6A6">journal article</mods:classification>
<mods:identifier id="3648731FAF8CBC790E86C8AFA0EB615B" type="CLB-Dataset">24019</mods:identifier>
<mods:identifier id="2866D7459EA0FC73D02E4A900BBB7A0D" type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
<mods:identifier id="3ECB49594E7164C8A8363EEE5943B1BB" type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
<mods:identifier id="57B8D0E9DA5C2D498E7663F849D48A05" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="37842365D2A23B60951B3F2E0D42790B" type="Zenodo-Dep">3670229</mods:identifier>
</mods:mods>
<treatment id="03BB2338E35BE22FFE6AA3B85E88FDCF" ID-DOI="http://doi.org/10.5281/zenodo.3671228" ID-GBIF-Taxon="162362490" ID-Zenodo-Dep="3671228" LSID="urn:lsid:plazi:treatment:03BB2338E35BE22FFE6AA3B85E88FDCF" httpUri="http://treatment.plazi.org/id/03BB2338E35BE22FFE6AA3B85E88FDCF" lastPageId="20" lastPageNumber="1552" pageId="19" pageNumber="1551">
<subSection id="E29D8BC5E35BE228FE6AA3B85955FCD8" box="[459,857,763,788]" pageId="19" pageNumber="1551" type="multiple">
<subSubSection id="C308C1A5E35BE228FE6AA3B85955FCD8" box="[459,857,763,788]" pageId="19" pageNumber="1551" type="nomenclature">
<paragraph id="8BAD922EE35BE228FE6AA3B85955FCD8" blockId="19.[459,857,763,823]" box="[459,857,763,788]" pageId="19" pageNumber="1551">
<heading id="D0E52542E35BE228FE6AA3B85955FCD8" box="[459,857,763,788]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="1551" reason="2">
<taxonomicName id="4C12E9ADE35BE228FE6AA3B858FFFCD8" authority="DeVries" box="[459,755,763,788]" class="Bivalvia" family="Mactridae" genus="Mulinoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="19" pageNumber="1551" phylum="Mollusca" rank="species" species="plenicostatus">
<emphasis id="B9664E3CE35BE228FE6AA3B858FFFCD8" bold="true" box="[459,755,763,788]" italics="true" pageId="19" pageNumber="1551">Mulinoidesplenicostatus</emphasis>
</taxonomicName>
sp. nov.
</heading>
</paragraph>
</subSubSection>
</subSection>
<subSection id="E29D8BC5E35BE22FFDE1A25D5E88FDCF" lastPageId="20" lastPageNumber="1552" pageId="19" pageNumber="1551" type="multiple">
<subSubSection id="C308C1A5E35BE228FDE1A25D58EFFCFB" box="[576,739,798,823]" pageId="19" pageNumber="1551" type="description">
<paragraph id="8BAD922EE35BE228FDE1A25D58EFFCFB" blockId="19.[459,857,763,823]" box="[576,739,798,823]" pageId="19" pageNumber="1551">
<heading id="D0E52542E35BE228FDE1A25D58EFFCFB" box="[576,739,798,823]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="1551" reason="5">
(
<figureCitation id="13298EABE35BE228FDE9A25D58A0FCFB" box="[584,684,798,823]" captionStart="Figure 3" captionStartId="11.[160,225,1261,1284]" captionTargetBox="[200,1113,144,1228]" captionTargetId="figure@11.[199,1117,142,1230]" captionTargetPageId="11" captionText="Figure 3. Bivalves from the Caballas Formation. (a, b) Twig moulds on attachment scars of Ambigostrea sechura. (a) UWBM 107557, latex cast. (b) UWBM 107564, oyster length 11 mm. (c–e) Carolia (Parinomya) parinensis Olsson, 1928, on oyster attachment scars. (c) MUSM INV 243, latex cast of external mould on Ambigostrea sechura (MUSM INV 237), Carolia H 11.7 mm (top to bottom). (d) UWBM 107569, external mould on Ambigostrea sechura (UWBM 107559), Carolia H 22.9 mm (left to right). (e) UWBM 107570, external mould on Ambigostrea sechura (UWBM 107560), Carolia H 15.3 mm (left to right). (f–k) Incatina collinsae sp. nov., B8772. (f) UWBM 107637, holotype, L 13.7 mm. Right valve of pair, exterior. (g) UWBM 107637, left valve of pair, exterior. (h) UWBM 107637, paired valves, dorsal view. (i) UWBM 107638, paratype, L 10.0 mm. Right valve of pair. (j) UWBM 107639, paratype, L 16.4 mm. Right valve of pair. (k) UWBM 107638, left valve of pair. (l–q) Pleuriocardia (Incacardium) schneideri sp. nov. (l) UWBM 107575, paratype, B8769, L 13.7 mm. Paired valves, dorsal view, anterior to right. (m) UWBM 107574, paratype, B8770, L 25.0 mm. Right valve, exterior. (n) MUSM INV 247, paratype, B8772, L 21.1 mm. Right valve, exterior. (o) UWBM 107576, paratype, B8769, L 25.5 mm. External mould of left valve, oblique view. (p) MUSM INV 246, paratype, B8772, L 20.4 mm. External mould of right valve. (q) 107573, holotype, B8772, L 21.5 mm. Left valve, exterior. (r–t) Mulinoides plenicostatus sp. nov., UWBM 107571, holotype, B8769, L 28.1 mm. (r) Left valve, exterior. (s) Dorsal view, anterior on right. (t) Right valve, exterior." figureDoi="http://doi.org/10.5281/zenodo.3670235" httpUri="https://zenodo.org/record/3670235/files/figure.png" pageId="19" pageNumber="1551">Figure 3</figureCitation>
(r
<emphasis id="B9664E3CE35BE228FD1DA25D58C6FCFB" box="[700,714,798,823]" italics="true" pageId="19" pageNumber="1551">–</emphasis>
t))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35BE228FF01A2205B11FC29" pageId="19" pageNumber="1551" type="diagnosis">
<paragraph id="8BAD922EE35BE228FF01A2205B18FCB1" blockId="19.[160,1155,867,997]" box="[160,276,867,893]" pageId="19" pageNumber="1551">
<emphasis id="B9664E3CE35BE228FF01A2205B18FCB1" bold="true" box="[160,276,867,893]" italics="true" pageId="19" pageNumber="1551">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD922EE35BE228FF01A2CB5B11FC29" blockId="19.[160,1155,867,997]" pageId="19" pageNumber="1551">Highly arched valves, barely elongated posteriorly; sharply angled posteroventral and anteroventral umbonal ridges. Well defined and regular commarginal ribs across most of specimens.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35BE228FF01A551592AFAA8" pageId="19" pageNumber="1551" type="description">
<paragraph id="8BAD922EE35BE228FF01A5515B2AFBE0" blockId="19.[160,1155,1042,1380]" box="[160,294,1042,1068]" pageId="19" pageNumber="1551">
<emphasis id="B9664E3CE35BE228FF01A5515B2AFBE0" bold="true" box="[160,294,1042,1068]" italics="true" pageId="19" pageNumber="1551">Description</emphasis>
</paragraph>
<paragraph id="8BAD922EE35BE228FF01A575592AFAA8" blockId="19.[160,1155,1042,1380]" pageId="19" pageNumber="1551">
Shell about
<quantity id="4CEA3FCBE35BE228FE8AA5755B77FB82" box="[299,379,1078,1102]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="1551" unit="mm" value="30.0">30 mm</quantity>
long, trapezoidal, subequilateral, inflated medially, posteroventrally truncate; for adult specimens, H:L = 0.9, W:H = 0.7. Beaks low, prominent, prosogyrate, extending nearly to hingeline. Small lunular-like area with change in sculpture; posterior counterpart not visible. Commarginal sculpture of low, bevelled, flattened, narrow ribs, rarely bifurcating, with interspaces narrower than commarginal ribs; ribs extending at least three-quarters distance from ventral margin to beak. Anteroventral umbonal ridge angular; commarginal ribs on anterodorsal area strong, half as numerous as across middle of valve. Posteroventral umbonal ridge strongly angular with broad and shallow anteriorly adjacent sulcus. Hinge and interior not visible.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35BE228FF01A4D3594DF973" pageId="19" pageNumber="1551" type="discussion">
<paragraph id="8BAD922EE35BE228FF01A4D35B0AFA66" blockId="19.[160,1156,1424,1727]" box="[160,262,1424,1450]" pageId="19" pageNumber="1551">
<emphasis id="B9664E3CE35BE228FF01A4D35B0AFA66" bold="true" box="[160,262,1424,1450]" italics="true" pageId="19" pageNumber="1551">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE35BE228FF01A4F6594DF973" blockId="19.[160,1156,1424,1727]" pageId="19" pageNumber="1551">
Specimens of
<emphasis id="B9664E3CE35BE228FEE0A4F65849FA01" box="[321,581,1461,1485]" italics="true" pageId="19" pageNumber="1551">Mulinoidesplenicostatus</emphasis>
sp. nov. are more highly arched and have a more centrally located beak than specimens of the
<typeStatus id="54A92C8CE35BE228FD1AA49458E1FA23" box="[699,749,1495,1519]" pageId="19" pageNumber="1551">type</typeStatus>
species,
<taxonomicName id="4C12E9ADE35BE228FCFBA49459B3FA23" authorityName="Olssoni" authorityYear="1944" box="[858,959,1495,1519]" class="Bivalvia" family="Mactridae" genus="Mulinoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="19" pageNumber="1551" phylum="Mollusca" rank="species" species="chilca">
<emphasis id="B9664E3CE35BE228FCFBA49459B3FA23" box="[858,959,1495,1519]" italics="true" pageId="19" pageNumber="1551">M. chilca</emphasis>
</taxonomicName>
, from the upper Maastrichtian
<emphasis id="B9664E3CE35BE228FE9FA4B95B4FF9DE" box="[318,323,1530,1554]" italics="true" pageId="19" pageNumber="1551">‘</emphasis>
Baculites
<emphasis id="B9664E3CE35BE228FE06A4B95BA1F9DE" box="[423,429,1530,1554]" italics="true" pageId="19" pageNumber="1551">’</emphasis>
beds of the
<collectingCountry id="F305D2BEE35BE228FD9DA4B95898F9DE" box="[572,660,1530,1554]" name="Haiti" pageId="19" pageNumber="1551">Tortuga</collectingCountry>
Formation in the Sechura Basin of northern
<collectingCountry id="F305D2BEE35BE228FF01A75E5ADFF9F9" box="[160,211,1565,1589]" name="Peru" pageId="19" pageNumber="1551">Peru</collectingCountry>
(
<bibRefCitation id="EF83EFDFE35BE228FF7EA75E5B66F9F9" author="Olsson AA" box="[223,362,1565,1589]" pageId="19" pageNumber="1551" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson 1944</bibRefCitation>
). In these characters, the Caballas Formation species greatly resembles
<taxonomicName id="4C12E9ADE35BE228FF01A77C5BB4F99B" authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[160,440,1599,1623]" class="Bivalvia" family="Mactridae" genus="Mulinoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="19" pageNumber="1551" phylum="Mollusca" rank="species" species="chicama">
<emphasis id="B9664E3CE35BE228FF01A77C5B13F99B" box="[160,287,1599,1623]" italics="true" pageId="19" pageNumber="1551">M. chicama</emphasis>
Olsson, 1944
</taxonomicName>
, from the upper Campanian portion of the
<collectingCountry id="F305D2BEE35BE228FC0EA77C5E0BF99B" box="[943,1031,1599,1623]" name="Haiti" pageId="19" pageNumber="1551">Tortuga</collectingCountry>
Formation (
<bibRefCitation id="EF83EFDFE35BE228FF07A7215B39F9B6" author="Olsson AA" box="[166,309,1634,1658]" pageId="19" pageNumber="1551" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson 1944</bibRefCitation>
). Specimens of
<emphasis id="B9664E3CE35BE228FE55A7215803F9B6" box="[500,527,1634,1658]" italics="true" pageId="19" pageNumber="1551">M.</emphasis>
plenicostatus differ from the figured specimen of
<taxonomicName id="4C12E9ADE35BE228FBC8A7215AF0F951" authorityName="Olssoni" authorityYear="1944" class="Bivalvia" family="Mactridae" genus="Mulinoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="19" pageNumber="1551" phylum="Mollusca" rank="species" species="chicama">
<emphasis id="B9664E3CE35BE228FBC8A7215AF0F951" italics="true" pageId="19" pageNumber="1551">M. chicama</emphasis>
</taxonomicName>
by having a truncate posterior margin and commarginal ribs developed over most, if not all, the area between the two umbonal ridges.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22FFF01A1CE5B87FF20" pageId="20" pageNumber="1552" type="etymology">
<paragraph id="8BAD922EE35CE22FFF01A1CE5B11FF6B" blockId="20.[160,1155,141,236]" box="[160,285,141,167]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A1CE5B11FF6B" bold="true" box="[160,285,141,167]" italics="true" pageId="20" pageNumber="1552">Etymology</emphasis>
</paragraph>
<paragraph id="8BAD922EE35CE22FFF01A1F15B87FF20" blockId="20.[160,1155,141,236]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A1F15AAAFF06" box="[160,166,178,202]" italics="true" pageId="20" pageNumber="1552">‘</emphasis>
Plenicostatus
<emphasis id="B9664E3CE35CE22FFE98A1F15B33FF06" box="[313,319,178,202]" italics="true" pageId="20" pageNumber="1552">’</emphasis>
, Latin for
<emphasis id="B9664E3CE35CE22FFE16A1F15BB0FF06" box="[439,444,178,202]" italics="true" pageId="20" pageNumber="1552">‘</emphasis>
fully ribbed
<emphasis id="B9664E3CE35CE22FFD9EA1F15848FF06" box="[575,580,178,202]" italics="true" pageId="20" pageNumber="1552">’</emphasis>
, referring to commarginal ribs crossing the entire valve of this species.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22FFF01A05B5838FE56" pageId="20" pageNumber="1552" type="materials_examined">
<paragraph id="8BAD922EE35CE22FFF01A05B5B0FFEFE" blockId="20.[160,1155,280,410]" box="[160,259,280,306]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A05B5B0FFEFE" bold="true" box="[160,259,280,306]" italics="true" pageId="20" pageNumber="1552">Material</emphasis>
</paragraph>
<paragraph id="8BAD922EE35CE22FFF01A07E5825FE56" blockId="20.[160,1155,280,410]" pageId="20" pageNumber="1552">
<materialsCitation id="3B7A9873E35CE22FFF01A07E5E24FE99" ID-GBIF-Occurrence="2571180954" box="[160,1064,317,341]" collectionCode="UWBM" longitude="-17.2" pageId="20" pageNumber="1552" specimenCode="UWBM 107571" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB43A55E35CE22FFF01A07E5B41FE99" box="[160,333,317,341]" pageId="20" pageNumber="1552">UWBM 107571</specimenCode>
,
<typeStatus id="54A92C8CE35CE22FFEFDA07E5BCAFE99" box="[348,454,317,341]" pageId="20" pageNumber="1552" type="holotype">holotype,</typeStatus>
B8772 (type locality), L 28.1, H 25.7, W 17.2 (pair)
</materialsCitation>
;
<materialsCitation id="3B7A9873E35CE22FFB96A07E5958FEB4" ID-GBIF-Occurrence="2571180957" collectionCode="UWBM" pageId="20" pageNumber="1552" specimenCode="UWBM 107572" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB43A55E35CE22FFB96A07E5AF9FEB4" pageId="20" pageNumber="1552">UWBM 107572</specimenCode>
,
<typeStatus id="54A92C8CE35CE22FFEA1A0235B65FEB4" box="[256,361,352,376]" pageId="20" pageNumber="1552" type="paratype">paratype,</typeStatus>
left valve, B8769, L (18.2), H (15.9), W (8.2)
</materialsCitation>
;
<materialsCitation id="3B7A9873E35CE22FFCC0A0235825FE56" ID-GBIF-Occurrence="2571180953" collectionCode="MUSM" longitude="-19.8" pageId="20" pageNumber="1552" specimenCode="INV 244" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED030AEBE35CE22FFCC0A02359A6FEB4" box="[865,938,352,376]" pageId="20" pageNumber="1552">MUSM</collectionCode>
<specimenCode id="DBB43A55E35CE22FFC12A0235E03FEB4" box="[947,1039,352,376]" pageId="20" pageNumber="1552">INV 244</specimenCode>
,
<typeStatus id="54A92C8CE35CE22FFBBBA0235E8FFEB4" box="[1050,1155,352,376]" pageId="20" pageNumber="1552" type="paratype">paratype,</typeStatus>
B8772, L 31.7, H 29.0, W 19.8 (pair
</materialsCitation>
</paragraph>
<paragraph id="8BAD922EE35CE22FFD88A0C15838FE56" blockId="20.[160,1155,280,410]" box="[553,564,386,410]" pageId="20" pageNumber="1552">).</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E35CE22FFF01A0855E88FDCF" box="[160,1156,454,515]" pageId="20" pageNumber="1552" type="distribution">
<paragraph id="8BAD922EE35CE22FFF01A0855B28FE2C" blockId="20.[160,1156,454,515]" box="[160,292,454,480]" pageId="20" pageNumber="1552">
<emphasis id="B9664E3CE35CE22FFF01A0855B28FE2C" bold="true" box="[160,292,454,480]" italics="true" pageId="20" pageNumber="1552">Occurrence</emphasis>
</paragraph>
<paragraph id="8BAD922EE35CE22FFF01A0A85E88FDCF" blockId="20.[160,1156,454,515]" box="[160,1156,491,515]" pageId="20" pageNumber="1552">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry id="F305D2BEE35CE22FFBEAA0A85E8DFDCF" box="[1099,1153,491,515]" name="Peru" pageId="20" pageNumber="1552">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>
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